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<document id="593FF6DE0C515ACB55B23EE6D070C963" ID-CLB-Dataset="22067" ID-DOI="10.5852/ejt.2017.337" ID-GBIF-Dataset="5a7f73c5-5646-446f-a72c-3ea18d8f634f" ID-ISSN="2118-9773" ID-Zenodo-Dep="3832483" ID-ZooBank="2669CB69-5064-4B5D-AC25-856680E3B9AA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589830819450" checkinUser="carolina" docAuthor="Budha, Prem B., Naggs, Fred &amp; Backeljau, Thierry" docDate="2017" docId="03C6CF41FFE3FFE3FE5542C8D112FBBE" docLanguage="en" docName="ejt-2017-337.pdf.imf" docOrigin="European Journal of Taxonomy 337" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Diplommatina fistulata Budha &amp; Naggs &amp; Backeljau 2017, sp. nov." docType="treatment" docUuid="CC722581-EC59-42AC-AD0F-EFD6247BD1A0" docUuidSource="ZooBank" docVersion="6" lastPageNumber="9" masterDocId="FFFFB739FFE4FFEBFFAE4762D552FF83" masterDocTitle="The genus Diplommatina Benson, 1849 (Gastropoda: Caenogastropoda: Diplommatinidae) in Nepal, with the description of seven new species" masterLastPageNumber="30" masterPageNumber="1" pageNumber="8" updateTime="1698840088040" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="64640871CDE2A3D9D600D0A32BC0B2E6" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E3B7CBD896D2967B5E53F887C3CBAE17">
<mods:title id="C2EF845C27D9F55CCE89E9DB49930AE5">The genus Diplommatina Benson, 1849 (Gastropoda: Caenogastropoda: Diplommatinidae) in Nepal, with the description of seven new species</mods:title>
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<mods:name id="306ADF3016C9F28A688C2F311A5B7604" type="personal">
<mods:role id="3701B644B692E859115130E2B859ECD2">
<mods:roleTerm id="693FCAAD9C627117824920031253A7B0">Author</mods:roleTerm>
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<mods:namePart id="485EB596A177B08136D8C795A79F5448">Budha, Prem B.</mods:namePart>
<mods:affiliation id="63F582EF411D6C16663B53695AE2176A">University of Antwerp, Evolutionary Ecology Group, Universiteitsplein 1, B- 2610, Antwerp, Belgium. &amp; Central Department of Zoology, Tribhuvan University, Kirtipur, Kathmandu, Nepal. &amp; Corresponding author: prembudha @ yahoo. com</mods:affiliation>
<mods:nameIdentifier id="E6C5691CFF82A3D438D9927D255A3583" type="email">prembudha@yahoo.com</mods:nameIdentifier>
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<mods:name id="505753BCA38E536365BE6F569E8348E9" type="personal">
<mods:role id="C4EE592E587597E66EDBC71131744AA8">
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<mods:namePart id="56909E64540371EAA9222DA2B79A6C66">Naggs, Fred</mods:namePart>
<mods:affiliation id="E380C9F021ACDEADB429BDE81BBB61A3">Natural History Museum, Cromwell Road, London, SW 7 5 BD, UK. &amp; Email: f. naggs @ nhm. ac. uk</mods:affiliation>
<mods:nameIdentifier id="3D2C0D67706A46005720E0F6D8612EEE" type="email">f.naggs@nhm.ac.uk</mods:nameIdentifier>
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<mods:name id="A029D6A6EED36C92CF42C9907B472E78" type="personal">
<mods:role id="FEC5DECF52750F5962166388EC77B715">
<mods:roleTerm id="E7DFAA8266554719739784D4CC772922">Author</mods:roleTerm>
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<mods:namePart id="81732D764F6601E6939DD3C2E7BB9D88">Backeljau, Thierry</mods:namePart>
<mods:affiliation id="CB9B76414920AD7B7614E6DECD6F1F2A">University of Antwerp, Evolutionary Ecology Group, Universiteitsplein 1, B- 2610, Antwerp, Belgium. &amp; Royal Belgian Institute of Natural Sciences, Vautierstraat 29, B- 1000, Brussels, Belgium. &amp; Email: thierry. backeljau @ naturalsciences. be</mods:affiliation>
<mods:nameIdentifier id="1950F5AE5B3FEAC86E2A5B81F9F5BA8F" type="email">thierry.backeljau@naturalsciences.be</mods:nameIdentifier>
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<mods:title id="B509FDD27D94BE0D23D86E03079C6956">European Journal of Taxonomy</mods:title>
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<mods:part id="B3DBC001C7E6F02C54C5C403ACE77899">
<mods:date id="0BB32CE7B8732BF04FD671A1C67DAAF3">2017</mods:date>
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<mods:number id="8C827613722D5D34C94603C320344D4E">2017-07-21</mods:number>
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<treatment id="03C6CF41FFE3FFE3FE5542C8D112FBBE" ID-DOI="http://doi.org/10.5281/zenodo.3850666" ID-GBIF-Taxon="164167136" ID-Zenodo-Dep="3850666" LSID="urn:lsid:zoobank.org:act:CC722581-EC59-42AC-AD0F-EFD6247BD1A0" httpUri="http://treatment.plazi.org/id/03C6CF41FFE3FFE3FE5542C8D112FBBE" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
<subSubSection id="C3752DDCFFE3FFECFE5542C8D16BFA46" box="[507,1081,1450,1477]" pageId="7" pageNumber="8" type="nomenclature">
<paragraph id="8BD07E57FFE3FFECFE5542C8D16BFA46" blockId="7.[356,1231,1450,1549]" box="[507,1081,1450,1477]" pageId="7" pageNumber="8">
<heading id="D098C93BFFE3FFECFE5542C8D16BFA46" box="[507,1081,1450,1477]" centered="true" fontSize="11" level="2" pageId="7" pageNumber="8" reason="2">
<taxonomicName id="4C6F05D4FFE3FFECFE5542C8D685FA46" ID-CoL="36M29" authority="Budha &amp; Naggs" authorityName="Budha &amp; Naggs &amp; Backeljau" authorityYear="2017" box="[507,983,1450,1477]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="fistulata" status="sp. nov.">
<emphasis id="B91BA245FFE3FFECFE5542C8D641FA46" bold="true" box="[507,787,1450,1477]" italics="true" pageId="7" pageNumber="8">Diplommatina fistulata</emphasis>
Budha &amp; Naggs
</taxonomicName>
<taxonomicNameLabel id="A2281F3EFFE3FFECFC7042C9D16BFA46" box="[990,1081,1451,1477]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="C3752DDCFFE3FFECFECA42ACD632F98E" pageId="7" pageNumber="8" type="description">
<paragraph id="8BD07E57FFE3FFECFECA42ACD19DFA6A" blockId="7.[356,1231,1450,1549]" box="[356,1231,1486,1513]" pageId="7" pageNumber="8">
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urn:lsid:zoobank.org:act:
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</paragraph>
<paragraph id="8BD07E57FFE3FFECFD7D4291D632F98E" blockId="7.[356,1231,1450,1549]" box="[723,864,1523,1549]" pageId="7" pageNumber="8">
<figureCitation id="135462D2FFE3FFECFD7D4291D663F98E" box="[723,817,1523,1549]" captionStart="Fig" captionStartId="6.[189,232,1930,1956]" captionTargetBox="[189,1393,274,1888]" captionTargetId="figure@6.[189,1398,269,1893]" captionTargetPageId="6" captionText="Fig. 3. Diplommatina Benson, 1849 of Nepal. A. D. abiesiana sp. nov., holotype (CDZMTU020). B. D. exserta Godwin-Austen, 1886 (CDZMTU024). C. D. fistulata sp. nov., holotype (CDZMTU025). The yellow arrow indicates the dorsal fold. Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832490" httpUri="https://zenodo.org/record/3832490/files/figure.png" pageId="7" pageNumber="8">Figs 3C</figureCitation>
,
<figureCitation id="135462D2FFE3FFECFC904291D632F98E" box="[830,864,1523,1549]" captionStart="Fig" captionStartId="24.[189,232,1789,1815]" captionTargetBox="[297,1297,294,1724]" captionTargetId="figure@24.[247,1340,265,1748]" captionTargetPageId="24" captionText="Fig. 9. Palatal lamellae (palatalis) of species of Diplommatina. Dotted lines indicate the position of the constriction. Dark lines indicate the palatalis. A. D. abiesiana sp. nov. B. D. exserta Godwin- Austen, 1886. C. D. fistulata sp. nov. D. D. folliculus (L. Pfeiffer, 1846). E. D. godawariensis sp. nov. F. D. maipokhariensis sp. nov. G. D. miriensis Godwin-Austen, 1917. H. D. munipurensis Godwin- Austen, 1892. I. D. oviformis Fulton, 1901. J. D. regularis Fulton, 1901. K. D. salgharica sp. nov. L. D. shivapuriensis sp. nov. M. D. silvicola Godwin-Austen, 1886. N. D. sperata W.T. Blanford, 1862. O. D. syabrubesiensis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3832502" httpUri="https://zenodo.org/record/3832502/files/figure.png" pageId="7" pageNumber="8">9C</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3752DDCFFE3FFE3FF134157D6AFFEC8" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="diagnosis">
<paragraph id="8BD07E57FFE3FFECFF134157D464F9CC" blockId="7.[189,310,1589,1615]" box="[189,310,1589,1615]" pageId="7" pageNumber="8">
<heading id="D098C93BFFE3FFECFF134157D464F9CC" bold="true" box="[189,310,1589,1615]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="B91BA245FFE3FFECFF134157D464F9CC" bold="true" box="[189,310,1589,1615]" pageId="7" pageNumber="8">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BD07E57FFE3FFE3FF134104D6AFFEC8" blockId="7.[189,1399,1637,2027]" lastBlockId="8.[189,1398,267,331]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
Measurements (n = 10): SH 3.0–
<quantity id="4C97D3B2FFE3FFECFDE14107D7FCF903" box="[591,686,1637,1664]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="7" pageNumber="8" unit="mm" value="3.3">3.3 mm</quantity>
, SW
<quantity id="4C97D3B2FFE3FFECFD5C4107D6D4F903" box="[754,902,1637,1664]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" pageId="7" pageNumber="8" unit="mm" value="1.65" valueMax="1.8" valueMin="1.5">1.5–1.8 mm</quantity>
, Wh 6.5–7.5;
<typeStatus id="54D4C0F5FFE3FFECFB9D4107D1CCF9FC" box="[1075,1182,1637,1663]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
: SH
<quantity id="4C97D3B2FFE3FFECFB724107D06FF903" box="[1244,1341,1637,1664]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="7" pageNumber="8" unit="mm" value="3.3">3.3 mm</quantity>
, SW 2.0 mm, Wh 7.5. Dextral, body whorl narrower than penultimate whorl. Constriction at the middle of the parietal lip (
<figureCitation id="135462D2FFE3FFECFEE241CCD4FAF94B" box="[332,424,1710,1736]" captionStart="Fig" captionStartId="24.[189,232,1789,1815]" captionTargetBox="[297,1297,294,1724]" captionTargetId="figure@24.[247,1340,265,1748]" captionTargetPageId="24" captionText="Fig. 9. Palatal lamellae (palatalis) of species of Diplommatina. Dotted lines indicate the position of the constriction. Dark lines indicate the palatalis. A. D. abiesiana sp. nov. B. D. exserta Godwin- Austen, 1886. C. D. fistulata sp. nov. D. D. folliculus (L. Pfeiffer, 1846). E. D. godawariensis sp. nov. F. D. maipokhariensis sp. nov. G. D. miriensis Godwin-Austen, 1917. H. D. munipurensis Godwin- Austen, 1892. I. D. oviformis Fulton, 1901. J. D. regularis Fulton, 1901. K. D. salgharica sp. nov. L. D. shivapuriensis sp. nov. M. D. silvicola Godwin-Austen, 1886. N. D. sperata W.T. Blanford, 1862. O. D. syabrubesiensis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3832502" httpUri="https://zenodo.org/record/3832502/files/figure.png" pageId="7" pageNumber="8">Fig. 9C</figureCitation>
). Parietalis long and very strong (
<figureCitation id="135462D2FFE3FFECFC9A41CCD6F2F94B" box="[820,928,1710,1736]" captionStart="Fig" captionStartId="6.[189,232,1930,1956]" captionTargetBox="[189,1393,274,1888]" captionTargetId="figure@6.[189,1398,269,1893]" captionTargetPageId="6" captionText="Fig. 3. Diplommatina Benson, 1849 of Nepal. A. D. abiesiana sp. nov., holotype (CDZMTU020). B. D. exserta Godwin-Austen, 1886 (CDZMTU024). C. D. fistulata sp. nov., holotype (CDZMTU025). The yellow arrow indicates the dorsal fold. Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832490" httpUri="https://zenodo.org/record/3832490/files/figure.png" pageId="7" pageNumber="8">Fig. 3C3</figureCitation>
). Two vertical palatalis: a stronger Vpt1 along the constriction and a short Vpt2 running parallel to Vpt1 above the parietal wall (
<figureCitation id="135462D2FFE3FFECFB7041B0D06EF96F" box="[1246,1340,1746,1772]" captionStart="Fig" captionStartId="24.[189,232,1789,1815]" captionTargetBox="[297,1297,294,1724]" captionTargetId="figure@24.[247,1340,265,1748]" captionTargetPageId="24" captionText="Fig. 9. Palatal lamellae (palatalis) of species of Diplommatina. Dotted lines indicate the position of the constriction. Dark lines indicate the palatalis. A. D. abiesiana sp. nov. B. D. exserta Godwin- Austen, 1886. C. D. fistulata sp. nov. D. D. folliculus (L. Pfeiffer, 1846). E. D. godawariensis sp. nov. F. D. maipokhariensis sp. nov. G. D. miriensis Godwin-Austen, 1917. H. D. munipurensis Godwin- Austen, 1892. I. D. oviformis Fulton, 1901. J. D. regularis Fulton, 1901. K. D. salgharica sp. nov. L. D. shivapuriensis sp. nov. M. D. silvicola Godwin-Austen, 1886. N. D. sperata W.T. Blanford, 1862. O. D. syabrubesiensis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3832502" httpUri="https://zenodo.org/record/3832502/files/figure.png" pageId="7" pageNumber="8">Fig. 9C</figureCitation>
). No horizontal palatalis. Columellaris prominent. Columellar tooth strong. Protoconch smooth, Wh 2.0, with minute pits scattered over the surface (
<figureCitation id="135462D2FFE3FFECFD294079D7B5F8B6" box="[647,743,1819,1845]" captionStart="Fig" captionStartId="6.[189,232,1930,1956]" captionTargetBox="[189,1393,274,1888]" captionTargetId="figure@6.[189,1398,269,1893]" captionTargetPageId="6" captionText="Fig. 3. Diplommatina Benson, 1849 of Nepal. A. D. abiesiana sp. nov., holotype (CDZMTU020). B. D. exserta Godwin-Austen, 1886 (CDZMTU024). C. D. fistulata sp. nov., holotype (CDZMTU025). The yellow arrow indicates the dorsal fold. Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832490" httpUri="https://zenodo.org/record/3832490/files/figure.png" pageId="7" pageNumber="8">Fig. 3C</figureCitation>
<figureCitation id="135462D2FFE3FFECFD494079D7A0F8B6" box="[743,754,1819,1845]" captionStart="Fig" captionStartId="2.[189,235,1999,2025]" captionTargetBox="[189,1399,1129,1958]" captionTargetId="figure@2.[187,1401,1126,1960]" captionTargetPageId="2" captionText="Fig.1. Map of the localities and areas surveyed for Diplommatina spp. in Nepal." figureDoi="http://doi.org/10.5281/zenodo.3832486" httpUri="https://zenodo.org/record/3832486/files/figure.png" pageId="7" pageNumber="8">1</figureCitation>
). Teleoconch with dense radial ribs; 9–11 ribs/
<quantity id="4C97D3B2FFE3FFECFAB34079D025F8B5" box="[1309,1399,1819,1846]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="8" unit="mm" value="0.5">0.5 mm</quantity>
on the penultimate whorl; 15–20 ribs/
<quantity id="4C97D3B2FFE3FFECFD2E405DD788F8D9" box="[640,730,1855,1882]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="8" unit="mm" value="0.5">0.5 mm</quantity>
on the body whorl towards the aperture (
<figureCitation id="135462D2FFE3FFECFB64405DD074F8DA" box="[1226,1318,1855,1881]" captionStart="Fig" captionStartId="6.[189,232,1930,1956]" captionTargetBox="[189,1393,274,1888]" captionTargetId="figure@6.[189,1398,269,1893]" captionTargetPageId="6" captionText="Fig. 3. Diplommatina Benson, 1849 of Nepal. A. D. abiesiana sp. nov., holotype (CDZMTU020). B. D. exserta Godwin-Austen, 1886 (CDZMTU024). C. D. fistulata sp. nov., holotype (CDZMTU025). The yellow arrow indicates the dorsal fold. Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832490" httpUri="https://zenodo.org/record/3832490/files/figure.png" pageId="7" pageNumber="8">Fig. 3C</figureCitation>
<figureCitation id="135462D2FFE3FFECFA8B405DD067F8DA" box="[1317,1333,1855,1881]" captionStart="Fig" captionStartId="4.[189,232,1895,1921]" captionTargetBox="[189,1398,285,1853]" captionTargetId="figure@4.[189,1398,285,1853]" captionTargetPageId="4" captionText="Fig. 2. Shell terminology. A. D. regularis Fulton, 1901. B. D. folliculus (L. Pfeiffer, 1846). C. General outline of the lower part of a shell from Diplommatina species. Abbreviations: Prt = Parietalis (Prt1 = inner parietalis, Prt2 = outer parietalis); Pt = Palatalis; Hpt = Horizontal palatalis (Hpt1 = lower, Hpt2 = upper in case of two); Vpt = Vertical palatalis (Vpt1 = left side, Vpt2 = right side in case of two)." figureDoi="http://doi.org/10.5281/zenodo.3832488" httpUri="https://zenodo.org/record/3832488/files/figure.png" pageId="7" pageNumber="8">2</figureCitation>
, C
<figureCitation id="135462D2FFE3FFECFAF7405DD038F8DA" box="[1369,1386,1855,1881]" captionStart="Fig" captionStartId="9.[189,232,1930,1956]" captionTargetBox="[189,1398,274,1888]" captionTargetId="figure@9.[189,1398,266,1891]" captionTargetPageId="9" captionText="Fig. 4. Diplommatina Benson, 1849 of Nepal. A. D. folliculus (L. Pfeiffer, 1846) (CDZMTU028). B. D. godawariensis sp. nov., holotype (CDZMTU032). C. D. maipokhariensis sp. nov., holotype (CDZMTU034). Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832492" httpUri="https://zenodo.org/record/3832492/files/figure.png" pageId="7" pageNumber="8">4</figureCitation>
). Distinct dorsal fold at base of body whorl, running towards the aperture (
<figureCitation id="135462D2FFE3FFECFB914006D1CFF8FD" box="[1087,1181,1892,1918]" captionStart="Fig" captionStartId="6.[189,232,1930,1956]" captionTargetBox="[189,1393,274,1888]" captionTargetId="figure@6.[189,1398,269,1893]" captionTargetPageId="6" captionText="Fig. 3. Diplommatina Benson, 1849 of Nepal. A. D. abiesiana sp. nov., holotype (CDZMTU020). B. D. exserta Godwin-Austen, 1886 (CDZMTU024). C. D. fistulata sp. nov., holotype (CDZMTU025). The yellow arrow indicates the dorsal fold. Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832490" httpUri="https://zenodo.org/record/3832490/files/figure.png" pageId="7" pageNumber="8">Fig. 3C</figureCitation>
<figureCitation id="135462D2FFE3FFECFB334006D1FCF8FD" box="[1181,1198,1892,1918]" captionStart="Fig" captionStartId="9.[189,232,1930,1956]" captionTargetBox="[189,1398,274,1888]" captionTargetId="figure@9.[189,1398,266,1891]" captionTargetPageId="9" captionText="Fig. 4. Diplommatina Benson, 1849 of Nepal. A. D. folliculus (L. Pfeiffer, 1846) (CDZMTU028). B. D. godawariensis sp. nov., holotype (CDZMTU032). C. D. maipokhariensis sp. nov., holotype (CDZMTU034). Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832492" httpUri="https://zenodo.org/record/3832492/files/figure.png" pageId="7" pageNumber="8">4</figureCitation>
). Spiral striation present. Umbilical perforation visible, but umbilicus seems to be closed by the reflected edge of the columellar lip. Aperture ear-shaped, parietal wall oblique with regard to the coiling axis; peristome with two close lips; inner lip thickened and forming a thick callus over the parietal wall; under high magnification 12–15 highly dense ribs visible between the lips. Operculum paucispiral, corneous, thin, outer surface smooth, inner surface with thick peripheral circular edge.
</paragraph>
</subSubSection>
<subSubSection id="C3752DDCFFECFFE3FF134617D477FE66" pageId="8" pageNumber="9" type="etymology">
<paragraph id="8BD07E57FFECFFE3FF134617D416FE0C" blockId="8.[189,324,373,399]" box="[189,324,373,399]" pageId="8" pageNumber="9">
<heading id="D098C93BFFECFFE3FF134617D416FE0C" bold="true" box="[189,324,373,399]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="B91BA245FFECFFE3FF134617D416FE0C" bold="true" box="[189,324,373,399]" pageId="8" pageNumber="9">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BD07E57FFECFFE3FF1346C7D477FE66" blockId="8.[189,1399,421,485]" pageId="8" pageNumber="9">The species name refers to the tube-like structure of the dorsal fold on the body whorl just behind the aperture.</paragraph>
</subSubSection>
<subSubSection id="C3752DDCFFECFFE3FF13456DD025FC27" pageId="8" pageNumber="9" type="materials_examined">
<paragraph id="8BD07E57FFECFFE3FF13456DD43EFDAB" blockId="8.[189,364,526,553]" box="[189,364,526,553]" pageId="8" pageNumber="9">
<heading id="D098C93BFFECFFE3FF13456DD43EFDAB" bold="true" box="[189,364,526,553]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="B91BA245FFECFFE3FF13456DD43EFDAB" bold="true" box="[189,364,526,553]" pageId="8" pageNumber="9">Type material</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B07740AFFECFFE3FF44455DD76FFD26" ID-GBIF-Occurrence="2620517306" collectingDate="2007-05-04" collectionCode="CDZMTU" collectorName="P. B. Budha" country="Nepal" elevation="2396" latitude="27.574556" location="Phulchowki Hill" longLatPrecision="1" longitude="85.40084" municipality="Lalitpur District" pageId="8" pageNumber="9" specimenCode="CDZMTU025" specimenCount="1" typeStatus="holotype">
<paragraph id="8BD07E57FFECFFE3FF44455DD40EFDDA" blockId="8.[189,1398,575,677]" box="[234,348,575,601]" pageId="8" pageNumber="9">
<emphasis id="B91BA245FFECFFE3FF44455DD40EFDDA" bold="true" box="[234,348,575,601]" pageId="8" pageNumber="9">
<typeStatus id="54D4C0F5FFECFFE3FF44455DD40EFDDA" box="[234,348,575,601]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BD07E57FFECFFE3FF134507D76FFD26" blockId="8.[189,1398,575,677]" pageId="8" pageNumber="9">
<collectingCountry id="F3783EC7FFECFFE3FF134507D44CFDFC" box="[189,286,613,639]" name="Nepal" pageId="8" pageNumber="9">NEPAL</collectingCountry>
:
<location id="8EB0288CFFECFFE3FE854507D4A3FDFC" LSID="urn:lsid:plazi:treatment:03C6CF41FFE3FFE3FE5542C8D112FBBE:8EB0288CFFECFFE3FE854507D4A3FDFC" box="[299,497,613,639]" country="Nepal" latitude="27.574556" longLatPrecision="1" longitude="85.40084" municipality="Lalitpur District" name="Phulchowki Hill" pageId="8" pageNumber="9">Phulchowki Hill</location>
,
<collectingMunicipality id="6BB4E42DFFECFFE3FE514507D791FDFC" box="[511,707,613,639]" pageId="8" pageNumber="9">Lalitpur District</collectingMunicipality>
,
<quantity id="4C97D3B2FFECFFE3FD7F4507D625FDFC" box="[721,887,613,639]" metricMagnitude="3" metricUnit="m" metricValue="2.3955" metricValueMax="2.483" metricValueMin="2.308" pageId="8" pageNumber="9" unit="m" value="2395.5" valueMax="2483.0" valueMin="2308.0">
<elevation id="00429964FFECFFE3FD7F4507D625FDFC" box="[721,887,613,639]" metricMagnitude="3" metricUnit="m" metricValue="2.3955" metricValueMax="2.483" metricValueMin="2.308" pageId="8" pageNumber="9" unit="m" value="2395.5" valueMax="2483.0" valueMin="2308.0">2308–2483 m</elevation>
</quantity>
,
<geoCoordinate id="EE5B1890FFECFFE3FC284506D177FDFC" box="[902,1061,612,639]" degrees="27.574557" direction="north" orientation="latitude" pageId="8" pageNumber="9" precision="1" value="27.574556">27.574557 N</geoCoordinate>
,
<geoCoordinate id="EE5B1890FFECFFE3FB9D4506D19CFDFC" box="[1075,1230,612,639]" degrees="85.400842" direction="east" orientation="longitude" pageId="8" pageNumber="9" precision="1" value="85.40084">85.400842 E</geoCoordinate>
,
<date id="FFD15897FFECFFE3FB734507D020FDFC" box="[1245,1394,613,639]" pageId="8" pageNumber="9" value="2007-05-04">
<collectingDate id="EF95A17FFFECFFE3FB734507D020FDFC" box="[1245,1394,613,639]" pageId="8" pageNumber="9" value="2007-05-04">4 May 2007</collectingDate>
</date>
,
<collectorName id="269A1B81FFECFFE3FF1345E9D413FD27" box="[189,321,650,677]" pageId="8" pageNumber="9">P.B. Budha</collectorName>
leg. (
<specimenCode id="DBC9D62CFFECFFE3FE2C45E8D763FD26" box="[386,561,650,677]" collectionCode="CDZMTU" pageId="8" pageNumber="9">CDZMTU025</specimenCode>
).
</paragraph>
</materialsCitation>
<paragraph id="8BD07E57FFECFFE3FF4445ACD435FD6B" blockId="8.[189,1399,718,932]" box="[234,359,718,744]" pageId="8" pageNumber="9">
<emphasis id="B91BA245FFECFFE3FF4445ACD435FD6B" bold="true" box="[234,359,718,744]" pageId="8" pageNumber="9">
<typeStatus id="54D4C0F5FFECFFE3FF4445ACD435FD6B" box="[234,359,718,744]" pageId="8" pageNumber="9" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BD07E57FFECFFE3FF134591D025FC27" blockId="8.[189,1399,718,932]" pageId="8" pageNumber="9">
<collectingCountry id="F3783EC7FFECFFE3FF134591D44CFC8E" box="[189,286,755,781]" name="Nepal" pageId="8" pageNumber="9">NEPAL</collectingCountry>
: 5 shells from the type locality, same data as
<typeStatus id="54D4C0F5FFECFFE3FCE24591D6E1FC8E" box="[844,947,755,781]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED7EE692FFECFFE3FC6B4591D115FC8D" box="[965,1095,755,782]" pageId="8" pageNumber="9">CDZMTU</collectionCode>
026); 4 shells, Baghdwar, Shivapuri-Nagarjun National Park,
<quantity id="4C97D3B2FFECFFE3FDC3447BD79EFCB0" box="[621,716,793,819]" metricMagnitude="3" metricUnit="m" metricValue="2.3259999999999996" pageId="8" pageNumber="9" unit="m" value="2326.0">2326 m</quantity>
,
<geoCoordinate id="EE5B1890FFECFFE3FD70447BD633FCB0" box="[734,865,793,819]" degrees="27.8131" direction="north" orientation="latitude" pageId="8" pageNumber="9" precision="5" value="27.8131">27.8131 N</geoCoordinate>
,
<geoCoordinate id="EE5B1890FFECFFE3FCDC447BD6A0FCB0" box="[882,1010,793,820]" degrees="85.4217" direction="east" orientation="longitude" pageId="8" pageNumber="9" precision="5" value="85.4217">85.4217 E</geoCoordinate>
,
<date id="FFD15897FFECFFE3FBAD447BD1F5FCB0" box="[1027,1191,793,819]" pageId="8" pageNumber="9" value="2007-06-22">22 Jun. 2007</date>
, P.B. Budha leg. (
<collectionCode id="ED7EE692FFECFFE3FF6B445CD41AFCDB" box="[197,328,830,856]" pageId="8" pageNumber="9">CDZMTU</collectionCode>
027a); 8 shells, Baghdwar area, Shivapuri-Nagarjun National Park,
<quantity id="4C97D3B2FFECFFE3FBE4445CD1BEFCDA" box="[1098,1260,830,857]" metricMagnitude="3" metricUnit="m" metricValue="2.457" metricValueMax="2.502" metricValueMin="2.412" pageId="8" pageNumber="9" unit="m" value="2457.0" valueMax="2502.0" valueMin="2412.0">2412–2502 m</quantity>
,
<geoCoordinate id="EE5B1890FFECFFE3FB58445CD020FCDA" box="[1270,1394,830,857]" degrees="27.8233" direction="north" orientation="latitude" pageId="8" pageNumber="9" precision="5" value="27.8233">27.8233 N</geoCoordinate>
,
<geoCoordinate id="EE5B1890FFECFFE3FF134401D469FCFD" box="[189,315,867,894]" degrees="85.3877" direction="east" orientation="longitude" pageId="8" pageNumber="9" precision="5" value="85.3877">85.3877 E</geoCoordinate>
,
<date id="FFD15897FFECFFE3FEE54401D748FCFD" box="[331,538,867,894]" pageId="8" pageNumber="9" value="2008-04-24" valueMax="2008-04-25" valueMin="2008-04-24">24–25 Apr. 2008</date>
, R. Devkota and P.B. Budha leg. (
<collectionCode id="ED7EE692FFECFFE3FC6C4406D116FCFC" box="[962,1092,868,895]" pageId="8" pageNumber="9">CDZMTU</collectionCode>
027b); 2 shells, Dhunche, Langtang National Park,
<quantity id="4C97D3B2FFECFFE3FE7144EBD76AFC27" box="[479,568,905,932]" metricMagnitude="3" metricUnit="m" metricValue="2.056" pageId="8" pageNumber="9" unit="m" value="2056.0">2056 m</quantity>
,
<geoCoordinate id="EE5B1890FFECFFE3FDED44EBD7EDFC27" box="[579,703,905,932]" degrees="28.1087" direction="north" orientation="latitude" pageId="8" pageNumber="9" precision="5" value="28.1087">28.1087 N</geoCoordinate>
,
<geoCoordinate id="EE5B1890FFECFFE3FD6444EBD610FC27" box="[714,834,905,932]" degrees="85.2916" direction="east" orientation="longitude" pageId="8" pageNumber="9" precision="5" value="85.2916">85.2916 E</geoCoordinate>
,
<date id="FFD15897FFECFFE3FCE344EBD6A0FC20" box="[845,1010,905,932]" pageId="8" pageNumber="9" value="2007-06-03" valueMax="2007-06-04" valueMin="2007-06-03">3–4 Jun. 2007</date>
, P.B. Budha leg. (
<collectionCode id="ED7EE692FFECFFE3FB1244EBD06CFC27" box="[1212,1342,905,932]" pageId="8" pageNumber="9">CDZMTU</collectionCode>
057).
</paragraph>
</subSubSection>
<subSubSection id="C3752DDCFFECFFE3FF1344AFD112FBBE" pageId="8" pageNumber="9" type="discussion">
<paragraph id="8BD07E57FFECFFE3FF1344AFD47DFC64" blockId="8.[189,303,973,999]" box="[189,303,973,999]" pageId="8" pageNumber="9">
<heading id="D098C93BFFECFFE3FF1344AFD47DFC64" bold="true" box="[189,303,973,999]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="B91BA245FFECFFE3FF1344AFD47DFC64" bold="true" box="[189,303,973,999]" pageId="8" pageNumber="9">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BD07E57FFECFFE3FF13449CD112FBBE" blockId="8.[189,1399,1022,1086]" pageId="8" pageNumber="9">
<taxonomicName id="4C6F05D4FFECFFE3FF13449CD49CFB9B" authority="Budha &amp; Naggs &amp; Backeljau, 2017" authorityName="Budha &amp; Naggs &amp; Backeljau" authorityYear="2017" box="[189,462,1022,1048]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="fistulata" status="sp. nov.">
<emphasis id="B91BA245FFECFFE3FF13449CD49CFB9B" box="[189,462,1022,1048]" italics="true" pageId="8" pageNumber="9">Diplommatina fistulata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2281F3EFFECFFE3FE78449CD760FB9B" box="[470,562,1022,1048]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
resembles the NE Indian
<taxonomicName id="4C6F05D4FFECFFE3FCC9449CD174FB9B" authorityName="Benson" authorityYear="1857" box="[871,1062,1022,1048]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="pachycheilus">
<emphasis id="B91BA245FFECFFE3FCC9449CD174FB9B" box="[871,1062,1022,1048]" italics="true" pageId="8" pageNumber="9">D. pachycheilus</emphasis>
</taxonomicName>
, from which it differs by its distinct dorsal fold behind the aperture, smaller size and more slender shape.
</paragraph>
</subSubSection>
</treatment>
</document>
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