300 lines
33 KiB
XML
300 lines
33 KiB
XML
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<mods:title id="637FB98BE7D7110C9CC1376DC1146848">From the mud to the tree: phylogeny of Austrolebias killifishes, new generic structure and description of a new species (Cyprinodontiformes: Rivulidae)</mods:title>
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<mods:namePart id="C986639C8FC255DA5ABC9F14C963C613">Alonso, Felipe</mods:namePart>
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<mods:affiliation id="87F9B3692E57DDCD5218FF9191A545F0">Instituto de Bio y Geociencias del NOA (IBIGEO), National Scientific and Technical Research Council (CONICET) - UNSa, Av. 9 de Julio 14, A 4405 BBA Rosario de Lerma, Salta, Argentina & Killifish Foundation, La Plata, Buenos Aires Province, Argentina</mods:affiliation>
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<mods:namePart id="2BE78E7FC84949259739362360FEC4A5">Terán, Guillermo Enrique</mods:namePart>
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<mods:affiliation id="839D47D472F30F046A55853E2A38BCBA">Killifish Foundation, La Plata, Buenos Aires Province, Argentina & Fundación Miguel Lillo - Unidad Ejecutora Lillo (CONICET), San Miguel de Tucumán, Tucumán, Tucuman Province, Argentina</mods:affiliation>
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<mods:namePart id="A11FBE98447D54DA0F47E1228B6FA973">Alanís, Wilson Sebastián Serra</mods:namePart>
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<mods:affiliation id="F07DAD41E0D8933A1D7DAA4AEE3FCF67">Killifish Foundation, La Plata, Buenos Aires Province, Argentina & Sección Ictiología, Departamento de Zoología, Museo Nacional de Historia Natural, Montevideo, Uruguay & Centro Universitario Regional del Este (CURE) Sede Rocha, Rocha, Uruguay</mods:affiliation>
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<mods:namePart id="C91721460AEC0558DF104AD97E96D831">Calviño, Pablo</mods:namePart>
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<mods:affiliation id="FC5B48F1B8644680859DD496A4939A0D">Killifish Foundation, La Plata, Buenos Aires Province, Argentina</mods:affiliation>
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<mods:namePart id="039A369906B40F14AB4BBD662DB1B7D5">Montes, Martin Miguel</mods:namePart>
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<mods:affiliation id="B350A64651D8AF5A386825CBA450FC80">Killifish Foundation, La Plata, Buenos Aires Province, Argentina & Centro de Estudios Parasitológicos y de Vectores - [CCT-CONICET - La Plata] Centro Científico Tecnológico Conicet, La Plata - [Conicet] Consejo Nacional De Investigaciones Científicas y Técnicas, La Plata, Buenos Aires Province, Argentina</mods:affiliation>
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<mods:namePart id="EFF679646C244B4D8008A8E68573E196">García, Ignacio Daniel</mods:namePart>
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<mods:affiliation id="317E316E71747CD730B4DD7D7AD225AB">Killifish Foundation, La Plata, Buenos Aires Province, Argentina & Instituto de Limnología ‘ Dr. Raúl Ringuelet’ (ILPLA) - UNLP-CONICET, La Plata, Buenos Aires Province, Argentina</mods:affiliation>
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<mods:namePart id="43E7495B4C46F8DB2C2315A94F48FA40">Barneche, Jorge Adrián</mods:namePart>
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<mods:affiliation id="DACD81DB657C149CDC74C72D41FB38C5">Killifish Foundation, La Plata, Buenos Aires Province, Argentina & Centro de Estudios Parasitológicos y de Vectores - [CCT-CONICET - La Plata] Centro Científico Tecnológico Conicet, La Plata - [Conicet] Consejo Nacional De Investigaciones Científicas y Técnicas, La Plata, Buenos Aires Province, Argentina</mods:affiliation>
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<mods:namePart id="1DE004A2DCDF18E342F31470CFE787CD">Almirón, Adriana</mods:namePart>
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<mods:affiliation id="81E89E41DAFA7EE65202C7135627F883">Universidad Nacional de La Plata, Facultad de Ciencias Naturales y Museo, División Zoología Vertebrados, La Plata, Buenos Aires Province, Argentina</mods:affiliation>
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<mods:namePart id="E0D47B8C8EB25E593C1C1096CD9A6862">Ciotek, Liliana</mods:namePart>
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<mods:affiliation id="59F64909324145C485D651B47B7EDFA0">Administración de Parques Nacionales, Parque Nacional Campos del Tuyú (7103), General Lavalle, Buenos Aires Province, Argentina</mods:affiliation>
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<mods:namePart id="BA092E6908080EAE1F9241332AEFAB0E">Giorgis, Pablo</mods:namePart>
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<mods:affiliation id="A8CA115D607D6C0AD805DB98CA895166">Administración de Parques Nacionales, Parque Nacional Campos del Tuyú (7103), General Lavalle, Buenos Aires Province, Argentina</mods:affiliation>
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<mods:namePart id="2BA412AE7DD76119CA245784AAB56DF0">Cascioưa, Jorge</mods:namePart>
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<mods:affiliation id="71819A5A17D377BA772117B7EC3C981B">Universidad Nacional de La Plata, Facultad de Ciencias Naturales y Museo, División Zoología Vertebrados, La Plata, Buenos Aires Province, Argentina</mods:affiliation>
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<mods:title id="58F4F1D6B1FB61441D42F232C26C10A4">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="AA72CB280B856639A7F3FC201E8BE1A9">2023</mods:date>
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<mods:number id="F73A57BE49CF8F8C9ADC4D18FFDD0FF9">2023-07-10</mods:number>
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<treatment id="03CA879AFFAEFFFFFBC4FD09FF4DFD15" LSID="urn:lsid:plazi:treatment:03CA879AFFAEFFFFFBC4FD09FF4DFD15" httpUri="http://treatment.plazi.org/id/03CA879AFFAEFFFFFBC4FD09FF4DFD15" lastPageId="11" lastPageNumber="291" pageId="10" pageNumber="290">
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<subSubSection id="C3796507FFAEFFFEFBC4FD09FB71FD79" box="[1024,1245,642,669]" pageId="10" pageNumber="290" type="nomenclature">
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<paragraph id="8BDC368CFFAEFFFEFBC4FD09FB71FD79" blockId="10.[810,1459,642,737]" box="[1024,1245,642,669]" pageId="10" pageNumber="290">
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<emphasis id="B917EA9EFFAEFFFEFBC4FD09FB71FD79" bold="true" box="[1024,1245,642,669]" pageId="10" pageNumber="290">
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<taxonomicName id="4C634D0FFFAEFFFEFBC4FD09FBD1FD79" authority="Alonso & Terán & Alanís & Calviño & Montes & García & Barneche & Almirón & Ciotek & Giorgis & Cascioưa, 2023" authorityName="Alonso & Terán & Alanís & Calviño & Montes & García & Barneche & Almirón & Ciotek & Giorgis & Cascioưa" authorityYear="2023" box="[1024,1149,642,669]" family="Rivulidae" genus="Titanolebias" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="290" phylum="Chordata" rank="genus" status="gen. nov.">
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<emphasis id="B917EA9EFFAEFFFEFBC4FD09FBD1FD79" bold="true" box="[1024,1149,642,669]" italics="true" pageId="10" pageNumber="290">Titanolebias</emphasis>
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<taxonomicNameLabel id="A22457E5FFAEFFFEFB45FD08FB71FD79" box="[1153,1245,643,669]" pageId="10" pageNumber="290" rank="genus">gen. nov.</taxonomicNameLabel>
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<subSubSection id="C3796507FFAEFFFEFCEEFD22FC67FD05" pageId="10" pageNumber="290" type="description">
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LSID
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<paragraph id="8BDC368CFFAEFFFEFCEEFC88FAE8FCFE" blockId="10.[810,1348,770,794]" box="[810,1348,770,794]" pageId="10" pageNumber="290">
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<emphasis id="B917EA9EFFAEFFFEFCEEFC88FC0AFCFE" box="[810,934,771,794]" italics="true" pageId="10" pageNumber="290">Type species:</emphasis>
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<taxonomicName id="4C634D0FFFAEFFFEFC6FFC89FAECFCFE" authority="Steindachner, 1881" authorityName="Steindachner" authorityYear="1881" box="[939,1344,770,794]" family="Rivulidae" genus="Cynolebias" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="290" phylum="Chordata" rank="species" species="elongatus">
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<emphasis id="B917EA9EFFAEFFFEFC6FFC89FBD8FCFE" box="[939,1140,770,794]" italics="true" pageId="10" pageNumber="290">Cynolebias elongatus</emphasis>
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Steindachner, 1881
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</taxonomicName>
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.
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<subSubSection id="C3796507FFAEFFFEFCEEFCB7FAD9FA25" pageId="10" pageNumber="290" type="diagnosis">
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<paragraph id="8BDC368CFFAEFFFEFCEEFCB7FB70FBA9" blockId="10.[810,1459,827,1102]" pageId="10" pageNumber="290">
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<emphasis id="B917EA9EFFAEFFFEFCEEFCB7FC3CFCB7" box="[810,912,828,851]" italics="true" pageId="10" pageNumber="290">Diagnosis:</emphasis>
|
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The new genus
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<taxonomicName id="4C634D0FFFAEFFFEFB88FCB0FB68FCB7" authorityName="Alonso & Terán & Alanís & Calviño & Montes & García & Barneche & Almirón & Ciotek & Giorgis & Cascioưa" authorityYear="2023" box="[1100,1220,827,851]" family="Rivulidae" genus="Titanolebias" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="290" phylum="Chordata" rank="genus">
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<emphasis id="B917EA9EFFAEFFFEFB88FCB0FB68FCB7" box="[1100,1220,827,851]" italics="true" pageId="10" pageNumber="290">Titanolebias</emphasis>
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</taxonomicName>
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differs from all other genera of the
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<taxonomicName id="4C634D0FFFAEFFFEFC7CFCD0FB83FC97" authorityName="Costa" authorityYear="1998" box="[952,1071,859,883]" family="Rivulidae" genus="Austrolebias" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="290" phylum="Chordata" rank="genus">
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<emphasis id="B917EA9EFFAEFFFEFC7CFCD0FB83FC97" box="[952,1071,859,883]" italics="true" pageId="10" pageNumber="290">Austrolebias</emphasis>
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</taxonomicName>
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genus group by the following unique combination of characters: (1) contact organs on most pectoral fin rays, in males; (2)> 40 scales on the lateral line; (3) small size of scales; (4) centre of most scales on ventral half of flank, above anal fin, in dominant mature males (not considering bands, bars and iridescent lines) pastel yellow to golden; (5) contact organs on the anal fin of males; and (6) absence of scales in the anterior portion adjacent to the rostral neuromasts.
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</paragraph>
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<paragraph id="8BDC368CFFAEFFFEFCEEFBE4FAD9FA25" blockId="10.[810,1459,1135,1473]" pageId="10" pageNumber="290">
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<emphasis id="B917EA9EFFAEFFFEFCEEFBE4FC53FB63" box="[810,1023,1135,1159]" italics="true" pageId="10" pageNumber="290">Differential diagnosis:</emphasis>
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||
In addition to the unique combination of character states mentioned in the
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<emphasis id="B917EA9EFFAEFFFEFB5CFB04FB5BFB42" box="[1176,1271,1167,1190]" italics="true" pageId="10" pageNumber="290">Diagnosis</emphasis>
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||
above, the genus
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||
<taxonomicName id="4C634D0FFFAEFFFEFCEEFB25FC0EFB22" authorityName="Alonso & Terán & Alanís & Calviño & Montes & García & Barneche & Almirón & Ciotek & Giorgis & Cascioưa" authorityYear="2023" box="[810,930,1198,1222]" family="Rivulidae" genus="Titanolebias" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="290" phylum="Chordata" rank="genus">
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<emphasis id="B917EA9EFFAEFFFEFCEEFB25FC0EFB22" box="[810,930,1198,1222]" italics="true" pageId="10" pageNumber="290">Titanolebias</emphasis>
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</taxonomicName>
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can be distinguished from all other genera of the
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||
<taxonomicName id="4C634D0FFFAEFFFEFC91FB46FC60FB01" authorityName="Costa" authorityYear="1998" box="[853,972,1229,1253]" family="Rivulidae" genus="Austrolebias" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="290" phylum="Chordata" rank="genus">
|
||
<emphasis id="B917EA9EFFAEFFFEFC91FB46FC60FB01" box="[853,972,1229,1253]" italics="true" pageId="10" pageNumber="290">Austrolebias</emphasis>
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</taxonomicName>
|
||
genus group by having a posteriorly directed suborbital bar and the anterior region of the head, near the rostral neuromasts, without scales (vs. directed vertically, anteriorly or absent and the anterior region of the head, near rostral neuromasts, scaled in
|
||
<taxonomicName id="4C634D0FFFAEFFFEFB80FAC0FB05FA87" authorityName="sensu Costa" authorityYear="2008" box="[1092,1193,1355,1379]" family="Rivulidae" genus="Argolebias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="290" phylum="Chordata" rank="genus">
|
||
<emphasis id="B917EA9EFFAEFFFEFB80FAC0FB05FA87" box="[1092,1193,1355,1379]" italics="true" pageId="10" pageNumber="290">Argolebias</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C634D0FFFAEFFFEFB7DFAC0FAB3FA87" authorityName="Alonso & Terán & Alanís & Calviño & Montes & García & Barneche & Almirón & Ciotek & Giorgis & Cascioưa" authorityYear="2023" box="[1209,1311,1355,1379]" family="Rivulidae" genus="Matilebias" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="290" phylum="Chordata" rank="genus">
|
||
<emphasis id="B917EA9EFFAEFFFEFB7DFAC0FAB3FA87" box="[1209,1311,1355,1379]" italics="true" pageId="10" pageNumber="290">Matilebias</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C634D0FFFAEFFFEFAEBFAC0FA01FA87" authorityName="Costa" authorityYear="2008" box="[1327,1453,1355,1379]" family="Rivulidae" genus="Gymnolebias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="290" phylum="Chordata" rank="genus">
|
||
<emphasis id="B917EA9EFFAEFFFEFAEBFAC0FA01FA87" box="[1327,1453,1355,1379]" italics="true" pageId="10" pageNumber="290">Gymnolebias</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C634D0FFFAEFFFEFCEEFAE1FC1EFA66" authorityName="Costa" authorityYear="2008" box="[810,946,1386,1410]" family="Rivulidae" genus="Acantholebias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="290" phylum="Chordata" rank="genus">
|
||
<emphasis id="B917EA9EFFAEFFFEFCEEFAE1FC1EFA66" box="[810,946,1386,1410]" italics="true" pageId="10" pageNumber="290">Acantholebias</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C634D0FFFAEFFFEFC04FAE1FB9AFA66" authorityName="Costa" authorityYear="2008" box="[960,1078,1386,1410]" family="Rivulidae" genus="Cypholebias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="290" phylum="Chordata" rank="genus">
|
||
<emphasis id="B917EA9EFFAEFFFEFC04FAE1FB9AFA66" box="[960,1078,1386,1410]" italics="true" pageId="10" pageNumber="290">Cypholebias</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C634D0FFFAEFFFEFB80FAE1FB04FA66" authorityName="Costa" authorityYear="2008" box="[1092,1192,1386,1410]" family="Rivulidae" genus="Acrolebias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="290" phylum="Chordata" rank="genus">
|
||
<emphasis id="B917EA9EFFAEFFFEFB80FAE1FB04FA66" box="[1092,1192,1386,1410]" italics="true" pageId="10" pageNumber="290">Acrolebias</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C634D0FFFAEFFFEFB71FAE1FA82FA66" authorityName="Alonso & Terán & Alanís & Calviño & Montes & García & Barneche & Almirón & Ciotek & Giorgis & Cascioưa" authorityYear="2023" box="[1205,1326,1386,1410]" family="Rivulidae" genus="Garcialebias" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="290" phylum="Chordata" rank="genus">
|
||
<emphasis id="B917EA9EFFAEFFFEFB71FAE1FA82FA66" box="[1205,1326,1386,1410]" italics="true" pageId="10" pageNumber="290">Garcialebias</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C634D0FFFAEFFFEFAF8FAE1FA1FFA66" authorityName="Costa" authorityYear="1998" box="[1340,1459,1386,1410]" family="Rivulidae" genus="Austrolebias" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="290" phylum="Chordata" rank="genus">
|
||
<emphasis id="B917EA9EFFAEFFFEFAF8FAE1FA1FFA66" box="[1340,1459,1386,1410]" italics="true" pageId="10" pageNumber="290">Austrolebias</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C634D0FFFAEFFFEFC93FA02FC7CFA45" authorityName="Alonso & Terán & Alanís & Calviño & Montes & García & Barneche & Almirón & Ciotek & Giorgis & Cascioưa" authorityYear="2023" box="[855,976,1417,1441]" family="Rivulidae" genus="Amatolebias" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="290" phylum="Chordata" rank="genus">
|
||
<emphasis id="B917EA9EFFAEFFFEFC93FA02FC7CFA45" box="[855,976,1417,1441]" italics="true" pageId="10" pageNumber="290">Amatolebias</emphasis>
|
||
</taxonomicName>
|
||
); and by the presence of contact organs in the anal fin of males (vs. contact organs absent in
|
||
<taxonomicName id="4C634D0FFFAEFFFEFB3CFA22FAC8FA25" authorityName="Costa" authorityYear="1998" box="[1272,1380,1449,1473]" class="Actinopterygii" family="Rivulidae" genus="Megalebias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="290" phylum="Chordata" rank="genus">
|
||
<emphasis id="B917EA9EFFAEFFFEFB3CFA22FAC8FA25" box="[1272,1380,1449,1473]" italics="true" pageId="10" pageNumber="290">Megalebias</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3796507FFAEFFFEFCEEFA69FBB7F8B7" pageId="10" pageNumber="290" type="discussion">
|
||
<paragraph id="8BDC368CFFAEFFFEFCEEFA69FBB7F8B7" blockId="10.[810,1459,1506,1875]" pageId="10" pageNumber="290">
|
||
<emphasis id="B917EA9EFFAEFFFEFCEEFA69FC7EFA1E" box="[810,978,1506,1530]" italics="true" pageId="10" pageNumber="290">Synapomorphies:</emphasis>
|
||
The new genus
|
||
<taxonomicName id="4C634D0FFFAEFFFEFBB9FA69FB59FA1E" authorityName="Alonso & Terán & Alanís & Calviño & Montes & García & Barneche & Almirón & Ciotek & Giorgis & Cascioưa" authorityYear="2023" box="[1149,1269,1506,1530]" family="Rivulidae" genus="Titanolebias" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="290" phylum="Chordata" rank="genus">
|
||
<emphasis id="B917EA9EFFAEFFFEFBB9FA69FB59FA1E" box="[1149,1269,1506,1530]" italics="true" pageId="10" pageNumber="290">Titanolebias</emphasis>
|
||
</taxonomicName>
|
||
was supported by 30 molecular character states and the following seven morphological character states (see Supporting Information, Appendix S5): centre of most scales on ventral half of flank, above anal fin, in dominant mature males (not considering bands, bars and iridescent lines) pastel yellow to golden (1:2); centre of most scales on dorsal half of flank, below dorsal fin, in dominant mature males (not considering bands, bars and iridescent lines) light grey (6:6); grey spots on head present (23:1);> 40 scales on lateral line (69:2); small scales (70:1); contact organs on anal fin of males present (86:1); and contact organs on most pectoral fin rays of males (91:1).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3796507FFAEFFFFFCEEF8FFFE63FF23" lastPageId="11" lastPageNumber="291" pageId="10" pageNumber="290" type="taxon_list">
|
||
<paragraph id="8BDC368CFFAEFFFFFCEEF8FFFE63FF23" blockId="10.[810,1459,1908,1995]" lastBlockId="11.[129,777,144,199]" lastPageId="11" lastPageNumber="291" pageId="10" pageNumber="290">
|
||
<emphasis id="B917EA9EFFAEFFFEFCEEF8FFFABEF868" box="[810,1298,1908,1932]" italics="true" pageId="10" pageNumber="290">
|
||
Included species:
|
||
<taxonomicName id="4C634D0FFFAEFFFEFBC4F8FFFABEF868" box="[1024,1298,1908,1932]" family="Rivulidae" genus="Titanolebias" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="290" phylum="Chordata" rank="species" species="cheradophilus">Titanolebias cheradophilus</taxonomicName>
|
||
</emphasis>
|
||
(Vaz-Ferreira, Sierra de
|
||
<collectingRegion id="49A7F86EFFAEFFFEFC5CF818FC44F84F" box="[920,1000,1939,1963]" country="Uruguay" name="Soriano" pageId="10" pageNumber="290">Soriano</collectingRegion>
|
||
& Scaglia de Paulete, 1965) comb. nov.,
|
||
<taxonomicName id="4C634D0FFFAEFFFEFCEEF839FB14F82E" authority="(Amato, 1986)" authorityName="Alonso & Terán & Alanís & Calviño & Montes & García & Barneche & Almirón & Ciotek & Giorgis & Cascioưa" authorityYear="2023" baseAuthorityName="Amato" baseAuthorityYear="1986" box="[810,1208,1970,1995]" family="Rivulidae" genus="Titanolebias" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="290" phylum="Chordata" rank="species" species="prognathus" status="comb. nov.">
|
||
<emphasis id="B917EA9EFFAEFFFEFCEEF839FBBAF82E" box="[810,1046,1970,1994]" italics="true" pageId="10" pageNumber="290">Titanolebias prognathus</emphasis>
|
||
(Amato, 1986)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22457E5FFAEFFFEFB7BF839FA9DF82E" box="[1215,1329,1970,1994]" pageId="10" pageNumber="290" rank="species">comb. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C634D0FFFAEFFFFFAFFF839FE26FF4C" authority="(Huber, 1995)" authorityName="Alonso & Terán & Alanís & Calviño & Montes & García & Barneche & Almirón & Ciotek & Giorgis & Cascioưa" authorityYear="2023" baseAuthorityName="Huber" baseAuthorityYear="1995" family="Rivulidae" genus="Titanolebias" kingdom="Animalia" lastPageId="11" lastPageNumber="291" order="Cyprinodontiformes" pageId="10" pageNumber="290" phylum="Chordata" rank="species" species="monstrosus" status="comb. nov.">
|
||
<emphasis id="B917EA9EFFAEFFFFFAFFF839FF41FF4C" italics="true" lastPageId="11" lastPageNumber="291" pageId="10" pageNumber="290">Titanolebias monstrosus</emphasis>
|
||
(Huber, 1995)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22457E5FFAFFFFFFE4BFF1BFDACFF43" box="[399,512,144,168]" pageId="11" pageNumber="291" rank="species">comb. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C634D0FFFAFFFFFFDF5FF1BFEF7FF23" authority="(Steindachner, 1881)" authorityName="Alonso & Terán & Alanís & Calviño & Montes & García & Barneche & Almirón & Ciotek & Giorgis & Cascioưa" authorityYear="2023" baseAuthorityName="Steindachner" baseAuthorityYear="1881" family="Rivulidae" genus="Titanolebias" kingdom="Animalia" order="Cyprinodontiformes" pageId="11" pageNumber="291" phylum="Chordata" rank="species" species="elongatus" status="comb. nov.">
|
||
<emphasis id="B917EA9EFFAFFFFFFDF5FF1BFCA5FF4C" box="[561,777,144,168]" italics="true" pageId="11" pageNumber="291">Titanolebias elongatus</emphasis>
|
||
(Steindachner, 1881)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22457E5FFAFFFFFFEA4FF24FE63FF23" box="[352,463,175,199]" pageId="11" pageNumber="291" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3796507FFAFFFFFFF45FF62FD31FDDE" pageId="11" pageNumber="291" type="etymology">
|
||
<paragraph id="8BDC368CFFAFFFFFFF45FF62FD31FDDE" blockId="11.[129,778,233,570]" pageId="11" pageNumber="291">
|
||
<emphasis id="B917EA9EFFAFFFFFFF45FF62FF5CFEE5" box="[129,240,233,257]" italics="true" pageId="11" pageNumber="291">Etymology:</emphasis>
|
||
The name of the genus is composed of
|
||
<taxonomicName id="4C634D0FFFAFFFFFFD60FF62FD76FEE4" box="[676,730,233,256]" class="Insecta" family="Ptiliidae" genus="Titan" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="291" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B917EA9EFFAFFFFFFD60FF62FD76FEE4" box="[676,730,233,256]" italics="true" pageId="11" pageNumber="291">Titan</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B917EA9EFFAFFFFFFF45FE83FF1BFEC4" box="[129,183,264,288]" italics="true" pageId="11" pageNumber="291">lebias</emphasis>
|
||
.
|
||
<emphasis id="B917EA9EFFAFFFFFFF03FE83FEAAFEC4" box="[199,262,264,288]" italics="true" pageId="11" pageNumber="291">Lebias</emphasis>
|
||
is from the Greek, meaning ‘small fish’, a name commonly used to compose generic names of cyprinodontiform fishes. In Greek mythology, the Titans (Greek: Tιτᾶνες, Titânes, singular: Tιτάν, -ήν,
|
||
<taxonomicName id="4C634D0FFFAFFFFFFE9BFEEDFE35FE9A" box="[351,409,358,382]" class="Insecta" family="Ptiliidae" genus="Titan" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="291" phylum="Arthropoda" rank="genus">Titán</taxonomicName>
|
||
) were the pre-Olympian gods; they were the 12 children of the primordial parents Uranus (Sky) and Gaia (Earth) and famous for their strength and huge size. Named in allusion to the relative size of the fishes of this genus within Aplocheiloidei, bearing the largest species in this suborder of fishes, with
|
||
<taxonomicName id="4C634D0FFFAFFFFFFF33FD88FEC5FDFF" authorityName="Alonso & Terán & Alanís & Calviño & Montes & García & Barneche & Almirón & Ciotek & Giorgis & Cascioưa" authorityYear="2023" baseAuthorityName="Steindachner" baseAuthorityYear="1881" box="[247,361,515,539]" family="Rivulidae" genus="Titanolebias" kingdom="Animalia" order="Cyprinodontiformes" pageId="11" pageNumber="291" phylum="Chordata" rank="species" species="elongatus">
|
||
<emphasis id="B917EA9EFFAFFFFFFF33FD88FEC5FDFF" box="[247,361,515,539]" italics="true" pageId="11" pageNumber="291">T. elongatus</emphasis>
|
||
</taxonomicName>
|
||
reaching
|
||
<quantity id="4C9B9B69FFAFFFFFFE0EFD89FDB0FDFE" box="[458,540,514,538]" metricMagnitude="-1" metricUnit="m" metricValue="1.55" pageId="11" pageNumber="291" unit="cm" value="15.5">15.5 cm</quantity>
|
||
SL (MACN 3826) and
|
||
<taxonomicName id="4C634D0FFFAFFFFFFF45FDA8FEA9FDDE" authorityName="Alonso & Terán & Alanís & Calviño & Montes & García & Barneche & Almirón & Ciotek & Giorgis & Cascioưa" authorityYear="2023" baseAuthorityName="Huber" baseAuthorityYear="1995" box="[129,261,547,570]" family="Rivulidae" genus="Titanolebias" kingdom="Animalia" order="Cyprinodontiformes" pageId="11" pageNumber="291" phylum="Chordata" rank="species" species="monstrosus">
|
||
<emphasis id="B917EA9EFFAFFFFFFF45FDA8FEA9FDDE" box="[129,261,547,570]" italics="true" pageId="11" pageNumber="291">T. monstrosus</emphasis>
|
||
</taxonomicName>
|
||
<quantity id="4C9B9B69FFAFFFFFFECEFDA9FEE5FDDD" box="[266,329,546,570]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="11" pageNumber="291" unit="cm" value="15.0">15 cm</quantity>
|
||
(MNK P-3301) (
|
||
<bibRefCitation id="EFF24B7DFFAFFFFFFDC4FDA9FD20FDDE" author="Osinaga K" box="[512,652,546,570]" pageId="11" pageNumber="291" pagination="60 - 2" refId="ref25029" refString="Osinaga K. Nuevo registro para Bolivia de Austrolebias monstrosus (Huber, 1995 Rivulidae). Kempffiana 2006; 2: 60 - 2." type="journal article" year="2006">Osinaga 2006</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3796507FFAFFFFFFF45FDD0FF4DFD15" pageId="11" pageNumber="291" type="distribution">
|
||
<paragraph id="8BDC368CFFAFFFFFFF45FDD0FF4DFD15" blockId="11.[129,778,603,753]" pageId="11" pageNumber="291">
|
||
<emphasis id="B917EA9EFFAFFFFFFF45FDD0FF52FD97" box="[129,254,603,627]" italics="true" pageId="11" pageNumber="291">Distribution:</emphasis>
|
||
Río de la Plata basin (excluding the middle and upper portions of the Paraná and
|
||
<collectingCountry id="F374761CFFAFFFFFFE39FDF0FDFAFD77" box="[509,598,635,659]" name="Uruguay" pageId="11" pageNumber="291">Uruguay</collectingCountry>
|
||
basins), Laguna Merín basin, Atlantic basins of eastern
|
||
<collectingCountry id="F374761CFFAFFFFFFDC8FD11FDC9FD56" box="[524,613,666,690]" name="Uruguay" pageId="11" pageNumber="291">Uruguay</collectingCountry>
|
||
and headwaters of the Mamoré basin, in
|
||
<collectingCountry id="F374761CFFAFFFFFFEBFFD32FE4FFD35" box="[379,483,697,721]" name="Argentina" pageId="11" pageNumber="291">Argentina</collectingCountry>
|
||
,
|
||
<collectingCountry id="F374761CFFAFFFFFFE28FD31FDE9FD36" box="[492,581,698,722]" name="Uruguay" pageId="11" pageNumber="291">Uruguay</collectingCountry>
|
||
,
|
||
<collectingCountry id="F374761CFFAFFFFFFD8BFD32FD20FD35" box="[591,652,697,721]" name="Brazil" pageId="11" pageNumber="291">Brazil</collectingCountry>
|
||
,
|
||
<collectingCountry id="F374761CFFAFFFFFFD51FD32FD71FD35" box="[661,733,697,721]" name="Bolivia" pageId="11" pageNumber="291">Bolivia</collectingCountry>
|
||
and
|
||
<collectingCountry id="F374761CFFAFFFFFFF45FD52FF72FD15" box="[129,222,729,753]" name="Paraguay" pageId="11" pageNumber="291">Paraguay</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |