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<document ID-DOI="http://doi.org/10.5281/zenodo.4762615" ID-GBIF-Dataset="564cead7-264a-4ee4-b5ee-15fd3ba81657" ID-GBIF-Taxon="182073487" ID-ISSN="1854-0392" ID-Zenodo-Dep="4762615" checkinTime="1620935982597" checkinUser="felipe" docAuthor="Stark, Bill P. &amp; Sivec, Ignac" docDate="2008" docId="03CA87B2FFA7FFC1FF112A727116FC70" docLanguage="en" docName="Illiesia.4.3.19-54.pdf" docOrigin="Illiesia 4 (3)" docStyle="DocumentStyle:1DEF1196F206C1C297CF0540A5C6768F.1:Illesia.2005-2008.journal_article" docStyleId="1DEF1196F206C1C297CF0540A5C6768F" docStyleName="Illesia.2005-2008.journal_article" docStyleVersion="1" docTitle="Neoperla FROM" docType="key" docVersion="4" lastPageId="34" lastPageNumber="53" masterDocId="FFF3FFCAFF86FFE3FFAB2954746AFFD5" masterDocTitle="New Species And Records Of Neoperla (Plecoptera: Perlidae) From Vietnam" masterLastPageNumber="54" masterPageNumber="19" pageId="33" pageNumber="52" updateTime="1632024248002" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>New Species And Records Of Neoperla (Plecoptera: Perlidae) From Vietnam</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Stark, Bill P.</mods:namePart>
<mods:affiliation>and Ignac Sivec &amp; Box 4045, Department of Biology, Mississippi College, Clinton, Mississippi, U. S. A. 39058 E-mail: stark @ mc. edu</mods:affiliation>
<mods:nameIdentifier type="email">stark@mc.edu</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sivec, Ignac</mods:namePart>
<mods:affiliation>Slovenian Museum of Natural History, Prešernova 20, P. O. Box 290, SLO- 1001 Ljubljana, Slovenia E-mail: isivec @ pms-lj. si</mods:affiliation>
<mods:nameIdentifier type="email">isivec@pms-lj.si</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Illiesia</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2008</mods:date>
<mods:detail type="pubDate">
<mods:number>2008-03-20</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>4</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>3</mods:number>
</mods:detail>
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<mods:start>19</mods:start>
<mods:end>54</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.4758856</mods:identifier>
<mods:identifier type="GBIF-Dataset">564cead7-264a-4ee4-b5ee-15fd3ba81657</mods:identifier>
<mods:identifier type="ISSN">1854-0392</mods:identifier>
<mods:identifier type="Zenodo-Dep">4758856</mods:identifier>
</mods:mods>
<subSection pageId="33" pageNumber="52" type="multiple">
<treatment ID-DOI="http://doi.org/10.5281/zenodo.4762615" ID-GBIF-Taxon="182073487" ID-Zenodo-Dep="4762615" LSID="urn:lsid:plazi:treatment:03CA87B2FFA7FFC1FF112A727116FC70" httpUri="http://treatment.plazi.org/id/03CA87B2FFA7FFC1FF112A727116FC70" lastPageId="34" lastPageNumber="53" pageId="33" pageNumber="52">
<subSubSection pageId="33" pageNumber="52" type="nomenclature">
<paragraph blockId="33.[186,773,806,865]" pageId="33" pageNumber="52">
<heading allCaps="true" bold="true" centered="true" fontSize="10" level="1" pageId="33" pageNumber="52" reason="1">
<emphasis bold="true" pageId="33" pageNumber="52">
IDENTIFICATION KEY FOR
<taxonomicName authority="FROM" authorityName="FROM" box="[543,773,806,830]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="33" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[543,686,806,830]" italics="true" pageId="33" pageNumber="52">NEOPERLA</emphasis>
FROM
</taxonomicName>
<collectingCountry box="[414,545,841,865]" name="Vietnam" pageId="33" pageNumber="52">VIETNAM</collectingCountry>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="33" pageNumber="52" type="discussion">
<paragraph blockId="33.[184,776,911,1142]" pageId="33" pageNumber="52">
The following provisional key is provided to aid in identification of male
<taxonomicName authority="from" authorityName="FROM" box="[493,657,945,969]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="33" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis box="[493,586,945,969]" italics="true" pageId="33" pageNumber="52">Neoperla</emphasis>
from
</taxonomicName>
<collectingCountry box="[671,772,945,969]" name="Vietnam" pageId="33" pageNumber="52">Vietnam</collectingCountry>
. Identifications obtained with this key should be confirmed by comparisions with figures and descriptions provided elsewhere in this paper or in other sources (
<bibRefCitation author="Uchida, S. &amp; T. Yamasaki" box="[348,632,1083,1107]" pageId="33" pageNumber="52" pagination="135 - 143" refId="ref16821" refString="Uchida, S. &amp; T. Yamasaki. 1989. Some Perlinae (Plecoptera: Perlidae) from the Malay Peninsula and Thailand with the redescription of Neoperla hamata from Assam. Bulletin of the Biogeographical Society of Japan, 44: 135 - 143." type="journal article" year="1989">Uchida &amp; Yamasaki 1989</bibRefCitation>
;
<bibRefCitation author="Zwick, P." box="[641,771,1083,1107]" pageId="33" pageNumber="52" pagination="167 - 204" refId="ref17050" refString="Zwick, P. 1983. The Neoperla of Sumatra and Java (Indonesia) (Plecoptera: Perlidae). Spixiana, 6: 167 - 204." type="journal article" year="1983">Zwick 1983</bibRefCitation>
,
<bibRefCitation author="Zwick, P." box="[184,240,1118,1142]" pageId="33" pageNumber="52" pagination="393 - 407" refId="ref17108" refString="Zwick, P. 1988. Species of Neoperla from the south- east Asian mainland (Plecoptera: Perlidae). Entomologica Scandinavica, 18: 393 - 407." type="journal article" year="1988">1988</bibRefCitation>
).
</paragraph>
<subSection lastPageId="34" lastPageNumber="53" pageId="33" pageNumber="52" type="multiple">
<paragraph blockId="33.[444,516,1188,1212]" box="[444,516,1188,1212]" pageId="33" pageNumber="52">
<heading bold="true" box="[444,516,1188,1212]" centered="true" fontSize="10" level="2" pageId="33" pageNumber="52" reason="2">
<emphasis bold="true" box="[444,516,1188,1212]" pageId="33" pageNumber="52">Males</emphasis>
</heading>
</paragraph>
<paragraph blockId="33.[184,776,1222,1281]" pageId="33" pageNumber="52">
(
<emphasis italics="true" pageId="33" pageNumber="52">
<taxonomicName box="[192,337,1222,1246]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="33" pageNumber="52" phylum="Arthropoda" rank="species" species="brachyura">N. brachyura</taxonomicName>
,
<taxonomicName box="[350,499,1222,1246]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="33" pageNumber="52" phylum="Arthropoda" rank="species" species="hoabihnica">N. hoabihnica</taxonomicName>
,
<taxonomicName baseAuthorityName="Navas" baseAuthorityYear="1919" box="[512,626,1223,1246]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="33" pageNumber="52" phylum="Arthropoda" rank="species" species="ramosa">N. ramosa</taxonomicName>
,
<taxonomicName box="[639,735,1223,1246]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="33" pageNumber="52" phylum="Arthropoda" rank="species" species="sungi">N. sungi</taxonomicName>
,
<taxonomicName class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="33" pageNumber="52" phylum="Arthropoda" rank="species" species="vitalisi">N. vitalisi</taxonomicName>
</emphasis>
not included)
</paragraph>
<subSubSection lastPageId="34" lastPageNumber="53" pageId="33" pageNumber="52" type="key">
<key lastPageId="34" lastPageNumber="53" pageId="33" pageNumber="52">
<keyStep pageId="33" pageNumber="52">
<paragraph blockId="33.[184,776,1326,1972]" pageId="33" pageNumber="52">
<keyLead pageId="33" pageNumber="52">
1 Tergum 7 bears a broadly truncate, elevated, plateau like process, tergum 8 bears a slender, erect, curved process (
<figureCitation box="[476,553,1395,1419]" captionStart="Figs" captionStartId="11.[184,233,1186,1210]" captionTargetBox="[205,1432,273,1147]" captionTargetId="figure-371@11.[594,940,259,756]" captionTargetPageId="11" captionText="Figs. 2933. Neoperla hubleyi. 29. Head and pronotum, 30. Male terminalia, 31. Aedeagus, 32. Female terminalia, 33. Vagina and spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4758874" httpUri="https://zenodo.org/record/4758874/files/figure.png" pageId="33" pageNumber="52">Fig. 30</figureCitation>
) …….……….…... 2
</keyLead>
</paragraph>
<paragraph blockId="33.[184,776,1326,1972]" pageId="33" pageNumber="52">
<keyLead pageId="33" pageNumber="52">1 Tergum 7 and tergum 8 processes variable, but not as described above.…………………............ 9</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="33" pageNumber="52">
<paragraph blockId="33.[184,776,1326,1972]" pageId="33" pageNumber="52">
<keyLead pageId="33" pageNumber="52">
2 Aedeagal tube bears a projecting, sometimes spiny or forked, slender membranous lobe (
<figureCitation captionStart="Figs" captionStartId="8.[184,234,1117,1141]" captionTargetBox="[275,1379,290,1078]" captionTargetId="figure-353@8.[656,890,275,900]" captionTargetPageId="8" captionText="Figs. 1922. Neoperla dao. 19. Head and pronotum, 20. Male terminalia, 21. Male terminalia, lateral aspect, 22. Aedeagus." figureDoi="http://doi.org/10.5281/zenodo.4758870" httpUri="https://zenodo.org/record/4758870/files/figure.png" pageId="33" pageNumber="52">Fig. 22</figureCitation>
) ……………………………………………….…. 3
</keyLead>
</paragraph>
<paragraph blockId="33.[184,776,1326,1972]" pageId="33" pageNumber="52">
<keyLead pageId="33" pageNumber="52">
2 Aedeagal tube without lobes, or if present, lobes are large and spherical; lobes present or absent on sac (
<figureCitation box="[271,347,1671,1695]" captionStart="Figs" captionStartId="16.[184,234,1912,1936]" captionTargetBox="[246,1392,1068,1862]" captionTargetId="figure-359@16.[628,959,1076,1612]" captionTargetPageId="16" captionText="Figs. 5156. Neoperla nebulosa. 51. Head and pronotum, 52. Male terminalia, 53. Aedeagus, 54. Aedeagal sac, dorsal, 55. Female terminalia, 56. Vagina and spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4758884" httpUri="https://zenodo.org/record/4758884/files/figure.png" pageId="33" pageNumber="52">Fig. 53</figureCitation>
) ……….……………………..…….…. 7
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="33" pageNumber="52">
<paragraph blockId="33.[184,776,1326,1972]" pageId="33" pageNumber="52">
<keyLead pageId="33" pageNumber="52">
3 Lobe of aedeagal tube a simple, nipple shaped structure (
<figureCitation box="[337,412,1741,1765]" captionStart="Figs" captionStartId="8.[184,234,1117,1141]" captionTargetBox="[275,1379,290,1078]" captionTargetId="figure-353@8.[656,890,275,900]" captionTargetPageId="8" captionText="Figs. 1922. Neoperla dao. 19. Head and pronotum, 20. Male terminalia, 21. Male terminalia, lateral aspect, 22. Aedeagus." figureDoi="http://doi.org/10.5281/zenodo.4758870" httpUri="https://zenodo.org/record/4758870/files/figure.png" pageId="33" pageNumber="52">Fig. 22</figureCitation>
) …………………………..…..
<taxonomicName authorityName="Stark &amp; Sivec" authorityYear="2008" box="[740,776,1741,1765]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="33" pageNumber="52" phylum="Arthropoda" rank="species" species="dao">
<emphasis box="[740,776,1741,1765]" italics="true" pageId="33" pageNumber="52">dao</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="33.[184,776,1326,1972]" box="[184,775,1775,1799]" pageId="33" pageNumber="52">
<keyLead box="[184,775,1775,1799]" pageId="33" pageNumber="52">
3 Lobe of aedeagal tube bifurcate (
<figureCitation box="[594,671,1775,1799]" captionStart="Figs" captionStartId="14.[184,234,1159,1183]" captionTargetBox="[227,1400,285,1106]" captionTargetId="figure-385@14.[579,947,275,814]" captionTargetPageId="14" captionText="Figs. 4044. Neoperla mnong. 40. Head and pronotum, 41. Male terminalia, 42. Aedeagus, 43. Female terminalia, 44. Vagina and spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4758878" httpUri="https://zenodo.org/record/4758878/files/figure.png" pageId="33" pageNumber="52">Fig. 42</figureCitation>
) …..…. 4
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="33" pageNumber="52">
<paragraph blockId="33.[184,776,1326,1972]" box="[184,775,1810,1834]" pageId="33" pageNumber="52">
<keyLead box="[184,775,1810,1834]" pageId="33" pageNumber="52">4 Lobe of aedeagal tube deeply forked ………..…. 5</keyLead>
</paragraph>
<paragraph blockId="33.[184,776,1326,1972]" pageId="33" pageNumber="52">
<keyLead pageId="33" pageNumber="52">
4 Lobe of aedeagal tube with long Y stalk and short arms (
<figureCitation box="[292,368,1879,1903]" captionStart="Figs" captionStartId="14.[184,234,1159,1183]" captionTargetBox="[227,1400,285,1106]" captionTargetId="figure-385@14.[579,947,275,814]" captionTargetPageId="14" captionText="Figs. 4044. Neoperla mnong. 40. Head and pronotum, 41. Male terminalia, 42. Aedeagus, 43. Female terminalia, 44. Vagina and spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4758878" httpUri="https://zenodo.org/record/4758878/files/figure.png" pageId="33" pageNumber="52">Fig. 42</figureCitation>
) …………………………………….. 6
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="33" pageNumber="52">
<paragraph blockId="33.[184,776,1326,1972]" lastBlockId="33.[812,1404,288,1970]" pageId="33" pageNumber="52">
<keyLead pageId="33" pageNumber="52">
5 Lobe of aedeagal tube forked to base and without spiny armature; arms of lobe less than three times long as wide (
<figureCitation box="[1007,1084,288,312]" captionStart="Figs" captionStartId="23.[184,234,1151,1175]" captionTargetBox="[267,1401,523,1082]" captionTargetId="figure-445@23.[672,1407,588,957]" captionTargetPageId="23" captionText="Figs. 7577. Neoperla yentu. 75. Male terminalia, 76. Aedeagus. 77, Detail of aedeagus lobing." figureDoi="http://doi.org/10.5281/zenodo.4758892" httpUri="https://zenodo.org/record/4758892/files/figure.png" pageId="33" pageNumber="52">Fig. 76</figureCitation>
) ….……………….…...
<taxonomicName box="[1343,1403,288,311]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="33" pageNumber="52" phylum="Arthropoda" rank="species" species="yentu">
<emphasis box="[1343,1403,288,311]" italics="true" pageId="33" pageNumber="52">yentu</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="33.[812,1404,288,1970]" pageId="33" pageNumber="52">
<keyLead pageId="33" pageNumber="52">
5 Lobe of aedeagal tube with a short basal stalk, and armed apically with minute spines; arms of lobe at least five times long as wide ………..…
<taxonomicName authorityName="Stark" authorityYear="1987" box="[1368,1404,392,415]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="33" pageNumber="52" phylum="Arthropoda" rank="species" species="yao">
<emphasis box="[1368,1404,392,415]" italics="true" pageId="33" pageNumber="52">yao</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="33" pageNumber="52">
<paragraph blockId="33.[812,1404,288,1970]" pageId="33" pageNumber="52">
<keyLead pageId="33" pageNumber="52">
6 Stalk of aedeagal lobe subequal to arms in length; aedeagal sac unarmed on dorsum for most of length proximal to subapical spiny mound (
<figureCitation captionStart="Figs" captionStartId="14.[184,234,1159,1183]" captionTargetBox="[227,1400,285,1106]" captionTargetId="figure-385@14.[579,947,275,814]" captionTargetPageId="14" captionText="Figs. 4044. Neoperla mnong. 40. Head and pronotum, 41. Male terminalia, 42. Aedeagus, 43. Female terminalia, 44. Vagina and spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4758878" httpUri="https://zenodo.org/record/4758878/files/figure.png" pageId="33" pageNumber="52">Fig. 42</figureCitation>
) ………………………………………..…..
<taxonomicName authorityName="Stark" authorityYear="1987" box="[1331,1403,530,553]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="33" pageNumber="52" phylum="Arthropoda" rank="species" species="mnong">
<emphasis box="[1331,1403,530,553]" italics="true" pageId="33" pageNumber="52">mnong</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="33.[812,1404,288,1970]" pageId="33" pageNumber="52">
<keyLead pageId="33" pageNumber="52">
6 Stalk of aedeagal lobe about twice as long as arms; aedeagal sac rather densely and completely armed for all but a short basal section (
<figureCitation box="[1316,1395,633,657]" captionStart="Figs" captionStartId="11.[184,233,1186,1210]" captionTargetBox="[205,1432,273,1147]" captionTargetId="figure-371@11.[594,940,259,756]" captionTargetPageId="11" captionText="Figs. 2933. Neoperla hubleyi. 29. Head and pronotum, 30. Male terminalia, 31. Aedeagus, 32. Female terminalia, 33. Vagina and spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4758874" httpUri="https://zenodo.org/record/4758874/files/figure.png" pageId="33" pageNumber="52">Fig. 31</figureCitation>
) …………………………………………….....
<taxonomicName authorityName="Stark &amp; Sivec" authorityYear="2008" box="[1327,1403,667,691]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="33" pageNumber="52" phylum="Arthropoda" rank="species" species="hubleyi">
<emphasis box="[1327,1403,667,691]" italics="true" pageId="33" pageNumber="52">hubleyi</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="33" pageNumber="52">
<paragraph blockId="33.[812,1404,288,1970]" pageId="33" pageNumber="52">
<keyLead pageId="33" pageNumber="52">
7 Aedeagal sac without spiny lobes but dorsal margin bearing a double row of 47 cultriform spines (
<figureCitation box="[936,1058,771,795]" captionStart="Figs" captionStartId="13.[184,234,1289,1313]" captionTargetBox="[221,1403,275,1214]" captionTargetId="figure-323@13.[594,940,275,862]" captionTargetPageId="13" captionText="Figs. 3439. Neoperla idella. 34. Head and pronotum. 35. Male terminalia. 36. Aedeagus. 37. Aedeagal sac, dorsal. 38. Female terminalia. 39. Vagina and spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4758876" httpUri="https://zenodo.org/record/4758876/files/figure.png" pageId="33" pageNumber="52">Figs. 3637</figureCitation>
).…………………….…...
<taxonomicName authorityName="Stark &amp; Sivec" authorityYear="2008" box="[1344,1400,771,795]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="33" pageNumber="52" phylum="Arthropoda" rank="species" species="idella">
<emphasis box="[1344,1400,771,795]" italics="true" pageId="33" pageNumber="52">idella</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="33.[812,1404,288,1970]" pageId="33" pageNumber="52">
<keyLead pageId="33" pageNumber="52">
7 Aedeagal sac with a pair of spiny lobes (
<figureCitation box="[1318,1396,806,830]" captionStart="Figs" captionStartId="16.[184,234,1912,1936]" captionTargetBox="[246,1392,1068,1862]" captionTargetId="figure-359@16.[628,959,1076,1612]" captionTargetPageId="16" captionText="Figs. 5156. Neoperla nebulosa. 51. Head and pronotum, 52. Male terminalia, 53. Aedeagus, 54. Aedeagal sac, dorsal, 55. Female terminalia, 56. Vagina and spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4758884" httpUri="https://zenodo.org/record/4758884/files/figure.png" pageId="33" pageNumber="52">Fig. 53</figureCitation>
) ………………………………..…………………..… 8
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="33" pageNumber="52">
<paragraph blockId="33.[812,1404,288,1970]" pageId="33" pageNumber="52">
<keyLead pageId="33" pageNumber="52">
8 Sac angled abruptly dorsad; sac unarmed on ventral margin for most of length distal to lobes (
<figureCitation box="[857,974,944,968]" captionStart="Figs" captionStartId="16.[184,234,1912,1936]" captionTargetBox="[246,1392,1068,1862]" captionTargetId="figure-359@16.[628,959,1076,1612]" captionTargetPageId="16" captionText="Figs. 5156. Neoperla nebulosa. 51. Head and pronotum, 52. Male terminalia, 53. Aedeagus, 54. Aedeagal sac, dorsal, 55. Female terminalia, 56. Vagina and spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4758884" httpUri="https://zenodo.org/record/4758884/files/figure.png" pageId="33" pageNumber="52">Figs. 5354</figureCitation>
) ……………………………..….
<taxonomicName authorityName="Stark &amp; Sivec" authorityYear="2008" box="[1315,1404,944,968]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="33" pageNumber="52" phylum="Arthropoda" rank="species" species="nebulosa">
<emphasis box="[1315,1404,944,968]" italics="true" pageId="33" pageNumber="52">nebulosa</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="33.[812,1404,288,1970]" pageId="33" pageNumber="52">
<keyLead pageId="33" pageNumber="52">
8 Sac curved gradually dorsad; ventral margin of sac armed for most of length distal to lobes (
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) ………………………………………...
<taxonomicName authorityName="Stark &amp; Sivec" authorityYear="2008" box="[1304,1402,1047,1071]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="33" pageNumber="52" phylum="Arthropoda" rank="species" species="spinaloba">
<emphasis box="[1304,1402,1047,1071]" italics="true" pageId="33" pageNumber="52">spinaloba</emphasis>
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9 Process of tergum 7 broad at tip, often rounded, truncate or notched (
<figureCitation box="[1080,1166,1117,1141]" captionStart="Figs" captionStartId="9.[184,234,1255,1279]" captionTargetBox="[277,1336,318,1168]" captionTargetId="figure-298@9.[267,929,676,1152]" captionTargetPageId="9" captionText="Figs. 2325. Neoperla erecta. 23. Head and pronotum, 24. Male terminalia, 25. Aedeagus." figureDoi="http://doi.org/10.5281/zenodo.4758872" httpUri="https://zenodo.org/record/4758872/files/figure.png" pageId="33" pageNumber="52">Figs. 24</figureCitation>
,
<figureCitation box="[1175,1202,1116,1140]" captionStart="Figs" captionStartId="18.[184,234,1186,1210]" captionTargetBox="[236,1435,276,1131]" captionTargetId="figure-355@18.[610,978,276,877]" captionText="Figs. 5761. Neoperla song. 57. Head and pronotum, 58. Male terminalia, 59. Aedeagus, 60. Female terminalia, 61. Vagina and spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4762648" httpUri="https://zenodo.org/record/4762648/files/figure.png" pageId="33" pageNumber="52">58</figureCitation>
) ………………. 10
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<paragraph blockId="33.[812,1404,288,1970]" pageId="33" pageNumber="52">
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9 Process of tergum 7 more or less triangular (
<figureCitation captionStart="Figs" captionStartId="15.[184,233,1912,1936]" captionTargetBox="[272,1422,1054,1850]" captionTargetId="figure-428@15.[579,903,1053,1586]" captionTargetPageId="15" captionText="Figs. 4550. Neoperla multispinosa.45. Head and pronotum, 46. Male terminalia, 47. Aedeagus, 48.Aedeagal sac, dorsal, 49. Female terminalia, 50. Vagina and spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4758882" httpUri="https://zenodo.org/record/4758882/files/figure.png" pageId="33" pageNumber="52">Fig. 46</figureCitation>
) ………………………………………..………. 14
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<paragraph blockId="33.[812,1404,288,1970]" pageId="33" pageNumber="52">
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10 Everted aedeagal sac extends ventrally at an approximate right angle to tube (
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) …..
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<keyLead pageId="33" pageNumber="52">10 Everted aedeagal sac curved ventrad but not oriented at a right angle to tube.………..……… 11</keyLead>
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<paragraph blockId="33.[812,1404,288,1970]" pageId="33" pageNumber="52">
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11 Aedeagus with one or more spiny lobes (
<figureCitation box="[1319,1396,1358,1382]" captionStart="Figs" captionStartId="18.[184,234,1186,1210]" captionTargetBox="[236,1435,276,1131]" captionTargetId="figure-355@18.[610,978,276,877]" captionText="Figs. 5761. Neoperla song. 57. Head and pronotum, 58. Male terminalia, 59. Aedeagus, 60. Female terminalia, 61. Vagina and spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4762648" httpUri="https://zenodo.org/record/4762648/files/figure.png" pageId="33" pageNumber="52">Fig. 59</figureCitation>
) …………………………………………….……..… 12
</keyLead>
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<paragraph blockId="33.[812,1404,288,1970]" box="[812,1404,1427,1452]" pageId="33" pageNumber="52">
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11 Aedeagus without spiny lobes (
<figureCitation box="[1210,1286,1427,1451]" captionStart="Figs" captionStartId="21.[184,234,1220,1244]" captionTargetBox="[236,1425,292,1168]" captionTargetId="figure-350@21.[594,962,292,820]" captionTargetPageId="21" captionText="Figs. 6771. Neoperla teresa. 67. Head and pronotum, 68.Male terminalia, 69. Aedeagus, 70. Vagina and spermatheca, 71. Female terminalia." figureDoi="http://doi.org/10.5281/zenodo.4758888" httpUri="https://zenodo.org/record/4758888/files/figure.png" pageId="33" pageNumber="52">Fig. 69</figureCitation>
).…..… 13
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<paragraph blockId="33.[812,1404,288,1970]" pageId="33" pageNumber="52">
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12 Aedeagal sac with a dorsobasal pair of prominent spiny lobes; sac armed with an irregular, subapical ring of cultriform spines (
<figureCitation box="[1309,1396,1531,1555]" captionStart="Figs" captionStartId="18.[184,234,1186,1210]" captionTargetBox="[236,1435,276,1131]" captionTargetId="figure-355@18.[610,978,276,877]" captionText="Figs. 5761. Neoperla song. 57. Head and pronotum, 58. Male terminalia, 59. Aedeagus, 60. Female terminalia, 61. Vagina and spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4762648" httpUri="https://zenodo.org/record/4762648/files/figure.png" pageId="33" pageNumber="52">Fig. 59</figureCitation>
) ………………………………………………..…
<taxonomicName authorityName="Stark &amp; Sivec" authorityYear="2008" box="[1355,1403,1566,1589]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="33" pageNumber="52" phylum="Arthropoda" rank="species" species="song">
<emphasis box="[1355,1403,1566,1589]" italics="true" pageId="33" pageNumber="52">song</emphasis>
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12 Aedeagal sac without dorsobasal spiny lobes; sac armed at midlength by a large, ventral lobe and subapically by patches of cultriform spines …………………………………………...…...
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<emphasis box="[1328,1403,1704,1728]" italics="true" pageId="33" pageNumber="52">tamdao</emphasis>
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13 Hemitergal finger lobes curved dorsad near tips; process of tergum 7 truncate; aedeagal sac armature larger on dorsal margin, but not cultriform …………………………………...
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13 Hemitergal finger lobes curved inward; process of tergum 7 rounded (
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); aedeagal sac armature with a double row of dorsal cultriform spines on apical third (
<figureCitation box="[362,436,288,312]" captionStart="Figs" captionStartId="21.[184,234,1220,1244]" captionTargetBox="[236,1425,292,1168]" captionTargetId="figure-350@21.[594,962,292,820]" captionTargetPageId="21" captionText="Figs. 6771. Neoperla teresa. 67. Head and pronotum, 68.Male terminalia, 69. Aedeagus, 70. Vagina and spermatheca, 71. Female terminalia." figureDoi="http://doi.org/10.5281/zenodo.4758888" httpUri="https://zenodo.org/record/4758888/files/figure.png" pageId="34" pageNumber="53">Fig. 69</figureCitation>
) ……………………….…
<taxonomicName authorityName="Stark &amp; Sivec" authorityYear="2008" box="[716,776,288,311]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="34" pageNumber="53" phylum="Arthropoda" rank="species" species="teresa">
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14 Aedeagal sac with two or more spiny lobes (
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) …………………………………………….…. 15
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14 Aedeagal sac without spiny lobes (
<figureCitation box="[615,690,391,415]" captionStart="Figs" captionStartId="6.[184,234,1047,1071]" captionTargetBox="[211,1377,256,1011]" captionTargetId="figure-43@6.[608,903,279,1020]" captionTargetPageId="6" captionText="Figs. 1214. Neoperla monacha. 12. Head and pronotum, 13. Male terminalia, 14. Aedeagus." figureDoi="http://doi.org/10.5281/zenodo.4758866" httpUri="https://zenodo.org/record/4758866/files/figure.png" pageId="34" pageNumber="53">Fig. 14</figureCitation>
) ….. 16
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<paragraph blockId="34.[184,776,288,1970]" box="[184,775,426,450]" pageId="34" pageNumber="53">
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15 Aedeagal sac trilobed (
<figureCitation box="[475,550,426,450]" captionStart="Figs" captionStartId="24.[184,234,944,968]" captionTargetBox="[236,1405,212,922]" captionTargetId="figure-449@24.[656,878,321,687]" captionTargetPageId="24" captionText="Figs. 7881. Neoperla zonata. 78. Head and pronotum, 79. Male terminalia, 80. Aedeagus, 81. Aedeagal sac, ventral." figureDoi="http://doi.org/10.5281/zenodo.4758896" httpUri="https://zenodo.org/record/4758896/files/figure.png" pageId="34" pageNumber="53">Fig. 80</figureCitation>
) ………….….
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<paragraph blockId="34.[184,776,288,1970]" pageId="34" pageNumber="53">
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15 Aedeagal sac with three ventral lobes arranged linearly (
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) ……………….…….
<taxonomicName authorityName="Stark &amp; Sivec" authorityYear="2008" box="[640,775,495,519]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="34" pageNumber="53" phylum="Arthropoda" rank="species" species="multispinosa">
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16 Aedeagal tube armed with one or more spines in apical half (
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) ……………………………... 17
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<paragraph blockId="34.[184,776,288,1970]" pageId="34" pageNumber="53">
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16 Aedeagal tube armature absent apically or reduced to small spicules (
<figureCitation box="[524,586,633,657]" captionStart="Figs" captionStartId="2.[184,234,978,1002]" captionTargetBox="[211,1392,249,953]" captionTargetId="figure-448@2.[579,910,244,750]" captionTargetPageId="2" captionText="Figs. 15. Neoperla clara. 1. Head and pronotum, 2. Male terminalia, 3. Aedeagus, 4. Female terminalia, 5. Vagina and spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4758858" httpUri="https://zenodo.org/record/4758858/files/figure.png" pageId="34" pageNumber="53">Fig. 3</figureCitation>
) …………….. 19
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<paragraph blockId="34.[184,776,288,1970]" pageId="34" pageNumber="53">
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17 Dorsal margin of aedeagal tube armed from near midlength to apical third with a few spines; sac armature extends from near base to apex.….
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17 Dorsal margin of aedeagal tube armed near apex with one or more spines; ventral margin of tube with additional spines (
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); sac armature begins near midlength (
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) …..………….. 18
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18 Aedeagal tube armed at midlength with a pair of small unsclerotized spiny patches; venter with a small patch of about five apical spines; sac armature consists of scattered cultriform spines primarily along dorsal margin (
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).
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18 Aedeagal tube without midlength armature; venter with a large apical patch of spines; sac armature covers most of surface at midlength (
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) ……………………………………..
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19 Base of aedeagal sac with a complete ring of long close set spines; process of tergum 7 very slender and acute at tip ……………………………
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19 Base of aedeagal sac without grouping of long spines; process of tergum 7 broadly triangular and rounded or slightly forked at tip (
<figureCitation box="[655,719,1393,1417]" captionStart="Figs" captionStartId="2.[184,234,978,1002]" captionTargetBox="[211,1392,249,953]" captionTargetId="figure-448@2.[579,910,244,750]" captionTargetPageId="2" captionText="Figs. 15. Neoperla clara. 1. Head and pronotum, 2. Male terminalia, 3. Aedeagus, 4. Female terminalia, 5. Vagina and spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4758858" httpUri="https://zenodo.org/record/4758858/files/figure.png" pageId="34" pageNumber="53">Fig. 2</figureCitation>
). 20
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20 Aedeagal tube somewhat plump and lacking sclerotization except basally and along dorsal margin …………………………….………...…
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20 Aedeagal tube slender and sclerotized along dorsal and ventral margins (
<figureCitation box="[543,606,1566,1590]" captionStart="Figs" captionStartId="2.[184,234,978,1002]" captionTargetBox="[211,1392,249,953]" captionTargetId="figure-448@2.[579,910,244,750]" captionTargetPageId="2" captionText="Figs. 15. Neoperla clara. 1. Head and pronotum, 2. Male terminalia, 3. Aedeagus, 4. Female terminalia, 5. Vagina and spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4758858" httpUri="https://zenodo.org/record/4758858/files/figure.png" pageId="34" pageNumber="53">Fig. 3</figureCitation>
) ……….….. 21
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<paragraph blockId="34.[184,776,288,1970]" pageId="34" pageNumber="53">
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21 Basal patch of small spines of aedeagal sac in contact with tube apex (
<figureCitation box="[494,557,1635,1659]" captionStart="Figs" captionStartId="2.[184,234,978,1002]" captionTargetBox="[211,1392,249,953]" captionTargetId="figure-448@2.[579,910,244,750]" captionTargetPageId="2" captionText="Figs. 15. Neoperla clara. 1. Head and pronotum, 2. Male terminalia, 3. Aedeagus, 4. Female terminalia, 5. Vagina and spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4758858" httpUri="https://zenodo.org/record/4758858/files/figure.png" pageId="34" pageNumber="53">Fig. 3</figureCitation>
) ……………….. 22
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21 Basal patch of small spines of aedeagal sac clearly separated from tube apex by unarmed membrane (
<figureCitation box="[229,291,1738,1762]" captionStart="Figs" captionStartId="3.[184,234,944,968]" captionTargetBox="[338,1377,265,914]" captionTargetId="figure-41@3.[796,1385,259,920]" captionTargetPageId="3" captionText="Figs. 67. Neoperla daklak. 6. Male terminalia, 7. Aedeagus." figureDoi="http://doi.org/10.5281/zenodo.4758860" httpUri="https://zenodo.org/record/4758860/files/figure.png" pageId="34" pageNumber="53">Fig. 7</figureCitation>
) ………………………………………..….. 23
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<paragraph blockId="34.[184,776,288,1970]" pageId="34" pageNumber="53">
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22 Basal patch of small spines on aedeagal sac restricted to dorsolateral surfaces; distal sac armature includes irregular rows of moderately sized spines on dorsal and ventral margins …………………..…………………………
<taxonomicName authorityName="Stark" authorityYear="1983" box="[680,775,1911,1935]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="34" pageNumber="53" phylum="Arthropoda" rank="species" species="gordonae">
<emphasis box="[680,775,1911,1935]" italics="true" pageId="34" pageNumber="53">gordonae</emphasis>
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22 Basal patch of small spines completely rings aedeagal sac; distal sac armature not organized into irregular rows of spines (
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) ………
<taxonomicName authorityName="Stark &amp; Sivec" authorityYear="2008" box="[1349,1400,322,346]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="34" pageNumber="53" phylum="Arthropoda" rank="species" species="clara">
<emphasis box="[1349,1400,322,346]" italics="true" pageId="34" pageNumber="53">clara</emphasis>
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<paragraph blockId="34.[812,1404,288,933]" pageId="34" pageNumber="53">
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23 Base of aedeagal sac bears a dorsal cushion of small spines and longer spines along dorsal margin; ventral sac margin bears a patch of more prominent spines near midlength, apical sac armature reduced on both margins to minute spicules …………………………………..
<taxonomicName authority="Enderlein, 1909" authorityName="Navas" authorityYear="1922" box="[1303,1401,529,553]" class="Insecta" family="Perlidae" genus="Ochthopetina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="34" pageNumber="53" phylum="Arthropoda" rank="species" species="cavaleriei">
<emphasis box="[1303,1401,529,553]" italics="true" pageId="34" pageNumber="53">cavaleriei</emphasis>
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<keyLead pageId="34" pageNumber="53">23 Base of aedeagal sac without dorsal cushion of spines ……………………………………….…… 24</keyLead>
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24 Aedeagal sac base completely ringed with small spines but discretely separated from tube apex by narrow band of membrane; margins of sac without larger spines (
<figureCitation box="[1103,1164,737,761]" captionStart="Figs" captionStartId="3.[184,234,944,968]" captionTargetBox="[338,1377,265,914]" captionTargetId="figure-41@3.[796,1385,259,920]" captionTargetPageId="3" captionText="Figs. 67. Neoperla daklak. 6. Male terminalia, 7. Aedeagus." figureDoi="http://doi.org/10.5281/zenodo.4758860" httpUri="https://zenodo.org/record/4758860/files/figure.png" pageId="34" pageNumber="53">Fig. 7</figureCitation>
) ………………
<taxonomicName authorityName="Stark &amp; Sivec" authorityYear="2008" box="[1337,1403,737,761]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="34" pageNumber="53" phylum="Arthropoda" rank="species" species="daklak">
<emphasis box="[1337,1403,737,761]" italics="true" pageId="34" pageNumber="53">daklak</emphasis>
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24 Aedeagal sac unarmed for at least basal 20% of sac length; armature on dorsal margin consists of irregular row of small peg shaped spines; ventral armature begins distally to origin of dorsal armature (
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) ……….……………...
<taxonomicName authorityName="Stark &amp; Sivec" authorityYear="2008" box="[1284,1404,909,933]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="34" pageNumber="53" phylum="Arthropoda" rank="species" species="leptacantha">
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