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<document id="146852DFB9ED5B2E4532B3736CBAC6E9" ID-DOI="10.11646/zootaxa.4810.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="3936945" ID-ZooBank="BBD6D565-039A-47C7-9110-653DD75EBDBF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1594292067076" checkinUser="plazi" docAuthor="Ariyama, Hiroyuki" docDate="2020" docId="0387413C4C7AFFB9FF2E62083474FE4F" docLanguage="en" docName="zootaxa.4810.1.1.pdf" docOrigin="Zootaxa 4810 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Grandidierella contigua Ariyama 2020, sp. nov." docType="treatment" docVersion="4" lastPageNumber="10" masterDocId="FFBE39444C78FFB0FFB9654C3061FFC3" masterDocTitle="Six species of Grandidierella collected from the Ryukyu Archipelago in Japan with descriptions of four new species (Crustacea: Amphipoda: Aoridae)" masterLastPageNumber="44" masterPageNumber="1" pageNumber="3" updateTime="1698779345914" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="21EBA343563C9C49E3C220647295308C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="598BEE434E81F8074595DCD1E3E7C8BF">
<mods:title id="4EE35D29E7498D5FE55EFCF7C088A950">Six species of Grandidierella collected from the Ryukyu Archipelago in Japan with descriptions of four new species (Crustacea: Amphipoda: Aoridae)</mods:title>
</mods:titleInfo>
<mods:name id="9C440639AC9729B6C1662AA91EB08A61" type="personal">
<mods:role id="045A611C51F2FCBBD378442776D4CEA3">
<mods:roleTerm id="3EE4AC2B166D7B1A192F88272EC6AC26">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2A4440556B00BF286A0018085F1E5119">Ariyama, Hiroyuki</mods:namePart>
</mods:name>
<mods:typeOfResource id="BD18F9CB3CECB8E6CDCA0565652FC9DD">text</mods:typeOfResource>
<mods:relatedItem id="9C23CBB711118901E3E2D4711326DC47" type="host">
<mods:titleInfo id="AA284D8B105874B32FD958D937000205">
<mods:title id="409590CD40A6FA9ACE6A1713C07D2A68">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="010ED65868FAB2FB2A5CE0500548F222">
<mods:date id="5936CA828AA5D003C641338C81E68DA4">2020</mods:date>
<mods:detail id="E35CC79B6CBF44F015570A60FD34B939" type="pubDate">
<mods:number id="C7CC3EAC7250D53CAAFF1C8B237BA003">2020-07-09</mods:number>
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<mods:detail id="23FCCE7FD6C78F3DBD8A516E8569D174" type="volume">
<mods:number id="45D3C14977B47FC1164D0AE466780AFB">4810</mods:number>
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<mods:classification id="E234E6BBB5DF7829A54E4D099D09FE92">journal article</mods:classification>
<mods:identifier id="A19BF86EE612DF5E2A96DACE80ACC966" type="DOI">10.11646/zootaxa.4810.1.1</mods:identifier>
<mods:identifier id="54BBF035BC4B8D430B0F62C531972822" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="604E689B4E4C9563209EDDA997FA6CD5" type="Zenodo-Dep">3936945</mods:identifier>
<mods:identifier id="FA15D5043ACAF7A3090BF301917DBEF7" type="ZooBank">BBD6D565-039A-47C7-9110-653DD75EBDBF</mods:identifier>
</mods:mods>
<treatment id="0387413C4C7AFFB9FF2E62083474FE4F" LSID="urn:lsid:plazi:treatment:0387413C4C7AFFB9FF2E62083474FE4F" httpUri="http://treatment.plazi.org/id/0387413C4C7AFFB9FF2E62083474FE4F" lastPageId="9" lastPageNumber="10" pageId="2" pageNumber="3">
<subSubSection id="C334A3A14C7AFFB2FF2E620832F0F841" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="8B91F02A4C7AFFB2FF2E6208327EF89D" blockId="2.[151,657,1860,1958]" box="[151,543,1860,1887]" pageId="2" pageNumber="3">
<heading id="D0D947464C7AFFB2FF2E6208327EF89D" bold="true" box="[151,543,1860,1887]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
<emphasis id="B95A2C384C7AFFB2FF2E6208327EF89D" bold="true" box="[151,543,1860,1887]" pageId="2" pageNumber="3">
<taxonomicName id="4C2E8BA94C7AFFB2FF2E620831DBF89D" ID-CoL="7ZKDG" authority="Ariyama, 2020" authorityName="Ariyama" authorityYear="2020" box="[151,442,1860,1887]" class="Malacostraca" family="Aoridae" genus="Grandidierella" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="contigua" status="sp. nov.">
<emphasis id="B95A2C384C7AFFB2FF2E620831DBF89D" bold="true" box="[151,442,1860,1887]" italics="true" pageId="2" pageNumber="3">Grandidierella contigua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26991434C7AFFB2FE786208327EF89D" box="[449,543,1860,1886]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B91F02A4C7AFFB2FF2E622532F0F841" blockId="2.[151,657,1860,1958]" box="[151,657,1895,1923]" pageId="2" pageNumber="3">[Japanese name: Rinsetsu-dorosokoebi, new]</paragraph>
</subSubSection>
<subSubSection id="C334A3A14C7AFFB2FF2E62C0316AF865" box="[151,267,1932,1958]" pageId="2" pageNumber="3" type="description">
<paragraph id="8B91F02A4C7AFFB2FF2E62C0316AF865" blockId="2.[151,657,1860,1958]" box="[151,267,1932,1958]" pageId="2" pageNumber="3">
(
<figureCitation id="1315ECAF4C7AFFB2FF2662C03165F865" box="[159,260,1932,1958]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6" captionStart-5="FIGURE 7" captionStartId-0="4.[151,250,1630,1655]" captionStartId-1="5.[151,250,1994,2019]" captionStartId-2="6.[151,250,1991,2016]" captionStartId-3="7.[151,250,1945,1970]" captionStartId-4="8.[151,250,1684,1709]" captionStartId-5="9.[151,250,926,951]" captionTargetBox-0="[165,1426,607,1597]" captionTargetBox-1="[197,1392,193,1949]" captionTargetBox-2="[188,1396,193,1955]" captionTargetBox-3="[204,1384,195,1906]" captionTargetBox-4="[151,1436,885,1660]" captionTargetBox-5="[151,1437,417,901]" captionTargetId-0="figure@4.[151,1436,597,1605]" captionTargetId-1="figure@5.[175,1413,181,1969]" captionTargetId-2="figure@6.[177,1411,181,1965]" captionTargetId-3="figure@7.[195,1393,181,1919]" captionTargetId-4="figure@8.[151,1436,885,1660]" captionTargetId-5="figure@9.[151,1437,417,901]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="8" captionTargetPageId-5="9" captionText-0="FIGURE 2. Grandidierella contigua sp. nov. Holotype, male, 9.1 mm (OMNH-Ar-11513). Habitus." captionText-1="FIGURE 3. Grandidierella contigua sp. nov. Holotype, male, 9.1 mm (OMNH-Ar-11513). Scales: 0.1 mm." captionText-2="FIGURE 4. Grandidierella contigua sp. nov. Holotype, male, 9.1 mm (OMNH-Ar-11513). Scales: 0.2 mm." captionText-3="FIGURE 5. Grandidierella contigua sp. nov. All except for F, M1 and M2: holotype, male, 9.1 mm (OMNH-Ar-11513); F, paratype, ovigerous female, 8.4 mm (OMNH-Ar-11514); M1, paratype, male, 6.7 mm (OMNH-Ar-11516); M2, paratype, male, ca. 3.7 mm (OMNH-Ar-11518). Scales: 0.2 mm." captionText-4="FIGURE 6. Grandidierella contigua sp. nov. Photograph of paratype, male, 3.5 mm (OMNH-Ar-11519), fixed specimen (6.5 hours after fixation)." captionText-5="FIGURE 7. Grandidierella contigua sp. nov. Photographs of holotype, male, 9.1 mm (OMNH-Ar-11513, left) and paratype, ovigerous female, 8.4 mm (OMNH-Ar-11514, right), preserved specimens." figureDoi-0="http://doi.org/10.5281/zenodo.3936949" figureDoi-1="http://doi.org/10.5281/zenodo.3936951" figureDoi-2="http://doi.org/10.5281/zenodo.3936953" figureDoi-3="http://doi.org/10.5281/zenodo.3936955" figureDoi-4="http://doi.org/10.5281/zenodo.3936957" figureDoi-5="http://doi.org/10.5281/zenodo.3936961" httpUri-0="https://zenodo.org/record/3936949/files/figure.png" httpUri-1="https://zenodo.org/record/3936951/files/figure.png" httpUri-2="https://zenodo.org/record/3936953/files/figure.png" httpUri-3="https://zenodo.org/record/3936955/files/figure.png" httpUri-4="https://zenodo.org/record/3936957/files/figure.png" httpUri-5="https://zenodo.org/record/3936961/files/figure.png" pageId="2" pageNumber="3">Figs 27</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C334A3A14C7AFFB3FF2E6298345CFD30" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="materials_examined">
<paragraph id="8B91F02A4C7AFFB2FF2E629835FDF82D" blockId="2.[151,1436,2003,2031]" box="[151,1436,2003,2031]" pageId="2" pageNumber="3">
<emphasis id="B95A2C384C7AFFB2FF2E62983127F82D" bold="true" box="[151,326,2003,2030]" pageId="2" pageNumber="3">Type material.</emphasis>
<materialsCitation id="3B46FA774C7AFFB2FEF7629835FDF82D" box="[334,1436,2003,2031]" location="Yakushima Island" municipality="Harutahama" pageId="2" pageNumber="3" specimenCount="1" stateProvince="Kagoshima" typeStatus="holotype">
<typeStatus id="54954E884C7AFFB2FEF7629831DAF82D" box="[334,443,2004,2030]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
: male,
<quantity id="4CD65DCF4C7AFFB2FDB262983203F82C" box="[523,610,2004,2031]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" pageId="2" pageNumber="3" unit="mm" value="9.1">9.1 mm</quantity>
(OMNH-Ar-11513),
<collectingMunicipality id="6BF56A504C7AFFB2FCEC62983380F82D" box="[853,993,2004,2030]" pageId="2" pageNumber="3">Harutahama</collectingMunicipality>
,
<location id="8EF1A6F14C7AFFB2FC55629834D9F82D" LSID="urn:lsid:plazi:treatment:0387413C4C7AFFB9FF2E62083474FE4F:8EF1A6F14C7AFFB2FC55629834D9F82D" box="[1004,1208,2004,2030]" municipality="Harutahama" name="Yakushima Island" pageId="2" pageNumber="3" stateProvince="Kagoshima">Yakushima Island</location>
,
<collectingRegion id="49EA3EC84C7AFFB2FB7D62983525F82D" box="[1220,1348,2004,2030]" country="Japan" name="Kagoshima" pageId="2" pageNumber="3">Kagoshima</collectingRegion>
Prefec-
</materialsCitation>
</paragraph>
<paragraph id="8B91F02A4C7BFFB3FF2E65D5313BFE6C" blockId="3.[151,1437,152,2015]" pageId="3" pageNumber="4">
ture,
<geoCoordinate id="EE1A96ED4C7BFFB3FF6965D4310DFF70" box="[208,364,152,179]" degrees="30" direction="north" minutes="18" orientation="latitude" pageId="3" pageNumber="4" precision="1" seconds="25.9" value="30.307194">30°18'25.9&quot;N</geoCoordinate>
,
<geoCoordinate id="EE1A96ED4C7BFFB3FECE65D5327BFF70" box="[375,538,153,179]" degrees="130" direction="east" minutes="39" orientation="longitude" pageId="3" pageNumber="4" precision="1" seconds="24.0" value="130.65666">130°39'24.0&quot;E</geoCoordinate>
(
<figureCitation id="1315ECAF4C7BFFB3FD9065D532E3FF70" box="[553,642,153,179]" captionStart="FIGURE 1" captionStartId="1.[151,250,1778,1803]" captionTargetBox="[151,1436,561,1752]" captionTargetId="figure@1.[151,1436,561,1752]" captionTargetPageId="1" captionText="FIGURE 1. Map showing collecting sites of the examined specimens in the Ryukyu Archipelago." figureDoi="http://doi.org/10.5281/zenodo.3936947" httpUri="https://zenodo.org/record/3936947/files/figure.png" pageId="3" pageNumber="4">Fig. 1A</figureCitation>
), coarse sand bottom in stream, brackish,
<date id="FF90D6EA4C7BFFB3FBEC65D53562FF70" box="[1109,1283,153,179]" pageId="3" pageNumber="4" value="2017-03-27">27 March 2017</date>
, coll. K. Kan.
<typeStatus id="54954E884C7BFFB3FF2E65F13169FF14" box="[151,264,189,215]" pageId="3" pageNumber="4" type="paratype">Paratypes</typeStatus>
:
<materialsCitation id="3B46FA774C7BFFB3FEAD65F135E0FEDC" collectingDate="2017-03-27" collectorName="K. Kan" latitude="30.303444" location="Yakushima Island" longLatPrecision="1" longitude="130.65463" municipality="Harutahama" pageId="3" pageNumber="4" specimenCount="2" specimenCount-male="2" stateProvince="Kagoshima">
ovigerous female,
<quantity id="4CD65DCF4C7BFFB3FE5F65F1325CFF14" box="[486,573,189,215]" metricMagnitude="-3" metricUnit="m" metricValue="8.4" pageId="3" pageNumber="4" unit="mm" value="8.4">8.4 mm</quantity>
(OMNH-Ar-11514), same data as holotype;
<specimenCount id="9D283BA34C7BFFB3FB8565F134F7FF14" box="[1084,1174,189,215]" pageId="3" pageNumber="4" type="male">2 males</specimenCount>
, 8.8,
<quantity id="4CD65DCF4C7BFFB3FB6C65F1354DFF14" box="[1237,1324,189,215]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="3" pageNumber="4" unit="mm" value="6.7">6.7 mm</quantity>
(OMNH- Ar-11515, 11516) and female,
<quantity id="4CD65DCF4C7BFFB3FE5465AC3223FF38" box="[493,578,224,251]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="3" pageNumber="4" unit="mm" value="6.5">6.5 mm</quantity>
(OMNH-Ar-11517),
<collectingMunicipality id="6BF56A504C7BFFB3FC8B65AD33DFFF38" box="[818,958,225,251]" pageId="3" pageNumber="4">Harutahama</collectingMunicipality>
,
<location id="8EF1A6F14C7BFFB3FC7E65AD34F0FF38" LSID="urn:lsid:plazi:treatment:0387413C4C7AFFB9FF2E62083474FE4F:8EF1A6F14C7BFFB3FC7E65AD34F0FF38" box="[967,1169,225,251]" latitude="30.303444" longLatPrecision="1" longitude="130.65463" municipality="Harutahama" name="Yakushima Island" pageId="3" pageNumber="4" stateProvince="Kagoshima">Yakushima Island</location>
,
<collectingRegion id="49EA3EC84C7BFFB3FB2565AD35F9FF38" box="[1180,1432,225,251]" country="Japan" name="Kagoshima" pageId="3" pageNumber="4">Kagoshima Prefecture</collectingRegion>
,
<geoCoordinate id="EE1A96ED4C7BFFB3FF2E64493152FEDC" box="[151,307,261,287]" degrees="30" direction="north" minutes="18" orientation="latitude" pageId="3" pageNumber="4" precision="1" seconds="12.4" value="30.303444">30°18'12.4&quot;N</geoCoordinate>
,
<geoCoordinate id="EE1A96ED4C7BFFB3FEF964493185FEDC" box="[320,484,261,287]" degrees="130" direction="east" minutes="39" orientation="longitude" pageId="3" pageNumber="4" precision="1" seconds="16.7" value="130.65463">130°39'16.7&quot;E</geoCoordinate>
(
<figureCitation id="1315ECAF4C7BFFB3FE4C6449322EFEDC" box="[501,591,261,287]" captionStart="FIGURE 1" captionStartId="1.[151,250,1778,1803]" captionTargetBox="[151,1436,561,1752]" captionTargetId="figure@1.[151,1436,561,1752]" captionTargetPageId="1" captionText="FIGURE 1. Map showing collecting sites of the examined specimens in the Ryukyu Archipelago." figureDoi="http://doi.org/10.5281/zenodo.3936947" httpUri="https://zenodo.org/record/3936947/files/figure.png" pageId="3" pageNumber="4">Fig. 1A</figureCitation>
), coarse sand bottom in stream, brackish,
<date id="FF90D6EA4C7BFFB3FB8A64493485FEDC" box="[1075,1252,261,287]" pageId="3" pageNumber="4" value="2017-03-27">
<collectingDate id="EFD42F024C7BFFB3FB8A64493485FEDC" box="[1075,1252,261,287]" pageId="3" pageNumber="4" value="2017-03-27">27 March 2017</collectingDate>
</date>
, coll.
<collectorName id="26DB95FC4C7BFFB3FA92644935E0FEDC" box="[1323,1409,261,287]" pageId="3" pageNumber="4">K. Kan</collectorName>
</materialsCitation>
;
<materialsCitation id="3B46FA774C7BFFB3FA376449313BFE6C" collectingDate="2015-07-16" collectorName="H. Ariyama." latitude="24.353056" location="Iriomote Island" longLatPrecision="21" longitude="123.753334" municipality="Nakara River" pageId="3" pageNumber="4" specimenCount="3" specimenCount-male="3" stateProvince="Okinawa">
<specimenCount id="9D283BA34C7BFFB3FA37644930BAFE80" pageId="3" pageNumber="4" type="male">3 males</specimenCount>
, ca. 3.7 (damaged), 3.5,
<quantity id="4CD65DCF4C7BFFB3FE4A64653228FE80" box="[499,585,297,323]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="3" pageNumber="4" unit="mm" value="2.9">2.9 mm</quantity>
(OMNH-Ar-1151811520), ovigerous female,
<quantity id="4CD65DCF4C7BFFB3FBD8646434D9FE80" box="[1121,1208,296,323]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="3" pageNumber="4" unit="mm" value="3.5">3.5 mm</quantity>
(OMNH-Ar-11521) and female,
<quantity id="4CD65DCF4C7BFFB3FEA664013115FEA4" box="[287,372,333,359]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="3" pageNumber="4" unit="mm" value="3.4">3.4 mm</quantity>
(OMNH-Ar-11522), mouth of
<collectingMunicipality id="6BF56A504C7BFFB3FD7764013304FEA4" box="[718,869,333,359]" pageId="3" pageNumber="4">Nakara River</collectingMunicipality>
,
<location id="8EF1A6F14C7BFFB3FCD66401347CFEA4" LSID="urn:lsid:plazi:treatment:0387413C4C7AFFB9FF2E62083474FE4F:8EF1A6F14C7BFFB3FCD66401347CFEA4" box="[879,1053,333,359]" latitude="24.353056" longLatPrecision="21" longitude="123.753334" municipality="Nakara River" name="Iriomote Island" pageId="3" pageNumber="4" stateProvince="Okinawa">Iriomote Island</location>
,
<collectingRegion id="49EA3EC84C7BFFB3FB9E64013569FEA4" box="[1063,1288,333,359]" country="Japan" name="Okinawa" pageId="3" pageNumber="4">Okinawa Prefecture</collectingRegion>
,
<geoCoordinate id="EE1A96ED4C7BFFB3FAAB640135F9FEA4" box="[1298,1432,333,359]" degrees="24" direction="north" minutes="21" orientation="latitude" pageId="3" pageNumber="4" precision="15" seconds="11" value="24.353056">24°21'11&quot;N</geoCoordinate>
,
<geoCoordinate id="EE1A96ED4C7BFFB3FF2E643C3147FE48" box="[151,294,368,395]" degrees="123" direction="east" minutes="45" orientation="longitude" pageId="3" pageNumber="4" precision="15" seconds="12" value="123.753334">123°45'12&quot;E</geoCoordinate>
(
<figureCitation id="1315ECAF4C7BFFB3FE8F643D31EFFE48" box="[310,398,369,395]" captionStart="FIGURE 1" captionStartId="1.[151,250,1778,1803]" captionTargetBox="[151,1436,561,1752]" captionTargetId="figure@1.[151,1436,561,1752]" captionTargetPageId="1" captionText="FIGURE 1. Map showing collecting sites of the examined specimens in the Ryukyu Archipelago." figureDoi="http://doi.org/10.5281/zenodo.3936947" httpUri="https://zenodo.org/record/3936947/files/figure.png" pageId="3" pageNumber="4">Fig. 1H</figureCitation>
), uppermiddle intertidal, sandy mud bottom under stones and soft sandstone,
<date id="FF90D6EA4C7BFFB3FABD643D35F9FE48" box="[1284,1432,368,395]" pageId="3" pageNumber="4" value="2015-07-16">
<collectingDate id="EFD42F024C7BFFB3FABD643D35F9FE48" box="[1284,1432,368,395]" pageId="3" pageNumber="4" value="2015-07-16">16 July 2015</collectingDate>
</date>
, coll.
<collectorName id="26DB95FC4C7BFFB3FF7664D9313BFE6C" box="[207,346,405,431]" pageId="3" pageNumber="4">H. Ariyama.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="8B91F02A4C7BFFB3FF7E64F4346EFD0C" blockId="3.[151,1437,152,2015]" pageId="3" pageNumber="4">
<emphasis id="B95A2C384C7BFFB3FF7E64F43381FE10" bold="true" box="[199,992,440,467]" pageId="3" pageNumber="4">
Material for comparison (
<taxonomicName id="4C2E8BA94C7BFFB3FDBF64F433B2FE10" authority="Ariyama, 1996" authorityName="Ariyama" authorityYear="1996" box="[518,979,440,467]" class="Malacostraca" family="Aoridae" genus="Grandidierella" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B95A2C384C7BFFB3FDBF64F43375FE10" bold="true" box="[518,788,440,467]" italics="true" pageId="3" pageNumber="4">Grandidierella fasciata</emphasis>
<bibRefCitation id="EFBF8DDB4C7BFFB3FCA464F433B2FE10" author="Ariyama, H." box="[797,979,440,467]" pageId="3" pageNumber="4" pagination="167 - 191" refId="ref16158" refString="Ariyama, H. (1996) Four species of the genus Grandidierella (Crustacea: Amphipoda: Aoridae) from Osaka Bay and the northern part of the Kii Channel, central Japan. Publications of the Seto Marine Biological Laboratory, 37, 167 - 191. https: // doi. org / 10.5134 / 176253" type="journal article" year="1996">Ariyama, 1996</bibRefCitation>
</taxonomicName>
).
</emphasis>
<materialsCitation id="3B46FA774C7BFFB3FC5364F533ECFDD8" collectingDate="1992-02-20" collectorName="H. Ariyama." latitude="34.316666" location="Misaki Town" longLatPrecision="1200" longitude="135.11667" municipality="Higashi River" pageId="3" pageNumber="4" specimenCount="1" stateProvince="Osaka" typeStatus="holotype">
<typeStatus id="54954E884C7BFFB3FC5364F53434FE10" box="[1002,1109,441,467]" pageId="3" pageNumber="4" type="holotype">Holotype</typeStatus>
, male,
<quantity id="4CD65DCF4C7BFFB3FB1264F53565FE10" box="[1195,1284,441,467]" metricMagnitude="-3" metricUnit="m" metricValue="8.4" pageId="3" pageNumber="4" unit="mm" value="8.4">8.4 mm</quantity>
(OMNH-Ar- 3840), mouth of
<collectingMunicipality id="6BF56A504C7BFFB3FEEF64913194FE34" box="[342,501,477,503]" pageId="3" pageNumber="4">Higashi River</collectingMunicipality>
,
<location id="8EF1A6F14C7BFFB3FDB8649132F9FE34" LSID="urn:lsid:plazi:treatment:0387413C4C7AFFB9FF2E62083474FE4F:8EF1A6F14C7BFFB3FDB8649132F9FE34" box="[513,664,477,503]" latitude="34.316666" longLatPrecision="1200" longitude="135.11667" municipality="Higashi River" name="Misaki Town" pageId="3" pageNumber="4" stateProvince="Osaka">Misaki Town</location>
,
<collectingRegion id="49EA3EC84C7BFFB3FD1C64913309FE34" box="[677,872,477,503]" country="Japan" name="Osaka" pageId="3" pageNumber="4">Osaka Prefecture</collectingRegion>
,
<geoCoordinate id="EE1A96ED4C7BFFB3FCCD649133B2FE34" box="[884,979,477,503]" degrees="34" direction="north" minutes="19" orientation="latitude" pageId="3" pageNumber="4" precision="925" value="34.316666">34°19'N</geoCoordinate>
,
<geoCoordinate id="EE1A96ED4C7BFFB3FC5964903428FE34" box="[992,1097,476,503]" degrees="135" direction="east" minutes="07" orientation="longitude" pageId="3" pageNumber="4" precision="925" value="135.11667">135°07'E</geoCoordinate>
, intertidal, sandy mud bottom mixed with gravel, brackish,
<date id="FF90D6EA4C7BFFB3FE51674D32D9FDD8" box="[488,696,513,539]" pageId="3" pageNumber="4" value="1992-02-20">
<collectingDate id="EFD42F024C7BFFB3FE51674D32D9FDD8" box="[488,696,513,539]" pageId="3" pageNumber="4" value="1992-02-20">20 February 1992</collectingDate>
</date>
, coll.
<collectorName id="26DB95FC4C7BFFB3FCB9674D33ECFDD8" box="[768,909,513,539]" pageId="3" pageNumber="4">H. Ariyama.</collectorName>
</materialsCitation>
<typeStatus id="54954E884C7BFFB3FC2D674D3398FDD8" box="[916,1017,513,539]" pageId="3" pageNumber="4" type="allotype">Allotype</typeStatus>
, female,
<quantity id="4CD65DCF4C7BFFB3FBDD674C34DDFDD8" box="[1124,1212,512,539]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="3" pageNumber="4" unit="mm" value="6.5">6.5 mm</quantity>
(OMNH-Ar-3841), same data as holotype.
<typeStatus id="54954E884C7BFFB3FE256769326CFDFC" box="[412,525,549,575]" pageId="3" pageNumber="4" type="paratype">Paratypes</typeStatus>
: male,
<quantity id="4CD65DCF4C7BFFB3FDE5676932D3FDFC" box="[604,690,549,575]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="3" pageNumber="4" unit="mm" value="7.4">7.4 mm</quantity>
(OMNH-Ar-3842) and female,
<quantity id="4CD65DCF4C7BFFB3FBA06769340EFDFC" box="[1049,1135,549,575]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="3" pageNumber="4" unit="mm" value="6.7">6.7 mm</quantity>
(OMNH-Ar-3843),
<date id="FF90D6EA4C7BFFB3FAED67693175FDA0" pageId="3" pageNumber="4" value="1989-02-06">6 February 1989</date>
, same place as
<typeStatus id="54954E884C7BFFB3FE0667053245FDA0" box="[447,548,585,611]" pageId="3" pageNumber="4" type="holotype">holotype</typeStatus>
, coll. H. Ariyama; male,
<quantity id="4CD65DCF4C7BFFB3FC82670533F1FDA0" box="[827,912,585,611]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="3" pageNumber="4" unit="mm" value="7.7">7.7 mm</quantity>
(OMNH-Ar-3844) and ovigerous female,
<quantity id="4CD65DCF4C7BFFB3FAD2670530A2FD44" metricMagnitude="-2" metricUnit="m" metricValue="1.01" pageId="3" pageNumber="4" unit="mm" value="10.1">10.1 mm</quantity>
(OMNH-Ar-3845),
<date id="FF90D6EA4C7BFFB3FE1E67213251FD44" box="[423,560,621,647]" pageId="3" pageNumber="4" value="1990-06-07">7 June 1990</date>
, same place as
<typeStatus id="54954E884C7BFFB3FD636721335EFD44" box="[730,831,621,647]" pageId="3" pageNumber="4" type="holotype">holotype</typeStatus>
, coll. H. Ariyama; male,
<quantity id="4CD65DCF4C7BFFB3FBEC672134CAFD44" box="[1109,1195,621,647]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="3" pageNumber="4" unit="mm" value="8.2">8.2 mm</quantity>
(OMNH-Ar-3846),
<date id="FF90D6EA4C7BFFB3FA376721316DFD68" pageId="3" pageNumber="4" value="1991-05-04">4 May 1991</date>
, same place as
<typeStatus id="54954E884C7BFFB3FE0567DD3240FD68" box="[444,545,657,683]" pageId="3" pageNumber="4" type="holotype">holotype</typeStatus>
, coll. H. Ariyama (
<bibRefCitation id="EFBF8DDB4C7BFFB3FD4167DD33C1FD6F" author="Ariyama, H." box="[760,928,657,684]" pageId="3" pageNumber="4" pagination="167 - 191" refId="ref16158" refString="Ariyama, H. (1996) Four species of the genus Grandidierella (Crustacea: Amphipoda: Aoridae) from Osaka Bay and the northern part of the Kii Channel, central Japan. Publications of the Seto Marine Biological Laboratory, 37, 167 - 191. https: // doi. org / 10.5134 / 176253" type="journal article" year="1996">Ariyama 1996</bibRefCitation>
).
<materialsCitation id="3B46FA774C7BFFB3FC0A67DD346AFD0C" collectingDate="1992-02-20" collectorName="H. Ariyama." latitude="34.316666" location="Three" longLatPrecision="1200" longitude="135.11667" municipality="Higashi River" pageId="3" pageNumber="4" specimenCount="1" stateProvince="Osaka" typeStatus="paratype">
<location id="8EF1A6F14C7BFFB3FC0A67DD3395FD68" LSID="urn:lsid:plazi:treatment:0387413C4C7AFFB9FF2E62083474FE4F:8EF1A6F14C7BFFB3FC0A67DD3395FD68" box="[947,1012,657,683]" name="Three" pageId="3" pageNumber="4">Three</location>
males, 5.3, 4.6,
<quantity id="4CD65DCF4C7BFFB3FB0967DD3567FD68" box="[1200,1286,657,683]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="3" pageNumber="4" unit="mm" value="4.1">4.1 mm</quantity>
(OMNH-Ar- 1152311525) and female,
<quantity id="4CD65DCF4C7BFFB3FE7067F8327EFD0C" box="[457,543,692,719]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="3" pageNumber="4" unit="mm" value="5.1">5.1 mm</quantity>
(OMNH-Ar-11526), same data as holotype
</materialsCitation>
.
</paragraph>
<paragraph id="8B91F02A4C7BFFB3FF7E6795345CFD30" blockId="3.[151,1437,152,2015]" box="[199,1085,728,755]" pageId="3" pageNumber="4">
<materialsCitation id="3B46FA774C7BFFB3FF7E67953458FD30" box="[199,1081,728,755]" country="Japan" location="Harutahama in Yakushima Island" pageId="3" pageNumber="4" specimenCount="1" stateProvince="Kagoshima" typeStatus="holotype">
<emphasis id="B95A2C384C7BFFB3FF7E67953107FD30" bold="true" box="[199,358,728,755]" pageId="3" pageNumber="4">
<typeStatus id="54954E884C7BFFB3FF7E67953163FD30" box="[199,258,729,755]" pageId="3" pageNumber="4">Type</typeStatus>
locality.
</emphasis>
<location id="8EF1A6F14C7BFFB3FED567953282FD30" LSID="urn:lsid:plazi:treatment:0387413C4C7AFFB9FF2E62083474FE4F:8EF1A6F14C7BFFB3FED567953282FD30" box="[364,739,729,755]" country="Japan" name="Harutahama in Yakushima Island" pageId="3" pageNumber="4" stateProvince="Kagoshima">Harutahama in Yakushima Island</location>
,
<collectingRegion id="49EA3EC84C7BFFB3FD576795338AFD30" box="[750,1003,729,755]" country="Japan" name="Kagoshima" pageId="3" pageNumber="4">Kagoshima Prefecture</collectingRegion>
,
<collectingCountry id="F339B0BA4C7BFFB3FC4F67953458FD30" box="[1014,1081,729,755]" name="Japan" pageId="3" pageNumber="4">Japan</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C334A3A14C7BFFB3FF7E67B1311BFCF8" pageId="3" pageNumber="4" type="etymology">
<paragraph id="8B91F02A4C7BFFB3FF7E67B1311BFCF8" blockId="3.[151,1437,152,2015]" pageId="3" pageNumber="4">
<emphasis id="B95A2C384C7BFFB3FF7E67B1312FFCD4" bold="true" box="[199,334,765,791]" pageId="3" pageNumber="4">Etymology.</emphasis>
From the Latin
<emphasis id="B95A2C384C7BFFB3FDB967B2320CFCD4" box="[512,621,766,791]" italics="true" pageId="3" pageNumber="4">contiguus</emphasis>
, -
<emphasis id="B95A2C384C7BFFB3FD3D67B232F3FCD4" box="[644,658,766,791]" italics="true" pageId="3" pageNumber="4">a</emphasis>
, -
<emphasis id="B95A2C384C7BFFB3FD1167B232ABFCD4" box="[680,714,766,791]" italics="true" pageId="3" pageNumber="4">um</emphasis>
(= contiguous), referring to the middle and proximal teeth on the gnathopod 1 carpus.
</paragraph>
</subSubSection>
<subSubSection id="C334A3A14C7BFFB3FF7E660832ADFBD0" pageId="3" pageNumber="4" type="diagnosis">
<paragraph id="8B91F02A4C7BFFB3FF7E660832ADFBD0" blockId="3.[151,1437,152,2015]" pageId="3" pageNumber="4">
<emphasis id="B95A2C384C7BFFB3FF7E66083123FC9C" bold="true" box="[199,322,836,863]" pageId="3" pageNumber="4">Diagnosis.</emphasis>
<emphasis id="B95A2C384C7BFFB3FEF3660931B8FC9C" box="[330,473,837,863]" italics="true" pageId="3" pageNumber="4">Male pereon</emphasis>
, ventral process absent.
<emphasis id="B95A2C384C7BFFB3FD48660933A0FC9C" box="[753,961,837,863]" italics="true" pageId="3" pageNumber="4">Male gnathopod 1</emphasis>
, ischium lacking posterodistal projection, merus posterodistal corner projected, posterior surface densely rugose; carpus with 1 small posterodistal tooth and 2 contiguous posterior teeth (large and small); propodus almost straight, posterodistal corner slightly swollen.
<emphasis id="B95A2C384C7BFFB3FADA66C13145FC08" italics="true" pageId="3" pageNumber="4">Male gnathopod 2</emphasis>
, coxa without posteromedial projection; basis straight; carpus longer than propodus, posterior margin and medial surface heavily setose; propodus with several robust setae on posterodistal corner and posterior margin.
<emphasis id="B95A2C384C7BFFB3FF2E66B53163FBD0" box="[151,258,1017,1043]" italics="true" pageId="3" pageNumber="4">Uropod 1</emphasis>
peduncle with short inter-ramal process.
</paragraph>
</subSubSection>
<subSubSection id="C334A3A14C7BFFB8FF7E61503315FC9C" lastPageId="8" lastPageNumber="9" pageId="3" pageNumber="4" type="description">
<paragraph id="8B91F02A4C7BFFB3FF7E61503443FBF4" blockId="3.[151,1437,152,2015]" box="[199,1058,1052,1079]" pageId="3" pageNumber="4">
<emphasis id="B95A2C384C7BFFB3FF7E615031D6FBF4" bold="true" box="[199,439,1052,1079]" pageId="3" pageNumber="4">Description of male.</emphasis>
Based on
<typeStatus id="54954E884C7BFFB3FD94615132F3FBF4" box="[557,658,1053,1079]" pageId="3" pageNumber="4" type="holotype">holotype</typeStatus>
, male,
<quantity id="4CD65DCF4C7BFFB3FD5861513356FBF4" box="[737,823,1053,1079]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" pageId="3" pageNumber="4" unit="mm" value="9.1">9.1 mm</quantity>
(OMNH-Ar-11513).
</paragraph>
<paragraph id="8B91F02A4C7BFFB3FF7E610D31FCF9EC" blockId="3.[151,1437,152,2015]" pageId="3" pageNumber="4">
<emphasis id="B95A2C384C7BFFB3FF7E610D3166FB98" bold="true" box="[199,263,1089,1115]" pageId="3" pageNumber="4">Head</emphasis>
(
<figureCitation id="1315ECAF4C7BFFB3FEA1610D3102FB98" box="[280,355,1089,1115]" captionStart="FIGURE 2" captionStartId="4.[151,250,1630,1655]" captionTargetBox="[165,1426,607,1597]" captionTargetId="figure@4.[151,1436,597,1605]" captionTargetPageId="4" captionText="FIGURE 2. Grandidierella contigua sp. nov. Holotype, male, 9.1 mm (OMNH-Ar-11513). Habitus." figureDoi="http://doi.org/10.5281/zenodo.3936949" httpUri="https://zenodo.org/record/3936949/files/figure.png" pageId="3" pageNumber="4">Figs 2</figureCitation>
,
<figureCitation id="1315ECAF4C7BFFB3FEC8610D311EFB98" box="[369,383,1089,1115]" captionStart="FIGURE 3" captionStartId="5.[151,250,1994,2019]" captionTargetBox="[197,1392,193,1949]" captionTargetId="figure@5.[175,1413,181,1969]" captionTargetPageId="5" captionText="FIGURE 3. Grandidierella contigua sp. nov. Holotype, male, 9.1 mm (OMNH-Ar-11513). Scales: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3936951" httpUri="https://zenodo.org/record/3936951/files/figure.png" pageId="3" pageNumber="4">3</figureCitation>
).
<emphasis id="B95A2C384C7BFFB3FE20610E31AFFB98" box="[409,462,1090,1115]" italics="true" pageId="3" pageNumber="4">Eyes</emphasis>
small, about 0.2 times length of head.
<emphasis id="B95A2C384C7BFFB3FC2B610D3467FB98" box="[914,1030,1089,1115]" italics="true" pageId="3" pageNumber="4">Antenna 1</emphasis>
slender, about 0.7 times as long as body; length ratio of peduncular articles 13 1:1.2:0.45, article 1 with 4 ventromedial robust setae; accessory flagellum very short, minutely biarticulate, tip with 5 setae; primary flagellum with 25 articles, terminal article minute.
<emphasis id="B95A2C384C7BFFB3FF2E61E1316AFB04" box="[151,267,1197,1223]" italics="true" pageId="3" pageNumber="4">Antenna 2</emphasis>
stout, length (peduncular articles 35 and flagellum) about 0.65 times as long as body; length ratio of peduncular articles 35 1:3.25:3.2, article 3 with 2 posterior and 1 medial robust setae; flagellum short, consisting of 7 normal and 1 minute articles, articles 2, 3, 57 with 1, 1, 1, 1, 2 robust setae, respectively.
<emphasis id="B95A2C384C7BFFB3FB7561BA3558FACC" box="[1228,1337,1269,1295]" italics="true" pageId="3" pageNumber="4">Upper lip</emphasis>
rounded ventrally, bearing many thin setae.
<emphasis id="B95A2C384C7BFFB3FD9B605532F9FAF0" box="[546,664,1305,1331]" italics="true" pageId="3" pageNumber="4">Mandibles</emphasis>
with length ratio of palp articles 13 1:1.4:1.4, article 1 with 5 short setae, article 2 bearing 68 setae, article 3 clavate, densely setose distally; incisor with 5 cusps, lacinia mobilis with 5 and 6 cusps in left and right, respectively, 9 accessory setae present.
<emphasis id="B95A2C384C7BFFB3FC7C602E3455FAB8" box="[965,1076,1377,1403]" italics="true" pageId="3" pageNumber="4">Lower lip</emphasis>
, apical parts of outer and inner lobes covered with thin setae.
<emphasis id="B95A2C384C7BFFB3FE5560C93238FA5C" box="[492,601,1413,1439]" italics="true" pageId="3" pageNumber="4">Maxilla 1</emphasis>
, outer plate with 10 distal robust setae, palp article 2 bearing 6 robust and 5 slender setae apically.
<emphasis id="B95A2C384C7BFFB3FE1560E53278FA00" box="[428,537,1449,1475]" italics="true" pageId="3" pageNumber="4">Maxilla 2</emphasis>
, inner plate bearing distal and medial setae and mediofacial row of setae, outer plate setose on distal margin.
<emphasis id="B95A2C384C7BFFB3FE5360813202FA24" box="[490,611,1485,1511]" italics="true" pageId="3" pageNumber="4">Maxilliped</emphasis>
, distal part of inner plate setose, with 3 quadrate and 1 triangular robust setae; outer plate bearing 14 long-to-short robust setae mediodistally; palp article 2 long, with many medial setae, article 3 setose distally.
</paragraph>
<paragraph id="8B91F02A4C7BFFB4FF7E637534DCFDFC" blockId="3.[151,1437,152,2015]" lastBlockId="4.[151,1437,153,575]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
<emphasis id="B95A2C384C7BFFB3FF7E6375317BF990" bold="true" box="[199,282,1593,1619]" pageId="3" pageNumber="4">Pereon</emphasis>
(
<figureCitation id="1315ECAF4C7BFFB3FE9563753113F990" box="[300,370,1593,1619]" captionStart="FIGURE 4" captionStartId="6.[151,250,1991,2016]" captionTargetBox="[188,1396,193,1955]" captionTargetId="figure@6.[177,1411,181,1965]" captionTargetPageId="6" captionText="FIGURE 4. Grandidierella contigua sp. nov. Holotype, male, 9.1 mm (OMNH-Ar-11513). Scales: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3936953" httpUri="https://zenodo.org/record/3936953/files/figure.png" pageId="3" pageNumber="4">Fig. 4</figureCitation>
).
<emphasis id="B95A2C384C7BFFB3FE3263753256F990" box="[395,567,1593,1619]" italics="true" pageId="3" pageNumber="4">Ventral surface</emphasis>
lacking process.
<emphasis id="B95A2C384C7BFFB3FCBB637533F6F990" box="[770,919,1593,1619]" italics="true" pageId="3" pageNumber="4">Gnathopod 1</emphasis>
, coxa trapezoidal; basis wide, lateral surface with ridge, posterior margin convex; ischium short, posterodistal corner not projected; merus posterodistal corner with projection, posterolateral and posteromedial surfaces densely rugose; carpus longish ovoid, length about 1.8 times width, posterodistal corner with small triangular tooth, posterior margin bearing 2 contiguous teeth (long and very short) in middle, sparsely setose; propodus almost straight, posterodistal corner a little swollen roundly, setose; dactylus short, curved posteriorly, posterior margin with 4 denticles.
<emphasis id="B95A2C384C7BFFB3FC4163A134EDF8C4" box="[1016,1164,1773,1799]" italics="true" pageId="3" pageNumber="4">Gnathopod 2</emphasis>
, coxa trapezoidal; basis slender, straight, anterior margin lined with setae; merus posterodistal margin densely setose; carpus long, relatively wide, posterior margin and medial surface heavily setose; propodus about 0.65 times as long as carpus, posterodistal corner and posterior margin with 2 and 3 short robust setae, respectively; dactylus short, curved posteriorly, posterior margin with many denticles.
<emphasis id="B95A2C384C7BFFB3FDF06231337DF854" box="[585,796,1917,1943]" italics="true" pageId="3" pageNumber="4">Pereopods 3 and 4</emphasis>
, pereopod 3 slightly longer than pereopod 4; coxae trapezoidal; bases long; meri slightly wide, straight, densely setose posteriorly and anterodistally; carpi shorter than meri; propodi and dactyli narrow.
<emphasis id="B95A2C384C7BFFB3FDA6628932C3F81C" box="[543,674,1989,2015]" italics="true" pageId="3" pageNumber="4">Pereopod 5</emphasis>
shorter than pereopod 4; coxa bilobed; basis roundish rectangular, posteroproximal corner roundly projected, anterior and posterior margins with several simple setae; merus with 4 long thick setae; carpus bearing 2 anterodistal thick setae and 3 posterior robust setae; propodus with 5 posterior robust setae; dactylus short, straight.
<emphasis id="B95A2C384C7CFFB4FD8265AD32DDFF38" box="[571,700,225,251]" italics="true" pageId="4" pageNumber="5">Pereopod 6</emphasis>
about 1.7 times as long as pereopod 5; coxa short, bilobed; basis rectangular, posteroproximal corner roundly projected, anterior margin with 1 plumose, 4 robust and several simple setae, posterior margin with many plumose and several simple setae; merus anterodistal corner with 2 thick setae, posterior margin bearing 3 thick setae, posterodistal corner with 2 robust setae; carpus with 5 lateral robust setae, posterodistal corner bearing 1 robust and 1 thick setae; propodus with 5 anterior robust setae, posterodistal corner setose; dactylus curved.
<emphasis id="B95A2C384C7CFFB4FE1E64D93249FE6C" box="[423,552,405,431]" italics="true" pageId="4" pageNumber="5">Pereopod 7</emphasis>
about 1.15 times as long as pereopod 6; coxa short, slightly bilobed; basis rectangular, posteroproximal corner not projected, anterior margin with 2 distal plumose setae and 2 robust and several simple setae, posterior margin bearing many plumose and several simple setae; merus posterior margin bearing 3 thick setae, posterodistal corner with 2 robust setae; carpus with 5 lateral robust setae, posterodistal corner bearing robust seta; propodus with 5 anterior robust setae, posterodistal corner setose; dactylus curved.
</paragraph>
<caption id="DF51A0A24C7CFFB4FF2E631234F5F9B4" ID-DOI="http://doi.org/10.5281/zenodo.3936949" ID-Zenodo-Dep="3936949" box="[151,1172,1630,1655]" httpUri="https://zenodo.org/record/3936949/files/figure.png" pageId="4" pageNumber="5" startId="4.[151,250,1630,1655]" targetBox="[165,1426,607,1597]" targetPageId="4">
<paragraph id="8B91F02A4C7CFFB4FF2E631234F5F9B4" blockId="4.[151,1172,1630,1655]" box="[151,1172,1630,1655]" pageId="4" pageNumber="5">
<emphasis id="B95A2C384C7CFFB4FF2E63123172F9B4" bold="true" box="[151,275,1630,1655]" pageId="4" pageNumber="5">FIGURE 2.</emphasis>
<taxonomicName id="4C2E8BA94C7CFFB4FEA063123271F9B5" authority="Ariyama, 2020" authorityName="Ariyama" authorityYear="2020" box="[281,528,1630,1654]" class="Malacostraca" family="Aoridae" genus="Grandidierella" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="contigua" status="sp. nov.">
<emphasis id="B95A2C384C7CFFB4FEA063123271F9B5" box="[281,528,1630,1654]" italics="true" pageId="4" pageNumber="5">Grandidierella contigua</emphasis>
</taxonomicName>
<emphasis id="B95A2C384C7CFFB4FDAF63123207F9B5" bold="true" box="[534,614,1630,1654]" pageId="4" pageNumber="5">
<taxonomicNameLabel id="A26991434C7CFFB4FDAF63123207F9B5" box="[534,614,1630,1654]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Holotype, male, 9.1 mm (OMNH-Ar-11513). Habitus.
</paragraph>
</caption>
<paragraph id="8B91F02A4C7CFFB8FF7E63E93559FF14" blockId="4.[151,1437,1700,2015]" lastBlockId="8.[151,1437,153,863]" lastPageId="8" lastPageNumber="9" pageId="4" pageNumber="5">
<emphasis id="B95A2C384C7CFFB4FF7E63E9316BF97C" bold="true" box="[199,266,1701,1727]" pageId="4" pageNumber="5">Pleon</emphasis>
(
<figureCitation id="1315ECAF4C7CFFB4FEA263E8313FF97C" box="[283,350,1700,1727]" captionStart="FIGURE 5" captionStartId="7.[151,250,1945,1970]" captionTargetBox="[204,1384,195,1906]" captionTargetId="figure@7.[195,1393,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Grandidierella contigua sp. nov. All except for F, M1 and M2: holotype, male, 9.1 mm (OMNH-Ar-11513); F, paratype, ovigerous female, 8.4 mm (OMNH-Ar-11514); M1, paratype, male, 6.7 mm (OMNH-Ar-11516); M2, paratype, male, ca. 3.7 mm (OMNH-Ar-11518). Scales: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3936955" httpUri="https://zenodo.org/record/3936955/files/figure.png" pageId="4" pageNumber="5">Fig. 5</figureCitation>
).
<emphasis id="B95A2C384C7CFFB4FECE63E9323DF97C" box="[375,604,1701,1727]" italics="true" pageId="4" pageNumber="5">Epimeral plates 13</emphasis>
rounded posteroventrally, each with long seta, ventral margins bare (
<figureCitation id="1315ECAF4C7CFFB4FAD663E830C9F920" captionStart="FIGURE 2" captionStartId="4.[151,250,1630,1655]" captionTargetBox="[165,1426,607,1597]" captionTargetId="figure@4.[151,1436,597,1605]" captionTargetPageId="4" captionText="FIGURE 2. Grandidierella contigua sp. nov. Holotype, male, 9.1 mm (OMNH-Ar-11513). Habitus." figureDoi="http://doi.org/10.5281/zenodo.3936949" httpUri="https://zenodo.org/record/3936949/files/figure.png" pageId="4" pageNumber="5">Fig. 2</figureCitation>
).
<emphasis id="B95A2C384C7CFFB4FF0263853142F920" box="[187,291,1737,1763]" italics="true" pageId="4" pageNumber="5">Pleopods</emphasis>
, each peduncle bearing many plumose setae and 2 coupling hooks; inner ramus longer than outer, outer rami with 1516 articles, inner rami with 1416 articles.
<emphasis id="B95A2C384C7CFFB4FCB263A13314F8C4" box="[779,885,1773,1799]" italics="true" pageId="4" pageNumber="5">Uropod 1</emphasis>
stout; dorsal surface of peduncle bearing 2 lateral and 4 medial robust setae, ventrodistal end with short inter-ramal process (about 9% length of peduncle); both rami shorter than peduncle, outer ramus subequal to inner ramus, former with 4 dorsal and 4 terminal robust setae, latter bearing 2 dorsolateral, 3 dorsomedial and 5 terminal robust setae.
<emphasis id="B95A2C384C7CFFB4FC3062153397F8B0" box="[905,1014,1881,1907]" italics="true" pageId="4" pageNumber="5">Uropod 2</emphasis>
about 0.6 times as long as uropod 1; peduncle dorsal surface with 2 lateral and 1 mediodistal robust setae; inner ramus almost same length as peduncle, outer ramus shorter than inner ramus, former with 2 lateral and 4 terminal robust setae, latter bearing 2 dorsolateral, 3 dorsomedial and 5 terminal robust setae.
<emphasis id="B95A2C384C7CFFB4FD3D62893290F81C" box="[644,753,1989,2015]" italics="true" pageId="4" pageNumber="5">Uropod 3</emphasis>
about half length of uropod 2; peduncle slightly wider than long, swollen mediodistally, lateral margin bare; single ramus slightly longer than peduncle, with tiny second article and 5 lateral, 2 medial and 5 terminal setae.
<taxonomicName id="4C2E8BA94C70FFB8FD3F65F132AFFF14" authorityName="Pearse" authorityYear="1952" box="[646,718,189,215]" class="Maxillopoda" family="Telsidae" genus="Telson" kingdom="Animalia" order="Poecilostomatoida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B95A2C384C70FFB8FD3F65F132AFFF14" box="[646,718,189,215]" italics="true" pageId="8" pageNumber="9">Telson</emphasis>
</taxonomicName>
wider than long, laterodistal corners each with 3 setae.
</paragraph>
<caption id="DF51A0A24C7DFFB5FF2E62863480F821" ID-DOI="http://doi.org/10.5281/zenodo.3936951" ID-Zenodo-Dep="3936951" box="[151,1249,1994,2019]" httpUri="https://zenodo.org/record/3936951/files/figure.png" pageId="5" pageNumber="6" startId="5.[151,250,1994,2019]" targetBox="[197,1392,193,1949]" targetPageId="5">
<paragraph id="8B91F02A4C7DFFB5FF2E62863480F821" blockId="5.[151,1249,1994,2019]" box="[151,1249,1994,2019]" pageId="5" pageNumber="6">
<emphasis id="B95A2C384C7DFFB5FF2E62863172F820" bold="true" box="[151,275,1994,2019]" pageId="5" pageNumber="6">FIGURE 3.</emphasis>
<taxonomicName id="4C2E8BA94C7DFFB5FEA062873271F820" authority="Ariyama, 2020" authorityName="Ariyama" authorityYear="2020" box="[281,528,1995,2019]" class="Malacostraca" family="Aoridae" genus="Grandidierella" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="contigua" status="sp. nov.">
<emphasis id="B95A2C384C7DFFB5FEA062873271F820" box="[281,528,1995,2019]" italics="true" pageId="5" pageNumber="6">Grandidierella contigua</emphasis>
</taxonomicName>
<emphasis id="B95A2C384C7DFFB5FDAF62873207F820" bold="true" box="[534,614,1995,2019]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="A26991434C7DFFB5FDAF62873207F820" box="[534,614,1995,2019]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Holotype, male, 9.1 mm (OMNH-Ar-11513). Scales: 0.1 mm.
</paragraph>
</caption>
<caption id="DF51A0A24C7EFFB6FF2E628B3480F81C" ID-DOI="http://doi.org/10.5281/zenodo.3936953" ID-Zenodo-Dep="3936953" box="[151,1249,1991,2016]" httpUri="https://zenodo.org/record/3936953/files/figure.png" pageId="6" pageNumber="7" startId="6.[151,250,1991,2016]" targetBox="[188,1396,193,1955]" targetPageId="6">
<paragraph id="8B91F02A4C7EFFB6FF2E628B3480F81C" blockId="6.[151,1249,1991,2016]" box="[151,1249,1991,2016]" pageId="6" pageNumber="7">
<emphasis id="B95A2C384C7EFFB6FF2E628B3172F823" bold="true" box="[151,275,1991,2016]" pageId="6" pageNumber="7">FIGURE 4.</emphasis>
<taxonomicName id="4C2E8BA94C7EFFB6FEA062843271F81C" authority="Ariyama, 2020" authorityName="Ariyama" authorityYear="2020" box="[281,528,1992,2016]" class="Malacostraca" family="Aoridae" genus="Grandidierella" kingdom="Animalia" order="Amphipoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="contigua" status="sp. nov.">
<emphasis id="B95A2C384C7EFFB6FEA062843271F81C" box="[281,528,1992,2016]" italics="true" pageId="6" pageNumber="7">Grandidierella contigua</emphasis>
</taxonomicName>
<emphasis id="B95A2C384C7EFFB6FDAF62843207F823" bold="true" box="[534,614,1992,2016]" pageId="6" pageNumber="7">
<taxonomicNameLabel id="A26991434C7EFFB6FDAF62843207F823" box="[534,614,1992,2016]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Holotype, male, 9.1 mm (OMNH-Ar-11513). Scales: 0.2 mm.
</paragraph>
</caption>
<caption id="DF51A0A24C7FFFB7FF2E62D532E4F83A" ID-DOI="http://doi.org/10.5281/zenodo.3936955" ID-Zenodo-Dep="3936955" httpUri="https://zenodo.org/record/3936955/files/figure.png" pageId="7" pageNumber="8" startId="7.[151,250,1945,1970]" targetBox="[204,1384,195,1906]" targetPageId="7">
<paragraph id="8B91F02A4C7FFFB7FF2E62D532E4F83A" blockId="7.[151,1436,1945,2042]" pageId="7" pageNumber="8">
<emphasis id="B95A2C384C7FFFB7FF2E62D53174F871" bold="true" box="[151,277,1945,1970]" pageId="7" pageNumber="8">FIGURE 5.</emphasis>
<taxonomicName id="4C2E8BA94C7FFFB7FEA462D63274F872" authority="Ariyama, 2020" authorityName="Ariyama" authorityYear="2020" box="[285,533,1946,1970]" class="Malacostraca" family="Aoridae" genus="Grandidierella" kingdom="Animalia" order="Amphipoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="contigua" status="sp. nov.">
<emphasis id="B95A2C384C7FFFB7FEA462D63274F872" box="[285,533,1946,1970]" italics="true" pageId="7" pageNumber="8">Grandidierella contigua</emphasis>
</taxonomicName>
<emphasis id="B95A2C384C7FFFB7FDA462D63211F871" bold="true" box="[541,624,1946,1970]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="A26991434C7FFFB7FDA462D63211F871" box="[541,624,1946,1970]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
All except for F, M1 and M2: holotype, male, 9.1 mm (OMNH-Ar-11513); F, paratype, ovigerous female, 8.4 mm (OMNH-Ar-11514); M1, paratype, male, 6.7 mm (OMNH-Ar-11516); M2, paratype, male, ca. 3.7 mm (OMNH-Ar-11518). Scales: 0.2 mm.
</paragraph>
</caption>
<paragraph id="8B91F02A4C70FFB8FF7E65AC31E9FE6C" blockId="8.[151,1437,153,863]" pageId="8" pageNumber="9">
<emphasis id="B95A2C384C70FFB8FF7E65AC31AFFF38" bold="true" box="[199,462,224,251]" pageId="8" pageNumber="9">Description of female.</emphasis>
Based on
<typeStatus id="54954E884C70FFB8FDFD65AD32C6FF38" box="[580,679,225,251]" pageId="8" pageNumber="9" type="paratype">paratype</typeStatus>
, ovigerous female,
<quantity id="4CD65DCF4C70FFB8FC3A65AD33BBFF38" box="[899,986,225,251]" metricMagnitude="-3" metricUnit="m" metricValue="8.4" pageId="8" pageNumber="9" unit="mm" value="8.4">8.4 mm</quantity>
(OMNH-Ar-11514).
<emphasis id="B95A2C384C70FFB8FB7565AD353EFF38" box="[1228,1375,225,251]" italics="true" pageId="8" pageNumber="9">Gnathopod 1</emphasis>
(
<figureCitation id="1315ECAF4C70FFB8FAD665AD30C4FEDC" captionStart="FIGURE 5" captionStartId="7.[151,250,1945,1970]" captionTargetBox="[204,1384,195,1906]" captionTargetId="figure@7.[195,1393,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Grandidierella contigua sp. nov. All except for F, M1 and M2: holotype, male, 9.1 mm (OMNH-Ar-11513); F, paratype, ovigerous female, 8.4 mm (OMNH-Ar-11514); M1, paratype, male, 6.7 mm (OMNH-Ar-11516); M2, paratype, male, ca. 3.7 mm (OMNH-Ar-11518). Scales: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3936955" httpUri="https://zenodo.org/record/3936955/files/figure.png" pageId="8" pageNumber="9">Fig. 5</figureCitation>
F-G1), coxa trapezoidal; basis almost straight; ischium and merus short, both lacking projection; carpus wide, posterior margin with robust seta and many simple setae; propodus about 0.85 times as long as carpus, posterior margin with 5 robust setae; dactylus slightly curved posteriorly, posterior margin with 6 denticles.
<emphasis id="B95A2C384C70FFB8FABE640135FDFEA4" box="[1287,1436,333,359]" italics="true" pageId="8" pageNumber="9">Gnathopod 2</emphasis>
(
<figureCitation id="1315ECAF4C70FFB8FF19643D3082FE48" box="[160,227,368,395]" captionStart="FIGURE 5" captionStartId="7.[151,250,1945,1970]" captionTargetBox="[204,1384,195,1906]" captionTargetId="figure@7.[195,1393,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Grandidierella contigua sp. nov. All except for F, M1 and M2: holotype, male, 9.1 mm (OMNH-Ar-11513); F, paratype, ovigerous female, 8.4 mm (OMNH-Ar-11514); M1, paratype, male, 6.7 mm (OMNH-Ar-11516); M2, paratype, male, ca. 3.7 mm (OMNH-Ar-11518). Scales: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3936955" httpUri="https://zenodo.org/record/3936955/files/figure.png" pageId="8" pageNumber="9">Fig. 5</figureCitation>
F-G2) almost same as that of
<typeStatus id="54954E884C70FFB8FDF8643D32C7FE48" box="[577,678,369,395]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
, male,
<quantity id="4CD65DCF4C70FFB8FD42643D3332FE48" box="[763,851,369,395]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" pageId="8" pageNumber="9" unit="mm" value="9.1">9.1 mm</quantity>
(OMNH-Ar-11513) except for oostegite, but basis wider, carpus shorter.
</paragraph>
<paragraph id="8B91F02A4C70FFB8FF7E64F431B9FD44" blockId="8.[151,1437,153,863]" pageId="8" pageNumber="9">
<emphasis id="B95A2C384C70FFB8FF7E64F4315EFE10" bold="true" box="[199,319,440,467]" pageId="8" pageNumber="9">Variation.</emphasis>
<typeStatus id="54954E884C70FFB8FEFD64F531C8FE10" box="[324,425,441,467]" pageId="8" pageNumber="9" type="paratype">Paratype</typeStatus>
, middle-sized male,
<quantity id="4CD65DCF4C70FFB8FD3464F53280FE10" box="[653,737,441,467]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="8" pageNumber="9" unit="mm" value="6.7">6.7 mm</quantity>
(OMNH-Ar-11516).
<emphasis id="B95A2C384C70FFB8FC6964F53403FE10" box="[976,1122,441,467]" italics="true" pageId="8" pageNumber="9">Gnathopod 1</emphasis>
(
<figureCitation id="1315ECAF4C70FFB8FBC964F534AEFE10" box="[1136,1231,440,467]" captionStart="FIGURE 5" captionStartId="7.[151,250,1945,1970]" captionTargetBox="[204,1384,195,1906]" captionTargetId="figure@7.[195,1393,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Grandidierella contigua sp. nov. All except for F, M1 and M2: holotype, male, 9.1 mm (OMNH-Ar-11513); F, paratype, ovigerous female, 8.4 mm (OMNH-Ar-11514); M1, paratype, male, 6.7 mm (OMNH-Ar-11516); M2, paratype, male, ca. 3.7 mm (OMNH-Ar-11518). Scales: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3936955" httpUri="https://zenodo.org/record/3936955/files/figure.png" pageId="8" pageNumber="9">Fig. 5M</figureCitation>
<figureCitation id="1315ECAF4C70FFB8FB7664F43565FE10" box="[1231,1284,440,467]" captionStart="FIGURE 1" captionStartId="1.[151,250,1778,1803]" captionTargetBox="[151,1436,561,1752]" captionTargetId="figure@1.[151,1436,561,1752]" captionTargetPageId="1" captionText="FIGURE 1. Map showing collecting sites of the examined specimens in the Ryukyu Archipelago." figureDoi="http://doi.org/10.5281/zenodo.3936947" httpUri="https://zenodo.org/record/3936947/files/figure.png" pageId="8" pageNumber="9">1-G1</figureCitation>
) almost same as that of
<typeStatus id="54954E884C70FFB8FEBB64913106FE34" box="[258,359,477,503]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
, male,
<quantity id="4CD65DCF4C70FFB8FE0C6491326BFE34" box="[437,522,477,503]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" pageId="8" pageNumber="9" unit="mm" value="9.1">9.1 mm</quantity>
(OMNH-Ar-11513), but middle tooth on carpus posterior margin situated distally, separated from proximal short tooth, propodus short and wide, posterior margin of dactylus with 3 denticles.
<typeStatus id="54954E884C70FFB8FAD8674D30ABFDFC" pageId="8" pageNumber="9" type="paratype">Paratype</typeStatus>
, small male, ca.
<quantity id="4CD65DCF4C70FFB8FE3367693183FDFC" box="[394,482,549,575]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="8" pageNumber="9" unit="mm" value="3.7">3.7 mm</quantity>
(OMNH-Ar-11518).
<emphasis id="B95A2C384C70FFB8FD6F6769330AFDFC" box="[726,875,549,575]" italics="true" pageId="8" pageNumber="9">Gnathopod 1</emphasis>
(
<figureCitation id="1315ECAF4C70FFB8FCC2676933BDFDFC" box="[891,988,548,575]" captionStart="FIGURE 5" captionStartId="7.[151,250,1945,1970]" captionTargetBox="[204,1384,195,1906]" captionTargetId="figure@7.[195,1393,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Grandidierella contigua sp. nov. All except for F, M1 and M2: holotype, male, 9.1 mm (OMNH-Ar-11513); F, paratype, ovigerous female, 8.4 mm (OMNH-Ar-11514); M1, paratype, male, 6.7 mm (OMNH-Ar-11516); M2, paratype, male, ca. 3.7 mm (OMNH-Ar-11518). Scales: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3936955" httpUri="https://zenodo.org/record/3936955/files/figure.png" pageId="8" pageNumber="9">Fig. 5M</figureCitation>
<figureCitation id="1315ECAF4C70FFB8FC656768346AFDFC" box="[988,1035,548,575]" captionStart="FIGURE 2" captionStartId="4.[151,250,1630,1655]" captionTargetBox="[165,1426,607,1597]" captionTargetId="figure@4.[151,1436,597,1605]" captionTargetPageId="4" captionText="FIGURE 2. Grandidierella contigua sp. nov. Holotype, male, 9.1 mm (OMNH-Ar-11513). Habitus." figureDoi="http://doi.org/10.5281/zenodo.3936949" httpUri="https://zenodo.org/record/3936949/files/figure.png" pageId="8" pageNumber="9">2-G</figureCitation>
<figureCitation id="1315ECAF4C70FFB8FBB267683474FDFC" box="[1035,1045,548,575]" captionStart="FIGURE 1" captionStartId="1.[151,250,1778,1803]" captionTargetBox="[151,1436,561,1752]" captionTargetId="figure@1.[151,1436,561,1752]" captionTargetPageId="1" captionText="FIGURE 1. Map showing collecting sites of the examined specimens in the Ryukyu Archipelago." figureDoi="http://doi.org/10.5281/zenodo.3936947" httpUri="https://zenodo.org/record/3936947/files/figure.png" pageId="8" pageNumber="9">1</figureCitation>
), middle tooth on carpus posterior margin situated more distally, posterior tooth minute, propodus shorter and wider than
<typeStatus id="54954E884C70FFB8FBE3670534DCFDA0" box="[1114,1213,585,611]" pageId="8" pageNumber="9" type="paratype">paratype</typeStatus>
, middle-sized male,
<quantity id="4CD65DCF4C70FFB8FF2E6721308CFD44" box="[151,237,621,647]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="8" pageNumber="9" unit="mm" value="6.7">6.7 mm</quantity>
(OMNH-Ar-11516).
</paragraph>
<paragraph id="8B91F02A4C70FFB8FF7E67DC3315FC9C" blockId="8.[151,1437,153,863]" pageId="8" pageNumber="9">
<emphasis id="B95A2C384C70FFB8FF7E67DC312EFD68" bold="true" box="[199,335,656,683]" pageId="8" pageNumber="9">Coloration.</emphasis>
Recently fixed small specimen (
<figureCitation id="1315ECAF4C70FFB8FD7767DD3376FD6F" box="[718,791,657,684]" captionStart="FIGURE 6" captionStartId="8.[151,250,1684,1709]" captionTargetBox="[151,1436,885,1660]" captionTargetId="figure@8.[151,1436,885,1660]" captionTargetPageId="8" captionText="FIGURE 6. Grandidierella contigua sp. nov. Photograph of paratype, male, 3.5 mm (OMNH-Ar-11519), fixed specimen (6.5 hours after fixation)." figureDoi="http://doi.org/10.5281/zenodo.3936957" httpUri="https://zenodo.org/record/3936957/files/figure.png" pageId="8" pageNumber="9">Fig. 6</figureCitation>
): eyes dark brown; dorsal surface of head mottled dark brown, dorsal parts of pereonites 4 and 7 dark brown; other body parts yellowish white with indistinct brown speckles; antennae white and partly pale brown; gnathopods and pereopods white. Preserved large specimens (
<figureCitation id="1315ECAF4C70FFB8FAFE679535EFFD30" box="[1351,1422,729,755]" captionStart="FIGURE 7" captionStartId="9.[151,250,926,951]" captionTargetBox="[151,1437,417,901]" captionTargetId="figure@9.[151,1437,417,901]" captionTargetPageId="9" captionText="FIGURE 7. Grandidierella contigua sp. nov. Photographs of holotype, male, 9.1 mm (OMNH-Ar-11513, left) and paratype, ovigerous female, 8.4 mm (OMNH-Ar-11514, right), preserved specimens." figureDoi="http://doi.org/10.5281/zenodo.3936961" httpUri="https://zenodo.org/record/3936961/files/figure.png" pageId="8" pageNumber="9">Fig. 7</figureCitation>
): eyes dark brown; dorsal surface of head mottled dark brown, pereonites 4 and 7 dark brown, other pereonites, pleonites and urosomites in male brown, pereonites 3, 5 and 6, pleonites and urosomites in female pale brown; antenna 2 peduncle white and partly dark brown; other appendages white.
</paragraph>
</subSubSection>
<caption id="DF51A0A24C70FFB8FF2E63D83108F912" ID-DOI="http://doi.org/10.5281/zenodo.3936957" ID-Zenodo-Dep="3936957" httpUri="https://zenodo.org/record/3936957/files/figure.png" pageId="8" pageNumber="9" startId="8.[151,250,1684,1709]" targetBox="[151,1436,885,1660]" targetPageId="8">
<paragraph id="8B91F02A4C70FFB8FF2E63D83108F912" blockId="8.[151,1436,1684,1745]" pageId="8" pageNumber="9">
<emphasis id="B95A2C384C70FFB8FF2E63D83172F96E" bold="true" box="[151,275,1684,1709]" pageId="8" pageNumber="9">FIGURE 6.</emphasis>
<taxonomicName id="4C2E8BA94C70FFB8FEA363D93270F96E" authority="Ariyama, 2020" authorityName="Ariyama" authorityYear="2020" box="[282,529,1685,1709]" class="Malacostraca" family="Aoridae" genus="Grandidierella" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="contigua" status="sp. nov.">
<emphasis id="B95A2C384C70FFB8FEA363D93270F96E" box="[282,529,1685,1709]" italics="true" pageId="8" pageNumber="9">Grandidierella contigua</emphasis>
</taxonomicName>
<emphasis id="B95A2C384C70FFB8FDAE63D93209F96E" bold="true" box="[535,616,1685,1709]" pageId="8" pageNumber="9">
<taxonomicNameLabel id="A26991434C70FFB8FDAE63D93209F96E" box="[535,616,1685,1709]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Photograph of paratype, male, 3.5 mm (OMNH-Ar-11519), fixed specimen (6.5 hours after fixation).
</paragraph>
</caption>
<subSubSection id="C334A3A14C70FFB9FF7E624C3514FEA4" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="discussion">
<paragraph id="8B91F02A4C70FFB8FF7E624C35F4F86A" blockId="8.[151,1437,1791,2034]" pageId="8" pageNumber="9">
<emphasis id="B95A2C384C70FFB8FF7E624C315AF8D9" bold="true" box="[199,315,1792,1818]" pageId="8" pageNumber="9">Remarks.</emphasis>
<taxonomicName id="4C2E8BA94C70FFB8FEFC63B33239F8DA" authority="Ariyama, 2020" authorityName="Ariyama" authorityYear="2020" box="[325,600,1791,1817]" class="Malacostraca" family="Aoridae" genus="Grandidierella" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="contigua" status="sp. nov.">
<emphasis id="B95A2C384C70FFB8FEFC63B33239F8DA" box="[325,600,1791,1817]" italics="true" pageId="8" pageNumber="9">Grandidierella contigua</emphasis>
</taxonomicName>
<emphasis id="B95A2C384C70FFB8FDD8624C32DFF8D9" bold="true" box="[609,702,1792,1818]" pageId="8" pageNumber="9">
<taxonomicNameLabel id="A26991434C70FFB8FDD8624C32DFF8D9" box="[609,702,1792,1818]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has a male gnathopod 1 with a posterodistally projected merus and a carpus bearing three distinct teeth on the posterodistal margin, the proximal tooth of which is very small and contiguous with the large middle tooth (
<figureCitation id="1315ECAF4C70FFB8FDE3620B32D9F8A1" box="[602,696,1863,1890]" captionStart="FIGURE 4" captionStartId="6.[151,250,1991,2016]" captionTargetBox="[188,1396,193,1955]" captionTargetId="figure@6.[177,1411,181,1965]" captionTargetPageId="6" captionText="FIGURE 4. Grandidierella contigua sp. nov. Holotype, male, 9.1 mm (OMNH-Ar-11513). Scales: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3936953" httpUri="https://zenodo.org/record/3936953/files/figure.png" pageId="8" pageNumber="9">Fig. 4G</figureCitation>
<figureCitation id="1315ECAF4C70FFB8FD01620432A3F8A1" box="[696,706,1864,1890]" captionStart="FIGURE 1" captionStartId="1.[151,250,1778,1803]" captionTargetBox="[151,1436,561,1752]" captionTargetId="figure@1.[151,1436,561,1752]" captionTargetPageId="1" captionText="FIGURE 1. Map showing collecting sites of the examined specimens in the Ryukyu Archipelago." figureDoi="http://doi.org/10.5281/zenodo.3936947" httpUri="https://zenodo.org/record/3936947/files/figure.png" pageId="8" pageNumber="9">1</figureCitation>
). Such a gnathopod 1 is unique in the genus
<taxonomicName id="4C2E8BA94C70FFB8FB05620B3502F8A2" authorityName="Coutiere" authorityYear="1904" box="[1212,1379,1863,1889]" class="Malacostraca" family="Aoridae" genus="Grandidierella" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B95A2C384C70FFB8FB05620B3502F8A2" box="[1212,1379,1863,1889]" italics="true" pageId="8" pageNumber="9">Grandidierella</emphasis>
</taxonomicName>
. The rugose merus of the male gnathopod 1 is also characteristic; however,
<taxonomicName id="4C2E8BA94C70FFB8FC2262203474F845" authorityName="Ariyama" authorityYear="1996" box="[923,1045,1900,1926]" class="Malacostraca" family="Aoridae" genus="Grandidierella" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B95A2C384C70FFB8FC2262203474F845" box="[923,1045,1900,1926]" italics="true" pageId="8" pageNumber="9">G. fasciata</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2E8BA94C70FFB8FB9B622034DAF846" authorityName="Ariyama" authorityYear="1996" box="[1058,1211,1899,1926]" class="Malacostraca" family="Aoridae" genus="Grandidierella" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="osakaensis">
<emphasis id="B95A2C384C70FFB8FB9B622034DAF846" box="[1058,1211,1899,1926]" italics="true" pageId="8" pageNumber="9">G. osakaensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2E8BA94C70FFB8FB56622035FDF846" authorityName="Ariyama &amp; Taru" authorityYear="2017" box="[1263,1436,1899,1926]" class="Malacostraca" family="Aoridae" genus="Grandidierella" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="sanrikuensis">
<emphasis id="B95A2C384C70FFB8FB56622035FDF846" box="[1263,1436,1899,1926]" italics="true" pageId="8" pageNumber="9">G. sanrikuensis</emphasis>
</taxonomicName>
(reexamination of the
<typeStatus id="54954E884C70FFB8FE3662DC31DEF869" box="[399,447,1936,1962]" pageId="8" pageNumber="9">type</typeStatus>
specimens) and
<taxonomicName id="4C2E8BA94C70FFB8FDC262DC3339F86A" authority="Ariyama, 2020" authorityName="Ariyama" authorityYear="2020" box="[635,856,1935,1962]" class="Malacostraca" family="Aoridae" genus="Grandidierella" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="pseudosakaensis" status="sp. nov.">
<emphasis id="B95A2C384C70FFB8FDC262DC3339F86A" box="[635,856,1935,1962]" italics="true" pageId="8" pageNumber="9">G. pseudosakaensis</emphasis>
</taxonomicName>
<emphasis id="B95A2C384C70FFB8FCE662DC33D9F869" bold="true" box="[863,952,1936,1962]" pageId="8" pageNumber="9">
<taxonomicNameLabel id="A26991434C70FFB8FCE662DC33D9F869" box="[863,952,1936,1962]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
share this character with this new species.
</paragraph>
<paragraph id="8B91F02A4C70FFB9FF7E62FF3341FE80" blockId="8.[151,1437,1791,2034]" lastBlockId="9.[151,1437,153,396]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
The gnathopod 1 of the middle-sized male (
<figureCitation id="1315ECAF4C70FFB8FD1362FF336BF80D" box="[682,778,1971,1998]" captionStart="FIGURE 5" captionStartId="7.[151,250,1945,1970]" captionTargetBox="[204,1384,195,1906]" captionTargetId="figure@7.[195,1393,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Grandidierella contigua sp. nov. All except for F, M1 and M2: holotype, male, 9.1 mm (OMNH-Ar-11513); F, paratype, ovigerous female, 8.4 mm (OMNH-Ar-11514); M1, paratype, male, 6.7 mm (OMNH-Ar-11516); M2, paratype, male, ca. 3.7 mm (OMNH-Ar-11518). Scales: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3936955" httpUri="https://zenodo.org/record/3936955/files/figure.png" pageId="8" pageNumber="9">Fig. 5M</figureCitation>
<figureCitation id="1315ECAF4C70FFB8FCB362FF335EF80D" box="[778,831,1971,1998]" captionStart="FIGURE 1" captionStartId="1.[151,250,1778,1803]" captionTargetBox="[151,1436,561,1752]" captionTargetId="figure@1.[151,1436,561,1752]" captionTargetPageId="1" captionText="FIGURE 1. Map showing collecting sites of the examined specimens in the Ryukyu Archipelago." figureDoi="http://doi.org/10.5281/zenodo.3936947" httpUri="https://zenodo.org/record/3936947/files/figure.png" pageId="8" pageNumber="9">1-G1</figureCitation>
) is similar to that of
<taxonomicName id="4C2E8BA94C70FFB8FB9D62F834B0F80E" authorityName="Ariyama &amp; Taru" authorityYear="2017" box="[1060,1233,1971,1998]" class="Malacostraca" family="Aoridae" genus="Grandidierella" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="sanrikuensis">
<emphasis id="B95A2C384C70FFB8FB9D62F834B0F80E" box="[1060,1233,1971,1998]" italics="true" pageId="8" pageNumber="9">G. sanrikuensis</emphasis>
</taxonomicName>
which has a small proximal tooth on the carpus, but
<taxonomicName id="4C2E8BA94C70FFB8FDA4629432C2F832" authorityName="Ariyama" authorityYear="2020" box="[541,675,2008,2034]" class="Malacostraca" family="Aoridae" genus="Grandidierella" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="contigua">
<emphasis id="B95A2C384C70FFB8FDA4629432C2F832" box="[541,675,2008,2034]" italics="true" pageId="8" pageNumber="9">G. contigua</emphasis>
</taxonomicName>
is different from
<taxonomicName id="4C2E8BA94C70FFB8FCD66294347EF832" authorityName="Ariyama &amp; Taru" authorityYear="2017" box="[879,1055,2007,2034]" class="Malacostraca" family="Aoridae" genus="Grandidierella" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="sanrikuensis">
<emphasis id="B95A2C384C70FFB8FCD66294347EF832" box="[879,1055,2007,2034]" italics="true" pageId="8" pageNumber="9">G. sanrikuensis</emphasis>
</taxonomicName>
in the densely setose meri of the male pereopods 3 and 4 (
<figureCitation id="1315ECAF4C71FFB9FE0865D53267FF70" box="[433,518,153,179]" captionStart="FIGURE 4" captionStartId="6.[151,250,1991,2016]" captionTargetBox="[188,1396,193,1955]" captionTargetId="figure@6.[177,1411,181,1965]" captionTargetPageId="6" captionText="FIGURE 4. Grandidierella contigua sp. nov. Holotype, male, 9.1 mm (OMNH-Ar-11513). Scales: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3936953" httpUri="https://zenodo.org/record/3936953/files/figure.png" pageId="9" pageNumber="10">Fig. 4P</figureCitation>
<figureCitation id="1315ECAF4C71FFB9FDBF65D53272FF70" box="[518,531,153,179]" captionStart="FIGURE 3" captionStartId="5.[151,250,1994,2019]" captionTargetBox="[197,1392,193,1949]" captionTargetId="figure@5.[175,1413,181,1969]" captionTargetPageId="5" captionText="FIGURE 3. Grandidierella contigua sp. nov. Holotype, male, 9.1 mm (OMNH-Ar-11513). Scales: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3936951" httpUri="https://zenodo.org/record/3936951/files/figure.png" pageId="9" pageNumber="10">3</figureCitation>
, P
<figureCitation id="1315ECAF4C71FFB9FD9665D5325CFF70" box="[559,573,153,179]" captionStart="FIGURE 4" captionStartId="6.[151,250,1991,2016]" captionTargetBox="[188,1396,193,1955]" captionTargetId="figure@6.[177,1411,181,1965]" captionTargetPageId="6" captionText="FIGURE 4. Grandidierella contigua sp. nov. Holotype, male, 9.1 mm (OMNH-Ar-11513). Scales: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3936953" httpUri="https://zenodo.org/record/3936953/files/figure.png" pageId="9" pageNumber="10">4</figureCitation>
) and the coloration (
<figureCitation id="1315ECAF4C71FFB9FC9F65D5330EFF70" box="[806,879,153,179]" captionStart="FIGURE 6" captionStartId="8.[151,250,1684,1709]" captionTargetBox="[151,1436,885,1660]" captionTargetId="figure@8.[151,1436,885,1660]" captionTargetPageId="8" captionText="FIGURE 6. Grandidierella contigua sp. nov. Photograph of paratype, male, 3.5 mm (OMNH-Ar-11519), fixed specimen (6.5 hours after fixation)." figureDoi="http://doi.org/10.5281/zenodo.3936957" httpUri="https://zenodo.org/record/3936957/files/figure.png" pageId="9" pageNumber="10">Figs 6</figureCitation>
,
<figureCitation id="1315ECAF4C71FFB9FCC265D533EAFF70" box="[891,907,153,179]" captionStart="FIGURE 7" captionStartId="9.[151,250,926,951]" captionTargetBox="[151,1437,417,901]" captionTargetId="figure@9.[151,1437,417,901]" captionTargetPageId="9" captionText="FIGURE 7. Grandidierella contigua sp. nov. Photographs of holotype, male, 9.1 mm (OMNH-Ar-11513, left) and paratype, ovigerous female, 8.4 mm (OMNH-Ar-11514, right), preserved specimens." figureDoi="http://doi.org/10.5281/zenodo.3936961" httpUri="https://zenodo.org/record/3936961/files/figure.png" pageId="9" pageNumber="10">7</figureCitation>
). The small specimens of this species collected from Irimomote Island resemble
<taxonomicName id="4C2E8BA94C71FFB9FDBC65F132E1FF14" authorityName="Ariyama" authorityYear="1996" box="[517,640,189,215]" class="Malacostraca" family="Aoridae" genus="Grandidierella" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B95A2C384C71FFB9FDBC65F132E1FF14" box="[517,640,189,215]" italics="true" pageId="9" pageNumber="10">G. fasciata</emphasis>
</taxonomicName>
in the shape of the male gnathopod 1 (
<figureCitation id="1315ECAF4C71FFB9FB9365F134E6FF14" box="[1066,1159,188,215]" captionStart="FIGURE 5" captionStartId="7.[151,250,1945,1970]" captionTargetBox="[204,1384,195,1906]" captionTargetId="figure@7.[195,1393,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Grandidierella contigua sp. nov. All except for F, M1 and M2: holotype, male, 9.1 mm (OMNH-Ar-11513); F, paratype, ovigerous female, 8.4 mm (OMNH-Ar-11514); M1, paratype, male, 6.7 mm (OMNH-Ar-11516); M2, paratype, male, ca. 3.7 mm (OMNH-Ar-11518). Scales: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3936955" httpUri="https://zenodo.org/record/3936955/files/figure.png" pageId="9" pageNumber="10">Fig. 5M</figureCitation>
<figureCitation id="1315ECAF4C71FFB9FB3E65F034D4FF14" box="[1159,1205,188,215]" captionStart="FIGURE 2" captionStartId="4.[151,250,1630,1655]" captionTargetBox="[165,1426,607,1597]" captionTargetId="figure@4.[151,1436,597,1605]" captionTargetPageId="4" captionText="FIGURE 2. Grandidierella contigua sp. nov. Holotype, male, 9.1 mm (OMNH-Ar-11513). Habitus." figureDoi="http://doi.org/10.5281/zenodo.3936949" httpUri="https://zenodo.org/record/3936949/files/figure.png" pageId="9" pageNumber="10">2-G</figureCitation>
<figureCitation id="1315ECAF4C71FFB9FB0C65F034DFFF14" box="[1205,1214,188,215]" captionStart="FIGURE 1" captionStartId="1.[151,250,1778,1803]" captionTargetBox="[151,1436,561,1752]" captionTargetId="figure@1.[151,1436,561,1752]" captionTargetPageId="1" captionText="FIGURE 1. Map showing collecting sites of the examined specimens in the Ryukyu Archipelago." figureDoi="http://doi.org/10.5281/zenodo.3936947" httpUri="https://zenodo.org/record/3936947/files/figure.png" pageId="9" pageNumber="10">1</figureCitation>
) and the coloration. But
<taxonomicName id="4C2E8BA94C71FFB9FF7E65AD3123FF38" authorityName="Ariyama" authorityYear="1996" box="[199,322,225,251]" class="Malacostraca" family="Aoridae" genus="Grandidierella" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B95A2C384C71FFB9FF7E65AD3123FF38" box="[199,322,225,251]" italics="true" pageId="9" pageNumber="10">G. fasciata</emphasis>
</taxonomicName>
has a posterodistal projection on the male gnathopod 1 ischium and a longer inter-ramal process on the uropod 1 peduncle. The length of the inter-ramal process is 79% (average: 8%) of the peduncle length in
<taxonomicName id="4C2E8BA94C71FFB9FA3864493098FE80" authorityName="Ariyama" authorityYear="2020" class="Malacostraca" family="Aoridae" genus="Grandidierella" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="contigua">
<emphasis id="B95A2C384C71FFB9FA3864493098FE80" italics="true" pageId="9" pageNumber="10">G. contigua</emphasis>
</taxonomicName>
, whereas 1422% (average: 18%) in
<taxonomicName id="4C2E8BA94C71FFB9FD2764653378FE80" authorityName="Ariyama" authorityYear="1996" box="[670,793,297,323]" class="Malacostraca" family="Aoridae" genus="Grandidierella" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B95A2C384C71FFB9FD2764653378FE80" box="[670,793,297,323]" italics="true" pageId="9" pageNumber="10">G. fasciata</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B91F02A4C71FFB9FF7E64003514FEA4" blockId="9.[151,1437,153,396]" box="[199,1397,332,359]" pageId="9" pageNumber="10">
<emphasis id="B95A2C384C71FFB9FF7E6400314BFEA4" bold="true" box="[199,298,332,359]" pageId="9" pageNumber="10">Habitat.</emphasis>
Coarse sand bottom and sandy mud bottom under stones and soft sandstone in the intertidal zone.
</paragraph>
</subSubSection>
<subSubSection id="C334A3A14C71FFB9FF7E643C3474FE4F" box="[199,1045,368,396]" pageId="9" pageNumber="10" type="distribution">
<paragraph id="8B91F02A4C71FFB9FF7E643C3474FE4F" blockId="9.[151,1437,153,396]" box="[199,1045,368,396]" pageId="9" pageNumber="10">
<emphasis id="B95A2C384C71FFB9FF7E643C3100FE48" bold="true" box="[199,353,368,395]" pageId="9" pageNumber="10">Distribution.</emphasis>
<collectingCountry id="F339B0BA4C71FFB9FEDE643D31CAFE48" box="[359,427,369,395]" name="Japan" pageId="9" pageNumber="10">Japan</collectingCountry>
: Yakushima Island and Iriomote Island (present study).
</paragraph>
</subSubSection>
</treatment>
</document>