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<mods:title id="086D6EE3812B985C7DB49CEDB793B8E7">The genus Diplommatina Benson, 1849 (Gastropoda: Caenogastropoda: Diplommatinidae) in Nepal, with the description of seven new species</mods:title>
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<mods:affiliation id="8D3AB562D0BEDEDD0136FB33BE4CF5E3">University of Antwerp, Evolutionary Ecology Group, Universiteitsplein 1, B- 2610, Antwerp, Belgium. &amp; Central Department of Zoology, Tribhuvan University, Kirtipur, Kathmandu, Nepal. &amp; Corresponding author: prembudha @ yahoo. com</mods:affiliation>
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<mods:affiliation id="B6D3D86508B22D06D32DC627C85E2637">Natural History Museum, Cromwell Road, London, SW 7 5 BD, UK. &amp; Email: f. naggs @ nhm. ac. uk</mods:affiliation>
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<mods:affiliation id="94A3AF4899AB93D86880C0E913753B5E">University of Antwerp, Evolutionary Ecology Group, Universiteitsplein 1, B- 2610, Antwerp, Belgium. &amp; Royal Belgian Institute of Natural Sciences, Vautierstraat 29, B- 1000, Brussels, Belgium. &amp; Email: thierry. backeljau @ naturalsciences. be</mods:affiliation>
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(L. 
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)
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<figureCitation id="135462D2FFECFFE3FD1743C1D64AFB3E" box="[697,792,1187,1213]" captionStart="Fig" captionStartId="4.[189,232,1895,1921]" captionTargetBox="[189,1398,285,1853]" captionTargetId="figure@4.[189,1398,285,1853]" captionTargetPageId="4" captionText="Fig. 2. Shell terminology. A. D. regularis Fulton, 1901. B. D. folliculus (L. Pfeiffer, 1846). C. General outline of the lower part of a shell from Diplommatina species. Abbreviations: Prt = Parietalis (Prt1 = inner parietalis, Prt2 = outer parietalis); Pt = Palatalis; Hpt = Horizontal palatalis (Hpt1 = lower, Hpt2 = upper in case of two); Vpt = Vertical palatalis (Vpt1 = left side, Vpt2 = right side in case of two)." figureDoi="http://doi.org/10.5281/zenodo.3832488" httpUri="https://zenodo.org/record/3832488/files/figure.png" pageId="8" pageNumber="9">Figs 2B</figureCitation>
, 
<figureCitation id="135462D2FFECFFE3FC8A43C1D618FB3E" box="[804,842,1187,1213]" captionStart="Fig" captionStartId="9.[189,232,1930,1956]" captionTargetBox="[189,1398,274,1888]" captionTargetId="figure@9.[189,1398,266,1891]" captionTargetPageId="9" captionText="Fig. 4. Diplommatina Benson, 1849 of Nepal. A. D. folliculus (L. Pfeiffer, 1846) (CDZMTU028). B. D. godawariensis sp. nov., holotype (CDZMTU032). C. D. maipokhariensis sp. nov., holotype (CDZMTU034). Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832492" httpUri="https://zenodo.org/record/3832492/files/figure.png" pageId="8" pageNumber="9">4A</figureCitation>
, 
<figureCitation id="135462D2FFECFFE3FCF843C1D628FB3E" box="[854,890,1187,1213]" captionStart="Fig" captionStartId="24.[189,232,1789,1815]" captionTargetBox="[297,1297,294,1724]" captionTargetId="figure@24.[247,1340,265,1748]" captionTargetPageId="24" captionText="Fig. 9. Palatal lamellae (palatalis) of species of Diplommatina. Dotted lines indicate the position of the constriction. Dark lines indicate the palatalis. A. D. abiesiana sp. nov. B. D. exserta Godwin- Austen, 1886. C. D. fistulata sp. nov. D. D. folliculus (L. Pfeiffer, 1846). E. D. godawariensis sp. nov. F. D. maipokhariensis sp. nov. G. D. miriensis Godwin-Austen, 1917. H. D. munipurensis Godwin- Austen, 1892. I. D. oviformis Fulton, 1901. J. D. regularis Fulton, 1901. K. D. salgharica sp. nov. L. D. shivapuriensis sp. nov. M. D. silvicola Godwin-Austen, 1886. N. D. sperata W.T. Blanford, 1862. O. D. syabrubesiensis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3832502" httpUri="https://zenodo.org/record/3832502/files/figure.png" pageId="8" pageNumber="9">9D</figureCitation>
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– 
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– 
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<bibRefCitation id="EFFE03A6FFECFFE3FE504232D67DFAE9" author="Godwin-Austen H. H." box="[510,815,1360,1386]" pageId="8" pageNumber="9" pagination="83 - 84" refId="ref13218" refString="Godwin-Austen H. H. 1868. Figures of the species of Diplommatina, Benson, hitherto described as inhabiting the Himalayas, Khasi Hills and Burma, with some additional forms from Darjiling and the Khasi Hills. Journal of the Asiatic Society of Bengal 37: 83 - 84." type="journal article" year="1868">Godwin-Austen 1868: 83</bibRefCitation>
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, pl.1, figs 1–2. — 
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<bibRefCitation id="EFFE03A6FFECFFE3FBBE4232D020FAE8" author="Hanley S. &amp; Theobald W." box="[1040,1394,1360,1387]" pageId="8" pageNumber="9" refId="ref13613" refString="Hanley S. &amp; Theobald W. 1876. Conchologia Indica; being illustrations of the land and freshwater shells of British India. London. https: // doi. org / 10.5962 / bhl. title. 14456" type="book" year="1876">Hanley &amp; Theobald 1876: 56</bibRefCitation>
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, pl. CXL, figs 8–9. — 
<treatmentCitation id="0ACE5846FFECFFE3FE134217D72CFA0C" author="Gude G. K." box="[445,638,1397,1424]" page="314" pageId="8" pageNumber="9" year="1921">
<bibRefCitation id="EFFE03A6FFECFFE3FE134217D72CFA0C" author="Gude G. K." box="[445,638,1397,1424]" pageId="8" pageNumber="9" refId="ref13576" refString="Gude G. K. 1921. The fauna of British India. Mollusca-III (Land operculates: Cyclophoridae, Truncatellidae, Assimineidae, Helicinidae). Today &amp; Tomorrow's Printers &amp; Publishers, New Delhi." type="book" year="1921">Gude 1921: 314</bibRefCitation>
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. — 
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<bibRefCitation id="EFFE03A6FFECFFE3FD034214D639FA13" author="Naggs F." box="[685,875,1397,1424]" pageId="8" pageNumber="9" pagination="37 - 88" refId="ref13882" refString="Naggs F. 1997. William Benson and the early study of land snail in British India and Ceylon. Archives of Natural History 24: 37 - 88. https: // doi. org / 10.3366 / anh. 1997.24.1.37" type="journal article" year="1997">Naggs 1997: 58</bibRefCitation>
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<emphasis id="B91BA245FFECFFE3FF1342F9D434FA36" box="[189,358,1435,1461]" italics="true" pageId="8" pageNumber="9">Diplommatina</emphasis>
(
<emphasis id="B91BA245FFECFFE3FED642FED4C6FA36" box="[376,404,1436,1461]" italics="true" pageId="8" pageNumber="9">D.</emphasis>
) 
<emphasis id="B91BA245FFECFFE3FE0B42F9D740FA36" box="[421,530,1435,1461]" italics="true" pageId="8" pageNumber="9">folliculus</emphasis>
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– 
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<bibRefCitation id="EFFE03A6FFECFFE3FD9E42F9D62DFA36" author="Ramakrishna &amp; Mitra S. C. &amp; Dey A." box="[560,895,1435,1461]" pageId="8" pageNumber="9" pagination="1 - 359" refId="ref14255" refString="Ramakrishna, Mitra S. C. &amp; Dey A. 2010. Annotated checklist of Indian land molluscs. Records of Zoological Survey of India, Occasional Paper 306: 1 - 359." type="journal article" year="2010">
Ramakrishna 
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2010: 86
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<emphasis id="B91BA245FFECFFE3FF1342A2D434FA59" box="[189,358,1472,1498]" italics="true" pageId="8" pageNumber="9">Diplommatina</emphasis>
(
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– 
<treatmentCitation id="0ACE5846FFECFFE3FD1342A2D6DCFA58" box="[701,910,1472,1499]" pageId="8" pageNumber="9">
<bibRefCitation id="EFFE03A6FFECFFE3FD1342A2D6DCFA58" author="Budha P. B. &amp; Naggs F. &amp; Backeljau T." box="[701,910,1472,1499]" pageId="8" pageNumber="9" pagination="1 - 48" refId="ref13118" refString="Budha P. B., Naggs F. &amp; Backeljau T. 2015. Annotated checklist of the terrestrial gastropods of Nepal. ZooKeys 492: 1 - 48. https: // doi. org / 10.3897 / zookeys. 492.9175" type="journal article" year="2015">
Budha 
<emphasis id="B91BA245FFECFFE3FCBF42A3D619FA59" box="[785,843,1472,1498]" italics="true" pageId="8" pageNumber="9">et al.</emphasis>
2015
</bibRefCitation>
</treatmentCitation>
: p. 6.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3752DDCFFECFFE3FF134166D445F868" pageId="8" pageNumber="9" type="diagnosis">
<paragraph id="8BD07E57FFECFFE3FF134166D464F99D" blockId="8.[189,310,1540,1566]" box="[189,310,1540,1566]" pageId="8" pageNumber="9">
<heading id="D098C93BFFECFFE3FF134166D464F99D" bold="true" box="[189,310,1540,1566]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="B91BA245FFECFFE3FF134166D464F99D" bold="true" box="[189,310,1540,1566]" pageId="8" pageNumber="9">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BD07E57FFECFFE3FF134157D445F868" blockId="8.[189,1399,1588,2027]" pageId="8" pageNumber="9">
Measurements (n = 15): SH 2.6–3.0 mm, SW 
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, Wh 6.5–7.0. Dextral, penultimate whorl slightly wider than the body whorl. Constriction at the middle of the parietal wall (
<figureCitation id="135462D2FFECFFE3FB3E4138D1BDF9F7" box="[1168,1263,1626,1652]" captionStart="Fig" captionStartId="24.[189,232,1789,1815]" captionTargetBox="[297,1297,294,1724]" captionTargetId="figure@24.[247,1340,265,1748]" captionTargetPageId="24" captionText="Fig. 9. Palatal lamellae (palatalis) of species of Diplommatina. Dotted lines indicate the position of the constriction. Dark lines indicate the palatalis. A. D. abiesiana sp. nov. B. D. exserta Godwin- Austen, 1886. C. D. fistulata sp. nov. D. D. folliculus (L. Pfeiffer, 1846). E. D. godawariensis sp. nov. F. D. maipokhariensis sp. nov. G. D. miriensis Godwin-Austen, 1917. H. D. munipurensis Godwin- Austen, 1892. I. D. oviformis Fulton, 1901. J. D. regularis Fulton, 1901. K. D. salgharica sp. nov. L. D. shivapuriensis sp. nov. M. D. silvicola Godwin-Austen, 1886. N. D. sperata W.T. Blanford, 1862. O. D. syabrubesiensis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3832502" httpUri="https://zenodo.org/record/3832502/files/figure.png" pageId="8" pageNumber="9">Fig. 9D</figureCitation>
). Internally with two parietal lamellae, both Prt1 and Prt2 very short, prominent at the lodging edge of the operculum (
<figureCitation id="135462D2FFECFFE3FF6B41C7D474F93C" box="[197,294,1701,1727]" captionStart="Fig" captionStartId="4.[189,232,1895,1921]" captionTargetBox="[189,1398,285,1853]" captionTargetId="figure@4.[189,1398,285,1853]" captionTargetPageId="4" captionText="Fig. 2. Shell terminology. A. D. regularis Fulton, 1901. B. D. folliculus (L. Pfeiffer, 1846). C. General outline of the lower part of a shell from Diplommatina species. Abbreviations: Prt = Parietalis (Prt1 = inner parietalis, Prt2 = outer parietalis); Pt = Palatalis; Hpt = Horizontal palatalis (Hpt1 = lower, Hpt2 = upper in case of two); Vpt = Vertical palatalis (Vpt1 = left side, Vpt2 = right side in case of two)." figureDoi="http://doi.org/10.5281/zenodo.3832488" httpUri="https://zenodo.org/record/3832488/files/figure.png" pageId="8" pageNumber="9">Figs 2B</figureCitation>
, 
<figureCitation id="135462D2FFECFFE3FE9D41C7D409F93C" box="[307,347,1701,1727]" captionStart="Fig" captionStartId="9.[189,232,1930,1956]" captionTargetBox="[189,1398,274,1888]" captionTargetId="figure@9.[189,1398,266,1891]" captionTargetPageId="9" captionText="Fig. 4. Diplommatina Benson, 1849 of Nepal. A. D. folliculus (L. Pfeiffer, 1846) (CDZMTU028). B. D. godawariensis sp. nov., holotype (CDZMTU032). C. D. maipokhariensis sp. nov., holotype (CDZMTU034). Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832492" httpUri="https://zenodo.org/record/3832492/files/figure.png" pageId="8" pageNumber="9">4A</figureCitation>
<figureCitation id="135462D2FFECFFE3FEF241C7D438F93C" box="[348,362,1701,1727]" captionStart="Fig" captionStartId="6.[189,232,1930,1956]" captionTargetBox="[189,1393,274,1888]" captionTargetId="figure@6.[189,1398,269,1893]" captionTargetPageId="6" captionText="Fig. 3. Diplommatina Benson, 1849 of Nepal. A. D. abiesiana sp. nov., holotype (CDZMTU020). B. D. exserta Godwin-Austen, 1886 (CDZMTU024). C. D. fistulata sp. nov., holotype (CDZMTU025). The yellow arrow indicates the dorsal fold. Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832490" httpUri="https://zenodo.org/record/3832490/files/figure.png" pageId="8" pageNumber="9">3</figureCitation>
). Two vertical palatal lamellae: Vpt1 along the constriction and Vpt2 short, but running parallel to Vpt1. No horizontal palatal lamella (
<figureCitation id="135462D2FFECFFE3FD5541A8D60EF966" box="[763,860,1738,1765]" captionStart="Fig" captionStartId="24.[189,232,1789,1815]" captionTargetBox="[297,1297,294,1724]" captionTargetId="figure@24.[247,1340,265,1748]" captionTargetPageId="24" captionText="Fig. 9. Palatal lamellae (palatalis) of species of Diplommatina. Dotted lines indicate the position of the constriction. Dark lines indicate the palatalis. A. D. abiesiana sp. nov. B. D. exserta Godwin- Austen, 1886. C. D. fistulata sp. nov. D. D. folliculus (L. Pfeiffer, 1846). E. D. godawariensis sp. nov. F. D. maipokhariensis sp. nov. G. D. miriensis Godwin-Austen, 1917. H. D. munipurensis Godwin- Austen, 1892. I. D. oviformis Fulton, 1901. J. D. regularis Fulton, 1901. K. D. salgharica sp. nov. L. D. shivapuriensis sp. nov. M. D. silvicola Godwin-Austen, 1886. N. D. sperata W.T. Blanford, 1862. O. D. syabrubesiensis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3832502" httpUri="https://zenodo.org/record/3832502/files/figure.png" pageId="8" pageNumber="9">Fig. 9D</figureCitation>
). Columellaris thin. Columellar tooth visible in the aperture. Protoconch smooth, Wh 1.5–2.0, with fine pits. Teleoconch with radial ribs visible only after the second whorl; widely spaced radial ribs; 6–7 ribs/ 
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on the penultimate whorl; 4–6 ribs/ 
<quantity id="4C97D3B2FFECFFE3FF5C4059D419F8D6" box="[242,331,1851,1878]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="9" unit="mm" value="0.5">0.5 mm</quantity>
on the body whorl towards the aperture (
<figureCitation id="135462D2FFECFFE3FC814059D6DFF8D6" box="[815,909,1851,1877]" captionStart="Fig" captionStartId="9.[189,232,1930,1956]" captionTargetBox="[189,1398,274,1888]" captionTargetId="figure@9.[189,1398,266,1891]" captionTargetPageId="9" captionText="Fig. 4. Diplommatina Benson, 1849 of Nepal. A. D. folliculus (L. Pfeiffer, 1846) (CDZMTU028). B. D. godawariensis sp. nov., holotype (CDZMTU032). C. D. maipokhariensis sp. nov., holotype (CDZMTU034). Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832492" httpUri="https://zenodo.org/record/3832492/files/figure.png" pageId="8" pageNumber="9">Fig. 4A</figureCitation>
<figureCitation id="135462D2FFECFFE3FC234059D6CEF8D6" box="[909,924,1851,1877]" captionStart="Fig" captionStartId="4.[189,232,1895,1921]" captionTargetBox="[189,1398,285,1853]" captionTargetId="figure@4.[189,1398,285,1853]" captionTargetPageId="4" captionText="Fig. 2. Shell terminology. A. D. regularis Fulton, 1901. B. D. folliculus (L. Pfeiffer, 1846). C. General outline of the lower part of a shell from Diplommatina species. Abbreviations: Prt = Parietalis (Prt1 = inner parietalis, Prt2 = outer parietalis); Pt = Palatalis; Hpt = Horizontal palatalis (Hpt1 = lower, Hpt2 = upper in case of two); Vpt = Vertical palatalis (Vpt1 = left side, Vpt2 = right side in case of two)." figureDoi="http://doi.org/10.5281/zenodo.3832488" httpUri="https://zenodo.org/record/3832488/files/figure.png" pageId="8" pageNumber="9">2</figureCitation>
, A 
<figureCitation id="135462D2FFECFFE3FC104059D69CF8D6" box="[958,974,1851,1877]" captionStart="Fig" captionStartId="9.[189,232,1930,1956]" captionTargetBox="[189,1398,274,1888]" captionTargetId="figure@9.[189,1398,266,1891]" captionTargetPageId="9" captionText="Fig. 4. Diplommatina Benson, 1849 of Nepal. A. D. folliculus (L. Pfeiffer, 1846) (CDZMTU028). B. D. godawariensis sp. nov., holotype (CDZMTU032). C. D. maipokhariensis sp. nov., holotype (CDZMTU034). Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832492" httpUri="https://zenodo.org/record/3832492/files/figure.png" pageId="8" pageNumber="9">4</figureCitation>
). Spiral striation distinct. Umbilicus closed. Aperture circular. Peristome with two lips and expanded; lips widely spaced, sinuous at the angular edge. Outer lip slightly reflected. Operculum paucispiral, corneous, thin, transparent, slightly concave, outer surface smooth, inner surface with peripheral circular edge which is thicker at columellar margin.
</paragraph>
</subSubSection>
<caption id="DF102EDFFFEDFFE2FF1340E8D15DF868" ID-DOI="http://doi.org/10.5281/zenodo.3832492" ID-Zenodo-Dep="3832492" httpUri="https://zenodo.org/record/3832492/files/figure.png" pageId="9" pageNumber="10" startId="9.[189,232,1930,1956]" targetBox="[189,1398,274,1888]" targetPageId="9">
<paragraph id="8BD07E57FFEDFFE2FF1340E8D15DF868" blockId="9.[189,1399,1930,2027]" pageId="9" pageNumber="10">
<emphasis id="B91BA245FFEDFFE2FF1340E8D45DF827" bold="true" box="[189,271,1930,1956]" pageId="9" pageNumber="10">Fig. 4.</emphasis>
<taxonomicName id="4C6F05D4FFEDFFE2FEB540E8D724F827" authority="Benson, 1849" authorityName="Benson" authorityYear="1849" box="[283,630,1930,1956]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="9" pageNumber="10" phylum="Mollusca" rank="genus">
<emphasis id="B91BA245FFEDFFE2FEB540E8D496F827" box="[283,452,1930,1956]" italics="true" pageId="9" pageNumber="10">Diplommatina</emphasis>
<bibRefCitation id="EFFE03A6FFEDFFE2FE7E40E8D724F827" author="Benson W. H." box="[464,630,1930,1956]" pageId="9" pageNumber="10" pagination="193 - 195" refId="ref12623" refString="Benson W. H. 1849. Characters of Diplommatina, a new genus of terrestrial mollusks belonging to the family of Carychiadae, and of a second species contained in it; also of a new species of Carychium inhabiting the Western Himalaya. Annals and Magazine of Natural History, Series 2 4: 193 - 195. https: // doi. org / 10.1080 / 03745486009494813" type="journal article" year="1849">Benson, 1849</bibRefCitation>
</taxonomicName>
of Nepal. 
<emphasis id="B91BA245FFEDFFE2FCAE40E8D647F827" bold="true" box="[768,789,1930,1956]" pageId="9" pageNumber="10">A</emphasis>
. 
<taxonomicName id="4C6F05D4FFEDFFE2FC8640E9D1FAF826" authority="(L. Pfeiffer, 1846)" baseAuthorityName="L. Pfeiffer" baseAuthorityYear="1846" box="[808,1192,1930,1957]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="folliculus">
<emphasis id="B91BA245FFEDFFE2FC8640E9D6EFF827" box="[808,957,1930,1956]" italics="true" pageId="9" pageNumber="10">D. folliculus</emphasis>
(L. 
<bibRefCitation id="EFFE03A6FFEDFFE2FC5640E8D1CDF826" author="Pfeiffer L." box="[1016,1183,1930,1957]" pageId="9" pageNumber="10" refId="ref14017" refString="Pfeiffer L. 1846. Symbolae ad Historiam Heliceorum. Fischer Verlag, Kassel, Germany. https: // doi. org / 10.5962 / bhl. title. 11903" type="book" year="1846">Pfeiffer, 1846</bibRefCitation>
)
</taxonomicName>
(CDZMTU028). 
<emphasis id="B91BA245FFEDFFE2FF1340CFD583F844" bold="true" box="[189,209,1965,1991]" pageId="9" pageNumber="10">B</emphasis>
. 
<taxonomicName id="4C6F05D4FFEDFFE2FF4B40CCD4EBF844" authority="Budha &amp; Naggs &amp; Backeljau, 2017" authorityName="Budha &amp; Naggs &amp; Backeljau" authorityYear="2017" box="[229,441,1965,1991]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="godawariensis" status="sp. nov.">
<emphasis id="B91BA245FFEDFFE2FF4B40CCD4EBF844" box="[229,441,1965,1991]" italics="true" pageId="9" pageNumber="10">D. godawariensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2281F3EFFEDFFE2FE6B40CCD77AF84B" box="[453,552,1966,1992]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
, holotype (CDZMTU032). 
<emphasis id="B91BA245FFEDFFE2FCD540CFD6C2F844" bold="true" box="[891,912,1965,1991]" pageId="9" pageNumber="10">C</emphasis>
. 
<taxonomicName id="4C6F05D4FFEDFFE2FC0B40CCD1C2F844" authority="Budha &amp; Naggs &amp; Backeljau, 2017" authorityName="Budha &amp; Naggs &amp; Backeljau" authorityYear="2017" box="[933,1168,1965,1991]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="maipokhariensis" status="sp. nov.">
<emphasis id="B91BA245FFEDFFE2FC0B40CCD1C2F844" box="[933,1168,1965,1991]" italics="true" pageId="9" pageNumber="10">D. maipokhariensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2281F3EFFEDFFE2FB3240CCD1ADF84B" box="[1180,1279,1966,1992]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
, holotype (CDZMTU034). Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm.
</paragraph>
</caption>
<subSubSection id="C3752DDCFFEEFFE1FF134669D4E1FDE5" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="8BD07E57FFEEFFE1FF134669D4F8FEA6" blockId="10.[189,426,267,293]" box="[189,426,267,293]" pageId="10" pageNumber="11">
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<emphasis id="B91BA245FFEEFFE1FF134669D4F8FEA6" bold="true" box="[189,426,267,293]" pageId="10" pageNumber="11">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BD07E57FFEEFFE1FF13465FD4E1FDE5" blockId="10.[189,1399,317,614]" pageId="10" pageNumber="11">
<collectingCountry id="F3783EC7FFEEFFE1FF13465FD44CFED4" box="[189,286,317,343]" name="Nepal" pageId="10" pageNumber="11">NEPAL</collectingCountry>
: 80+ shells, Phulchowki Hill, Lalitpur District, 
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, 
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, 
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, 
<date id="FFD15897FFEEFFE1FF134606D429FEFD" box="[189,379,356,382]" pageId="10" pageNumber="11" value="2007-05-03" valueMax="2007-05-11" valueMin="2007-05-03">3–11 May 2007</date>
, P.B. Budha leg. (
<collectionCode id="ED7EE692FFEEFFE1FDE04606D783FEFD" box="[590,721,356,382]" pageId="10" pageNumber="11">CDZMTU</collectionCode>
028); 95 shells, Shivapuri-Nagarjun National Park, 
<quantity id="4C97D3B2FFEEFFE1FA834606D448FE26" metricMagnitude="3" metricUnit="m" metricValue="2.5149999999999997" metricValueMax="2.716" metricValueMin="2.314" pageId="10" pageNumber="11" unit="m" value="2515.0" valueMax="2716.0" valueMin="2314.0">2314– 2716 m</quantity>
, 
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, 
<geoCoordinate id="EE5B1890FFEEFFE1FE7846E8D723FE26" box="[470,625,394,421]" degrees="85.378807" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="1" value="85.37881">85.378807 E</geoCoordinate>
, 
<date id="FFD15897FFEEFFE1FD2E46E9D618FE26" box="[640,842,395,421]" pageId="10" pageNumber="11" value="2007-06-24" valueMax="2007-06-21" valueMin="2007-06-24">24–21 Jun. 2007</date>
, P.B. Budha leg. (
<collectionCode id="ED7EE692FFEEFFE1FB8B46E9D1F5FE26" box="[1061,1191,395,421]" pageId="10" pageNumber="11">CDZMTU</collectionCode>
029); 100+ shells, Shivapuri-Nagarjun National Park, 
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, 
<geoCoordinate id="EE5B1890FFEEFFE1FCB746D3D6E5FE4F" box="[793,951,433,460]" degrees="27.812591" direction="north" orientation="latitude" pageId="10" pageNumber="11" precision="1" value="27.812592">27.812591 N</geoCoordinate>
, 
<geoCoordinate id="EE5B1890FFEEFFE1FC6B46D3D10DFE4F" box="[965,1119,433,460]" degrees="85.378807" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="1" value="85.37881">85.378807 E</geoCoordinate>
, 
<date id="FFD15897FFEEFFE1FBC046D3D068FE48" box="[1134,1338,433,459]" pageId="10" pageNumber="11" value="2008-04-24" valueMax="2008-04-26" valueMin="2008-04-24">24–26 Apr. 2008</date>
, P.B. Budha leg. (
<collectionCode id="ED7EE692FFEEFFE1FEFF46BAD481FE70" box="[337,467,472,499]" pageId="10" pageNumber="11">CDZMTU</collectionCode>
029b); 2 shells, Raniban, Balaju, Kathmandu, 
<quantity id="4C97D3B2FFEEFFE1FBAE46BAD10CFE70" box="[1024,1118,472,499]" metricMagnitude="3" metricUnit="m" metricValue="1.373" pageId="10" pageNumber="11" unit="m" value="1373.0">1373 m</quantity>
, 
<geoCoordinate id="EE5B1890FFEEFFE1FBC346BAD1BDFE70" box="[1133,1263,472,499]" degrees="27.7426" direction="north" orientation="latitude" pageId="10" pageNumber="11" precision="5" value="27.7426">27.7426 N</geoCoordinate>
, 
<geoCoordinate id="EE5B1890FFEEFFE1FB5046BAD020FE70" box="[1278,1394,472,499]" degrees="85.2847" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="5" value="85.2847">85.2847E</geoCoordinate>
, 
<date id="FFD15897FFEEFFE1FF13469DD434FD9A" box="[189,358,511,537]" pageId="10" pageNumber="11" value="2009-03-12">12 Mar. 2009</date>
, Srijana Khanal leg. (
<collectionCode id="ED7EE692FFEEFFE1FDDC469DD7A6FD9A" box="[626,756,511,537]" pageId="10" pageNumber="11">CDZMTU</collectionCode>
030); 100+ shells, Dhunche, Langtang National Park, 
<quantity id="4C97D3B2FFEEFFE1FF134547D430FDC3" box="[189,354,549,576]" metricMagnitude="3" metricUnit="m" metricValue="2.3594999999999997" metricValueMax="2.663" metricValueMin="2.056" pageId="10" pageNumber="11" unit="m" value="2359.5" valueMax="2663.0" valueMin="2056.0">2056–2663 m</quantity>
, 
<geoCoordinate id="EE5B1890FFEEFFE1FEDE4547D75EFDC3" box="[368,524,549,576]" degrees="28.106128" direction="north" orientation="latitude" pageId="10" pageNumber="11" precision="1" value="28.106129">28.106128 N</geoCoordinate>
, 
<geoCoordinate id="EE5B1890FFEEFFE1FDB74547D7E0FDC3" box="[537,690,549,576]" degrees="85.303259" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="1" value="85.30326">85.303259 E</geoCoordinate>
, 
<date id="FFD15897FFEEFFE1FD6E4544D618FDC3" box="[704,842,550,576]" pageId="10" pageNumber="11" value="2007-06-02">2 Jun. 2007</date>
and 
<date id="FFD15897FFEEFFE1FC2A4544D177FDC3" box="[900,1061,550,576]" pageId="10" pageNumber="11" value="2008-05-11">11 May 2008</date>
, P.B. Budha and R. Devkota leg. (
<collectionCode id="ED7EE692FFEEFFE1FF56452ED42EFDE5" box="[248,380,588,614]" pageId="10" pageNumber="11">CDZMTU</collectionCode>
031).
</paragraph>
</subSubSection>
<subSubSection id="C3752DDCFFEEFFE1FF1345F0D1E9FCD1" pageId="10" pageNumber="11" type="discussion">
<paragraph id="8BD07E57FFEEFFE1FF1345F0D47DFD2F" blockId="10.[189,303,658,684]" box="[189,303,658,684]" pageId="10" pageNumber="11">
<heading id="D098C93BFFEEFFE1FF1345F0D47DFD2F" bold="true" box="[189,303,658,684]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="B91BA245FFEEFFE1FF1345F0D47DFD2F" bold="true" box="[189,303,658,684]" pageId="10" pageNumber="11">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BD07E57FFEEFFE1FF1345A1D1E9FCD1" blockId="10.[189,1399,707,850]" pageId="10" pageNumber="11">
<taxonomicName id="4C6F05D4FFEEFFE1FF1345A1D488FD5E" baseAuthorityName="L. Pfeiffer" baseAuthorityYear="1846" box="[189,474,707,733]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="folliculus">
<emphasis id="B91BA245FFEEFFE1FF1345A1D488FD5E" box="[189,474,707,733]" italics="true" pageId="10" pageNumber="11">Diplommatina folliculus</emphasis>
</taxonomicName>
is a very common, abundant and distinct species that can be distinguished from the others by its strong, widely-spaced lips and costulated radial ribs. It is distributed from the western to the eastern Himalaya. The most similar species is 
<taxonomicName id="4C6F05D4FFEEFFE1FC8F4470D1E1FCA8" authority="Godwin-Austen, 1886" authorityName="Godwin-Austen" authorityYear="1886" box="[801,1203,785,811]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="silvicola">
<emphasis id="B91BA245FFEEFFE1FC8F4470D6FAFCA8" box="[801,936,785,811]" italics="true" pageId="10" pageNumber="11">D. silvicola</emphasis>
<bibRefCitation id="EFFE03A6FFEEFFE1FC014473D1E1FCA8" author="Godwin-Austen H. H." box="[943,1203,785,811]" pageId="10" pageNumber="11" pagination="165 - 185" refId="ref13270" refString="Godwin-Austen H. H. 1886. Land and freshwater Mollusca of India. Conchologia Indica 1 (5): 165 - 185." type="journal article" year="1886">Godwin-Austen, 1886</bibRefCitation>
</taxonomicName>
(pl. LXV, figs 3, 3a), but 
<taxonomicName id="4C6F05D4FFEEFFE1FEB0445BD4FDFCD1" baseAuthorityName="L. Pfeiffer" baseAuthorityYear="1846" box="[286,431,824,850]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="folliculus">
<emphasis id="B91BA245FFEEFFE1FEB0445BD4FDFCD1" box="[286,431,824,850]" italics="true" pageId="10" pageNumber="11">D. folliculus</emphasis>
</taxonomicName>
differs by its more slender shell and less distinct columellar tooth.
</paragraph>
</subSubSection>
</treatment>
</document>