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<document id="57C2FA53193AFDCADDB139B625784A39" ID-CLB-Dataset="22067" ID-DOI="10.5852/ejt.2017.337" ID-GBIF-Dataset="5a7f73c5-5646-446f-a72c-3ea18d8f634f" ID-ISSN="2118-9773" ID-Zenodo-Dep="3832483" ID-ZooBank="2669CB69-5064-4B5D-AC25-856680E3B9AA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589830819450" checkinUser="carolina" docAuthor="Budha, Prem B., Naggs, Fred &amp; Backeljau, Thierry" docDate="2017" docId="03C6CF41FFE8FFE7FDAF4669D485FA69" docLanguage="en" docName="ejt-2017-337.pdf.imf" docOrigin="European Journal of Taxonomy 337" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Diplommatina miriensis Godwin-Austen 1917" docType="treatment" docVersion="6" lastPageNumber="13" masterDocId="FFFFB739FFE4FFEBFFAE4762D552FF83" masterDocTitle="The genus Diplommatina Benson, 1849 (Gastropoda: Caenogastropoda: Diplommatinidae) in Nepal, with the description of seven new species" masterLastPageNumber="30" masterPageNumber="1" pageNumber="13" updateTime="1698840088040" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="A7E9E5704FFA8A8FC7D5F0193850C4E7" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="9CB336B18EE70B90E36D4118A0149B63">The genus Diplommatina Benson, 1849 (Gastropoda: Caenogastropoda: Diplommatinidae) in Nepal, with the description of seven new species</mods:title>
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<mods:roleTerm id="B1E81D10ABD883AFFE0BD9EE8F2A3825">Author</mods:roleTerm>
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<mods:namePart id="3B22B7E2D0A156160AFA9317AFB02411">Budha, Prem B.</mods:namePart>
<mods:affiliation id="07100FE35974D8850BBA0397489A9708">University of Antwerp, Evolutionary Ecology Group, Universiteitsplein 1, B- 2610, Antwerp, Belgium. &amp; Central Department of Zoology, Tribhuvan University, Kirtipur, Kathmandu, Nepal. &amp; Corresponding author: prembudha @ yahoo. com</mods:affiliation>
<mods:nameIdentifier id="AED8C2D65B151C5E574606FADB8BDFFB" type="email">prembudha@yahoo.com</mods:nameIdentifier>
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<mods:namePart id="31F39FBA78EC2FF747987113E041B81E">Naggs, Fred</mods:namePart>
<mods:affiliation id="826EB4CAECD4D648C8972006D82B05F0">Natural History Museum, Cromwell Road, London, SW 7 5 BD, UK. &amp; Email: f. naggs @ nhm. ac. uk</mods:affiliation>
<mods:nameIdentifier id="E50A522463CC7688A026E15D144EBD64" type="email">f.naggs@nhm.ac.uk</mods:nameIdentifier>
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<mods:namePart id="A9483608722916B72D3A3B78FA3663DF">Backeljau, Thierry</mods:namePart>
<mods:affiliation id="B3DEC475C239F3E9A53E42028C562B0C">University of Antwerp, Evolutionary Ecology Group, Universiteitsplein 1, B- 2610, Antwerp, Belgium. &amp; Royal Belgian Institute of Natural Sciences, Vautierstraat 29, B- 1000, Brussels, Belgium. &amp; Email: thierry. backeljau @ naturalsciences. be</mods:affiliation>
<mods:nameIdentifier id="BD9B50DD237CE76FB04A6368A67ED296" type="email">thierry.backeljau@naturalsciences.be</mods:nameIdentifier>
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<mods:title id="C20774077FBCEFFF848C67E07F90A2F8">European Journal of Taxonomy</mods:title>
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<mods:date id="45970D9B25155F834697C08321443A94">2017</mods:date>
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<mods:number id="FF3A9FE422B461EF52337658154A5503">2017-07-21</mods:number>
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<treatment id="03C6CF41FFE8FFE7FDAF4669D485FA69" ID-DOI="http://doi.org/10.5281/zenodo.3850706" ID-GBIF-Taxon="164167132" ID-Zenodo-Dep="3850706" LSID="urn:lsid:plazi:treatment:03C6CF41FFE8FFE7FDAF4669D485FA69" httpUri="http://treatment.plazi.org/id/03C6CF41FFE8FFE7FDAF4669D485FA69" lastPageNumber="13" pageId="12" pageNumber="13">
<subSubSection id="C3752DDCFFE8FFE7FDAF4669D160FEA6" box="[513,1074,267,294]" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="8BD07E57FFE8FFE7FDAF4669D160FEA6" blockId="12.[513,1074,267,330]" box="[513,1074,267,294]" pageId="12" pageNumber="13">
<heading id="D098C93BFFE8FFE7FDAF4669D160FEA6" box="[513,1074,267,294]" centered="true" fontSize="11" level="2" pageId="12" pageNumber="13" reason="2">
<taxonomicName id="4C6F05D4FFE8FFE7FDAF4669D160FEA6" ID-CoL="36M37" authority="Godwin-Austen, 1917" authorityName="Godwin-Austen" authorityYear="1917" box="[513,1074,267,294]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="miriensis">
<emphasis id="B91BA245FFE8FFE7FDAF4669D674FEA6" bold="true" box="[513,806,267,294]" italics="true" pageId="12" pageNumber="13">Diplommatina miriensis</emphasis>
<bibRefCitation id="EFFE03A6FFE8FFE7FC834669D160FEA6" author="Godwin-Austen H. H." box="[813,1074,267,293]" pageId="12" pageNumber="13" pagination="547 - 559" refId="ref13391" refString="Godwin-Austen H. H. 1917. Zoological results of the Abor Expedition, Mollusca 6. Records of the Indian Museum 8: 547 - 559." type="journal article" year="1917">Godwin-Austen, 1917</bibRefCitation>
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<subSubSection id="C3752DDCFFE8FFE7FD7C464DD630FECA" box="[722,866,303,330]" pageId="12" pageNumber="13" type="description">
<paragraph id="8BD07E57FFE8FFE7FD7C464DD630FECA" blockId="12.[513,1074,267,330]" box="[722,866,303,330]" pageId="12" pageNumber="13">
<figureCitation id="135462D2FFE8FFE7FD7C464DD660FEC9" box="[722,818,303,330]" captionStart="Fig" captionStartId="13.[189,232,1930,1956]" captionTargetBox="[189,1398,265,1879]" captionTargetId="figure@13.[189,1398,261,1881]" captionTargetPageId="13" captionText="Fig. 5. Diplommatina Benson, 1849 of Nepal. A. D. miriensis Godwin-Austen, 1917 (CDZMTU036). B. D.munipurensis Godwin-Austen, 1892 (CDZMTU039).C. D.oviformis Fulton, 1901 (CDZMTU042). Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832494" httpUri="https://zenodo.org/record/3832494/files/figure.png" pageId="12" pageNumber="13">Figs 5A</figureCitation>
,
<figureCitation id="135462D2FFE8FFE7FC90464DD630FECA" box="[830,866,303,329]" captionStart="Fig" captionStartId="24.[189,232,1789,1815]" captionTargetBox="[297,1297,294,1724]" captionTargetId="figure@24.[247,1340,265,1748]" captionTargetPageId="24" captionText="Fig. 9. Palatal lamellae (palatalis) of species of Diplommatina. Dotted lines indicate the position of the constriction. Dark lines indicate the palatalis. A. D. abiesiana sp. nov. B. D. exserta Godwin- Austen, 1886. C. D. fistulata sp. nov. D. D. folliculus (L. Pfeiffer, 1846). E. D. godawariensis sp. nov. F. D. maipokhariensis sp. nov. G. D. miriensis Godwin-Austen, 1917. H. D. munipurensis Godwin- Austen, 1892. I. D. oviformis Fulton, 1901. J. D. regularis Fulton, 1901. K. D. salgharica sp. nov. L. D. shivapuriensis sp. nov. M. D. silvicola Godwin-Austen, 1886. N. D. sperata W.T. Blanford, 1862. O. D. syabrubesiensis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3832502" httpUri="https://zenodo.org/record/3832502/files/figure.png" pageId="12" pageNumber="13">9G</figureCitation>
</paragraph>
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<paragraph id="8BD07E57FFE8FFE7FF134613D6DBFE08" blockId="12.[189,905,369,396]" box="[189,905,369,396]" pageId="12" pageNumber="13">
<treatmentCitationGroup id="AB7F5979FFE8FFE7FF134613D6DBFE08" box="[189,905,369,396]" pageId="12" pageNumber="13">
<treatmentCitation id="0ACE5846FFE8FFE7FF134613D676FE0F" author="Godwin-Austen H. H." box="[189,804,369,396]" page="577" pageId="12" pageNumber="13" year="1917">
<taxonomicName id="4C6F05D4FFE8FFE7FF134613D676FE0F" ID-CoL="36M37" authority="Godwin-Austen, 1917: 577" authorityName="Godwin-Austen" authorityPageNumber="577" authorityYear="1917" box="[189,804,369,396]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="miriensis">
<emphasis id="B91BA245FFE8FFE7FF134613D488FE08" box="[189,474,369,395]" italics="true" pageId="12" pageNumber="13">Diplommatina miriensis</emphasis>
<bibRefCitation id="EFFE03A6FFE8FFE7FE4F4613D676FE0F" author="Godwin-Austen H. H." box="[481,804,369,396]" pageId="12" pageNumber="13" pagination="547 - 559" refId="ref13391" refString="Godwin-Austen H. H. 1917. Zoological results of the Abor Expedition, Mollusca 6. Records of the Indian Museum 8: 547 - 559." type="journal article" year="1917">Godwin-Austen, 1917: 577</bibRefCitation>
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, fig. 4D.
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</paragraph>
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<treatmentCitationGroup id="AB7F5979FFE8FFE7FF1346D1D4E6FE72" pageId="12" pageNumber="13">
<taxonomicName id="4C6F05D4FFE8FFE7FF1346D1D7C9FE4E" ID-CoL="36M37" authorityName="Godwin-Austen" authorityYear="1917" box="[189,667,435,461]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="miriensis" subGenus="Metadiancta">
<emphasis id="B91BA245FFE8FFE7FF1346D1D434FE4E" box="[189,358,435,461]" italics="true" pageId="12" pageNumber="13">Diplommatina</emphasis>
(
<emphasis id="B91BA245FFE8FFE7FE2E46D1D746FE4E" box="[384,532,435,461]" italics="true" pageId="12" pageNumber="13">Metadiancta</emphasis>
)
<emphasis id="B91BA245FFE8FFE7FD8146D6D7C9FE4E" box="[559,667,436,461]" italics="true" pageId="12" pageNumber="13">miriensis</emphasis>
</taxonomicName>
–
<treatmentCitation id="0ACE5846FFE8FFE7FD6446D1D6CEFE4E" box="[714,924,435,461]" pageId="12" pageNumber="13">
<bibRefCitation id="EFFE03A6FFE8FFE7FD6446D1D6CEFE4E" author="Gude G. K." box="[714,924,435,461]" pageId="12" pageNumber="13" refId="ref13576" refString="Gude G. K. 1921. The fauna of British India. Mollusca-III (Land operculates: Cyclophoridae, Truncatellidae, Assimineidae, Helicinidae). Today &amp; Tomorrow's Printers &amp; Publishers, New Delhi." type="book" year="1921">Gude 1921, 342</bibRefCitation>
</treatmentCitation>
.
<emphasis id="B91BA245FFE8FFE7FC0146D6D69BFE4E" box="[943,969,436,461]" italics="true" pageId="12" pageNumber="13">—</emphasis>
<treatmentCitation id="0ACE5846FFE8FFE7FC7446D1D01AFE4D" author="Ramakrishna &amp; Mitra S. C. &amp; Dey A." box="[986,1352,435,462]" page="95" pageId="12" pageNumber="13" year="2010">
<bibRefCitation id="EFFE03A6FFE8FFE7FC7446D1D01AFE4D" author="Ramakrishna &amp; Mitra S. C. &amp; Dey A." box="[986,1352,435,462]" pageId="12" pageNumber="13" pagination="1 - 359" refId="ref14255" refString="Ramakrishna, Mitra S. C. &amp; Dey A. 2010. Annotated checklist of Indian land molluscs. Records of Zoological Survey of India, Occasional Paper 306: 1 - 359." type="journal article" year="2010">
Ramakrishna
<emphasis id="B91BA245FFE8FFE7FB2B46D6D19AFE4E" box="[1157,1224,435,461]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
2010: 95
</bibRefCitation>
</treatmentCitation>
—
<treatmentCitation id="0ACE5846FFE8FFE7FF1346B5D4FCFE72" author="Budha P. B. &amp; Naggs F. &amp; Backeljau T." box="[189,430,470,497]" page="7" pageId="12" pageNumber="13" year="2015">
<bibRefCitation id="EFFE03A6FFE8FFE7FF1346B5D4FCFE72" author="Budha P. B. &amp; Naggs F. &amp; Backeljau T." box="[189,430,470,497]" pageId="12" pageNumber="13" pagination="1 - 48" refId="ref13118" refString="Budha P. B., Naggs F. &amp; Backeljau T. 2015. Annotated checklist of the terrestrial gastropods of Nepal. ZooKeys 492: 1 - 48. https: // doi. org / 10.3897 / zookeys. 492.9175" type="journal article" year="2015">
Budha
<emphasis id="B91BA245FFE8FFE7FEBF46BAD41EFE72" box="[273,332,471,497]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
2015: 7
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3752DDCFFE8FFE7FF13457AD194FC44" pageId="12" pageNumber="13" type="diagnosis">
<paragraph id="8BD07E57FFE8FFE7FF13457AD464FDB1" blockId="12.[189,310,536,562]" box="[189,310,536,562]" pageId="12" pageNumber="13">
<heading id="D098C93BFFE8FFE7FF13457AD464FDB1" bold="true" box="[189,310,536,562]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="B91BA245FFE8FFE7FF13457AD464FDB1" bold="true" box="[189,310,536,562]" pageId="12" pageNumber="13">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BD07E57FFE8FFE7FF13452AD194FC44" blockId="12.[189,1399,583,967]" pageId="12" pageNumber="13">
Measurements (n = 5): SH
<quantity id="4C97D3B2FFE8FFE7FE554525D7D9FDE1" box="[507,651,583,610]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="12" pageNumber="13" unit="mm" value="1.55" valueMax="1.6" valueMin="1.5">1.5–1.6 mm</quantity>
, SW 0.8–1.0 mm, Wh 5.5. Dextral, whorls regularly increasing, penultimate and body whorl nearly equal in width. Constriction at the middle of the parietal wall (
<figureCitation id="135462D2FFE8FFE7FF6845EDD476FD2A" box="[198,292,655,681]" captionStart="Fig" captionStartId="24.[189,232,1789,1815]" captionTargetBox="[297,1297,294,1724]" captionTargetId="figure@24.[247,1340,265,1748]" captionTargetPageId="24" captionText="Fig. 9. Palatal lamellae (palatalis) of species of Diplommatina. Dotted lines indicate the position of the constriction. Dark lines indicate the palatalis. A. D. abiesiana sp. nov. B. D. exserta Godwin- Austen, 1886. C. D. fistulata sp. nov. D. D. folliculus (L. Pfeiffer, 1846). E. D. godawariensis sp. nov. F. D. maipokhariensis sp. nov. G. D. miriensis Godwin-Austen, 1917. H. D. munipurensis Godwin- Austen, 1892. I. D. oviformis Fulton, 1901. J. D. regularis Fulton, 1901. K. D. salgharica sp. nov. L. D. shivapuriensis sp. nov. M. D. silvicola Godwin-Austen, 1886. N. D. sperata W.T. Blanford, 1862. O. D. syabrubesiensis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3832502" httpUri="https://zenodo.org/record/3832502/files/figure.png" pageId="12" pageNumber="13">Fig. 9G</figureCitation>
). Parietalis very weak, nearly invisible (
<figureCitation id="135462D2FFE8FFE7FD5645EDD605FD29" box="[760,855,655,682]" captionStart="Fig" captionStartId="13.[189,232,1930,1956]" captionTargetBox="[189,1398,265,1879]" captionTargetId="figure@13.[189,1398,261,1881]" captionTargetPageId="13" captionText="Fig. 5. Diplommatina Benson, 1849 of Nepal. A. D. miriensis Godwin-Austen, 1917 (CDZMTU036). B. D.munipurensis Godwin-Austen, 1892 (CDZMTU039).C. D.oviformis Fulton, 1901 (CDZMTU042). Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832494" httpUri="https://zenodo.org/record/3832494/files/figure.png" pageId="12" pageNumber="13">Fig. 5A</figureCitation>
<figureCitation id="135462D2FFE8FFE7FCF945EDD637FD29" box="[855,869,655,682]" captionStart="Fig" captionStartId="6.[189,232,1930,1956]" captionTargetBox="[189,1393,274,1888]" captionTargetId="figure@6.[189,1398,269,1893]" captionTargetPageId="6" captionText="Fig. 3. Diplommatina Benson, 1849 of Nepal. A. D. abiesiana sp. nov., holotype (CDZMTU020). B. D. exserta Godwin-Austen, 1886 (CDZMTU024). C. D. fistulata sp. nov., holotype (CDZMTU025). The yellow arrow indicates the dorsal fold. Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832490" httpUri="https://zenodo.org/record/3832490/files/figure.png" pageId="12" pageNumber="13">3</figureCitation>
). One vertical palatalis along the constriction and one very strong horizontal palatalis just above the columella (
<figureCitation id="135462D2FFE8FFE7FC1245D1D14BFD4E" box="[956,1049,691,717]" captionStart="Fig" captionStartId="24.[189,232,1789,1815]" captionTargetBox="[297,1297,294,1724]" captionTargetId="figure@24.[247,1340,265,1748]" captionTargetPageId="24" captionText="Fig. 9. Palatal lamellae (palatalis) of species of Diplommatina. Dotted lines indicate the position of the constriction. Dark lines indicate the palatalis. A. D. abiesiana sp. nov. B. D. exserta Godwin- Austen, 1886. C. D. fistulata sp. nov. D. D. folliculus (L. Pfeiffer, 1846). E. D. godawariensis sp. nov. F. D. maipokhariensis sp. nov. G. D. miriensis Godwin-Austen, 1917. H. D. munipurensis Godwin- Austen, 1892. I. D. oviformis Fulton, 1901. J. D. regularis Fulton, 1901. K. D. salgharica sp. nov. L. D. shivapuriensis sp. nov. M. D. silvicola Godwin-Austen, 1886. N. D. sperata W.T. Blanford, 1862. O. D. syabrubesiensis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3832502" httpUri="https://zenodo.org/record/3832502/files/figure.png" pageId="12" pageNumber="13">Fig. 9G</figureCitation>
). Columellaris thin and weak. Columellar tooth very weak to nearly absent. Protoconch smooth, Wh approx. 1.5, with very distinct small pits evenly distributed all over, no radial ribs (
<figureCitation id="135462D2FFE8FFE7FCB04598D62DFC96" box="[798,895,762,789]" captionStart="Fig" captionStartId="13.[189,232,1930,1956]" captionTargetBox="[189,1398,265,1879]" captionTargetId="figure@13.[189,1398,261,1881]" captionTargetPageId="13" captionText="Fig. 5. Diplommatina Benson, 1849 of Nepal. A. D. miriensis Godwin-Austen, 1917 (CDZMTU036). B. D.munipurensis Godwin-Austen, 1892 (CDZMTU039).C. D.oviformis Fulton, 1901 (CDZMTU042). Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832494" httpUri="https://zenodo.org/record/3832494/files/figure.png" pageId="12" pageNumber="13">Fig. 5A</figureCitation>
<figureCitation id="135462D2FFE8FFE7FCD04598D6DBFC96" box="[894,905,762,789]" captionStart="Fig" captionStartId="2.[189,235,1999,2025]" captionTargetBox="[189,1399,1129,1958]" captionTargetId="figure@2.[187,1401,1126,1960]" captionTargetPageId="2" captionText="Fig.1. Map of the localities and areas surveyed for Diplommatina spp. in Nepal." figureDoi="http://doi.org/10.5281/zenodo.3832486" httpUri="https://zenodo.org/record/3832486/files/figure.png" pageId="12" pageNumber="13">1</figureCitation>
). Teleoconch with dense radial ribs; about 10–13 ribs/
<quantity id="4C97D3B2FFE8FFE7FEED447CD4CFFCBA" box="[323,413,798,825]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="13" unit="mm" value="0.5">0.5 mm</quantity>
on the penultimate whorl; 8–9 ribs/
<quantity id="4C97D3B2FFE8FFE7FCEB447CD6CDFCBA" box="[837,927,798,825]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="13" unit="mm" value="0.5">0.5 mm</quantity>
on the body whorl towards the aperture (
<figureCitation id="135462D2FFE8FFE7FF6B4420D475FCDE" box="[197,295,834,861]" captionStart="Fig" captionStartId="13.[189,232,1930,1956]" captionTargetBox="[189,1398,265,1879]" captionTargetId="figure@13.[189,1398,261,1881]" captionTargetPageId="13" captionText="Fig. 5. Diplommatina Benson, 1849 of Nepal. A. D. miriensis Godwin-Austen, 1917 (CDZMTU036). B. D.munipurensis Godwin-Austen, 1892 (CDZMTU039).C. D.oviformis Fulton, 1901 (CDZMTU042). Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832494" httpUri="https://zenodo.org/record/3832494/files/figure.png" pageId="12" pageNumber="13">Figs 5A</figureCitation>
<figureCitation id="135462D2FFE8FFE7FE894420D464FCDE" box="[295,310,834,861]" captionStart="Fig" captionStartId="4.[189,232,1895,1921]" captionTargetBox="[189,1398,285,1853]" captionTargetId="figure@4.[189,1398,285,1853]" captionTargetPageId="4" captionText="Fig. 2. Shell terminology. A. D. regularis Fulton, 1901. B. D. folliculus (L. Pfeiffer, 1846). C. General outline of the lower part of a shell from Diplommatina species. Abbreviations: Prt = Parietalis (Prt1 = inner parietalis, Prt2 = outer parietalis); Pt = Palatalis; Hpt = Horizontal palatalis (Hpt1 = lower, Hpt2 = upper in case of two); Vpt = Vertical palatalis (Vpt1 = left side, Vpt2 = right side in case of two)." figureDoi="http://doi.org/10.5281/zenodo.3832488" httpUri="https://zenodo.org/record/3832488/files/figure.png" pageId="12" pageNumber="13">2</figureCitation>
,
<figureCitation id="135462D2FFE8FFE7FEEC4420D43BFCDE" box="[322,361,834,861]" captionStart="Fig" captionStartId="13.[189,232,1930,1956]" captionTargetBox="[189,1398,265,1879]" captionTargetId="figure@13.[189,1398,261,1881]" captionTargetPageId="13" captionText="Fig. 5. Diplommatina Benson, 1849 of Nepal. A. D. miriensis Godwin-Austen, 1917 (CDZMTU036). B. D.munipurensis Godwin-Austen, 1892 (CDZMTU039).C. D.oviformis Fulton, 1901 (CDZMTU042). Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832494" httpUri="https://zenodo.org/record/3832494/files/figure.png" pageId="12" pageNumber="13">5A</figureCitation>
<figureCitation id="135462D2FFE8FFE7FEC74420D42BFCDE" box="[361,377,834,861]" captionStart="Fig" captionStartId="9.[189,232,1930,1956]" captionTargetBox="[189,1398,274,1888]" captionTargetId="figure@9.[189,1398,266,1891]" captionTargetPageId="9" captionText="Fig. 4. Diplommatina Benson, 1849 of Nepal. A. D. folliculus (L. Pfeiffer, 1846) (CDZMTU028). B. D. godawariensis sp. nov., holotype (CDZMTU032). C. D. maipokhariensis sp. nov., holotype (CDZMTU034). Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832492" httpUri="https://zenodo.org/record/3832492/files/figure.png" pageId="12" pageNumber="13">4</figureCitation>
). Very fine spiral striation present. Umbilicus closed. Aperture circular. Peristome with two lips; under high magnification 6–10 ribs visible between the lips (
<figureCitation id="135462D2FFE8FFE7FC5F4407D11CFC03" box="[1009,1102,869,896]" captionStart="Fig" captionStartId="13.[189,232,1930,1956]" captionTargetBox="[189,1398,265,1879]" captionTargetId="figure@13.[189,1398,261,1881]" captionTargetPageId="13" captionText="Fig. 5. Diplommatina Benson, 1849 of Nepal. A. D. miriensis Godwin-Austen, 1917 (CDZMTU036). B. D.munipurensis Godwin-Austen, 1892 (CDZMTU039).C. D.oviformis Fulton, 1901 (CDZMTU042). Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832494" httpUri="https://zenodo.org/record/3832494/files/figure.png" pageId="12" pageNumber="13">Fig. 5A</figureCitation>
<figureCitation id="135462D2FFE8FFE7FBE04407D10CFC03" box="[1102,1118,869,896]" captionStart="Fig" captionStartId="9.[189,232,1930,1956]" captionTargetBox="[189,1398,274,1888]" captionTargetId="figure@9.[189,1398,266,1891]" captionTargetPageId="9" captionText="Fig. 4. Diplommatina Benson, 1849 of Nepal. A. D. folliculus (L. Pfeiffer, 1846) (CDZMTU028). B. D. godawariensis sp. nov., holotype (CDZMTU032). C. D. maipokhariensis sp. nov., holotype (CDZMTU034). Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832492" httpUri="https://zenodo.org/record/3832492/files/figure.png" pageId="12" pageNumber="13">4</figureCitation>
). Parietal wall with thin callus. Angular edge of the peristome curved (
<figureCitation id="135462D2FFE8FFE7FD7944EBD667FC27" box="[727,821,905,932]" captionStart="Fig" captionStartId="13.[189,232,1930,1956]" captionTargetBox="[189,1398,265,1879]" captionTargetId="figure@13.[189,1398,261,1881]" captionTargetPageId="13" captionText="Fig. 5. Diplommatina Benson, 1849 of Nepal. A. D. miriensis Godwin-Austen, 1917 (CDZMTU036). B. D.munipurensis Godwin-Austen, 1892 (CDZMTU039).C. D.oviformis Fulton, 1901 (CDZMTU042). Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832494" httpUri="https://zenodo.org/record/3832494/files/figure.png" pageId="12" pageNumber="13">Fig. 5A</figureCitation>
<figureCitation id="135462D2FFE8FFE7FC9B44EBD611FC27" box="[821,835,905,932]" captionStart="Fig" captionStartId="6.[189,232,1930,1956]" captionTargetBox="[189,1393,274,1888]" captionTargetId="figure@6.[189,1398,269,1893]" captionTargetPageId="6" captionText="Fig. 3. Diplommatina Benson, 1849 of Nepal. A. D. abiesiana sp. nov., holotype (CDZMTU020). B. D. exserta Godwin-Austen, 1886 (CDZMTU024). C. D. fistulata sp. nov., holotype (CDZMTU025). The yellow arrow indicates the dorsal fold. Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832490" httpUri="https://zenodo.org/record/3832490/files/figure.png" pageId="12" pageNumber="13">3</figureCitation>
, A
<figureCitation id="135462D2FFE8FFE7FCCB44EBD627FC20" box="[869,885,905,931]" captionStart="Fig" captionStartId="9.[189,232,1930,1956]" captionTargetBox="[189,1398,274,1888]" captionTargetId="figure@9.[189,1398,266,1891]" captionTargetPageId="9" captionText="Fig. 4. Diplommatina Benson, 1849 of Nepal. A. D. folliculus (L. Pfeiffer, 1846) (CDZMTU028). B. D. godawariensis sp. nov., holotype (CDZMTU032). C. D. maipokhariensis sp. nov., holotype (CDZMTU034). Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832492" httpUri="https://zenodo.org/record/3832492/files/figure.png" pageId="12" pageNumber="13">4</figureCitation>
); basal edge of peristome rounded; outer lip thin, inner lip thick. Parietal wall of the aperture hardly reaching half of the body whorl.
</paragraph>
</subSubSection>
<subSubSection id="C3752DDCFFE8FFE7FF13448DD185FA8C" pageId="12" pageNumber="13" type="materials_examined">
<paragraph id="8BD07E57FFE8FFE7FF13448DD4F8FB8A" blockId="12.[189,426,1007,1033]" box="[189,426,1007,1033]" pageId="12" pageNumber="13">
<heading id="D098C93BFFE8FFE7FF13448DD4F8FB8A" bold="true" box="[189,426,1007,1033]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="B91BA245FFE8FFE7FF13448DD4F8FB8A" bold="true" box="[189,426,1007,1033]" pageId="12" pageNumber="13">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BD07E57FFE8FFE7FF13437CD185FA8C" blockId="12.[189,1399,1054,1295]" pageId="12" pageNumber="13">
<collectingCountry id="F3783EC7FFE8FFE7FF13437CD44CFBBB" box="[189,286,1054,1080]" name="Nepal" pageId="12" pageNumber="13">NEPAL</collectingCountry>
: 1 shell, Deurali, Syabru, Langtang National Park,
<quantity id="4C97D3B2FFE8FFE7FC3E437CD6BDFBBB" box="[912,1007,1054,1080]" metricMagnitude="3" metricUnit="m" metricValue="2.412" pageId="12" pageNumber="13" unit="m" value="2412.0">2412 m</quantity>
,
<geoCoordinate id="EE5B1890FFE8FFE7FC51437CD1CCFBBB" box="[1023,1182,1054,1081]" degrees="27.805037" direction="north" orientation="latitude" pageId="12" pageNumber="13" precision="1" value="27.805037">27.805037 N</geoCoordinate>
,
<geoCoordinate id="EE5B1890FFE8FFE7FB03437CD01BFBBB" box="[1197,1353,1054,1081]" degrees="85.387862" direction="east" orientation="longitude" pageId="12" pageNumber="13" precision="1" value="85.38786">85.387862 E</geoCoordinate>
,
<date id="FFD15897FFE8FFE7FAF7437CD468FBDF" pageId="12" pageNumber="13" value="2008-04-25">25 Apr. 2008</date>
, P.B. Budha leg. (
<collectionCode id="ED7EE692FFE8FFE7FDB24320D7CCFBDE" box="[540,670,1090,1117]" pageId="12" pageNumber="13">CDZMTU</collectionCode>
036); 3 shells, Okhareni, Shivapuri-Nagarjun National Park,
<quantity id="4C97D3B2FFE8FFE7FF134307D44AFB03" box="[189,280,1125,1152]" metricMagnitude="3" metricUnit="m" metricValue="2.3259999999999996" pageId="12" pageNumber="13" unit="m" value="2326.0">2326 m</quantity>
,
<geoCoordinate id="EE5B1890FFE8FFE7FE8A4307D4EDFB03" box="[292,447,1125,1152]" degrees="27.813051" direction="north" orientation="latitude" pageId="12" pageNumber="13" precision="1" value="27.813051">27.813051 N</geoCoordinate>
,
<geoCoordinate id="EE5B1890FFE8FFE7FE644307D717FB03" box="[458,581,1125,1152]" degrees="85.4217" direction="east" orientation="longitude" pageId="12" pageNumber="13" precision="5" value="85.4217">85.4217 E</geoCoordinate>
,
<date id="FFD15897FFE8FFE7FDFF4304D7B8FB03" box="[593,746,1126,1152]" pageId="12" pageNumber="13" value="2007-06-22">22 Jun. 2007</date>
, R. Devkota and N. Kohar leg. (
<collectionCode id="ED7EE692FFE8FFE7FBCE4304D1B0FB03" box="[1120,1250,1126,1152]" pageId="12" pageNumber="13">CDZMTU</collectionCode>
037); 1 shell, Baghdwar, Shivapuri-Nagarjun National Park,
<quantity id="4C97D3B2FFE8FFE7FD5743EBD60BFB27" box="[761,857,1161,1188]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="12" pageNumber="13" unit="m" value="2500.0">2500 m</quantity>
,
<geoCoordinate id="EE5B1890FFE8FFE7FCC443EBD6BCFB27" box="[874,1006,1161,1188]" degrees="27.8133" direction="north" orientation="latitude" pageId="12" pageNumber="13" precision="5" value="27.8133">27.8133 N</geoCoordinate>
,
<geoCoordinate id="EE5B1890FFE8FFE7FC5143EBD12DFB27" box="[1023,1151,1161,1188]" degrees="85.3877" direction="east" orientation="longitude" pageId="12" pageNumber="13" precision="5" value="85.3877">85.3877 E</geoCoordinate>
,
<date id="FFD15897FFE8FFE7FB3F43EBD065FB20" box="[1169,1335,1161,1188]" pageId="12" pageNumber="13" value="2008-04-25">25 Apr. 2008</date>
, P.B. Budha leg. (
<collectionCode id="ED7EE692FFE8FFE7FEFC43CFD486FB4B" box="[338,468,1197,1224]" pageId="12" pageNumber="13">CDZMTU</collectionCode>
037b); 4 shells, Cholangpati, Langtang National Park,
<quantity id="4C97D3B2FFE8FFE7FBC843CFD196FB44" box="[1126,1220,1197,1223]" metricMagnitude="3" metricUnit="m" metricValue="3.443" pageId="12" pageNumber="13" unit="m" value="3443.0">3443 m</quantity>
,
<geoCoordinate id="EE5B1890FFE8FFE7FB7A43CFD020FB44" box="[1236,1394,1197,1224]" degrees="28.110154" direction="north" orientation="latitude" pageId="12" pageNumber="13" precision="1" value="28.110153">28.110154 N</geoCoordinate>
,
<geoCoordinate id="EE5B1890FFE8FFE7FF1343B2D405FB68" box="[189,343,1232,1259]" degrees="85.334238" direction="east" orientation="longitude" pageId="12" pageNumber="13" precision="1" value="85.334236">85.334238 E</geoCoordinate>
,
<date id="FFD15897FFE8FFE7FEC843B3D759FB68" box="[358,523,1233,1259]" pageId="12" pageNumber="13" value="2008-05-19">19 May 2008</date>
, P.B. Budha leg. (
<collectionCode id="ED7EE692FFE8FFE7FD4B43B3D63AFB68" box="[741,872,1233,1259]" pageId="12" pageNumber="13">CDZMTU</collectionCode>
038); 7 shells, Raniban, Balaju, Kathmandu,
<quantity id="4C97D3B2FFE8FFE7FF134396D433FA8C" box="[189,353,1268,1295]" metricMagnitude="3" metricUnit="m" metricValue="1.871" metricValueMax="1.902" metricValueMin="1.8399999999999999" pageId="12" pageNumber="13" unit="m" value="1871.0" valueMax="1902.0" valueMin="1840.0">1840–1902 m</quantity>
,
<geoCoordinate id="EE5B1890FFE8FFE7FEC04396D758FA8C" box="[366,522,1268,1295]" degrees="27.745997" direction="north" orientation="latitude" pageId="12" pageNumber="13" precision="1" value="27.745996">27.745997 N</geoCoordinate>
,
<geoCoordinate id="EE5B1890FFE8FFE7FDB84396D7FCFA8C" box="[534,686,1268,1295]" degrees="85.287240" direction="east" orientation="longitude" pageId="12" pageNumber="13" precision="1" value="85.28724">85.287240 E</geoCoordinate>
,
<date id="FFD15897FFE8FFE7FD144396D609FA8D" box="[698,859,1268,1295]" pageId="12" pageNumber="13" value="2009-03-12">12 Mar. 2009</date>
, S. Khanal leg. (
<collectionCode id="ED7EE692FFE8FFE7FBB24396D1CDFA8C" box="[1052,1183,1268,1295]" pageId="12" pageNumber="13">CDZMTU</collectionCode>
058).
</paragraph>
</subSubSection>
<subSubSection id="C3752DDCFFE8FFE7FF134254D485FA69" pageId="12" pageNumber="13" type="discussion">
<paragraph id="8BD07E57FFE8FFE7FF134254D47DFAD3" blockId="12.[189,303,1334,1360]" box="[189,303,1334,1360]" pageId="12" pageNumber="13">
<heading id="D098C93BFFE8FFE7FF134254D47DFAD3" bold="true" box="[189,303,1334,1360]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="B91BA245FFE8FFE7FF134254D47DFAD3" bold="true" box="[189,303,1334,1360]" pageId="12" pageNumber="13">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BD07E57FFE8FFE7FF134204D485FA69" blockId="12.[189,1399,1381,1515]" pageId="12" pageNumber="13">
<taxonomicName id="4C6F05D4FFE8FFE7FF134204D41CFAFC" authorityName="Godwin-Austen" authorityYear="1917" box="[189,334,1382,1407]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="miriensis">
<emphasis id="B91BA245FFE8FFE7FF134204D41CFAFC" box="[189,334,1382,1407]" italics="true" pageId="12" pageNumber="13">D. miriensis</emphasis>
</taxonomicName>
is the smallest species reported in the present study. Similar sized species are
<taxonomicName id="4C6F05D4FFE8FFE7FB5D4204D4CFFA20" authority="Mollendorff, 1898" authorityName="Mollendorff" authorityYear="1898" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="godwini">
<emphasis id="B91BA245FFE8FFE7FB5D4204D025FAFC" box="[1267,1399,1381,1407]" italics="true" pageId="12" pageNumber="13">D. godwini</emphasis>
Möllendorff, 1898
</taxonomicName>
from Naga Hill, NE
<collectingCountry id="F3783EC7FFE8FFE7FD1642EBD7A6FA20" box="[696,756,1417,1443]" name="India" pageId="12" pageNumber="13">India</collectingCountry>
, and the south Indian
<taxonomicName id="4C6F05D4FFE8FFE7FBB342E8D020FA27" authority="Beddome, 1875" authorityName="Beddome" authorityYear="1875" box="[1053,1394,1417,1444]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="minima">
<emphasis id="B91BA245FFE8FFE7FBB342E8D1F2FA20" box="[1053,1184,1418,1443]" italics="true" pageId="12" pageNumber="13">D. minima</emphasis>
Beddome, 1875
</taxonomicName>
.
<taxonomicName id="4C6F05D4FFE8FFE7FF1342CFD49CFA44" authorityName="Mollendorff" authorityYear="1898" box="[189,462,1453,1479]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="godwini">
<emphasis id="B91BA245FFE8FFE7FF1342CFD49CFA44" box="[189,462,1453,1479]" italics="true" pageId="12" pageNumber="13">Diplommatina godwini</emphasis>
</taxonomicName>
has more whorls (&gt;7) than
<taxonomicName id="4C6F05D4FFE8FFE7FCB642CCD6FBFA44" authorityName="Godwin-Austen" authorityYear="1917" box="[792,937,1454,1479]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="miriensis">
<emphasis id="B91BA245FFE8FFE7FCB642CCD6FBFA44" box="[792,937,1454,1479]" italics="true" pageId="12" pageNumber="13">D. miriensis</emphasis>
</taxonomicName>
, while the peristome of
<taxonomicName id="4C6F05D4FFE8FFE7FB6242CCD068FA44" box="[1228,1338,1454,1479]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="minim">
<emphasis id="B91BA245FFE8FFE7FB6242CCD068FA44" box="[1228,1338,1454,1479]" italics="true" pageId="12" pageNumber="13">D. minim</emphasis>
</taxonomicName>
a has no curved angular edge.
</paragraph>
</subSubSection>
</treatment>
</document>
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