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<document id="4B79C9F9F03ADD1B3F899A22BEF0CC2F" ID-CLB-Dataset="22067" ID-DOI="10.5852/ejt.2017.337" ID-GBIF-Dataset="5a7f73c5-5646-446f-a72c-3ea18d8f634f" ID-ISSN="2118-9773" ID-Zenodo-Dep="3832483" ID-ZooBank="2669CB69-5064-4B5D-AC25-856680E3B9AA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589830819450" checkinUser="carolina" docAuthor="Budha, Prem B., Naggs, Fred &amp; Backeljau, Thierry" docDate="2017" docId="03C6CF41FFE1FFEEFE5C4538D729F868" docLanguage="en" docName="ejt-2017-337.pdf.imf" docOrigin="European Journal of Taxonomy 337" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Diplommatina abiesiana Budha &amp; Naggs &amp; Backeljau 2017, sp. nov." docType="treatment" docUuid="F203B92F-6A52-4687-91DA-749554BB03AC" docUuidSource="ZooBank" docVersion="6" lastPageNumber="6" masterDocId="FFFFB739FFE4FFEBFFAE4762D552FF83" masterDocTitle="The genus Diplommatina Benson, 1849 (Gastropoda: Caenogastropoda: Diplommatinidae) in Nepal, with the description of seven new species" masterLastPageNumber="30" masterPageNumber="1" pageNumber="6" updateTime="1698840088040" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="B0CCF7B3B716F14CE3CDC0E018035A93" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C9BB122075FB3104F87D65939E59BC92">
<mods:title id="F2EA7E39860F738C567D91CEE845665A">The genus Diplommatina Benson, 1849 (Gastropoda: Caenogastropoda: Diplommatinidae) in Nepal, with the description of seven new species</mods:title>
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<mods:name id="821B825BF26A33D27561CFA2A35F6C76" type="personal">
<mods:role id="6C9D1A84078407F4D5FFA2505CD54B5A">
<mods:roleTerm id="AFDFA38DE165582A6983AB9E695178DC">Author</mods:roleTerm>
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<mods:namePart id="AED0EAC8384606F97F0610FC33DBA587">Budha, Prem B.</mods:namePart>
<mods:affiliation id="E648F779A01330EA34940C29D3B8A73F">University of Antwerp, Evolutionary Ecology Group, Universiteitsplein 1, B- 2610, Antwerp, Belgium. &amp; Central Department of Zoology, Tribhuvan University, Kirtipur, Kathmandu, Nepal. &amp; Corresponding author: prembudha @ yahoo. com</mods:affiliation>
<mods:nameIdentifier id="9A1651FCE05CE2605BCFA5350E8402E4" type="email">prembudha@yahoo.com</mods:nameIdentifier>
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<mods:name id="03718290638F2268DD91ADFF98F74EE5" type="personal">
<mods:role id="18D90000FD99F97DEB43399B7E1534FB">
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<mods:namePart id="0F428F643D0235B1051CF26EB90E2AD4">Naggs, Fred</mods:namePart>
<mods:affiliation id="EF6608ABF8B6BC898F46C9061F4CD096">Natural History Museum, Cromwell Road, London, SW 7 5 BD, UK. &amp; Email: f. naggs @ nhm. ac. uk</mods:affiliation>
<mods:nameIdentifier id="55D969D7E00F08313FEE00C525677C76" type="email">f.naggs@nhm.ac.uk</mods:nameIdentifier>
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<mods:name id="80813150615F513421D843AEF4B15D88" type="personal">
<mods:role id="D3D3E43BAAB8A20D3E34F3B724DC10E4">
<mods:roleTerm id="46DB30C49032CCFEE9533512D169BFD0">Author</mods:roleTerm>
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<mods:namePart id="0E16C359B75A7F7F8745D85FD05287A7">Backeljau, Thierry</mods:namePart>
<mods:affiliation id="2E485F635B36CC8D7C73FB7EF8F187DA">University of Antwerp, Evolutionary Ecology Group, Universiteitsplein 1, B- 2610, Antwerp, Belgium. &amp; Royal Belgian Institute of Natural Sciences, Vautierstraat 29, B- 1000, Brussels, Belgium. &amp; Email: thierry. backeljau @ naturalsciences. be</mods:affiliation>
<mods:nameIdentifier id="BFF29A3B4D729F78D13FA9C88F474ED4" type="email">thierry.backeljau@naturalsciences.be</mods:nameIdentifier>
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<mods:typeOfResource id="731BED62128D68F00B223EEB753702A4">text</mods:typeOfResource>
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<mods:title id="4809745C59B43BA164A6BB76A806535F">European Journal of Taxonomy</mods:title>
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<mods:part id="2EAEC7F17CA0FA50EA86592C342B987C">
<mods:date id="EF49711BF88A571D6EF6B93CFA199F3A">2017</mods:date>
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<mods:number id="F58A98D2836B28B7767FBDD8D3B24C29">2017-07-21</mods:number>
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<mods:classification id="F1610BF7C41A0946F448A04F6E9321B4">journal article</mods:classification>
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<mods:identifier id="DF7A64D6C80F84CE93BBD024F05FAE57" type="DOI">10.5852/ejt.2017.337</mods:identifier>
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<treatment id="03C6CF41FFE1FFEEFE5C4538D729F868" ID-DOI="http://doi.org/10.5281/zenodo.3850668" ID-GBIF-Taxon="164167137" ID-Zenodo-Dep="3850668" LSID="urn:lsid:zoobank.org:act:F203B92F-6A52-4687-91DA-749554BB03AC" httpUri="http://treatment.plazi.org/id/03C6CF41FFE1FFEEFE5C4538D729F868" lastPageNumber="6" pageId="5" pageNumber="6">
<subSubSection id="C3752DDCFFE1FFEEFE5C4538D110FDF6" box="[498,1090,602,629]" pageId="5" pageNumber="6" type="nomenclature">
<paragraph id="8BD07E57FFE1FFEEFE5C4538D110FDF6" blockId="5.[368,1219,602,708]" box="[498,1090,602,629]" pageId="5" pageNumber="6">
<heading id="D098C93BFFE1FFEEFE5C4538D110FDF6" box="[498,1090,602,629]" centered="true" fontSize="11" level="2" pageId="5" pageNumber="6" reason="2">
<taxonomicName id="4C6F05D4FFE1FFEEFE5C4538D6B2FDF6" ID-CoL="36LZD" authority="Budha &amp; Naggs" authorityName="Budha &amp; Naggs &amp; Backeljau" authorityYear="2017" box="[498,992,602,629]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="abiesiana" status="sp. nov.">
<emphasis id="B91BA245FFE1FFEEFE5C4538D64EFDF6" bold="true" box="[498,796,602,629]" italics="true" pageId="5" pageNumber="6">Diplommatina abiesiana</emphasis>
Budha &amp; Naggs
</taxonomicName>
<taxonomicNameLabel id="A2281F3EFFE1FFEEFC494539D110FDF6" box="[999,1090,603,629]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3752DDCFFE1FFEEFEDE45E0D631FD47" pageId="5" pageNumber="6" type="description">
<paragraph id="8BD07E57FFE1FFEEFEDE45E0D191FD1E" blockId="5.[368,1219,602,708]" box="[368,1219,642,669]" pageId="5" pageNumber="6">
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urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
<paragraph id="8BD07E57FFE1FFEEFD7C45C8D631FD47" blockId="5.[368,1219,602,708]" box="[722,867,682,708]" pageId="5" pageNumber="6">
<figureCitation id="135462D2FFE1FFEEFD7C45C8D660FD47" box="[722,818,682,708]" captionStart="Fig" captionStartId="6.[189,232,1930,1956]" captionTargetBox="[189,1393,274,1888]" captionTargetId="figure@6.[189,1398,269,1893]" captionTargetPageId="6" captionText="Fig. 3. Diplommatina Benson, 1849 of Nepal. A. D. abiesiana sp. nov., holotype (CDZMTU020). B. D. exserta Godwin-Austen, 1886 (CDZMTU024). C. D. fistulata sp. nov., holotype (CDZMTU025). The yellow arrow indicates the dorsal fold. Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832490" httpUri="https://zenodo.org/record/3832490/files/figure.png" pageId="5" pageNumber="6">Figs 3A</figureCitation>
,
<figureCitation id="135462D2FFE1FFEEFC9145C8D631FD47" box="[831,867,682,708]" captionStart="Fig" captionStartId="24.[189,232,1789,1815]" captionTargetBox="[297,1297,294,1724]" captionTargetId="figure@24.[247,1340,265,1748]" captionTargetPageId="24" captionText="Fig. 9. Palatal lamellae (palatalis) of species of Diplommatina. Dotted lines indicate the position of the constriction. Dark lines indicate the palatalis. A. D. abiesiana sp. nov. B. D. exserta Godwin- Austen, 1886. C. D. fistulata sp. nov. D. D. folliculus (L. Pfeiffer, 1846). E. D. godawariensis sp. nov. F. D. maipokhariensis sp. nov. G. D. miriensis Godwin-Austen, 1917. H. D. munipurensis Godwin- Austen, 1892. I. D. oviformis Fulton, 1901. J. D. regularis Fulton, 1901. K. D. salgharica sp. nov. L. D. shivapuriensis sp. nov. M. D. silvicola Godwin-Austen, 1886. N. D. sperata W.T. Blanford, 1862. O. D. syabrubesiensis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3832502" httpUri="https://zenodo.org/record/3832502/files/figure.png" pageId="5" pageNumber="6">9A</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3752DDCFFE1FFEEFF134592D775FB4C" pageId="5" pageNumber="6" type="diagnosis">
<paragraph id="8BD07E57FFE1FFEEFF134592D464FC89" blockId="5.[189,310,752,778]" box="[189,310,752,778]" pageId="5" pageNumber="6">
<heading id="D098C93BFFE1FFEEFF134592D464FC89" bold="true" box="[189,310,752,778]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="B91BA245FFE1FFEEFF134592D464FC89" bold="true" box="[189,310,752,778]" pageId="5" pageNumber="6">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BD07E57FFE1FFEEFF134447D775FB4C" blockId="5.[189,1399,804,1231]" pageId="5" pageNumber="6">
Measurements (n = 4): SH
<quantity id="4C97D3B2FFE1FFEEFE5F4446D72DFCBC" box="[497,639,804,831]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" metricValueMax="2.9" metricValueMin="2.7" pageId="5" pageNumber="6" unit="mm" value="2.8" valueMax="2.9" valueMin="2.7">2.7–2.9 mm</quantity>
, SW
<quantity id="4C97D3B2FFE1FFEEFD174446D615FCBC" box="[697,839,804,831]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="5" pageNumber="6" unit="mm" value="1.45" valueMax="1.5" valueMin="1.4">1.4–1.5 mm</quantity>
, Wh 6.0–6.5;
<typeStatus id="54D4C0F5FFE1FFEEFC4A4446D11DFCBD" box="[996,1103,804,830]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
: SH
<quantity id="4C97D3B2FFE1FFEEFB2D4446D18CFCBC" box="[1155,1246,804,831]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="5" pageNumber="6" unit="mm" value="2.8">2.8 mm</quantity>
, SW
<quantity id="4C97D3B2FFE1FFEEFAB64446D020FCBC" box="[1304,1394,804,831]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="6" unit="mm" value="1.5">1.5 mm</quantity>
, Wh 6.0. Dextral, penultimate whorl and body whorl nearly equal in width. Constriction at the middle of the parietal wall (
<figureCitation id="135462D2FFE1FFEEFE274416D4BAFC0D" box="[393,488,884,910]" captionStart="Fig" captionStartId="24.[189,232,1789,1815]" captionTargetBox="[297,1297,294,1724]" captionTargetId="figure@24.[247,1340,265,1748]" captionTargetPageId="24" captionText="Fig. 9. Palatal lamellae (palatalis) of species of Diplommatina. Dotted lines indicate the position of the constriction. Dark lines indicate the palatalis. A. D. abiesiana sp. nov. B. D. exserta Godwin- Austen, 1886. C. D. fistulata sp. nov. D. D. folliculus (L. Pfeiffer, 1846). E. D. godawariensis sp. nov. F. D. maipokhariensis sp. nov. G. D. miriensis Godwin-Austen, 1917. H. D. munipurensis Godwin- Austen, 1892. I. D. oviformis Fulton, 1901. J. D. regularis Fulton, 1901. K. D. salgharica sp. nov. L. D. shivapuriensis sp. nov. M. D. silvicola Godwin-Austen, 1886. N. D. sperata W.T. Blanford, 1862. O. D. syabrubesiensis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3832502" httpUri="https://zenodo.org/record/3832502/files/figure.png" pageId="5" pageNumber="6">Fig. 9A</figureCitation>
). Parietalis moderate, gradually elevated inner side of the body whorl. A short horizontal palatalis near the inner edge of the peristome. Columellaris moderately developed (
<figureCitation id="135462D2FFE1FFEEFB5344FED03BFC35" box="[1277,1385,924,950]" captionStart="Fig" captionStartId="6.[189,232,1930,1956]" captionTargetBox="[189,1393,274,1888]" captionTargetId="figure@6.[189,1398,269,1893]" captionTargetPageId="6" captionText="Fig. 3. Diplommatina Benson, 1849 of Nepal. A. D. abiesiana sp. nov., holotype (CDZMTU020). B. D. exserta Godwin-Austen, 1886 (CDZMTU024). C. D. fistulata sp. nov., holotype (CDZMTU025). The yellow arrow indicates the dorsal fold. Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832490" httpUri="https://zenodo.org/record/3832490/files/figure.png" pageId="5" pageNumber="6">Fig. 3A3</figureCitation>
). Columellar tooth weak. Protoconch smooth, Wh 1.5–2.0, with very minute evenly distributed pits, without radial ribs (
<figureCitation id="135462D2FFE1FFEEFE0B448ED755FB85" box="[421,519,1004,1030]" captionStart="Fig" captionStartId="6.[189,232,1930,1956]" captionTargetBox="[189,1393,274,1888]" captionTargetId="figure@6.[189,1398,269,1893]" captionTargetPageId="6" captionText="Fig. 3. Diplommatina Benson, 1849 of Nepal. A. D. abiesiana sp. nov., holotype (CDZMTU020). B. D. exserta Godwin-Austen, 1886 (CDZMTU024). C. D. fistulata sp. nov., holotype (CDZMTU025). The yellow arrow indicates the dorsal fold. Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832490" httpUri="https://zenodo.org/record/3832490/files/figure.png" pageId="5" pageNumber="6">Fig. 3A</figureCitation>
<figureCitation id="135462D2FFE1FFEEFDA9448ED740FB85" box="[519,530,1004,1030]" captionStart="Fig" captionStartId="2.[189,235,1999,2025]" captionTargetBox="[189,1399,1129,1958]" captionTargetId="figure@2.[187,1401,1126,1960]" captionTargetPageId="2" captionText="Fig.1. Map of the localities and areas surveyed for Diplommatina spp. in Nepal." figureDoi="http://doi.org/10.5281/zenodo.3832486" httpUri="https://zenodo.org/record/3832486/files/figure.png" pageId="5" pageNumber="6">1</figureCitation>
). Teleoconch with dense radial ribs; 9–12 ribs/
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on the penultimate whorl; 7–9 ribs/
<quantity id="4C97D3B2FFE1FFEEFED74376D480FBAC" box="[377,466,1044,1071]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="6" unit="mm" value="0.5">0.5 mm</quantity>
on the body whorl towards the aperture (
<figureCitation id="135462D2FFE1FFEEFC1A4376D142FBAC" box="[948,1040,1044,1071]" captionStart="Fig" captionStartId="6.[189,232,1930,1956]" captionTargetBox="[189,1393,274,1888]" captionTargetId="figure@6.[189,1398,269,1893]" captionTargetPageId="6" captionText="Fig. 3. Diplommatina Benson, 1849 of Nepal. A. D. abiesiana sp. nov., holotype (CDZMTU020). B. D. exserta Godwin-Austen, 1886 (CDZMTU024). C. D. fistulata sp. nov., holotype (CDZMTU025). The yellow arrow indicates the dorsal fold. Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832490" httpUri="https://zenodo.org/record/3832490/files/figure.png" pageId="5" pageNumber="6">Fig. 3A</figureCitation>
<figureCitation id="135462D2FFE1FFEEFBBE4377D14DFBAC" box="[1040,1055,1045,1071]" captionStart="Fig" captionStartId="4.[189,232,1895,1921]" captionTargetBox="[189,1398,285,1853]" captionTargetId="figure@4.[189,1398,285,1853]" captionTargetPageId="4" captionText="Fig. 2. Shell terminology. A. D. regularis Fulton, 1901. B. D. folliculus (L. Pfeiffer, 1846). C. General outline of the lower part of a shell from Diplommatina species. Abbreviations: Prt = Parietalis (Prt1 = inner parietalis, Prt2 = outer parietalis); Pt = Palatalis; Hpt = Horizontal palatalis (Hpt1 = lower, Hpt2 = upper in case of two); Vpt = Vertical palatalis (Vpt1 = left side, Vpt2 = right side in case of two)." figureDoi="http://doi.org/10.5281/zenodo.3832488" httpUri="https://zenodo.org/record/3832488/files/figure.png" pageId="5" pageNumber="6">2</figureCitation>
, A
<figureCitation id="135462D2FFE1FFEEFBEC4377D100FBAC" box="[1090,1106,1045,1071]" captionStart="Fig" captionStartId="9.[189,232,1930,1956]" captionTargetBox="[189,1398,274,1888]" captionTargetId="figure@9.[189,1398,266,1891]" captionTargetPageId="9" captionText="Fig. 4. Diplommatina Benson, 1849 of Nepal. A. D. folliculus (L. Pfeiffer, 1846) (CDZMTU028). B. D. godawariensis sp. nov., holotype (CDZMTU032). C. D. maipokhariensis sp. nov., holotype (CDZMTU034). Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832492" httpUri="https://zenodo.org/record/3832492/files/figure.png" pageId="5" pageNumber="6">4</figureCitation>
). Spiral striation present. Umbilicus open. Aperture circular. Peristome with two lips; inner lip thin; 8–10 thin ribs between the two lips (
<figureCitation id="135462D2FFE1FFEEFE844306D4D5FBFC" box="[298,391,1124,1151]" captionStart="Fig" captionStartId="6.[189,232,1930,1956]" captionTargetBox="[189,1393,274,1888]" captionTargetId="figure@6.[189,1398,269,1893]" captionTargetPageId="6" captionText="Fig. 3. Diplommatina Benson, 1849 of Nepal. A. D. abiesiana sp. nov., holotype (CDZMTU020). B. D. exserta Godwin-Austen, 1886 (CDZMTU024). C. D. fistulata sp. nov., holotype (CDZMTU025). The yellow arrow indicates the dorsal fold. Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832490" httpUri="https://zenodo.org/record/3832490/files/figure.png" pageId="5" pageNumber="6">Fig. 3A</figureCitation>
<figureCitation id="135462D2FFE1FFEEFE294307D4C5FBFC" box="[391,407,1125,1151]" captionStart="Fig" captionStartId="9.[189,232,1930,1956]" captionTargetBox="[189,1398,274,1888]" captionTargetId="figure@9.[189,1398,266,1891]" captionTargetPageId="9" captionText="Fig. 4. Diplommatina Benson, 1849 of Nepal. A. D. folliculus (L. Pfeiffer, 1846) (CDZMTU028). B. D. godawariensis sp. nov., holotype (CDZMTU032). C. D. maipokhariensis sp. nov., holotype (CDZMTU034). Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832492" httpUri="https://zenodo.org/record/3832492/files/figure.png" pageId="5" pageNumber="6">4</figureCitation>
). Parietal wall of the aperture extending to less than half the body whorl. Operculum paucispiral, corneous, thin, transparent, slightly concave, outer surface smooth, inner surface with weak ridge at the columellar margin.
</paragraph>
</subSubSection>
<subSubSection id="C3752DDCFFE1FFEEFF134399D100FACB" box="[189,1106,1275,1353]" pageId="5" pageNumber="6" type="etymology">
<paragraph id="8BD07E57FFE1FFEEFF134399D416FA96" blockId="5.[189,324,1275,1301]" box="[189,324,1275,1301]" pageId="5" pageNumber="6">
<heading id="D098C93BFFE1FFEEFF134399D416FA96" bold="true" box="[189,324,1275,1301]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="B91BA245FFE1FFEEFF134399D416FA96" bold="true" box="[189,324,1275,1301]" pageId="5" pageNumber="6">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BD07E57FFE1FFEEFF13424CD100FACB" blockId="5.[189,1106,1326,1353]" box="[189,1106,1326,1353]" pageId="5" pageNumber="6">
The species name refers to the habitat of the
<typeStatus id="54D4C0F5FFE1FFEEFD64424DD7AFFACA" box="[714,765,1327,1353]" pageId="5" pageNumber="6">type</typeStatus>
locality, i.e., an
<taxonomicName id="4C6F05D4FFE1FFEEFC10424CD6ADFACB" box="[958,1023,1326,1352]" class="Pinopsida" family="Pinaceae" genus="Abies" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="genus">
<emphasis id="B91BA245FFE1FFEEFC10424CD6ADFACB" box="[958,1023,1326,1352]" italics="true" pageId="5" pageNumber="6">Abies</emphasis>
</taxonomicName>
forest.
</paragraph>
</subSubSection>
<subSubSection id="C3752DDCFFE1FFEEFF134217D638F979" pageId="5" pageNumber="6" type="materials_examined">
<paragraph id="8BD07E57FFE1FFEEFF134217D43EFA0D" blockId="5.[189,364,1396,1423]" box="[189,364,1396,1423]" pageId="5" pageNumber="6">
<heading id="D098C93BFFE1FFEEFF134217D43EFA0D" bold="true" box="[189,364,1396,1423]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="B91BA245FFE1FFEEFF134217D43EFA0D" bold="true" box="[189,364,1396,1423]" pageId="5" pageNumber="6">Type material</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B07740AFFE1FFEEFF4442CAD66CF991" ID-GBIF-Occurrence="2620517305" collectingDate="2019-05" collectionCode="CDZMTU" collectorName="K. Saud" country="Nepal" elevation="3443" latitude="28.110153" location="Cholangpati" longLatPrecision="1" longitude="85.334236" municipality="Langtang National Park" pageId="5" pageNumber="6" specimenCode="CDZMTU020" specimenCount="1" typeStatus="holotype">
<paragraph id="8BD07E57FFE1FFEEFF4442CAD40EFA41" blockId="5.[189,1399,1448,1555]" box="[234,348,1448,1474]" pageId="5" pageNumber="6">
<emphasis id="B91BA245FFE1FFEEFF4442CAD40EFA41" bold="true" box="[234,348,1448,1474]" pageId="5" pageNumber="6">
<typeStatus id="54D4C0F5FFE1FFEEFF4442CAD40EFA41" box="[234,348,1448,1474]" pageId="5" pageNumber="6" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BD07E57FFE1FFEEFF1342B2D66CF991" blockId="5.[189,1399,1448,1555]" pageId="5" pageNumber="6">
<collectingCountry id="F3783EC7FFE1FFEEFF1342B2D44CFA69" box="[189,286,1488,1514]" name="Nepal" pageId="5" pageNumber="6">NEPAL</collectingCountry>
:
<location id="8EB0288CFFE1FFEEFE8042B2D492FA69" LSID="urn:lsid:plazi:treatment:03C6CF41FFE1FFEEFE5C4538D729F868:8EB0288CFFE1FFEEFE8042B2D492FA69" box="[302,448,1488,1514]" country="Nepal" latitude="28.110153" longLatPrecision="1" longitude="85.334236" municipality="Langtang National Park" name="Cholangpati" pageId="5" pageNumber="6">Cholangpati</location>
,
<collectingMunicipality id="6BB4E42DFFE1FFEEFE7C42B3D7AAFA69" box="[466,760,1488,1515]" pageId="5" pageNumber="6">Langtang National Park</collectingMunicipality>
,
<quantity id="4C97D3B2FFE1FFEEFCA742B2D638FA68" box="[777,874,1488,1515]" metricMagnitude="3" metricUnit="m" metricValue="3.443" pageId="5" pageNumber="6" unit="m" value="3443.0">
<elevation id="00429964FFE1FFEEFCA742B2D638FA68" box="[777,874,1488,1515]" metricMagnitude="3" metricUnit="m" metricValue="3.443" pageId="5" pageNumber="6" unit="m" value="3443.0">3443 m</elevation>
</quantity>
,
<taxonomicName id="4C6F05D4FFE1FFEEFCD242B2D6EFFA69" box="[892,957,1488,1514]" class="Pinopsida" family="Pinaceae" genus="Abies" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="genus">
<emphasis id="B91BA245FFE1FFEEFCD242B2D6EFFA69" box="[892,957,1488,1514]" italics="true" pageId="5" pageNumber="6">Abies</emphasis>
</taxonomicName>
forest,
<geoCoordinate id="EE5B1890FFE1FFEEFB8F42B2D190FA68" box="[1057,1218,1488,1515]" degrees="28.110154" direction="north" orientation="latitude" pageId="5" pageNumber="6" precision="1" value="28.110153">28.110154 N</geoCoordinate>
,
<geoCoordinate id="EE5B1890FFE1FFEEFB7A42B2D023FA68" box="[1236,1393,1488,1515]" degrees="85.334238" direction="east" orientation="longitude" pageId="5" pageNumber="6" precision="1" value="85.334236">85.334238 E</geoCoordinate>
,
<date id="FFD15897FFE1FFEEFF13429AD44BF990" box="[189,281,1528,1555]" pageId="5" pageNumber="6" value="2019-05">
<collectingDate id="EF95A17FFFE1FFEEFF13429AD44BF990" box="[189,281,1528,1555]" pageId="5" pageNumber="6" value="2019-05">19 May</collectingDate>
</date>
, R. Devkota and
<collectorName id="269A1B81FFE1FFEEFE4D429BD710F991" box="[483,578,1528,1555]" pageId="5" pageNumber="6">K. Saud</collectorName>
leg. (
<specimenCode id="DBC9D62CFFE1FFEEFD2D429AD660F991" box="[643,818,1528,1554]" collectionCode="CDZMTU" pageId="5" pageNumber="6">CDZMTU020</specimenCode>
).
</paragraph>
</materialsCitation>
<paragraph id="8BD07E57FFE1FFEEFF44415DD435F9DA" blockId="5.[189,1399,1599,1786]" box="[234,359,1599,1625]" pageId="5" pageNumber="6">
<emphasis id="B91BA245FFE1FFEEFF44415DD435F9DA" bold="true" box="[234,359,1599,1625]" pageId="5" pageNumber="6">
<typeStatus id="54D4C0F5FFE1FFEEFF44415DD435F9DA" box="[234,359,1599,1625]" pageId="5" pageNumber="6" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BD07E57FFE1FFEEFF134105D638F979" blockId="5.[189,1399,1599,1786]" pageId="5" pageNumber="6">
<collectingCountry id="F3783EC7FFE1FFEEFF134105D44CF902" box="[189,286,1639,1665]" name="Nepal" pageId="5" pageNumber="6">NEPAL</collectingCountry>
: 3 shells from the type locality, same data as
<typeStatus id="54D4C0F5FFE1FFEEFCE24104D6E1F903" box="[844,947,1638,1664]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED7EE692FFE1FFEEFC6B4105D11BF902" box="[965,1097,1639,1665]" pageId="5" pageNumber="6">CDZMTU</collectionCode>
021); 8 shells, Baghdwar,
<quantity id="4C97D3B2FFE1FFEEFF1341EDD44EF92A" box="[189,284,1679,1705]" metricMagnitude="3" metricUnit="m" metricValue="2.314" pageId="5" pageNumber="6" unit="m" value="2314.0">2314 m</quantity>
, Shivapuri-Nagarjun National Park,
<geoCoordinate id="EE5B1890FFE1FFEEFD7741ECD609F92A" box="[729,859,1678,1705]" degrees="27.8052" direction="north" orientation="latitude" pageId="5" pageNumber="6" precision="5" value="27.8052">27.8052 N</geoCoordinate>
,
<geoCoordinate id="EE5B1890FFE1FFEEFCC541ECD6AAF92A" box="[875,1016,1678,1705]" degrees="85.38447" direction="east" orientation="longitude" pageId="5" pageNumber="6" precision="1" value="85.38447">85.38447 E</geoCoordinate>
,
<date id="FFD15897FFE1FFEEFBA741EDD1F9F92A" box="[1033,1195,1679,1705]" pageId="5" pageNumber="6" value="2007-06-24">24 Jun. 2007</date>
, P.B. Budha and R. Devkota leg. (
<collectionCode id="ED7EE692FFE1FFEEFE3C41D5D747F951" box="[402,533,1719,1746]" pageId="5" pageNumber="6">CDZMTU</collectionCode>
055); 1 shell, Dhunche,
<quantity id="4C97D3B2FFE1FFEEFC9541D5D6C9F952" box="[827,923,1719,1746]" metricMagnitude="3" metricUnit="m" metricValue="2.056" pageId="5" pageNumber="6" unit="m" value="2056.0">2056 m</quantity>
, Langtang National Park,
<geoCoordinate id="EE5B1890FFE1FFEEFB4E41D5D020F952" box="[1248,1394,1719,1745]" degrees="28.10894" direction="north" orientation="latitude" pageId="5" pageNumber="6" precision="1" value="28.10894">28.10894 N</geoCoordinate>
,
<geoCoordinate id="EE5B1890FFE1FFEEFF1341BDD414F97A" box="[189,326,1759,1786]" degrees="85.29155" direction="east" orientation="longitude" pageId="5" pageNumber="6" precision="1" value="85.29155">85.29155 E</geoCoordinate>
,
<date id="FFD15897FFE1FFEEFEFD41BDD48CF97A" box="[339,478,1759,1785]" pageId="5" pageNumber="6" value="2007-06-03">3 Jun. 2007</date>
, P.B. Budha leg. (
<collectionCode id="ED7EE692FFE1FFEEFD0141BDD663F979" box="[687,817,1759,1786]" pageId="5" pageNumber="6">CDZMTU</collectionCode>
056).
</paragraph>
</subSubSection>
<subSubSection id="C3752DDCFFE1FFEEFF134047D729F868" pageId="5" pageNumber="6" type="discussion">
<paragraph id="8BD07E57FFE1FFEEFF134047D47DF8BC" blockId="5.[189,303,1829,1855]" box="[189,303,1829,1855]" pageId="5" pageNumber="6">
<heading id="D098C93BFFE1FFEEFF134047D47DF8BC" bold="true" box="[189,303,1829,1855]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="B91BA245FFE1FFEEFF134047D47DF8BC" bold="true" box="[189,303,1829,1855]" pageId="5" pageNumber="6">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BD07E57FFE1FFEEFF13403AD729F868" blockId="5.[189,1398,1880,2027]" pageId="5" pageNumber="6">
The shell form of
<taxonomicName id="4C6F05D4FFE1FFEEFE0C403BD76EF8F1" authorityName="Budha &amp; Naggs &amp; Backeljau" authorityYear="2017" box="[418,572,1880,1906]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="abiesiana">
<emphasis id="B91BA245FFE1FFEEFE0C403BD76EF8F1" box="[418,572,1880,1906]" italics="true" pageId="5" pageNumber="6">D. abiesiana</emphasis>
</taxonomicName>
is similar to that of
<taxonomicName id="4C6F05D4FFE1FFEEFCED403BD1F9F8F0" authority="W. T. Blanford, 1865" authorityName="W. T. Blanford" authorityYear="1865" box="[835,1195,1880,1907]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="nana">
<emphasis id="B91BA245FFE1FFEEFCED403BD6F4F8F1" box="[835,934,1881,1906]" italics="true" pageId="5" pageNumber="6">D. nana</emphasis>
W.T.
<bibRefCitation id="EFFE03A6FFE1FFEEFC5A403AD1F9F8F0" author="Blanford W. T." box="[1012,1195,1880,1907]" pageId="5" pageNumber="6" pagination="66 - 105" refId="ref13030" refString="Blanford W. T. 1865. Contributions to Indian Malacology, No. V. Descriptions of new land shells from Arakan, Pegu and Ava; with notes on the distribution of described species. Journal of the Asiatic Society of Bengal 34: 66 - 105." type="journal article" year="1865">Blanford, 1865</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="EFFE03A6FFE1FFEEFB6E403AD03DF8F0" author="Blanford W. T." box="[1216,1391,1880,1907]" pageId="5" pageNumber="6" pagination="66 - 105" refId="ref13030" refString="Blanford W. T. 1865. Contributions to Indian Malacology, No. V. Descriptions of new land shells from Arakan, Pegu and Ava; with notes on the distribution of described species. Journal of the Asiatic Society of Bengal 34: 66 - 105." type="journal article" year="1865">Blanford 1865</bibRefCitation>
;
<bibRefCitation id="EFFE03A6FFE1FFEEFF1340E2D4A2F819" author="Hanley S. &amp; Theobald W." box="[189,496,1920,1947]" pageId="5" pageNumber="6" refId="ref13613" refString="Hanley S. &amp; Theobald W. 1876. Conchologia Indica; being illustrations of the land and freshwater shells of British India. London. https: // doi. org / 10.5962 / bhl. title. 14456" type="book" year="1876">Hanley &amp; Theobald 1876</bibRefCitation>
: pl. 140, fig.1;
<bibRefCitation id="EFFE03A6FFE1FFEEFD0040E2D6FCF819" author="Godwin-Austen H. H." box="[686,942,1920,1946]" pageId="5" pageNumber="6" pagination="165 - 185" refId="ref13270" refString="Godwin-Austen H. H. 1886. Land and freshwater Mollusca of India. Conchologia Indica 1 (5): 165 - 185." type="journal article" year="1886">Godwin-Austen 1886</bibRefCitation>
: pl. 49, fig. 6.), but differs by its less prominent columellar tooth, the more widely-separated radial ribs, the penultimate and body whorls of equal width, and the circular aperture.
</paragraph>
</subSubSection>
</treatment>
</document>
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