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<document id="5F00B68921FAF63398BC47D5478AA6EA" ID-DOI="10.5252/zoosystema2024v46a5" ID-ISSN="1638-9387" ID-Zenodo-Dep="10803267" ID-ZooBank="1B0779F3-FAE4-4105-8130-A26DE519721E" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1710145347001" checkinUser="plazi" docAuthor="Cardoso, Giovanna Monticelli, M, Diego de, Bento, edeiros &amp; Ferreira, Rodrigo Lopes" docDate="2024" docId="03C687BFC609FFD713D0F7EE69C3F856" docLanguage="en" docName="zoosystema.2024.46.5.pdf" docOrigin="Zoosystema 46 (5)" docSource="https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a5.pdf" docStyle="DocumentStyle:5CC576F744449C1860B4E72ABD884631.7:Zoosystema.2020-.journal_article" docStyleId="5CC576F744449C1860B4E72ABD884631" docStyleName="Zoosystema.2020-.journal_article" docStyleVersion="7" docTitle="Ctenorillo intertidalis Cardoso &amp; Ferreira 2024, n. sp." docType="treatment" docUuid="B61E5142-E53A-4AD7-AF85-B24944928069" docUuidSource="ZooBank" docVersion="2" lastPageNumber="110" masterDocId="FFFFFFC7C606FFC6136FF25C6C6BFFBA" masterDocTitle="Unveiling a hidden diversity: descriptions of nine new species of Ctenorillo Verhoeff, 1942 (Isopoda, Armadillidae) discovered in Brazilian caves and their importance for conservation" masterLastPageNumber="132" masterPageNumber="95" pageNumber="108" updateTime="1710145921992" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="9BC833DF16280EBCEAAEAF2DCF57C7D9">Unveiling a hidden diversity: descriptions of nine new species of Ctenorillo Verhoeff, 1942 (Isopoda, Armadillidae) discovered in Brazilian caves and their importance for conservation</mods:title>
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<mods:namePart id="4B37A4B5E2523BC9832914D9A9F521C9">Cardoso, Giovanna Monticelli</mods:namePart>
<mods:affiliation id="40816EFD650CB4713D4F25535B8A08EA">Universidade Federal de Lavras (UFLA), Centro de Estudos em Biologia Subterrânea, Departamento de Biologia, programa de Pós-graduação em Ecologia Aplicada, Campus Universitário, CP 3037, 37200 - 000 Lavras, Minas Gerais (Brasil) gmcardoso. bio @ gmail. com (corresponding author)</mods:affiliation>
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<mods:namePart id="68C5B08AFA0CA45CB01E8D05C289E5E1">M, Diego de</mods:namePart>
</mods:name>
<mods:name id="F501669D04EE92D2F8110B65B96C7977" type="personal">
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<mods:namePart id="0086B822FB254C69866A7629DCF77F43">Bento, edeiros</mods:namePart>
</mods:name>
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<mods:namePart id="5CE65F121C5EF3957B3EB6F7D3A91EBB">Ferreira, Rodrigo Lopes</mods:namePart>
<mods:affiliation id="AC093EFA745116F174F5AFE96A9511BC">Universidade Federal de Lavras (UFLA), Centro de Estudos em Biologia Subterrânea, Departamento de Biologia, programa de Pós-graduação em Ecologia Aplicada, Campus Universitário, CP 3037, 37200 - 000 Lavras, Minas Gerais (Brasil) drops @ ufla. br</mods:affiliation>
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<mods:title id="1EB0204330364F225653AC3E7A48462E">Zoosystema</mods:title>
</mods:titleInfo>
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<mods:date id="7568A15569FC7CED110CC05CD4CD3AB4">2024</mods:date>
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<mods:number id="062812B31677A826FBC07224438A7497">2024-03-05</mods:number>
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<mods:url id="371CCA5EBE1089AA49E1C98912D2A246">https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a5.pdf</mods:url>
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<subSubSection id="C3756522C609FFC913D0F7EE6EA7FA77" box="[191,716,1458,1485]" pageId="15" pageNumber="108" type="nomenclature">
<paragraph id="8BD036A9C609FFC913D0F7EE6EA7FA77" blockId="15.[191,716,1458,1517]" box="[191,716,1458,1485]" pageId="15" pageNumber="108">
<heading id="D09881C5C609FFC913D0F7EE6EA7FA77" box="[191,716,1458,1485]" centered="true" fontSize="11" level="2" pageId="15" pageNumber="108" reason="2">
<taxonomicName id="4C6F4D2AC609FFC913D0F7EE6EECFA77" authority="Cardoso &amp; Ferreira" authorityName="Cardoso &amp; Ferreira" authorityYear="2024" box="[191,647,1458,1485]" class="Malacostraca" family="Armadillidae" genus="Ctenorillo" kingdom="Animalia" order="Isopoda" pageId="15" pageNumber="108" phylum="Arthropoda" rank="species" species="intertidalis" status="sp. nov.">
<emphasis id="B91BEABBC609FFC913D0F7EE6DC0FA77" bold="true" box="[191,427,1458,1485]" italics="true" pageId="15" pageNumber="108">Ctenorillo intertidalis</emphasis>
Cardoso &amp; Ferreira
</taxonomicName>
,
<taxonomicNameLabel id="A22857C0C609FFC911FEF7EF6EA7FA77" box="[657,716,1459,1485]" pageId="15" pageNumber="108" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="C3756522C609FFC91251F78F6E82F98B" pageId="15" pageNumber="108" type="description">
<paragraph id="8BD036A9C609FFC91251F78F6E26FA57" blockId="15.[191,716,1458,1517]" box="[318,589,1491,1517]" pageId="15" pageNumber="108">
(
<figureCitation id="13542A2CC609FFC9122AF78F6DC4FA57" box="[325,431,1491,1517]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStartId-0="15.[132,143,1325,1343]" captionStartId-1="16.[132,143,1409,1427]" captionStartId-2="17.[131,142,1160,1178]" captionTargetBox-0="[137,1449,217,1272]" captionTargetBox-1="[146,1444,223,1365]" captionTargetBox-2="[132,1455,214,1111]" captionTargetId-0="figure-385@15.[132,1455,214,1281]" captionTargetId-1="figure-284@16.[132,1455,215,1366]" captionTargetId-2="figure-438@17.[132,1455,214,1111]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionTargetPageId-2="17" captionText-0="FIG. 8. — Ctenorillo intertidalis Cardoso &amp; Ferreira,n. sp., paratype (male,7.5 mm,ISLA96786):A, habitus,lateral view;B, disposition of dorsal tubercles;C, cephalon and pereonite 1, frontal view; D, cephalon and pereonite 1, dorsal view; E, pleonites 3-5, uropods and telson, dorsal view; F, antennula; G, antenna; H, right mandible; I, left mandible; J, maxillula; K, maxilla; L, maxilliped. Scale bar: 0.5 mm." captionText-1="FIG. 9. — Ctenorillo intertidalis Cardoso &amp; Ferreira, n. sp.paratype (male,7.5 mm, ISLA96786): A, uropod,B, pereopod 1; C, pereopod 7; D, pleopod 1 and genital papilla; E, pleopod 2; F, pleopod 3 exopod; G, pleopod 4 exopod; H, pleopod 5 exopod." captionText-2="FIG. 10. — Ctenorillo intertidalis Cardoso &amp; Ferreira, n. sp.: A, quartzite outcrops on the shoreline of Jericoacoara, with a cave entrance (yellow star); B, inner portion of a cave where specimens are found; C, Ctenorillo intertidalis Cardoso &amp; Ferreira, n. sp. specimen; D, habitus in natural condition sharing the habitat (under rocks) with other isopods." figureDoi-0="http://doi.org/10.5281/zenodo.10803285" figureDoi-1="http://doi.org/10.5281/zenodo.10803287" figureDoi-2="http://doi.org/10.5281/zenodo.10803289" httpUri-0="https://zenodo.org/record/10803285/files/figure.png" httpUri-1="https://zenodo.org/record/10803287/files/figure.png" httpUri-2="https://zenodo.org/record/10803289/files/figure.png" pageId="15" pageNumber="108">Figs 8-10</figureCitation>
;
<figureCitation id="13542A2CC609FFC912D5F78F6D85FA57" box="[442,494,1491,1517]" captionStart="FIG" captionStartId="35.[132,143,1225,1243]" captionTargetBox="[132,1455,216,1183]" captionTargetId="figure-480@35.[699,983,655,744]" captionTargetPageId="35" captionText="FIG. 27. — Habitus of the species of the genus Ctenorillo Verhoeff, 1942 discussed in the present paper:A, Ctenorillo ferrarai Campos-Filho, Araujo &amp; Taiti,2014; B, Ctenorillo pelado Cardoso &amp; Ferreira, n. sp.; C, Ctenorillo araguaia Cardoso &amp; Ferreira, n. sp.; D, Ctenorillo intertidalis Cardoso &amp; Ferreira, n. sp.; E, Ctenorillo ubajarensis Cardoso &amp; Ferreira,n. sp.; F, Ctenorillo cearensis Cardoso &amp; Ferreira,n. sp.; G, Ctenorillo quiteriensis Cardoso &amp; Ferreira,n. sp.; H, Ctenorillo potiguar Cardoso &amp; Ferreira, n. sp.; I, Ctenorillo iuiuensis n. sp.; J, Ctenorillo jequitinhonha Cardoso &amp; Ferreira, n. sp. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10803325" httpUri="https://zenodo.org/record/10803325/files/figure.png" pageId="15" pageNumber="108">27D</figureCitation>
;
<figureCitation id="13542A2CC609FFC91295F78F6E2EFA57" box="[506,581,1491,1517]" captionStart="FIG" captionStartId="36.[133,144,1473,1491]" captionTargetBox="[132,1455,216,1425]" captionTargetId="figure-264@36.[699,984,774,961]" captionTargetPageId="36" captionText="FIG. 28. — SEM images of Ctenorillo Verhoeff, 1942 new species: A-D, Ctenorillo pelado Cardoso &amp; Ferreira, n. sp.: cephalon, dorsal view (A); epimera 1-3, lateral view (B); pleopod 3 exopod (C); epimera 1 and 2, ventral view (D); E, Ctenorillo araguaia Cardoso &amp; Ferreira, n. sp.:, cephalon and epimera 1-3, lateral view; F, G, Ctenorillo intertidalis Cardoso &amp; Ferreira, n. sp.: cephalon, frontal view (F); epimera 1-3, lateral view (G); H, Ctenorillo ubajarensis Cardoso &amp; Ferreira,n. sp.: cephalon and epimera 1-3, lateral view;I, Ctenorillo potiguar Cardoso &amp; Ferreira, n. sp.: cephalon and epimera 1 and 2, lateral view; J, K, Ctenorillo jequitinhonha Cardoso &amp; Ferreira, n. sp.: cephalon, frontal view (J); cephalon and epimera 1 and 2, lateral view (K). Scale bars: A, B, D-K, 0.5 mm; C, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10803327" httpUri="https://zenodo.org/record/10803327/files/figure.png" pageId="15" pageNumber="108">28F, G</figureCitation>
)
</paragraph>
<paragraph id="8BD036A9C609FFC913CDF4436E82F98B" blockId="15.[162,745,1567,1585]" box="[162,745,1567,1585]" pageId="15" pageNumber="108">
<uri id="FFFE3AABC609FFC913CDF4436E82F98B" box="[162,745,1567,1585]" pageId="15" pageNumber="108">
urn:lsid:zoobank.org:act:
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</paragraph>
</subSubSection>
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.—
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.
<collectingCountry id="F3787639C609FFC912C2F4006D8CF9CE" box="[429,487,1628,1652]" name="Brazil" pageId="15" pageNumber="108">Brazil</collectingCountry>
</emphasis>
</materialsCitation>
•
<materialsCitation id="3B073CF4C609FFC91290F4066F6FF91D" collectingDate="2019-09-25" collectionCode="R" collectorName="L. Ferreira" country="Iceland" latitude="-2.7866666" location="Jijoca de Jericoacoara" longLatPrecision="21" longitude="-40.503334" municipality="Jeri" pageId="15" pageNumber="108" specimenCount="1" specimenCount-male="1" stateProvince="Ceara" typeStatus="holotype">
<specimenCount id="9D69FD20C609FFC91290F4066E48F9CB" box="[511,547,1626,1650]" count="1" pageId="15" pageNumber="108" type="male">1 ♂</specimenCount>
(
<quantity id="4C979B4CC609FFC9115DF4066E1BF9C8" box="[562,624,1626,1650]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="15" pageNumber="108" unit="mm" value="6.0">6 mm</quantity>
; parts in slide);
<collectingRegion id="49ABF84BC609FFC913EAF4296C91F936" box="[133,250,1653,1677]" country="Brazil" name="Ceara" pageId="15" pageNumber="108">Ceará state</collectingRegion>
;
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municipality;
<collectingMunicipality id="6BB4ACD3C609FFC911EDF4296EC3F937" box="[642,680,1653,1677]" pageId="15" pageNumber="108">Jeri</collectingMunicipality>
_10 cave;
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,
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;
<date id="FFD11069C609FFC9121DF4D36D89F91D" box="[370,482,1679,1703]" pageId="15" pageNumber="108" value="2019-09-25">
<collectingDate id="EF95E981C609FFC9121DF4D36D89F91D" box="[370,482,1679,1703]" pageId="15" pageNumber="108" value="2019-09-25">25.IX.2019</collectingDate>
</date>
;
<collectionCode id="ED7EAE6CC609FFC91284F4CC6D95F91D" box="[491,510,1680,1703]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="15" pageNumber="108">R</collectionCode>
.
<collectorName id="269A537FC609FFC9116EF4CC6E0AF91D" box="[513,609,1679,1703]" pageId="15" pageNumber="108">L. Ferreira</collectorName>
leg.;
<collectingCountry id="F3787639C609FFC911FEF4D36EACF91D" box="[657,711,1679,1703]" name="Iceland" pageId="15" pageNumber="108">ISLA</collectingCountry>
96786
</materialsCitation>
.
<materialsCitation id="3B073CF4C609FFC913EDF4F66D42F979" box="[130,297,1706,1731]" country="Brazil" location="Brazil" pageId="15" pageNumber="108" specimenCount="1" typeStatus="paratype">
<emphasis id="B91BEABBC609FFC913EDF4F66D42F979" bold="true" box="[130,297,1706,1731]" pageId="15" pageNumber="108">
<typeStatus id="54D4880BC609FFC913EDF4F66C8FF978" box="[130,228,1706,1730]" pageId="15" pageNumber="108" type="paratype">Paratypes</typeStatus>
.
<collectingCountry id="F3787639C609FFC91380F4F76D42F979" box="[239,297,1707,1731]" name="Brazil" pageId="15" pageNumber="108">Brazil</collectingCountry>
</emphasis>
</materialsCitation>
•
<materialsCitation id="3B073CF4C609FFC91250F4F66E9DF978" box="[319,758,1706,1730]" collectingDate="2019-09-25" collectionCode="R" collectorName="L. Ferreira" country="Iceland" latitude="-2.7866666" location="Jijoca de Jericoacoara" longLatPrecision="21" longitude="-40.503334" municipality="Jeri" pageId="15" pageNumber="108" specimenCount="11" specimenCount-female="9" specimenCount-male="2" stateProvince="Ceara" typeStatus="paratype">
<specimenCount id="9D69FD20C609FFC91250F4F66D0EF97B" box="[319,357,1706,1730]" count="2" pageId="15" pageNumber="108" type="male">2 ♂</specimenCount>
,
<specimenCount id="9D69FD20C609FFC9121DF4F66DF8F97B" box="[370,403,1706,1730]" count="9" pageId="15" pageNumber="108" type="female">9 ♀</specimenCount>
; same data as holotype;
<collectingCountry id="F3787639C609FFC911EEF4F66ED2F978" box="[641,697,1706,1730]" name="Iceland" pageId="15" pageNumber="108">ISLA</collectingCountry>
96787
</materialsCitation>
.
</paragraph>
<paragraph id="8BD036A9C609FFC913EBF4BE6ED0F852" blockId="15.[131,775,1762,2025]" pageId="15" pageNumber="108">
<materialsCitation id="3B073CF4C609FFC913EBF4BE6D68F8AF" collectionCode="OTHER, MATERIAL" country="Brazil" location="Jijoca de Jericoacoara" pageId="15" pageNumber="108" specimenCount="1" stateProvince="Ceara">
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. —
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<collectingCountry id="F3787639C609FFC91209F4B86DCFF946" box="[358,420,1764,1788]" name="Brazil" pageId="15" pageNumber="108">Brazil</collectingCountry>
.
</emphasis>
<collectingRegion id="49ABF84BC609FFC912DEF4BE6E48F940" box="[433,547,1762,1786]" country="Brazil" name="Ceara" pageId="15" pageNumber="108">Ceará state</collectingRegion>
;
<location id="8EB06072C609FFC91140F4BE6F6CF940" LSID="urn:lsid:plazi:treatment:03C687BFC609FFD713D0F7EE69C3F856:8EB06072C609FFC91140F4BE6F6CF940" box="[559,775,1762,1786]" country="Brazil" name="Jijoca de Jericoacoara" pageId="15" pageNumber="108" stateProvince="Ceara">Jijoca de Jericoacoara</location>
municipality
</materialsCitation>
•
<materialsCitation id="3B073CF4C609FFC91272F4A16C93F8F0" collectingDate="2019-09-25" collectionCode="R" collectorName="L. Ferreira &amp; Jeri" country="Brazil" latitude="-2.7883334" location="Jeri" longLatPrecision="21" longitude="-40.511665" pageId="15" pageNumber="108" specimenCount="23" specimenCount-female="12" specimenCount-male="11">
<specimenCount id="9D69FD20C609FFC91272F4A16D27F8AE" box="[285,332,1789,1813]" count="10" pageId="15" pageNumber="108" type="male">10♂</specimenCount>
,
<specimenCount id="9D69FD20C609FFC91234F4A16DE3F8AE" box="[347,392,1789,1813]" count="12" pageId="15" pageNumber="108" type="female">12 ♀</specimenCount>
;
<location id="8EB06072C609FFC912F8F4A16DD1F8AF" LSID="urn:lsid:plazi:treatment:03C687BFC609FFD713D0F7EE69C3F856:8EB06072C609FFC912F8F4A16DD1F8AF" box="[407,442,1789,1813]" country="Brazil" latitude="-2.7883334" longLatPrecision="21" longitude="-40.511665" name="Jeri" pageId="15" pageNumber="108">Jeri</location>
_4 cave;
<geoCoordinate id="EE5B506EC609FFC9117DF4A16E13F8AF" box="[530,632,1789,1813]" degrees="2" direction="south" minutes="47" orientation="latitude" pageId="15" pageNumber="108" precision="15" seconds="18" value="-2.7883334">2°47’18”S</geoCoordinate>
,
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;
<date id="FFD11069C609FFC913EBF54B6C93F895" box="[132,248,1815,1839]" pageId="15" pageNumber="108" value="2019-09-25">
<collectingDate id="EF95E981C609FFC913EBF54B6C93F895" box="[132,248,1815,1839]" pageId="15" pageNumber="108" value="2019-09-25">25.IX.2019</collectingDate>
</date>
;
<collectionCode id="ED7EAE6CC609FFC9126FF5446D78F895" box="[256,275,1816,1839]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="15" pageNumber="108">R</collectionCode>
.
<collectorName id="269A537FC609FFC91279F5446D11F895" box="[278,378,1815,1839]" pageId="15" pageNumber="108">L. Ferreira</collectorName>
leg.; ISLA96788
<specimenCount id="9D69FD20C609FFC9114EF54B6E2EF894" box="[545,581,1815,1839]" count="1" pageId="15" pageNumber="108" type="male">1 ♂</specimenCount>
in slide.
<collectorName id="269A537FC609FFC911F6F54B6ED7F895" box="[665,700,1815,1839]" pageId="15" pageNumber="108">Jeri</collectorName>
_4 cave; ISLA96789
</materialsCitation>
•
<materialsCitation id="3B073CF4C609FFC9127DF56E6E53F8DE" collectingDate="2019-09-24" collectionCode="R" collectorName="L. Ferreira" country="Brazil" latitude="-2.7877777" location="Jeri" longLatPrecision="21" longitude="-40.509445" pageId="15" pageNumber="108" specimenCount="21" specimenCount-female="11" specimenCount-male="10">
<specimenCount id="9D69FD20C609FFC9127DF56E6D2EF8F3" box="[274,325,1842,1866]" count="10" pageId="15" pageNumber="108" type="male">10 ♂</specimenCount>
,
<specimenCount id="9D69FD20C609FFC9123BF56E6DE8F8F3" box="[340,387,1842,1866]" count="11" pageId="15" pageNumber="108" type="female">11 ♀</specimenCount>
;
<location id="8EB06072C609FFC912FDF56E6DDDF8F0" LSID="urn:lsid:plazi:treatment:03C687BFC609FFD713D0F7EE69C3F856:8EB06072C609FFC912FDF56E6DDDF8F0" box="[402,438,1842,1866]" country="Brazil" latitude="-2.7877777" longLatPrecision="21" longitude="-40.509445" name="Jeri" pageId="15" pageNumber="108">Jeri</location>
_7 cave;
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,
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;
<date id="FFD11069C609FFC913EBF5106C90F8DE" box="[132,251,1868,1892]" pageId="15" pageNumber="108" value="2019-09-24">
<collectingDate id="EF95E981C609FFC913EBF5106C90F8DE" box="[132,251,1868,1892]" pageId="15" pageNumber="108" value="2019-09-24">24.IX.2019</collectingDate>
</date>
;
<collectionCode id="ED7EAE6CC609FFC91267F5116D70F8DE" box="[264,283,1869,1892]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="15" pageNumber="108">R</collectionCode>
.
<collectorName id="269A537FC609FFC91271F5116DE2F8DE" box="[286,393,1868,1892]" pageId="15" pageNumber="108">L. Ferreira</collectorName>
leg.; ISLA96790
</materialsCitation>
•
<materialsCitation id="3B073CF4C609FFC9113DF5106F6CF8C5" collectingDate="2019-09-24" collectionCode="R" collectorName="L. Ferreira" country="Brazil" latitude="-2.7880554" location="Jeri" longLatPrecision="21" longitude="-40.50889" pageId="15" pageNumber="108" specimenCount="18" specimenCount-female="7" specimenCount-male="11">
<specimenCount id="9D69FD20C609FFC9113DF5106EEFF8D9" box="[594,644,1868,1892]" count="11" pageId="15" pageNumber="108" type="male">11 ♂</specimenCount>
,
<specimenCount id="9D69FD20C609FFC911FCF5116EDDF8D9" box="[659,694,1869,1892]" count="7" pageId="15" pageNumber="108" type="female">7 ♀</specimenCount>
;
<location id="8EB06072C609FFC911ABF5106E81F8DE" LSID="urn:lsid:plazi:treatment:03C687BFC609FFD713D0F7EE69C3F856:8EB06072C609FFC911ABF5106E81F8DE" box="[708,746,1868,1892]" country="Brazil" latitude="-2.7880554" longLatPrecision="21" longitude="-40.50889" name="Jeri" pageId="15" pageNumber="108">Jeri</location>
_8;
<geoCoordinate id="EE5B506EC609FFC913E9F53B6C81F8C5" box="[134,234,1895,1919]" degrees="2" direction="south" minutes="47" orientation="latitude" pageId="15" pageNumber="108" precision="15" seconds="17" value="-2.7880554">2°47’17”S</geoCoordinate>
,
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;
<date id="FFD11069C609FFC9121BF53B6D8EF8C5" box="[372,485,1895,1919]" pageId="15" pageNumber="108" value="2019-09-24">
<collectingDate id="EF95E981C609FFC9121BF53B6D8EF8C5" box="[372,485,1895,1919]" pageId="15" pageNumber="108" value="2019-09-24">24.IX.2019</collectingDate>
</date>
;
<collectionCode id="ED7EAE6CC609FFC91281F53B6E6AF8C4" box="[494,513,1895,1918]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="15" pageNumber="108">R</collectionCode>
.
<collectorName id="269A537FC609FFC9116BF53B6E0DF8C5" box="[516,614,1895,1919]" pageId="15" pageNumber="108">L. Ferreira</collectorName>
leg.; ISLA96791
</materialsCitation>
•
<materialsCitation id="3B073CF4C609FFC913FDF5DD6E9CF823" box="[146,759,1921,1945]" collectingDate="2019-09-24" collectionCode="R" collectorName="L. Ferreira" country="Brazil" location="Jeri" pageId="15" pageNumber="108" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D69FD20C609FFC913FDF5DD6CD3F822" box="[146,184,1921,1945]" count="1" pageId="15" pageNumber="108" type="male">1 ♂</specimenCount>
in slide;
<location id="8EB06072C609FFC9127FF5DD6D5EF823" LSID="urn:lsid:plazi:treatment:03C687BFC609FFD713D0F7EE69C3F856:8EB06072C609FFC9127FF5DD6D5EF823" box="[272,309,1921,1945]" country="Brazil" name="Jeri" pageId="15" pageNumber="108">Jeri</location>
_8;
<date id="FFD11069C609FFC91238F5DD6DA1F823" box="[343,458,1921,1945]" pageId="15" pageNumber="108" value="2019-09-24">
<collectingDate id="EF95E981C609FFC91238F5DD6DA1F823" box="[343,458,1921,1945]" pageId="15" pageNumber="108" value="2019-09-24">24.IX.2019</collectingDate>
</date>
;
<collectionCode id="ED7EAE6CC609FFC912BAF5DE6D83F823" box="[469,488,1922,1945]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="15" pageNumber="108">R</collectionCode>
.
<collectorName id="269A537FC609FFC91284F5DE6E3AF823" box="[491,593,1921,1945]" pageId="15" pageNumber="108">L. Ferreira</collectorName>
leg.; ISLA96792
</materialsCitation>
•
<materialsCitation id="3B073CF4C609FFC913EBF5C06DC3F874" collectingDate="2019-09-24" collectionCode="R" collectorName="L. Ferreira" country="Brazil" latitude="-2.7877777" location="Duas" longLatPrecision="21" longitude="-40.509167" pageId="15" pageNumber="108" specimenCount="8" specimenCount-female="1" specimenCount-male="7">
<specimenCount id="9D69FD20C609FFC913EBF5C06CC3F809" box="[132,168,1948,1971]" count="7" pageId="15" pageNumber="108" type="male">7 ♂</specimenCount>
,
<specimenCount id="9D69FD20C609FFC913D9F5C06CBCF809" box="[182,215,1948,1972]" count="1" pageId="15" pageNumber="108" type="female">1 ♀</specimenCount>
;
<location id="8EB06072C609FFC9138AF5C06D73F809" LSID="urn:lsid:plazi:treatment:03C687BFC609FFD713D0F7EE69C3F856:8EB06072C609FFC9138AF5C06D73F809" box="[229,280,1948,1971]" country="Brazil" latitude="-2.7877777" longLatPrecision="21" longitude="-40.509167" name="Duas" pageId="15" pageNumber="108">Duas</location>
bocas cave;
<geoCoordinate id="EE5B506EC609FFC912FAF5C76D91F809" box="[405,506,1947,1971]" degrees="2" direction="south" minutes="47" orientation="latitude" pageId="15" pageNumber="108" precision="15" seconds="16" value="-2.7877777">2°47’16”S</geoCoordinate>
,
<geoCoordinate id="EE5B506EC609FFC91169F5C06EEBF80E" box="[518,640,1948,1972]" degrees="40" direction="west" minutes="30" orientation="longitude" pageId="15" pageNumber="108" precision="15" seconds="33" value="-40.509167">40°30’33”W</geoCoordinate>
;
<date id="FFD11069C609FFC911E0F5C06F69F80E" box="[655,770,1948,1972]" pageId="15" pageNumber="108" value="2019-09-24">
<collectingDate id="EF95E981C609FFC911E0F5C06F69F80E" box="[655,770,1948,1972]" pageId="15" pageNumber="108" value="2019-09-24">24.IX.2019</collectingDate>
</date>
;
<collectionCode id="ED7EAE6CC609FFC913EBF5EA6CFCF877" box="[132,151,1974,1997]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="15" pageNumber="108">R</collectionCode>
.
<collectorName id="269A537FC609FFC913F4F5EA6D6AF874" box="[155,257,1974,1998]" pageId="15" pageNumber="108">L. Ferreira</collectorName>
leg.; ISLA96793
</materialsCitation>
•
<materialsCitation id="3B073CF4C609FFC912D0F5EA6EDCF852" collectingDate="2019-09-25" collectionCode="R" collectorName="L. Ferreira" country="Brazil" latitude="-2.7875" location="Jeri Novo cave" longLatPrecision="21" longitude="-40.509167" pageId="15" pageNumber="108" specimenCount="1">
<specimenCount id="9D69FD20C609FFC912D0F5EA6D92F874" box="[447,505,1974,1998]" count="1" pageId="15" pageNumber="108" type="generic">1 juv.</specimenCount>
;
<location id="8EB06072C609FFC9116DF5EA6EF8F877" LSID="urn:lsid:plazi:treatment:03C687BFC609FFD713D0F7EE69C3F856:8EB06072C609FFC9116DF5EA6EF8F877" box="[514,659,1974,1998]" country="Brazil" latitude="-2.7875" longLatPrecision="21" longitude="-40.509167" name="Jeri Novo cave" pageId="15" pageNumber="108">Jeri Novo cave</location>
;
<geoCoordinate id="EE5B506EC609FFC911F2F5EA6F69F874" box="[669,770,1974,1998]" degrees="2" direction="south" minutes="47" orientation="latitude" pageId="15" pageNumber="108" precision="15" seconds="15" value="-2.7875">2°47’15”S</geoCoordinate>
,
<geoCoordinate id="EE5B506EC609FFC913EBF58D6C95F853" box="[132,254,2001,2025]" degrees="40" direction="west" minutes="30" orientation="longitude" pageId="15" pageNumber="108" precision="15" seconds="33" value="-40.509167">40°30’33”W</geoCoordinate>
;
<date id="FFD11069C609FFC91263F58C6DEBF852" box="[268,384,2000,2024]" pageId="15" pageNumber="108" value="2019-09-25">
<collectingDate id="EF95E981C609FFC91263F58C6DEBF852" box="[268,384,2000,2024]" pageId="15" pageNumber="108" value="2019-09-25">25.IX.2019</collectingDate>
</date>
;
<collectionCode id="ED7EAE6CC609FFC912E5F58D6DF6F852" box="[394,413,2001,2024]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="15" pageNumber="108">R</collectionCode>
.
<collectorName id="269A537FC609FFC912CFF58D6E6DF852" box="[416,518,2000,2024]" pageId="15" pageNumber="108">L. Ferreira</collectorName>
leg.; ISLA 96800
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3756522C609FFC91042F7E56FB2F9BF" pageId="15" pageNumber="108" type="etymology">
<paragraph id="8BD036A9C609FFC91042F7E56FB2F9BF" blockId="15.[813,1455,1465,1542]" pageId="15" pageNumber="108">
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. — The new species name is a noun in apposition that refers to area in where the caves are located, close to the beach, in an intertidal area.
</paragraph>
</subSubSection>
<subSubSection id="C3756522C609FFC91042F44269DBF900" pageId="15" pageNumber="108" type="diagnosis">
<paragraph id="8BD036A9C609FFC91042F44269DBF900" blockId="15.[811,1456,1565,1722]" pageId="15" pageNumber="108">
<smallCapsWord id="8D36A075C609FFC91042F4426FFEF98E" baselines="1583,1584" box="[813,917,1566,1589]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="15" pageNumber="108">DIAGNOSIS</smallCapsWord>
.— Dorsum covered with conical tubercles and lateral ribs with three rows on cephalon (4, 2, 6); four rows on pereonite 1 (4, 7, 6, 7); two rows on pereonites 2-7 (8 +7); one row of four tubercles on pleonites 3-5; and two paramedian tubercles on telson. Pleopod 1 exopod distal portion round, sinuous outer margin; endopod with distal portion slightly bent outward, three times longer than exopod.
</paragraph>
</subSubSection>
<subSubSection id="C3756522C609FFC91042F48E68F2F8B9" pageId="15" pageNumber="108" type="distribution">
<paragraph id="8BD036A9C609FFC91042F48E68F2F8B9" blockId="15.[811,1455,1745,1796]" pageId="15" pageNumber="108">
<smallCapsWord id="8D36A075C609FFC91042F48E6FDCF952" baselines="1763,1764" box="[813,951,1746,1769]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Distribution" pageId="15" pageNumber="108">DISTRIBUTION</smallCapsWord>
. — Jericoacoara National Park, municipality of Jijoca de Jericoacoara, North of
<collectingRegion id="49ABF84BC609FFC91748F4B068FEF8B9" box="[1063,1173,1772,1796]" country="Brazil" name="Ceara" pageId="15" pageNumber="108">Ceará state</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C3756522C609FFD71042F56E680FF8B6" lastPageId="17" lastPageNumber="110" pageId="15" pageNumber="108" type="description">
<paragraph id="8BD036A9C609FFC91042F56E6FD6F8F0" blockId="15.[812,1456,1842,2028]" box="[813,957,1842,1868]" pageId="15" pageNumber="108">
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</paragraph>
<paragraph id="8BD036A9C609FFD61042F50D68AFF956" blockId="15.[812,1456,1842,2028]" lastBlockId="16.[812,1457,1523,1772]" lastPageId="16" lastPageNumber="109" pageId="15" pageNumber="108">
Maximum size:
<specimenCount id="9D69FD20C609FFC910B0F5096F9DF8D1" box="[991,1014,1877,1899]" count="1" pageId="15" pageNumber="108" type="male">♂</specimenCount>
and
<specimenCount id="9D69FD20C609FFC91742F509682AF8D1" box="[1069,1089,1877,1899]" count="1" pageId="15" pageNumber="108" type="female">♀</specimenCount>
,
<quantity id="4C979B4CC609FFC91721F50D68C0F8D6" box="[1102,1195,1873,1900]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="15" pageNumber="108" unit="mm" value="7.5">7.5 mm</quantity>
. Color grey with lateral margins depigmented. Dorsum covered with conical tubercles and lateral ribs (
<figureCitation id="13542A2CC609FFC91741F5CD68CFF816" box="[1070,1188,1937,1964]" captionStart="FIG" captionStartId="15.[132,143,1325,1343]" captionTargetBox="[137,1449,217,1272]" captionTargetId="figure-385@15.[132,1455,214,1281]" captionTargetPageId="15" captionText="FIG. 8. — Ctenorillo intertidalis Cardoso &amp; Ferreira,n. sp., paratype (male,7.5 mm,ISLA96786):A, habitus,lateral view;B, disposition of dorsal tubercles;C, cephalon and pereonite 1, frontal view; D, cephalon and pereonite 1, dorsal view; E, pleonites 3-5, uropods and telson, dorsal view; F, antennula; G, antenna; H, right mandible; I, left mandible; J, maxillula; K, maxilla; L, maxilliped. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10803285" httpUri="https://zenodo.org/record/10803285/files/figure.png" pageId="15" pageNumber="108">Figs 8A, B</figureCitation>
;
<figureCitation id="13542A2CC609FFC917C0F5CD6895F811" box="[1199,1278,1937,1963]" captionStart="FIG" captionStartId="36.[133,144,1473,1491]" captionTargetBox="[132,1455,216,1425]" captionTargetId="figure-264@36.[699,984,774,961]" captionTargetPageId="36" captionText="FIG. 28. — SEM images of Ctenorillo Verhoeff, 1942 new species: A-D, Ctenorillo pelado Cardoso &amp; Ferreira, n. sp.: cephalon, dorsal view (A); epimera 1-3, lateral view (B); pleopod 3 exopod (C); epimera 1 and 2, ventral view (D); E, Ctenorillo araguaia Cardoso &amp; Ferreira, n. sp.:, cephalon and epimera 1-3, lateral view; F, G, Ctenorillo intertidalis Cardoso &amp; Ferreira, n. sp.: cephalon, frontal view (F); epimera 1-3, lateral view (G); H, Ctenorillo ubajarensis Cardoso &amp; Ferreira,n. sp.: cephalon and epimera 1-3, lateral view;I, Ctenorillo potiguar Cardoso &amp; Ferreira, n. sp.: cephalon and epimera 1 and 2, lateral view; J, K, Ctenorillo jequitinhonha Cardoso &amp; Ferreira, n. sp.: cephalon, frontal view (J); cephalon and epimera 1 and 2, lateral view (K). Scale bars: A, B, D-K, 0.5 mm; C, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10803327" httpUri="https://zenodo.org/record/10803327/files/figure.png" pageId="15" pageNumber="108">28F, G</figureCitation>
): cephalon with three rows (4, 2, and 6 from front to back of vertex); pereonite 1 with
<quantity id="4C979B4CC609FFC910D6F58D6F98F851" box="[953,1011,2001,2027]" metricMagnitude="-1" metricUnit="m" metricValue="6.096" pageId="15" pageNumber="108" unit="in" value="24.0">24 in</quantity>
four rows (4, 7, 6 and 7); pereonites 2-7 with 15 (8 + 7); pleonites 3-5 with one row of four tubercles; telson with two paramedian tubercles. Dorsal cuticle (
<figureCitation id="13542A2CC616FFD613E3F46E6C99F9F7" box="[140,242,1586,1613]" captionStart="FIG" captionStartId="36.[133,144,1473,1491]" captionTargetBox="[132,1455,216,1425]" captionTargetId="figure-264@36.[699,984,774,961]" captionTargetPageId="36" captionText="FIG. 28. — SEM images of Ctenorillo Verhoeff, 1942 new species: A-D, Ctenorillo pelado Cardoso &amp; Ferreira, n. sp.: cephalon, dorsal view (A); epimera 1-3, lateral view (B); pleopod 3 exopod (C); epimera 1 and 2, ventral view (D); E, Ctenorillo araguaia Cardoso &amp; Ferreira, n. sp.:, cephalon and epimera 1-3, lateral view; F, G, Ctenorillo intertidalis Cardoso &amp; Ferreira, n. sp.: cephalon, frontal view (F); epimera 1-3, lateral view (G); H, Ctenorillo ubajarensis Cardoso &amp; Ferreira,n. sp.: cephalon and epimera 1-3, lateral view;I, Ctenorillo potiguar Cardoso &amp; Ferreira, n. sp.: cephalon and epimera 1 and 2, lateral view; J, K, Ctenorillo jequitinhonha Cardoso &amp; Ferreira, n. sp.: cephalon, frontal view (J); cephalon and epimera 1 and 2, lateral view (K). Scale bars: A, B, D-K, 0.5 mm; C, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10803327" httpUri="https://zenodo.org/record/10803327/files/figure.png" pageId="16" pageNumber="109">Fig. 28G</figureCitation>
) with short triangular scale setae, pereonites 1-7 bearing one line of
<emphasis id="B91BEABBC616FFD61238F40E6D9EF9D6" box="[343,501,1618,1644]" italics="true" pageId="16" pageNumber="109">noduli lateralis</emphasis>
per side on outer surface of posterior tubercle of second tubercle. Cephalon with frontal shield protruding above vertex; eye consisting of 10 ommatidia (
<figureCitation id="13542A2CC616FFD61250F4EE6D87F977" box="[319,492,1714,1741]" captionStart="FIG" captionStartId="15.[132,143,1325,1343]" captionTargetBox="[137,1449,217,1272]" captionTargetId="figure-385@15.[132,1455,214,1281]" captionTargetPageId="15" captionText="FIG. 8. — Ctenorillo intertidalis Cardoso &amp; Ferreira,n. sp., paratype (male,7.5 mm,ISLA96786):A, habitus,lateral view;B, disposition of dorsal tubercles;C, cephalon and pereonite 1, frontal view; D, cephalon and pereonite 1, dorsal view; E, pleonites 3-5, uropods and telson, dorsal view; F, antennula; G, antenna; H, right mandible; I, left mandible; J, maxillula; K, maxilla; L, maxilliped. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10803285" httpUri="https://zenodo.org/record/10803285/files/figure.png" pageId="16" pageNumber="109">Figs 8A, C, D</figureCitation>
;
<figureCitation id="13542A2CC616FFD61294F4EE6E45F976" box="[507,558,1714,1740]" captionStart="FIG" captionStartId="36.[133,144,1473,1491]" captionTargetBox="[132,1455,216,1425]" captionTargetId="figure-264@36.[699,984,774,961]" captionTargetPageId="36" captionText="FIG. 28. — SEM images of Ctenorillo Verhoeff, 1942 new species: A-D, Ctenorillo pelado Cardoso &amp; Ferreira, n. sp.: cephalon, dorsal view (A); epimera 1-3, lateral view (B); pleopod 3 exopod (C); epimera 1 and 2, ventral view (D); E, Ctenorillo araguaia Cardoso &amp; Ferreira, n. sp.:, cephalon and epimera 1-3, lateral view; F, G, Ctenorillo intertidalis Cardoso &amp; Ferreira, n. sp.: cephalon, frontal view (F); epimera 1-3, lateral view (G); H, Ctenorillo ubajarensis Cardoso &amp; Ferreira,n. sp.: cephalon and epimera 1-3, lateral view;I, Ctenorillo potiguar Cardoso &amp; Ferreira, n. sp.: cephalon and epimera 1 and 2, lateral view; J, K, Ctenorillo jequitinhonha Cardoso &amp; Ferreira, n. sp.: cephalon, frontal view (J); cephalon and epimera 1 and 2, lateral view (K). Scale bars: A, B, D-K, 0.5 mm; C, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10803327" httpUri="https://zenodo.org/record/10803327/files/figure.png" pageId="16" pageNumber="109">28F</figureCitation>
). Pereonite 1 with posterior margin sinuous at sides; inner lobe of schisma rounded, extending beyond posterior margin of outer lobe; pereonite 2 with triangular ventral tooth reaching epimeron posterior margin (
<figureCitation id="13542A2CC616FFD61221F56D6DCCF8F6" box="[334,423,1841,1868]" captionStart="FIG" captionStartId="15.[132,143,1325,1343]" captionTargetBox="[137,1449,217,1272]" captionTargetId="figure-385@15.[132,1455,214,1281]" captionTargetPageId="15" captionText="FIG. 8. — Ctenorillo intertidalis Cardoso &amp; Ferreira,n. sp., paratype (male,7.5 mm,ISLA96786):A, habitus,lateral view;B, disposition of dorsal tubercles;C, cephalon and pereonite 1, frontal view; D, cephalon and pereonite 1, dorsal view; E, pleonites 3-5, uropods and telson, dorsal view; F, antennula; G, antenna; H, right mandible; I, left mandible; J, maxillula; K, maxilla; L, maxilliped. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10803285" httpUri="https://zenodo.org/record/10803285/files/figure.png" pageId="16" pageNumber="109">Fig. 8C</figureCitation>
). Pereonites 2-7 (
<figureCitation id="13542A2CC616FFD61102F56D6EA9F8F6" box="[621,706,1841,1868]" captionStart="FIG" captionStartId="15.[132,143,1325,1343]" captionTargetBox="[137,1449,217,1272]" captionTargetId="figure-385@15.[132,1455,214,1281]" captionTargetPageId="15" captionText="FIG. 8. — Ctenorillo intertidalis Cardoso &amp; Ferreira,n. sp., paratype (male,7.5 mm,ISLA96786):A, habitus,lateral view;B, disposition of dorsal tubercles;C, cephalon and pereonite 1, frontal view; D, cephalon and pereonite 1, dorsal view; E, pleonites 3-5, uropods and telson, dorsal view; F, antennula; G, antenna; H, right mandible; I, left mandible; J, maxillula; K, maxilla; L, maxilliped. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10803285" httpUri="https://zenodo.org/record/10803285/files/figure.png" pageId="16" pageNumber="109">Fig. 8A</figureCitation>
) with wide quadrangular epimera slightly bent outwards. Telson (
<figureCitation id="13542A2CC616FFD613E4F52D6CB7F831" box="[139,220,1905,1931]" captionStart="FIG" captionStartId="15.[132,143,1325,1343]" captionTargetBox="[137,1449,217,1272]" captionTargetId="figure-385@15.[132,1455,214,1281]" captionTargetPageId="15" captionText="FIG. 8. — Ctenorillo intertidalis Cardoso &amp; Ferreira,n. sp., paratype (male,7.5 mm,ISLA96786):A, habitus,lateral view;B, disposition of dorsal tubercles;C, cephalon and pereonite 1, frontal view; D, cephalon and pereonite 1, dorsal view; E, pleonites 3-5, uropods and telson, dorsal view; F, antennula; G, antenna; H, right mandible; I, left mandible; J, maxillula; K, maxilla; L, maxilliped. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10803285" httpUri="https://zenodo.org/record/10803285/files/figure.png" pageId="16" pageNumber="109">Fig. 8E</figureCitation>
) hourglass-shaped, proximal portion broader than distal portion. Antennula (
<figureCitation id="13542A2CC616FFD612C8F5CD6D9CF811" box="[423,503,1937,1963]" captionStart="FIG" captionStartId="15.[132,143,1325,1343]" captionTargetBox="[137,1449,217,1272]" captionTargetId="figure-385@15.[132,1455,214,1281]" captionTargetPageId="15" captionText="FIG. 8. — Ctenorillo intertidalis Cardoso &amp; Ferreira,n. sp., paratype (male,7.5 mm,ISLA96786):A, habitus,lateral view;B, disposition of dorsal tubercles;C, cephalon and pereonite 1, frontal view; D, cephalon and pereonite 1, dorsal view; E, pleonites 3-5, uropods and telson, dorsal view; F, antennula; G, antenna; H, right mandible; I, left mandible; J, maxillula; K, maxilla; L, maxilliped. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10803285" httpUri="https://zenodo.org/record/10803285/files/figure.png" pageId="16" pageNumber="109">Fig. 8F</figureCitation>
) of three articles, second article shorter than first and third, third article with four apical and five subapical aesthetascs. Antenna (
<figureCitation id="13542A2CC616FFD611CBF58D6E94F851" box="[676,767,2001,2027]" captionStart="FIG" captionStartId="15.[132,143,1325,1343]" captionTargetBox="[137,1449,217,1272]" captionTargetId="figure-385@15.[132,1455,214,1281]" captionTargetPageId="15" captionText="FIG. 8. — Ctenorillo intertidalis Cardoso &amp; Ferreira,n. sp., paratype (male,7.5 mm,ISLA96786):A, habitus,lateral view;B, disposition of dorsal tubercles;C, cephalon and pereonite 1, frontal view; D, cephalon and pereonite 1, dorsal view; E, pleonites 3-5, uropods and telson, dorsal view; F, antennula; G, antenna; H, right mandible; I, left mandible; J, maxillula; K, maxilla; L, maxilliped. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10803285" httpUri="https://zenodo.org/record/10803285/files/figure.png" pageId="16" pageNumber="109">Fig. 8G</figureCitation>
) short and stout, flagellum shorter than fifth article of peduncle; second article of flagellum about twice as long as first. Buccal pieces as
<taxonomicName id="4C6F4D2AC616FFD61772F46F690CF9F6" authority="Cardoso &amp; Ferreira" authorityName="Cardoso &amp; Ferreira" authorityYear="2024" box="[1053,1383,1586,1613]" class="Malacostraca" family="Armadillidae" genus="Ctenorillo" kingdom="Animalia" order="Isopoda" pageId="16" pageNumber="109" phylum="Arthropoda" rank="species" species="pelado" status="n. sp.">
<emphasis id="B91BEABBC616FFD61772F46F68EFF9F6" box="[1053,1156,1586,1613]" italics="true" pageId="16" pageNumber="109">C. pelado</emphasis>
Cardoso &amp; Ferreira
</taxonomicName>
,
<taxonomicNameLabel id="A22857C0C616FFD6161DF46F69C4F9F7" box="[1394,1455,1587,1613]" pageId="16" pageNumber="109" rank="species">n. sp.</taxonomicNameLabel>
(
<figureCitation id="13542A2CC616FFD6105BF40E6F1CF9D6" box="[820,887,1618,1644]" captionStart="FIG" captionStartId="15.[132,143,1325,1343]" captionTargetBox="[137,1449,217,1272]" captionTargetId="figure-385@15.[132,1455,214,1281]" captionTargetPageId="15" captionText="FIG. 8. — Ctenorillo intertidalis Cardoso &amp; Ferreira,n. sp., paratype (male,7.5 mm,ISLA96786):A, habitus,lateral view;B, disposition of dorsal tubercles;C, cephalon and pereonite 1, frontal view; D, cephalon and pereonite 1, dorsal view; E, pleonites 3-5, uropods and telson, dorsal view; F, antennula; G, antenna; H, right mandible; I, left mandible; J, maxillula; K, maxilla; L, maxilliped. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10803285" httpUri="https://zenodo.org/record/10803285/files/figure.png" pageId="16" pageNumber="109">Fig. 8</figureCitation>
H-L), except maxillula (
<figureCitation id="13542A2CC616FFD617E2F40E68B0F9D6" box="[1165,1243,1618,1644]" captionStart="FIG" captionStartId="15.[132,143,1325,1343]" captionTargetBox="[137,1449,217,1272]" captionTargetId="figure-385@15.[132,1455,214,1281]" captionTargetPageId="15" captionText="FIG. 8. — Ctenorillo intertidalis Cardoso &amp; Ferreira,n. sp., paratype (male,7.5 mm,ISLA96786):A, habitus,lateral view;B, disposition of dorsal tubercles;C, cephalon and pereonite 1, frontal view; D, cephalon and pereonite 1, dorsal view; E, pleonites 3-5, uropods and telson, dorsal view; F, antennula; G, antenna; H, right mandible; I, left mandible; J, maxillula; K, maxilla; L, maxilliped. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10803285" httpUri="https://zenodo.org/record/10803285/files/figure.png" pageId="16" pageNumber="109">Fig. 8J</figureCitation>
) with 4+ 6 simple teeth. All pleopod exopods with monospiracular covered lungs. Uropod (
<figureCitation id="13542A2CC616FFD6108EF4CE685CF916" box="[993,1079,1682,1708]" captionStart="FIG" captionStartId="16.[132,143,1409,1427]" captionTargetBox="[146,1444,223,1365]" captionTargetId="figure-284@16.[132,1455,215,1366]" captionTargetPageId="16" captionText="FIG. 9. — Ctenorillo intertidalis Cardoso &amp; Ferreira, n. sp.paratype (male,7.5 mm, ISLA96786): A, uropod,B, pereopod 1; C, pereopod 7; D, pleopod 1 and genital papilla; E, pleopod 2; F, pleopod 3 exopod; G, pleopod 4 exopod; H, pleopod 5 exopod." figureDoi="http://doi.org/10.5281/zenodo.10803287" httpUri="https://zenodo.org/record/10803287/files/figure.png" pageId="16" pageNumber="109">Fig. 9A</figureCitation>
) protopod flattened, enlarged on basal portion; exopod outward, short, inserted dorsally close to medial margin of protopod.
</paragraph>
<caption id="DF106621C616FFD613EBF7DD6F0CFA13" ID-DOI="http://doi.org/10.5281/zenodo.10803287" ID-Zenodo-Dep="10803287" httpUri="https://zenodo.org/record/10803287/files/figure.png" pageId="16" pageNumber="109" startId="16.[132,143,1409,1427]" targetBox="[146,1444,223,1365]" targetPageId="16" targetType="figure">
<paragraph id="8BD036A9C616FFD613EBF7DD6F0CFA13" blockId="16.[132,1455,1408,1449]" pageId="16" pageNumber="109">
<smallCapsWord id="8D36A075C616FFD613EBF7DD6CF6FA2B" baselines="1422,1422" box="[132,157,1409,1427]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="16" pageNumber="109">FIG</smallCapsWord>
. 9. —
<taxonomicName id="4C6F4D2AC616FFD613BDF7DC6E76FA29" authority="Cardoso &amp; Ferreira" authorityName="Cardoso &amp; Ferreira" authorityYear="2024" box="[210,541,1408,1427]" class="Malacostraca" family="Armadillidae" genus="Ctenorillo" kingdom="Animalia" order="Isopoda" pageId="16" pageNumber="109" phylum="Arthropoda" rank="species" species="intertidalis" status="sp. nov.">
<emphasis id="B91BEABBC616FFD613BDF7DC6D10FA29" box="[210,379,1408,1427]" italics="true" pageId="16" pageNumber="109">Ctenorillo intertidalis</emphasis>
Cardoso &amp; Ferreira
</taxonomicName>
,
<taxonomicNameLabel id="A22857C0C616FFD6114BF7DD6E38FA29" box="[548,595,1409,1427]" pageId="16" pageNumber="109" rank="species">n. sp.</taxonomicNameLabel>
paratype (male,7.5 mm, ISLA96786):
<emphasis id="B91BEABBC616FFD610E4F7DD6FF3FA29" bold="true" box="[907,920,1409,1427]" pageId="16" pageNumber="109">A</emphasis>
, uropod,
<emphasis id="B91BEABBC616FFD61089F7DD6F98FA29" bold="true" box="[998,1011,1409,1427]" pageId="16" pageNumber="109">B</emphasis>
, pereopod 1;
<emphasis id="B91BEABBC616FFD6170AF7DC6818FA28" bold="true" box="[1125,1139,1408,1426]" pageId="16" pageNumber="109">C</emphasis>
, pereopod 7;
<emphasis id="B91BEABBC616FFD6178AF7DD6898FA29" bold="true" box="[1253,1267,1409,1427]" pageId="16" pageNumber="109">D</emphasis>
, pleopod 1 and genital papilla;
<emphasis id="B91BEABBC616FFD613AAF7CB6CBAFA13" bold="true" box="[197,209,1431,1449]" pageId="16" pageNumber="109">E</emphasis>
, pleopod 2;
<emphasis id="B91BEABBC616FFD61254F7CB6D2DFA13" bold="true" box="[315,326,1431,1449]" pageId="16" pageNumber="109">F</emphasis>
, pleopod 3 exopod;
<emphasis id="B91BEABBC616FFD6129CF7CB6E6AFA13" bold="true" box="[499,513,1431,1449]" pageId="16" pageNumber="109">G</emphasis>
, pleopod 4 exopod;
<emphasis id="B91BEABBC616FFD611C0F7CB6ED6FA13" bold="true" box="[687,701,1431,1449]" pageId="16" pageNumber="109">H</emphasis>
, pleopod 5 exopod.
</paragraph>
</caption>
<paragraph id="8BD036A9C616FFD61042F54E6F09F896" blockId="16.[811,1457,1810,2028]" box="[813,866,1810,1836]" pageId="16" pageNumber="109">
<emphasis id="B91BEABBC616FFD61042F54E6F09F896" box="[813,866,1810,1836]" italics="true" pageId="16" pageNumber="109">Male</emphasis>
</paragraph>
<paragraph id="8BD036A9C616FFD61042F56D6F99F851" blockId="16.[811,1457,1810,2028]" pageId="16" pageNumber="109">
Pereopods without particular modifications (
<figureCitation id="13542A2CC616FFD61649F56D69CAF8F6" box="[1318,1441,1841,1868]" captionStart="FIG" captionStartId="16.[132,143,1409,1427]" captionTargetBox="[146,1444,223,1365]" captionTargetId="figure-284@16.[132,1455,215,1366]" captionTargetPageId="16" captionText="FIG. 9. — Ctenorillo intertidalis Cardoso &amp; Ferreira, n. sp.paratype (male,7.5 mm, ISLA96786): A, uropod,B, pereopod 1; C, pereopod 7; D, pleopod 1 and genital papilla; E, pleopod 2; F, pleopod 3 exopod; G, pleopod 4 exopod; H, pleopod 5 exopod." figureDoi="http://doi.org/10.5281/zenodo.10803287" httpUri="https://zenodo.org/record/10803287/files/figure.png" pageId="16" pageNumber="109">Fig. 9B, C</figureCitation>
). Pleopod 1 exopod (
<figureCitation id="13542A2CC616FFD6176EF50D683DF8D6" box="[1025,1110,1873,1900]" captionStart="FIG" captionStartId="16.[132,143,1409,1427]" captionTargetBox="[146,1444,223,1365]" captionTargetId="figure-284@16.[132,1455,215,1366]" captionTargetPageId="16" captionText="FIG. 9. — Ctenorillo intertidalis Cardoso &amp; Ferreira, n. sp.paratype (male,7.5 mm, ISLA96786): A, uropod,B, pereopod 1; C, pereopod 7; D, pleopod 1 and genital papilla; E, pleopod 2; F, pleopod 3 exopod; G, pleopod 4 exopod; H, pleopod 5 exopod." figureDoi="http://doi.org/10.5281/zenodo.10803287" httpUri="https://zenodo.org/record/10803287/files/figure.png" pageId="16" pageNumber="109">Fig. 9D</figureCitation>
) wider than long, distal portion round, sinuous outer margin; endopod three times longer than exopod, distal portion slightly bent outwards. Pleopod 2 endopod longer than exopod (
<figureCitation id="13542A2CC616FFD61709F5ED68DDF871" box="[1126,1206,1969,1995]" captionStart="FIG" captionStartId="16.[132,143,1409,1427]" captionTargetBox="[146,1444,223,1365]" captionTargetId="figure-284@16.[132,1455,215,1366]" captionTargetPageId="16" captionText="FIG. 9. — Ctenorillo intertidalis Cardoso &amp; Ferreira, n. sp.paratype (male,7.5 mm, ISLA96786): A, uropod,B, pereopod 1; C, pereopod 7; D, pleopod 1 and genital papilla; E, pleopod 2; F, pleopod 3 exopod; G, pleopod 4 exopod; H, pleopod 5 exopod." figureDoi="http://doi.org/10.5281/zenodo.10803287" httpUri="https://zenodo.org/record/10803287/files/figure.png" pageId="16" pageNumber="109">Fig. 9E</figureCitation>
). Pleopods 3-5 exopods as in
<figureCitation id="13542A2CC616FFD6100CF58D6FD7F851" box="[867,956,2001,2027]" captionStart="FIG" captionStartId="16.[132,143,1409,1427]" captionTargetBox="[146,1444,223,1365]" captionTargetId="figure-284@16.[132,1455,215,1366]" captionTargetPageId="16" captionText="FIG. 9. — Ctenorillo intertidalis Cardoso &amp; Ferreira, n. sp.paratype (male,7.5 mm, ISLA96786): A, uropod,B, pereopod 1; C, pereopod 7; D, pleopod 1 and genital papilla; E, pleopod 2; F, pleopod 3 exopod; G, pleopod 4 exopod; H, pleopod 5 exopod." figureDoi="http://doi.org/10.5281/zenodo.10803287" httpUri="https://zenodo.org/record/10803287/files/figure.png" pageId="16" pageNumber="109">Figure 9</figureCitation>
F-H.
</paragraph>
<caption id="DF106621C617FFD713ECF6D46DFDFB7D" ID-DOI="http://doi.org/10.5281/zenodo.10803289" ID-Zenodo-Dep="10803289" httpUri="https://zenodo.org/record/10803289/files/figure.png" pageId="17" pageNumber="110" startId="17.[131,142,1160,1178]" targetBox="[132,1455,214,1111]" targetPageId="17" targetType="figure">
<paragraph id="8BD036A9C617FFD713ECF6D46DFDFB7D" blockId="17.[131,1453,1160,1223]" pageId="17" pageNumber="110">
<smallCapsWord id="8D36A075C617FFD713ECF6D46CF0FB23" baselines="1173,1174" box="[131,155,1160,1178]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="17" pageNumber="110">FIG</smallCapsWord>
. 10. —
<taxonomicName id="4C6F4D2AC617FFD713B1F6D46E5BFB20" authority="Cardoso &amp; Ferreira" authorityName="Cardoso &amp; Ferreira" authorityYear="2024" box="[222,560,1160,1178]" class="Malacostraca" family="Armadillidae" genus="Ctenorillo" kingdom="Animalia" order="Isopoda" pageId="17" pageNumber="110" phylum="Arthropoda" rank="species" species="intertidalis" status="sp. nov.">
<emphasis id="B91BEABBC617FFD713B1F6D46DE2FB20" box="[222,393,1160,1178]" italics="true" pageId="17" pageNumber="110">Ctenorillo intertidalis</emphasis>
Cardoso &amp; Ferreira
</taxonomicName>
,
<taxonomicNameLabel id="A22857C0C617FFD71157F6D46E00FB20" box="[568,619,1160,1178]" pageId="17" pageNumber="110" rank="species">n. sp.</taxonomicNameLabel>
:
<emphasis id="B91BEABBC617FFD7111DF6D46E14FB20" bold="true" box="[626,639,1160,1178]" pageId="17" pageNumber="110">A</emphasis>
, quartzite outcrops on the shoreline of Jericoacoara, with a cave entrance (
<emphasis id="B91BEABBC617FFD71794F6D46937FB20" bold="true" box="[1275,1372,1160,1178]" pageId="17" pageNumber="110">yellow star</emphasis>
);
<emphasis id="B91BEABBC617FFD71603F6D46912FB20" bold="true" box="[1388,1401,1160,1178]" pageId="17" pageNumber="110">B</emphasis>
, inner portion of a cave where specimens are found;
<emphasis id="B91BEABBC617FFD71164F6C36E72FB0B" bold="true" box="[523,537,1183,1201]" pageId="17" pageNumber="110">C</emphasis>
,
<taxonomicName id="4C6F4D2AC617FFD7114CF6C36F1EFB0B" authority="Cardoso &amp; Ferreira" authorityName="Cardoso &amp; Ferreira" authorityYear="2024" box="[547,885,1183,1201]" class="Malacostraca" family="Armadillidae" genus="Ctenorillo" kingdom="Animalia" order="Isopoda" pageId="17" pageNumber="110" phylum="Arthropoda" rank="species" species="intertidalis" status="sp. nov.">
<emphasis id="B91BEABBC617FFD7114CF6C36EA5FB0B" box="[547,718,1183,1201]" italics="true" pageId="17" pageNumber="110">Ctenorillo intertidalis</emphasis>
Cardoso &amp; Ferreira
</taxonomicName>
,
<taxonomicNameLabel id="A22857C0C617FFD71012F6C36FC5FB0B" box="[893,942,1183,1201]" pageId="17" pageNumber="110" rank="species">n. sp.</taxonomicNameLabel>
specimen;
<emphasis id="B91BEABBC617FFD71761F6C36877FB0B" bold="true" box="[1038,1052,1183,1201]" pageId="17" pageNumber="110">D</emphasis>
, habitus in natural condition sharing the habitat (under rocks) with other isopods.
</paragraph>
</caption>
<paragraph id="8BD036A9C617FFD713EBF7486C8EFA96" blockId="17.[131,776,1300,2028]" box="[132,229,1300,1326]" pageId="17" pageNumber="110">
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</paragraph>
<paragraph id="8BD036A9C617FFD713EBF76F6D02F9F7" blockId="17.[131,776,1300,2028]" pageId="17" pageNumber="110">
Specimens of
<taxonomicName id="4C6F4D2AC617FFD71246F7686ED3FAF7" authority="Cardoso &amp; Ferreira" authorityName="Cardoso &amp; Ferreira" authorityYear="2024" box="[297,696,1331,1358]" class="Malacostraca" family="Armadillidae" genus="Ctenorillo" kingdom="Animalia" order="Isopoda" pageId="17" pageNumber="110" phylum="Arthropoda" rank="species" species="intertidalis" status="n. sp.">
<emphasis id="B91BEABBC617FFD71246F7686DACFAF7" box="[297,455,1331,1358]" italics="true" pageId="17" pageNumber="110">C. intertidalis</emphasis>
Cardoso &amp; Ferreira
</taxonomicName>
,
<taxonomicNameLabel id="A22857C0C617FFD711A9F7686F6CFAF4" box="[710,775,1332,1358]" pageId="17" pageNumber="110" rank="species">n. sp.</taxonomicNameLabel>
were found in six caves in quartzite outcrops on the shoreline (
<figureCitation id="13542A2CC617FFD713D1F72F6D2EFA37" box="[190,325,1395,1421]" captionStart="FIG" captionStartId="17.[131,142,1160,1178]" captionTargetBox="[132,1455,214,1111]" captionTargetId="figure-438@17.[132,1455,214,1111]" captionTargetPageId="17" captionText="FIG. 10. — Ctenorillo intertidalis Cardoso &amp; Ferreira, n. sp.: A, quartzite outcrops on the shoreline of Jericoacoara, with a cave entrance (yellow star); B, inner portion of a cave where specimens are found; C, Ctenorillo intertidalis Cardoso &amp; Ferreira, n. sp. specimen; D, habitus in natural condition sharing the habitat (under rocks) with other isopods." figureDoi="http://doi.org/10.5281/zenodo.10803289" httpUri="https://zenodo.org/record/10803289/files/figure.png" pageId="17" pageNumber="110">Fig. 10A, B</figureCitation>
), near the village of Jericoacoara, in the municipality of Jijoca de Jericoacoara. The climate is classified as hot and humid (Aw’sg. Köppen), with summer rains from January to July. Temperatures range from 25°C to 35°C, and the dry season lasts about six months and is often interrupted by heavy rains in October or September (
<bibRefCitation id="EFFE4B58C617FFD713E1F46E6D31F9F7" author="ALVARES C. A. &amp; STAPE J. L. &amp; SENTELHAS P. C. &amp; MORAES G. &amp; DE &amp; LEONARDO J. &amp; SPAROVEK G." box="[142,346,1586,1613]" pageId="17" pageNumber="110" pagination="711 - 728" refId="ref31163" refString="ALVARES C. A., STAPE J. L., SENTELHAS P. C., MORAES G. DE, LEONARDO J. &amp; SPAROVEK G. 2013. - Koppen's climate classification map for Brazil. Meteorologische Zeitschrift 22 (6): 711 - 728. https: // doi. org / 10.1127 / 0941 - 2948 / 2013 / 0507" type="journal article" year="2013">
Alvares
<emphasis id="B91BEABBC617FFD7138BF46F6D72F9F6" box="[228,281,1586,1612]" italics="true" pageId="17" pageNumber="110">et al.</emphasis>
2013
</bibRefCitation>
).
</paragraph>
<paragraph id="8BD036A9C617FFD713F4F40E6865F996" blockId="17.[131,776,1300,2028]" lastBlockId="17.[811,1458,1299,1804]" pageId="17" pageNumber="110">
The new species was found in six out of eight sampled caves in the area. The caves are quite small presenting, on average, 14.48 (± 7.48) meters of horizontal projection. Thus, most of them were devoid of aphotic zones. Specimens of
<taxonomicName id="4C6F4D2AC617FFD7138DF48E6E30F956" authority="Cardoso &amp; Ferreira" authorityName="Cardoso &amp; Ferreira" authorityYear="2024" box="[226,603,1746,1772]" class="Malacostraca" family="Armadillidae" genus="Ctenorillo" kingdom="Animalia" order="Isopoda" pageId="17" pageNumber="110" phylum="Arthropoda" rank="species" species="intertidalis" status="n. sp.">
<emphasis id="B91BEABBC617FFD7138DF48E6D12F956" box="[226,377,1746,1772]" italics="true" pageId="17" pageNumber="110">C. intertidalis</emphasis>
Cardoso &amp; Ferreira
</taxonomicName>
,
<taxonomicNameLabel id="A22857C0C617FFD7110AF48E6ECAF956" box="[613,673,1746,1772]" pageId="17" pageNumber="110" rank="species">n. sp.</taxonomicNameLabel>
were observed under rocks, sheltered from the wind off the coast (
<figureCitation id="13542A2CC617FFD713E3F54E6C9EF896" box="[140,245,1810,1836]" captionStart="FIG" captionStartId="17.[131,142,1160,1178]" captionTargetBox="[132,1455,214,1111]" captionTargetId="figure-438@17.[132,1455,214,1111]" captionTargetPageId="17" captionText="FIG. 10. — Ctenorillo intertidalis Cardoso &amp; Ferreira, n. sp.: A, quartzite outcrops on the shoreline of Jericoacoara, with a cave entrance (yellow star); B, inner portion of a cave where specimens are found; C, Ctenorillo intertidalis Cardoso &amp; Ferreira, n. sp. specimen; D, habitus in natural condition sharing the habitat (under rocks) with other isopods." figureDoi="http://doi.org/10.5281/zenodo.10803289" httpUri="https://zenodo.org/record/10803289/files/figure.png" pageId="17" pageNumber="110">Fig. 10C</figureCitation>
). The organic matter inside the caves consisted of plant debris (brought by the wind or the tides, in caves closer to the sea) and bat guano in some caves (produced by the piscivorous
<taxonomicName id="4C6F4D2AC617FFD7123CF52D6E8FF831" authority="(Linnaeus, 1758))" authorityName="" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[339,740,1905,1932]" class="Mammalia" family="Noctilionidae" genus="Noctilio" kingdom="Animalia" order="Chiroptera" pageId="17" pageNumber="110" phylum="Chordata" rank="species" species="leporinus">
<emphasis id="B91BEABBC617FFD7123CF52D6E7BF831" box="[339,528,1905,1931]" italics="true" pageId="17" pageNumber="110">Noctilio leporinus</emphasis>
(
<bibRefCitation id="EFFE4B58C617FFD7114FF52D6EA5F836" author="LINNAEUS C." box="[544,718,1905,1932]" pageId="17" pageNumber="110" pagination="1 - 824" refId="ref33262" refString="LINNAEUS C. 1758. - Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Holmiae, Stockholm, ed. 10, vol. 1: 1 - 824. https: // doi. org / 10.5962 / bhl. title. 542" type="journal article" year="1758">Linnaeus, 1758</bibRefCitation>
))
</taxonomicName>
. In some caves, individuals of
<taxonomicName id="4C6F4D2AC617FFD712DCF5CE6CD3F871" authority="Cardoso &amp; Ferreira" authorityName="Cardoso &amp; Ferreira" authorityYear="2024" class="Malacostraca" family="Armadillidae" genus="Ctenorillo" kingdom="Animalia" order="Isopoda" pageId="17" pageNumber="110" phylum="Arthropoda" rank="species" species="intertidalis" status="n. sp.">
<emphasis id="B91BEABBC617FFD712DCF5CE6E20F811" box="[435,587,1937,1964]" italics="true" pageId="17" pageNumber="110">C. intertidalis</emphasis>
Cardoso &amp; Ferreira
</taxonomicName>
,
<taxonomicNameLabel id="A22857C0C617FFD713AEF5EE6C90F876" box="[193,251,1970,1996]" pageId="17" pageNumber="110" rank="species">n. sp.</taxonomicNameLabel>
were observed co-occurring with another isopod species (
<figureCitation id="13542A2CC617FFD713B2F58D6D2CF851" box="[221,327,2001,2027]" captionStart="FIG" captionStartId="17.[131,142,1160,1178]" captionTargetBox="[132,1455,214,1111]" captionTargetId="figure-438@17.[132,1455,214,1111]" captionTargetPageId="17" captionText="FIG. 10. — Ctenorillo intertidalis Cardoso &amp; Ferreira, n. sp.: A, quartzite outcrops on the shoreline of Jericoacoara, with a cave entrance (yellow star); B, inner portion of a cave where specimens are found; C, Ctenorillo intertidalis Cardoso &amp; Ferreira, n. sp. specimen; D, habitus in natural condition sharing the habitat (under rocks) with other isopods." figureDoi="http://doi.org/10.5281/zenodo.10803289" httpUri="https://zenodo.org/record/10803289/files/figure.png" pageId="17" pageNumber="110">Fig. 10D</figureCitation>
). The external environment comprises a “restinga” ecosystem dominated by herbaceous vegetation, which grows along the dunes. Thus, there are few external shelters for these organisms. As the samplings that resulted in the discovery of this new species focused specifically on caves, the actual extent of
<taxonomicName id="4C6F4D2AC617FFD717E6F7CF6FE2FA77" authority="Cardoso &amp; Ferreira" authorityName="Cardoso &amp; Ferreira" authorityYear="2024" class="Malacostraca" family="Armadillidae" genus="Ctenorillo" kingdom="Animalia" order="Isopoda" pageId="17" pageNumber="110" phylum="Arthropoda" rank="species" species="intertidalis" status="n. sp.">
<emphasis id="B91BEABBC617FFD717E6F7CF694EFA17" box="[1161,1317,1427,1453]" italics="true" pageId="17" pageNumber="110">C. intertidalis</emphasis>
Cardoso &amp; Ferreira
</taxonomicName>
,
<taxonomicNameLabel id="A22857C0C617FFD710F8F7EF6FBCFA77" box="[919,983,1459,1485]" pageId="17" pageNumber="110" rank="species">n. sp.</taxonomicNameLabel>
habitat is currently unknown, but their distribution may be much wider than that observed in this work, especially considering the lack of troglomorphic traits in this species.
</paragraph>
<paragraph id="8BD036A9C617FFD7102CF46E680FF8B6" blockId="17.[811,1458,1299,1804]" pageId="17" pageNumber="110">The region is inserted in the Jericoacoara National Park, so the area and the caves are currently protected. Nonetheless, it is noteworthy that there is no regulation of visitor access to these caves, and due to the region’s popularity as a tourist destination, uncontrolled foot traffic may inadvertently pose a threat to the invertebrate populations, including those of the newly discovered species.</paragraph>
</subSubSection>
<subSubSection id="C3756522C617FFD71042F56E69C3F856" pageId="17" pageNumber="110" type="discussion">
<paragraph id="8BD036A9C617FFD71042F56E6FECF8F0" blockId="17.[811,1456,1842,2028]" box="[813,903,1842,1868]" pageId="17" pageNumber="110">
<smallCapsWord id="8D36A075C617FFD71042F56E6FECF8F0" baselines="1861,1861" box="[813,903,1842,1868]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Remark" pageId="17" pageNumber="110">REMARK</smallCapsWord>
</paragraph>
<paragraph id="8BD036A9C617FFD71042F50D69C3F856" blockId="17.[811,1456,1842,2028]" pageId="17" pageNumber="110">
<taxonomicName id="4C6F4D2AC617FFD71042F50D6899F8D1" authority="Cardoso &amp; Ferreira" authorityName="Cardoso &amp; Ferreira" authorityYear="2024" box="[813,1266,1873,1900]" class="Malacostraca" family="Armadillidae" genus="Ctenorillo" kingdom="Animalia" order="Isopoda" pageId="17" pageNumber="110" phylum="Arthropoda" rank="species" species="intertidalis" status="sp. nov.">
<emphasis id="B91BEABBC617FFD71042F50D6866F8D1" box="[813,1037,1873,1899]" italics="true" pageId="17" pageNumber="110">Ctenorillo intertidalis</emphasis>
Cardoso &amp; Ferreira
</taxonomicName>
,
<taxonomicNameLabel id="A22857C0C617FFD71790F50E6956F8D6" box="[1279,1341,1874,1900]" pageId="17" pageNumber="110" rank="species">n. sp.</taxonomicNameLabel>
resembles
<taxonomicName id="4C6F4D2AC617FFD71042F52E6803F831" authority="Cardoso &amp; Ferreira" authorityName="Cardoso &amp; Ferreira" authorityYear="2024" box="[813,1128,1905,1932]" class="Malacostraca" family="Armadillidae" genus="Ctenorillo" kingdom="Animalia" order="Isopoda" pageId="17" pageNumber="110" phylum="Arthropoda" rank="species" species="pelado" status="n. sp.">
<emphasis id="B91BEABBC617FFD71042F52E6FE4F831" box="[813,911,1905,1932]" italics="true" pageId="17" pageNumber="110">C. pelado</emphasis>
Cardoso &amp; Ferreira
</taxonomicName>
,
<taxonomicNameLabel id="A22857C0C617FFD7171DF52E68C7F836" box="[1138,1196,1906,1932]" pageId="17" pageNumber="110" rank="species">n. sp.</taxonomicNameLabel>
in the number of tubercles on pereonite 1 (24), however the disposition with the presence of a lateral rib is similar to
<taxonomicName id="4C6F4D2AC617FFD717CCF5EE6965F871" authorityName="Campos-Filho, Araujo &amp; Taiti" authorityYear="2014" box="[1187,1294,1969,1996]" class="Malacostraca" family="Armadillidae" genus="Ctenorillo" kingdom="Animalia" order="Isopoda" pageId="17" pageNumber="110" phylum="Arthropoda" rank="species" species="ferrarai">
<emphasis id="B91BEABBC617FFD717CCF5EE6965F871" box="[1187,1294,1969,1996]" italics="true" pageId="17" pageNumber="110">C. ferrarai</emphasis>
</taxonomicName>
, differing from both by the number of tubercles on pereonites 2 to 7 (15).
</paragraph>
</subSubSection>
</treatment>
</document>
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