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<mods:title id="5625A53527031E9C91FB87788BE5804B">Review of the Palaearctic genera of Saprininae (Coleoptera: Histeridae)</mods:title>
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<taxonomicName id="4C2C5BCBFF0709B9601CFC84CD98FC84" ID-CoL="7VHSY" authority="(Duftschmid, 1805)" baseAuthorityName="Duftschmid" baseAuthorityYear="1805" box="[331,946,866,891]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="144" pageNumber="141" phylum="Arthropoda" rank="species" species="rugiceps" subGenus="Hypocaccus">
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<emphasis id="B958FC5AFF0709B9601CFC84CFFAFC84" bold="true" box="[331,464,867,890]" italics="true" pageId="144" pageNumber="141">Hypocaccus</emphasis>
(
<emphasis id="B958FC5AFF0709B960B7FC84CC4FFC84" bold="true" box="[480,613,867,890]" italics="true" pageId="144" pageNumber="141">Hypocaccus</emphasis>
) 
<emphasis id="B958FC5AFF0709B96322FC84CCF8FC85" bold="true" box="[629,722,867,891]" italics="true" pageId="144" pageNumber="141">rugiceps</emphasis>
(Duftschmid, 1805)
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<subSubSection id="C33673C3FF0709B960A2FC66CD23FC6A" box="[501,777,897,916]" pageId="144" pageNumber="141" type="description">
<paragraph id="8B932048FF0709B960A2FC66CD23FC6A" blockId="144.[331,946,866,916]" box="[501,777,897,916]" pageId="144" pageNumber="141">
(
<figureCitation id="13173CCDFF0709B960ABFC66CC69FC6A" box="[508,579,897,916]" captionStart="Figs" captionStartId="25.[113,152,1384,1403]" captionTargetBox="[140,1062,219,1365]" captionTargetId="figure-12@25.[140,1062,219,1366]" captionTargetPageId="25" captionText="Figs. 9–28. Saprininae, sensory structures of the antennal club (vesicles marked with grey; sensory areas shaded): 9 – Saprinus (Saprinus) semistriatus (Scriba, 1790); 10 – Saprinus (Hemisaprinus) subvirescens (Ménétries, 1832); 11 – Myrmetes paykulli Kanaar, 1979; 12 – Styphrus corpulentus Motschulsky, 1845; 13 – Saprinus (Phaonius) pharao (Marseul, 1855); 14 – Ammostyphrus cerberus Reichardt, 1924; 15 – Chalcionellus amoenus (Erichson, 1834); 16 – Chivaenius kryzhanovskii Olexa,1980; 17 – Ctenophilothis chobauti (Théry,1900); 18 – Eopachylopus ripae (Lewis, 1885); 19 – Exaesiopus grossipes grossipes (Marseul, 1855); 20 – Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805); 21 – Hypocaccus (Baeckmanniolus) dimidiatus dimidiatus (Illiger, 1807); 22 – Hypocacculus (Hypocacculus) metallescens (Erichson, 1834); 23 – Hypocacculus (Colpellus) praecox (Erichson, 1834); 24 – Hypocacculus (Nessus) rubripes (Erichson, 1834); 25 – Paravolvulus lateristrius (Solskij, 1876); 26 – Pholioxenus phoenix (Reichardt, 1930); 27 – Reichardtiolus duriculus (Reitter, 1904); 28 – Xenonychus tridens (Jacquelin-Duval, 1852)." figureDoi="http://doi.org/10.5281/zenodo.4272139" httpUri="https://zenodo.org/record/4272139/files/figure.png" pageId="144" pageNumber="141">Figs. 20</figureCitation>
, 
<figureCitation id="13173CCDFF0709B9631BFC66CC49FC6A" box="[588,611,897,916]" captionStart="Figs" captionStartId="31.[122,161,1196,1215]" captionTargetId="figure-11@32.[151,1129,219,1531]" captionTargetPageId="32" captionText="Figs. 47–73. Saprininae, labral structures, dorsal view (left half showing external and right half showing internal structures): 47 – Alienocacculus neftensis (Olexa, 1984); 48 – Ammostyphrus cerberus Reichardt, 1924; 49 – Chalcionellus amoenus (Erichson, 1834); 50 – Chivaenius kryzhanovskii Olexa, 1980; 51 – Eopachylopus ripae (Lewis, 1885); 52 – Eremosaprinus vlasovi (Reichardt, 1941); 53 – Euspilotus (Neosaprinus) perrisi (Marseul, 1872); 54 – Exaesiopus grossipes grossipes (Marseul, 55); 55 – Gnathoncus rotundatus (Kugelann, 1792); 56 – Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805); 57 – Hypocacculus (Hypocacculus) metallescens (Erichson, 1834); 58 – Hypocacculus (Colpellus) praecox (Erichson, 1834); 59 – Hypocacculus (Nessus) rubripes (Erichson, 1834); 60 – Microsaprinus therondianus (Dahlgren, 1973); 61 – Myrmetes paykulli Kanaar, 1979; 62 – Paravolvulus lateristriatus (Solskij, 1876); 63 – Philothis (Philothis) arcanus Reichardt, 1930; 64 – Philothis (Atavinus) atavus (Reichardt, 1931); 65 – Philothis (Farabius) hexeris Reichardt, 1930; 66 – Pholioxenus phoenix (Reichardt, 1930); 67 – Reichardtiolus duriculus (Reitter, 1904); 68 – Saprinus (Saprinus) semistriatus (Scriba, 1790); 69 – Saprinus (Hemisaprinus) subvirescens (Ménétries, 1832); 70 – Saprinus (Phaonius) pharao (Marseul, 1855); 71 – Styphrus corpulentus Motschulsky, 1845; 72 – Xenonychus tridens (Jacquelin-Duval, 1852); 73 – Xenophilothis choumovitchi (Thérond, 1965)." figureDoi="http://doi.org/10.5281/zenodo.4288743" httpUri="https://zenodo.org/record/4288743/files/figure.png" pageId="144" pageNumber="141">56</figureCitation>
, 
<figureCitation id="13173CCDFF0709B9633BFC66CCA9FC6A" box="[620,643,897,916]" captionStart="Figs" captionStartId="33.[113,152,1542,1561]" captionTargetBox="[129,1076,1125,1526]" captionTargetId="figure-197@33.[129,1081,1125,1529]" captionTargetPageId="33" captionText="Figs.79–80. Saprininae, mandibular structures, ventral view: 79 – Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805); 80 – Microsaprinus therondianus (Dahlgren, 1973)." figureDoi="http://doi.org/10.5281/zenodo.4272149" httpUri="https://zenodo.org/record/4272149/files/figure.png" pageId="144" pageNumber="141">79</figureCitation>
, 
<figureCitation id="13173CCDFF0709B963DBFC66CC84FC6A" box="[652,686,897,916]" captionStart="Figs" captionStartId="38.[151,190,1224,1243]" captionTargetBox="[229,1046,222,1195]" captionTargetId="figure-11@38.[229,1046,219,1196]" captionTargetPageId="38" captionText="Figs. 109–141. Saprininae, anterior margin of mentum, ventral view: 109 – Alienocacculus neftensis (Olexa, 1984); 110 – Ammostyphrus cerberus Reichardt, 1924; 111 – Axelinus ghilarovi Kryzhanovskij, 1976; 112 – Chalcionellus amoenus (Erichson, 1834); 113 – Chivaenius kryzhanovskii Olexa, 1980; 114 – Ctenophilotis chobauti (Théry, 1900); 115 – Eopachylopus ripae (Lewis, 1885); 116 – Eremosaprinus vlasovi (Reichardt, 1941); 117 – Euspilotus (Neosaprinus) perrisi (Marseul, 1872); 118 – Exaesiopus grossipes grossipes (Marseul, 1855); 119 – Gnathoncus rotundatus (Kugelann, 1792); 120 – Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805); 121 – Hypocaccus (Baeckmanniolus) dimidiatus dimidiatus (Illiger, 1807); 122 – Hypocacculus (Hypocacculus) metallescens (Erichson, 1834); 123 – Hypocacculus (Colpellus) praecox (Erichson, 1834); 124 – Hypocacculus (Nessus) rubripes (Erichson, 1834); 125 – Microsaprinus therondianus (Dahlgren, 1973); 126 – Myrmetes paykulli Kanaar, 1979; 127 – Paravolvulus ovillum (Solskij, 1876); 128 – Philothis (Philothis) arcanus Reichardt, 1930; 129 – Philothis (Atavinus) atavus (Reichardt, 1931); 130 – Philothis (Farabius) hexeris Reichardt, 1930; 131 – Pholioxenus phoenix (Reichardt, 1930); 132 – Reichardtiolus duriculus (Reitter, 1904); 133 – Saprinillus paromaloides Kryzhanovskij, 1974; 134 – Saprinus (Saprinus) semistriatus (Scriba, 1790); 135 – Saprinus (Hemisaprinus) subvirescens (Ménétries, 1832); 136 – Saprinus (Phaonius) pharao (Marseul, 1855); 137 – Styphrus corpulentus Motschulsky, 1845; 138 – Turanostyphrus ignoratus Tishechkin, 2005; 139 – Xenonychus tridens (Jacquelin-Duval, 1852); 140 – Xenophilotis choumovitchi (Thérond, 1965); 141 – Zorius funereus (Schmidt, 1890)." figureDoi="http://doi.org/10.5281/zenodo.4272160" httpUri="https://zenodo.org/record/4272160/files/figure.png" pageId="144" pageNumber="141">120</figureCitation>
, 
<figureCitation id="13173CCDFF0709B963E0FC66CD29FC6A" box="[695,771,897,916]" captionStart-0="Figs" captionStart-1="Figs" captionStartId-0="146.[151,190,1489,1508]" captionStartId-1="147.[113,152,1070,1089]" captionTargetBox-0="[247,1029,217,1469]" captionTargetBox-1="[181,996,220,1036]" captionTargetId-0="figure-11@146.[246,1029,217,1470]" captionTargetId-1="figure-143@147.[167,1035,208,1066]" captionTargetPageId-0="146" captionTargetPageId-1="147" captionText-0="Figs. 433–443. Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805), SEM micrographs: 433 – habitus, dorsal view; 434 – ditto, ventral view; 435 – propygidium and pygidium; 436 – head, dorsal view; 437 – clypeus + labrum, dorsal view; 438 – head, ventral view; 439 – prosternum; 440 – antennal club, ventral view; 441 – lateral disc of metaventrite, metepisternum and fused metepimeron; 442 – protibia, dorsal view; 443 – ditto, ventral view." captionText-1="Figs. 444–452. Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805), male terminalia: 444 – 8th sternite and tergite, dorsal view; 445 – ditto, ventral view; 446 – ditto, lateral view; 447 – 9th tergite, 10th tergite, dorsal view; 448 – ditto, lateral view; 449 – aedeagus, dorsal view; 450 – ditto, lateral view; 451– spiculum gastrale, ventral view; 452 – ditto, lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.4272244" figureDoi-1="http://doi.org/10.5281/zenodo.4272246" httpUri-0="https://zenodo.org/record/4272244/files/figure.png" httpUri-1="https://zenodo.org/record/4272246/files/figure.png" pageId="144" pageNumber="141">433–452</figureCitation>
)
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<treatmentCitationGroup id="AB3C0766FF0709B961C0FC43CDA4FC48" box="[151,910,931,951]" pageId="144" pageNumber="141">
<treatmentCitation id="0A8D0659FF0709B961C0FC43CC25FC48" author="HOFFMANN J. J." box="[151,527,931,951]" page="85" pageId="144" pageNumber="141" year="1803">
<taxonomicName id="4C2C5BCBFF0709B961C0FC43CFDAFC48" ID-CoL="9MR63" authority=": HOFFMANN (1803)" authorityName=": HOFFMANN" authorityYear="1803" box="[151,496,931,951]" class="Insecta" family="Histeridae" genus="Hister" kingdom="Animalia" order="Coleoptera" pageId="144" pageNumber="141" phylum="Arthropoda" rank="species" species="quadristriatus">
<emphasis id="B958FC5AFF0709B961C0FC43CF61FC48" box="[151,331,932,950]" italics="true" pageId="144" pageNumber="141">Hister quadristriatus</emphasis>
: 
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: 85
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, nec 
<treatmentCitation id="0A8D0659FF0709B9636CFC43CCC9FC48" author="PAYKULL G." box="[571,739,931,951]" page="45" pageId="144" pageNumber="141" year="1798">
<bibRefCitation id="EFBD5DB9FF0709B9636CFC43CCECFC48" author="PAYKULL G." box="[571,710,931,951]" pageId="144" pageNumber="141" refId="ref119748" refString="PAYKULL G. 1798: Fauna Suecica. Insecta. Tomus I. Joh. F. Edman, Upsaliae, 10 + 358 + 2 pp." type="book" year="1798">PAYKULL (1798)</bibRefCitation>
: 45
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(misidentification).
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<taxonomicName id="4C2C5BCBFF0709B961C0FC58CFC2FC2F" authority="Duftschmid, 1805: 125" authorityName="Duftschmid" authorityPageNumber="125" authorityYear="1805" box="[151,488,958,977]" class="Insecta" family="Histeridae" genus="Hister" kingdom="Animalia" order="Coleoptera" pageId="144" pageNumber="141" phylum="Arthropoda" rank="species" species="rugiceps">
<emphasis id="B958FC5AFF0709B961C0FC58CF31FC2F" box="[151,283,959,977]" italics="true" pageId="144" pageNumber="141">Hister rugiceps</emphasis>
Duftschmid, 1805: 125
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.
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<paragraph id="8B932048FF0709B961C0FC3DCC01FC12" blockId="144.[151,1128,931,1217]" box="[151,555,985,1004]" pageId="144" pageNumber="141">
<treatmentCitationGroup id="AB3C0766FF0709B961C0FC3DCC01FC12" box="[151,555,985,1004]" pageId="144" pageNumber="141">
<treatmentCitation id="0A8D0659FF0709B961C0FC3DCC02FC12" author="ERICHSON W. F." box="[151,552,985,1004]" page="194" pageId="144" pageNumber="141" year="1834">
<taxonomicName id="4C2C5BCBFF0709B961C0FC3DCFD6FC12" authority=": ERICHSON (1834)" authorityName="ERICHSON" authorityYear="1834" box="[151,508,985,1004]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="144" pageNumber="141" phylum="Arthropoda" rank="species" species="quadristriatus">
<emphasis id="B958FC5AFF0709B961C0FC3DCF4BFC12" box="[151,353,986,1004]" italics="true" pageId="144" pageNumber="141">Saprinus quadristriatus</emphasis>
: 
<bibRefCitation id="EFBD5DB9FF0709B9603AFC3ECFD6FC12" author="ERICHSON W. F." box="[365,508,985,1004]" pageId="144" pageNumber="141" pagination="83 - 208" refId="ref114365" refString="ERICHSON W. F. 1834: Uebersich der Histeroides der Sammlung. Pp. 83 - 208. In: KLUG J. C. F.: Jahrbucher der Insectenkunde mit besonderer Rucksicht auf die Sammlung in Konigl. Museum in Berlin. Erster Band. Enslin, Berlin, viii + 296 pp." type="book chapter" year="1834">ERICHSON (1834)</bibRefCitation>
</taxonomicName>
: 194
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B932048FF0709B961C0FC13CDAEFBDF" blockId="144.[151,1128,931,1217]" pageId="144" pageNumber="141">
<treatmentCitationGroup id="AB3C0766FF0709B961C0FC13CDAEFBDF" pageId="144" pageNumber="141">
<treatmentCitation id="0A8D0659FF0709B961C0FC13CC03FBF8" author="REDTENBACHER L." box="[151,553,1011,1031]" page="238" pageId="144" pageNumber="141" year="1849">
<taxonomicName id="4C2C5BCBFF0709B961C0FC13CFD7FBF8" authority=": REDTENBACHER (1849)" authorityName="REDTENBACHER" authorityYear="1849" box="[151,509,1011,1031]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="144" pageNumber="141" phylum="Arthropoda" rank="species" species="rugiceps">
<emphasis id="B958FC5AFF0709B961C0FC13CF18FBF8" box="[151,306,1012,1030]" italics="true" pageId="144" pageNumber="141">Saprinus rugiceps</emphasis>
: 
<bibRefCitation id="EFBD5DB9FF0709B96068FC13CFD7FBF8" author="REDTENBACHER L." box="[319,509,1011,1031]" pageId="144" pageNumber="141" refId="ref120071" refString="REDTENBACHER L. 1849: Fauna Austriaca. Die Kafer. Nach dem analytischen Methode bearbeitet. C. Gerold, Wien, xxvii + 883 pp." type="book" year="1849">REDTENBACHER (1849)</bibRefCitation>
</taxonomicName>
: 238
</treatmentCitation>
; 
<treatmentCitation id="0A8D0659FF0709B96365FC13CCC7FBF8" author="MARSEUL S. A." box="[562,749,1011,1031]" page="703" pageId="144" pageNumber="141" year="1855">
<bibRefCitation id="EFBD5DB9FF0709B96365FC13CCE8FBF8" author="MARSEUL S. A." box="[562,706,1011,1031]" pageId="144" pageNumber="141" pagination="677 - 758" refId="ref117906" refString="MARSEUL S. A. 1855: Essai monographique sur la familie des Histerides. (Suite). Annales de Societe Entomologique de France, Serie 3 3: 83 - 165, 327 - 506, 677 - 758." type="journal article" year="1855">MARSEUL (1855)</bibRefCitation>
: 703
</treatmentCitation>
, t. XIX, 
<figureCitation id="13173CCDFF0709B9626EFC14CDA9FBF8" box="[825,899,1011,1030]" captionStart="Figs" captionStartId="38.[151,190,1224,1243]" captionTargetBox="[229,1046,222,1195]" captionTargetId="figure-11@38.[229,1046,219,1196]" captionTargetPageId="38" captionText="Figs. 109–141. Saprininae, anterior margin of mentum, ventral view: 109 – Alienocacculus neftensis (Olexa, 1984); 110 – Ammostyphrus cerberus Reichardt, 1924; 111 – Axelinus ghilarovi Kryzhanovskij, 1976; 112 – Chalcionellus amoenus (Erichson, 1834); 113 – Chivaenius kryzhanovskii Olexa, 1980; 114 – Ctenophilotis chobauti (Théry, 1900); 115 – Eopachylopus ripae (Lewis, 1885); 116 – Eremosaprinus vlasovi (Reichardt, 1941); 117 – Euspilotus (Neosaprinus) perrisi (Marseul, 1872); 118 – Exaesiopus grossipes grossipes (Marseul, 1855); 119 – Gnathoncus rotundatus (Kugelann, 1792); 120 – Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805); 121 – Hypocaccus (Baeckmanniolus) dimidiatus dimidiatus (Illiger, 1807); 122 – Hypocacculus (Hypocacculus) metallescens (Erichson, 1834); 123 – Hypocacculus (Colpellus) praecox (Erichson, 1834); 124 – Hypocacculus (Nessus) rubripes (Erichson, 1834); 125 – Microsaprinus therondianus (Dahlgren, 1973); 126 – Myrmetes paykulli Kanaar, 1979; 127 – Paravolvulus ovillum (Solskij, 1876); 128 – Philothis (Philothis) arcanus Reichardt, 1930; 129 – Philothis (Atavinus) atavus (Reichardt, 1931); 130 – Philothis (Farabius) hexeris Reichardt, 1930; 131 – Pholioxenus phoenix (Reichardt, 1930); 132 – Reichardtiolus duriculus (Reitter, 1904); 133 – Saprinillus paromaloides Kryzhanovskij, 1974; 134 – Saprinus (Saprinus) semistriatus (Scriba, 1790); 135 – Saprinus (Hemisaprinus) subvirescens (Ménétries, 1832); 136 – Saprinus (Phaonius) pharao (Marseul, 1855); 137 – Styphrus corpulentus Motschulsky, 1845; 138 – Turanostyphrus ignoratus Tishechkin, 2005; 139 – Xenonychus tridens (Jacquelin-Duval, 1852); 140 – Xenophilotis choumovitchi (Thérond, 1965); 141 – Zorius funereus (Schmidt, 1890)." figureDoi="http://doi.org/10.5281/zenodo.4272160" httpUri="https://zenodo.org/record/4272160/files/figure.png" pageId="144" pageNumber="141">Fig. 141</figureCitation>
; C. 
<treatmentCitation id="0A8D0659FF0709B962F0FC13CA49FBF8" author="THOMSON C. P." box="[935,1123,1011,1031]" page="238" pageId="144" pageNumber="141" year="1862">
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: 238
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<treatmentCitation id="0A8D0659FF0709B961E2FBE9CFA6FBDF" author="GANGLBAUER L." box="[181,396,1038,1057]" page="391" pageId="144" pageNumber="141" year="1899">
<bibRefCitation id="EFBD5DB9FF0709B961E2FBE9CF49FBDF" author="GANGLBAUER L." box="[181,355,1038,1057]" pageId="144" pageNumber="141" refId="ref114798" refString="GANGLBAUER L. 1899: Die Kafer von Mitteleuropa. Die Kafer der osterreichisch-ungarischen Monarchie, Deutschland, der Schweiz, sowie des franzosischen und italienischen Alpengebiets. III, 1. Familienreihe Staphylinoidea. 2 Theil: Scydmaenidae, Silphidae, Clambidae, Leptinidae, Platypsyllidae, Corylophidae, Sphaeritidae, Trichopterigydae, Hydroscaphidae, Scaphidiidae, Histeridae. Carl Gerolds Sohn, Wien, 408 pp." type="book" year="1899">GANGLBAUER (1899)</bibRefCitation>
: 391
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; 
<treatmentCitation id="0A8D0659FF0709B960C1FBE9CC69FBDF" author="REITTER E." box="[406,579,1038,1057]" page="294" pageId="144" pageNumber="141" year="1909">
<bibRefCitation id="EFBD5DB9FF0709B960C1FBE9CC33FBDF" author="REITTER E." box="[406,537,1038,1057]" pageId="144" pageNumber="141" refId="ref120554" refString="REITTER E. 1909: Fauna Germanica. Die Kafer des Deutschen Reiches. Nach den analytischen Methode bearbeitet. II Band. Schriften des Deutschen Lehrervereins fur Naturkunde. Vol. 24. K. G. Lutz, Stuttgart, 392 pp." type="book" year="1909">REITTER (1909)</bibRefCitation>
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</treatmentCitation>
, t. 67, 
<figureCitation id="13173CCDFF0709B9632BFBE9CC90FBDF" box="[636,698,1038,1057]" captionStart="Figs" captionStartId="25.[113,152,1384,1403]" captionTargetBox="[140,1062,219,1365]" captionTargetId="figure-12@25.[140,1062,219,1366]" captionTargetPageId="25" captionText="Figs. 9–28. Saprininae, sensory structures of the antennal club (vesicles marked with grey; sensory areas shaded): 9 – Saprinus (Saprinus) semistriatus (Scriba, 1790); 10 – Saprinus (Hemisaprinus) subvirescens (Ménétries, 1832); 11 – Myrmetes paykulli Kanaar, 1979; 12 – Styphrus corpulentus Motschulsky, 1845; 13 – Saprinus (Phaonius) pharao (Marseul, 1855); 14 – Ammostyphrus cerberus Reichardt, 1924; 15 – Chalcionellus amoenus (Erichson, 1834); 16 – Chivaenius kryzhanovskii Olexa,1980; 17 – Ctenophilothis chobauti (Théry,1900); 18 – Eopachylopus ripae (Lewis, 1885); 19 – Exaesiopus grossipes grossipes (Marseul, 1855); 20 – Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805); 21 – Hypocaccus (Baeckmanniolus) dimidiatus dimidiatus (Illiger, 1807); 22 – Hypocacculus (Hypocacculus) metallescens (Erichson, 1834); 23 – Hypocacculus (Colpellus) praecox (Erichson, 1834); 24 – Hypocacculus (Nessus) rubripes (Erichson, 1834); 25 – Paravolvulus lateristrius (Solskij, 1876); 26 – Pholioxenus phoenix (Reichardt, 1930); 27 – Reichardtiolus duriculus (Reitter, 1904); 28 – Xenonychus tridens (Jacquelin-Duval, 1852)." figureDoi="http://doi.org/10.5281/zenodo.4272139" httpUri="https://zenodo.org/record/4272139/files/figure.png" pageId="144" pageNumber="141">Fig. 12</figureCitation>
; 
<treatmentCitation id="0A8D0659FF0709B96393FBE9CDABFBDF" author="JAKOBSON G. G." box="[708,897,1038,1057]" page="651" pageId="144" pageNumber="141" year="1911">
<bibRefCitation id="EFBD5DB9FF0709B96393FBE9CD7CFBDF" author="JAKOBSON G. G." box="[708,854,1038,1057]" pageId="144" pageNumber="141" pagination="641 - 720" refId="ref116036" refString="JAKOBSON G. G. 1911: Zhuki Rosii i zapadnoy Evropy. Rukovodstvo k opredeleniyu zhukov. 9. [Beetles of Russia and western Europe. Manual for the beetles' determination. 9]. A. F. Devrien, St. Petersburg, pp. 1 + 641 - 720 (in Russian)." type="journal article" year="1911">JAKOBSON (1911)</bibRefCitation>
: 651
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.
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</paragraph>
<paragraph id="8B932048FF0709B961C0FBCDCC4AFBC2" blockId="144.[151,1128,931,1217]" box="[151,608,1065,1084]" pageId="144" pageNumber="141">
<treatmentCitationGroup id="AB3C0766FF0709B961C0FBCDCC4AFBC2" box="[151,608,1065,1084]" pageId="144" pageNumber="141">
<treatmentCitation id="0A8D0659FF0709B961C0FBCDCC76FBC2" author="THOMSON C. P." box="[151,604,1065,1084]" page="401" pageId="144" pageNumber="141" year="1867">
<taxonomicName id="4C2C5BCBFF0709B961C0FBCDCC1BFBC2" authority=": C. THOMSON (1867)" authorityName="C. THOMSON" authorityYear="1867" box="[151,561,1065,1084]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="144" pageNumber="141" phylum="Arthropoda" rank="species" species="quadristriatus">
<emphasis id="B958FC5AFF0709B961C0FBCDCF54FBC2" box="[151,382,1066,1084]" italics="true" pageId="144" pageNumber="141">Hypocaccus quadristriatus</emphasis>
: C. 
<bibRefCitation id="EFBD5DB9FF0709B960F5FBCECC1BFBC2" author="THOMSON C. P." box="[418,561,1065,1084]" pageId="144" pageNumber="141" refId="ref122003" refString="THOMSON C. P. 1867: Skandinaviens Coleoptera, synoptiskt bearbetade, IX. Berlingska Bocktryckeriet, Lund, 407 pp." type="book" year="1867">THOMSON (1867)</bibRefCitation>
</taxonomicName>
: 401
</treatmentCitation>
.
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</paragraph>
<paragraph id="8B932048FF0709B961C0FBA3CC83FBA8" blockId="144.[151,1128,931,1217]" box="[151,681,1091,1111]" pageId="144" pageNumber="141">
<treatmentCitationGroup id="AB3C0766FF0709B961C0FBA3CC83FBA8" box="[151,681,1091,1111]" pageId="144" pageNumber="141">
<treatmentCitation id="0A8D0659FF0709B961C0FBA3CC8FFBA8" author="SCHMIDT J." box="[151,677,1091,1111]" page="316" pageId="144" pageNumber="141" year="1885">
<taxonomicName id="4C2C5BCBFF0709B961C0FBA3CC53FBA8" authority=": SCHMIDT (1885 a)" authorityName="SCHMIDT" authorityYear="1885" box="[151,633,1091,1111]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="144" pageNumber="141" phylum="Arthropoda" rank="species" species="quadristriatus" subGenus="Hypocaccus">
<emphasis id="B958FC5AFF0709B961C0FBA3CEC9FBA8" box="[151,227,1092,1110]" italics="true" pageId="144" pageNumber="141">Saprinus</emphasis>
(
<emphasis id="B958FC5AFF0709B961A7FBA3CF72FBA8" box="[240,344,1092,1110]" italics="true" pageId="144" pageNumber="141">Hypocaccus</emphasis>
) 
<emphasis id="B958FC5AFF0709B96033FBA3CFF7FBA8" box="[356,477,1092,1110]" italics="true" pageId="144" pageNumber="141">quadristriatus</emphasis>
: 
<bibRefCitation id="EFBD5DB9FF0709B960BFFBA4CC53FBA8" author="SCHMIDT J." box="[488,633,1091,1110]" pageId="144" pageNumber="141" pagination="279 - 330" refId="ref120947" refString="SCHMIDT J. 1885 a: Bestimmungs-Tabellen der europaischen Coleopteren. XIV. Histeridae. Berliner Entomologische Zeitung 29: 279 - 330." type="journal article" year="1885">SCHMIDT (1885a)</bibRefCitation>
</taxonomicName>
: 316
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<paragraph id="8B932048FF0709B961C0FBB8CA1EFB3F" blockId="144.[151,1128,931,1217]" pageId="144" pageNumber="141">
<treatmentCitationGroup id="AB3C0766FF0709B961C0FBB8CA1EFB3F" pageId="144" pageNumber="141">
<treatmentCitation id="0A8D0659FF0709B961C0FBB8CC0BFB8F" author="BICKHARDT H." box="[151,545,1118,1137]" page="100" pageId="144" pageNumber="141" year="1916">
<taxonomicName id="4C2C5BCBFF0709B961C0FBB8CFDCFB8F" ID-CoL="7VHSY" authority=": BICKHARDT (1916)" authorityName="BICKHARDT" authorityYear="1916" box="[151,502,1118,1137]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="144" pageNumber="141" phylum="Arthropoda" rank="species" species="rugiceps">
<emphasis id="B958FC5AFF0709B961C0FBB8CF64FB8F" box="[151,334,1119,1137]" italics="true" pageId="144" pageNumber="141">Hypocaccus rugiceps</emphasis>
: 
<bibRefCitation id="EFBD5DB9FF0709B9600DFBB9CFDCFB8F" author="BICKHARDT H." box="[346,502,1118,1137]" pageId="144" pageNumber="141" refId="ref112724" refString="BICKHARDT H. 1916 - 1917: Histeridae. In: WYTSMAN P. (ed.): Genera Insectorum, Fasc. 166 a, b. La Haye, 302 pp." type="book" year="1916">BICKHARDT (1916)</bibRefCitation>
</taxonomicName>
: 100
</treatmentCitation>
; 
<treatmentCitation id="0A8D0659FF0709B9637CFBB9CCDAFB8F" author="REICHARDT A." box="[555,752,1118,1137]" page="308" pageId="144" pageNumber="141" year="1941">
<bibRefCitation id="EFBD5DB9FF0709B9637CFBB9CCECFB8F" author="REICHARDT A." box="[555,710,1118,1137]" pageId="144" pageNumber="141" refId="ref120382" refString="REICHARDT A. 1941: Semeystva Sphaeritidae i Histeridae (Vol. 1). [Families Sphaeritidae and Histeridae]. Fauna SSSR, Nasekomye Zhestokrylye, V, 3. Nauka, Moskva-Leningrad, xiii + 419 pp." type="book" year="1941">REICHARDT (1941)</bibRefCitation>
: 308
</treatmentCitation>
, 316, 
<figureCitation id="13173CCDFF0709B96273FBB9CD47FB8F" box="[804,877,1118,1137]" captionStart="Figs" captionStartId="56.[152,191,1557,1576]" captionTargetBox="[227,1045,933,1531]" captionTargetId="figure-254@56.[220,1054,928,1534]" captionTargetPageId="56" captionText="Figs. 162–167.Euspilotus (Neosaprinus) perrisi (Marseul, 1872), variability of female pygidial sulci. Figs. 162–165 after OLEXA (1975)." figureDoi="http://doi.org/10.5281/zenodo.4272181" httpUri="https://zenodo.org/record/4272181/files/figure.png" pageId="144" pageNumber="141">Fig. 165</figureCitation>
; 
<treatmentCitation id="0A8D0659FF0709B96220FBB9CA08FB8F" author="HORION A." box="[887,1058,1118,1137]" page="342" pageId="144" pageNumber="141" year="1949">
<bibRefCitation id="EFBD5DB9FF0709B96220FBB9CDDDFB8F" author="HORION A." box="[887,1015,1118,1137]" pageId="144" pageNumber="141" refId="ref115598" refString="HORION A. 1949: Faunistik der Mitteleuropaischen Kafer. 2. Palpicornia-Staphylinoidea (ausser Staphylinidae). Klostermann, Frankfurt-am-Main, xxiii + 388 pp." type="book" year="1949">HORION (1949)</bibRefCitation>
: 342
</treatmentCitation>
; 
<treatmentCitation id="0A8D0659FF0709B9657CFBB9CF25FB72" author="MAZUR S." page="40" pageId="144" pageNumber="141" year="1973">
<bibRefCitation id="EFBD5DB9FF0709B9657CFBB9CED8FB72" author="MAZUR S." pageId="144" pageNumber="141" refId="ref118330" refString="MAZUR S. 1973: Sphaeritidae i gniliki Histeridae. [Sphaeritidae and Histeridae]. Klucze do oznaczania owadow Polski. [Key to the identification of Polish insects]. Vol. XIX, part 11 - 12. Polish Entomological Society, Warszawa, 74 pp (in Polish)" type="book" year="1973">MAZUR (1973)</bibRefCitation>
: 40
</treatmentCitation>
, 
<figureCitation id="13173CCDFF0709B96040FB9ECF7EFB72" box="[279,340,1145,1164]" captionStart="Figs" captionStartId="33.[113,152,1542,1561]" captionTargetBox="[129,1076,1125,1526]" captionTargetId="figure-197@33.[129,1081,1125,1529]" captionTargetPageId="33" captionText="Figs.79–80. Saprininae, mandibular structures, ventral view: 79 – Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805); 80 – Microsaprinus therondianus (Dahlgren, 1973)." figureDoi="http://doi.org/10.5281/zenodo.4272149" httpUri="https://zenodo.org/record/4272149/files/figure.png" pageId="144" pageNumber="141">Fig. 80</figureCitation>
, 
<treatmentCitation id="0A8D0659FF0709B9600CFB9ECC92FB72" author="KRYZHANOVSKIJ O. L. &amp; REICHARDT A. N." box="[347,696,1145,1164]" page="218" pageId="144" pageNumber="141" year="1976">
<bibRefCitation id="EFBD5DB9FF0709B9600CFB9ECCA4FB72" author="KRYZHANOVSKIJ O. L. &amp; REICHARDT A. N." box="[347,654,1145,1164]" pageId="144" pageNumber="141" refId="ref117064" refString="KRYZHANOVSKIJ O. L. &amp; REICHARDT A. N. 1976: Zhuki Nadsemeystva Histeroidea (semeystva Sphaeritidae, Histeridae, Synteliidae). [Beetles of the superfamily Histeroidea (families Sphaeritidae, Histeridae, Syntelidae)]. Fauna SSSR, Zhestokrylye, Vyp. 4. Nauka, Leningrad, 434 pp (in Russian)." type="book" year="1976">KRYZHANOVSKIJ &amp; REICHARDT (1976)</bibRefCitation>
: 218
</treatmentCitation>
, 220, 
<figureCitation id="13173CCDFF0709B963BFFB9ECD12FB72" box="[744,824,1145,1164]" captionStart="Figs" captionStartId="141.[113,152,1271,1290]" captionTargetBox="[171,1034,218,1252]" captionTargetId="figure-72@141.[145,1057,211,1260]" captionTargetPageId="141" captionText="Figs. 426–432. Hypocacculus (Nessus) rubripes (Erichson, 1834), male terminalia: 426 – 8th sternite and tergite, ventral view; 427 – ditto, dorsal view; 428 – ditto, lateral view; 429 – aedeagus, dorsal view; 430 – ditto, lateral view; 431 – 9th tergite, 10th tergite (dorsal view) and spiculum gastrale (ventral view); 432 – 9th tergite, 10th tergite and spiculum gastrale, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4272242" httpUri="https://zenodo.org/record/4272242/files/figure.png" pageId="144" pageNumber="141">Figs.428</figureCitation>
, 
<figureCitation id="13173CCDFF0709B96217FB9FCDF5FB72" box="[832,991,1144,1164]" captionStart="Figs" captionStartId="146.[151,190,1489,1508]" captionTargetBox="[247,1029,217,1469]" captionTargetId="figure-11@146.[246,1029,217,1470]" captionTargetPageId="146" captionText="Figs. 433–443. Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805), SEM micrographs: 433 – habitus, dorsal view; 434 – ditto, ventral view; 435 – propygidium and pygidium; 436 – head, dorsal view; 437 – clypeus + labrum, dorsal view; 438 – head, ventral view; 439 – prosternum; 440 – antennal club, ventral view; 441 – lateral disc of metaventrite, metepisternum and fused metepimeron; 442 – protibia, dorsal view; 443 – ditto, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4272244" httpUri="https://zenodo.org/record/4272244/files/figure.png" pageId="144" pageNumber="141">435, 436, 437, 442</figureCitation>
; 
<treatmentCitation id="0A8D0659FF0709B962B1FB9ECEFDFB58" author="VIENNA P." page="185" pageId="144" pageNumber="141" year="1980">
<bibRefCitation id="EFBD5DB9FF0709B962B1FB9ECA49FB72" author="VIENNA P." box="[998,1123,1145,1164]" pageId="144" pageNumber="141" refId="ref122194" refString="VIENNA P. 1980: Fauna d'Italia. Vol. XVI. Coleoptera Histeridae. Bologna, Calderini Press, ix + 386 pp." type="book" year="1980">
<collectingRegion id="49E8EEAAFF0709B962B1FB9ECA09FB74" box="[998,1059,1145,1164]" country="Austria" name="Wien" pageId="144" pageNumber="141">VIENNA</collectingRegion>
(1980)
</bibRefCitation>
: 185
</treatmentCitation>
, 190; 
<treatmentCitation id="0A8D0659FF0709B96059FB73CC20FB58" author="MAZUR S. &amp; KASZAB Z." box="[270,522,1171,1191]" page="58" pageId="144" pageNumber="141" year="1980">
<bibRefCitation id="EFBD5DB9FF0709B96059FB73CFC3FB58" author="MAZUR S. &amp; KASZAB Z." box="[270,489,1171,1191]" pageId="144" pageNumber="141" refId="ref118755" refString="MAZUR S. &amp; KASZAB Z. 1980: Sutabogarak - Histeridae. 14. Fuzet. Fauna Hungariae. 138. VII Kotet. Academic Press, Budapest, 123 pp (In Hungarian)" type="book" year="1980">MAZUR &amp; KASZAB (1980)</bibRefCitation>
: 58
</treatmentCitation>
, 
<figureCitation id="13173CCDFF0709B96343FB74CCBFFB58" box="[532,661,1171,1190]" captionStart="Figs" captionStartId="25.[113,152,1384,1403]" captionTargetBox="[140,1062,219,1365]" captionTargetId="figure-12@25.[140,1062,219,1366]" captionTargetPageId="25" captionText="Figs. 9–28. Saprininae, sensory structures of the antennal club (vesicles marked with grey; sensory areas shaded): 9 – Saprinus (Saprinus) semistriatus (Scriba, 1790); 10 – Saprinus (Hemisaprinus) subvirescens (Ménétries, 1832); 11 – Myrmetes paykulli Kanaar, 1979; 12 – Styphrus corpulentus Motschulsky, 1845; 13 – Saprinus (Phaonius) pharao (Marseul, 1855); 14 – Ammostyphrus cerberus Reichardt, 1924; 15 – Chalcionellus amoenus (Erichson, 1834); 16 – Chivaenius kryzhanovskii Olexa,1980; 17 – Ctenophilothis chobauti (Théry,1900); 18 – Eopachylopus ripae (Lewis, 1885); 19 – Exaesiopus grossipes grossipes (Marseul, 1855); 20 – Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805); 21 – Hypocaccus (Baeckmanniolus) dimidiatus dimidiatus (Illiger, 1807); 22 – Hypocacculus (Hypocacculus) metallescens (Erichson, 1834); 23 – Hypocacculus (Colpellus) praecox (Erichson, 1834); 24 – Hypocacculus (Nessus) rubripes (Erichson, 1834); 25 – Paravolvulus lateristrius (Solskij, 1876); 26 – Pholioxenus phoenix (Reichardt, 1930); 27 – Reichardtiolus duriculus (Reitter, 1904); 28 – Xenonychus tridens (Jacquelin-Duval, 1852)." figureDoi="http://doi.org/10.5281/zenodo.4272139" httpUri="https://zenodo.org/record/4272139/files/figure.png" pageId="144" pageNumber="141">Figs. 26J, 28K</figureCitation>
, 
<figureCitation id="13173CCDFF0709B963C8FB74CD00FB5B" box="[671,810,1171,1191]" captionStart="Figs" captionStartId="26.[150,189,1085,1104]" captionTargetBox="[202,1075,219,1057]" captionTargetId="figure-156@26.[202,1076,219,1058]" captionTargetPageId="26" captionText="Figs. 29–38. Saprininae, sensory structures of the antennal club (vesicles marked with grey; sensory areas shaded): 29 – Xenophilothis choumovitchi (Thérond, 1965); 30 – Zorius funereus (Schmidt, 1890); 31 – Alienocacculus neftensis (Olexa, 1984); 32 – Philothis (Philothis) arcanus Reichardt, 1930; 33 – Philothis (Atavinus) atavus (Reichardt, 1931); 34 – Philothis (Farabius) hexeris Reichardt, 1930; 35 – Euspilotus (Neosaprinus) perrisi (Marseul, 1872); 36 – Gnathoncus rotundatus (Kugelann, 1792); 37 – Eremosaprinus vlasovi (Reichardt, 1941); 38 – Microsaprinus therondianus (Dahlgren, 1973)." figureDoi="http://doi.org/10.5281/zenodo.4272141" httpUri="https://zenodo.org/record/4272141/files/figure.png" pageId="144" pageNumber="141">
29, 30F; 
<treatmentCitation id="0A8D0659FF0709B963B9FB73CD00FB5B" author="MAZUR S." box="[750,810,1172,1191]" page="100" pageId="144" pageNumber="141" year="1981">
<bibRefCitation id="EFBD5DB9FF0709B963B9FB73CD00FB5B" author="MAZUR S." box="[750,810,1172,1191]" pageId="144" pageNumber="141" refId="ref118463" refString="MAZUR S. 1981 a: Histeridae - Gnilikowate (Insecta: Coleoptera). Fauna Polski, Tom 9. Polish Entomological Society, Warszawa, 206 pp (In Polish)" type="book" year="1981">MAZUR</bibRefCitation>
</treatmentCitation>
</figureCitation>
(1981a): 100, 103, 
<figureCitation id="13173CCDFF0709B96281FB74CA0AFB58" box="[982,1056,1171,1190]" captionStart="Figs" captionStartId="38.[151,190,1224,1243]" captionTargetBox="[229,1046,222,1195]" captionTargetId="figure-11@38.[229,1046,219,1196]" captionTargetPageId="38" captionText="Figs. 109–141. Saprininae, anterior margin of mentum, ventral view: 109 – Alienocacculus neftensis (Olexa, 1984); 110 – Ammostyphrus cerberus Reichardt, 1924; 111 – Axelinus ghilarovi Kryzhanovskij, 1976; 112 – Chalcionellus amoenus (Erichson, 1834); 113 – Chivaenius kryzhanovskii Olexa, 1980; 114 – Ctenophilotis chobauti (Théry, 1900); 115 – Eopachylopus ripae (Lewis, 1885); 116 – Eremosaprinus vlasovi (Reichardt, 1941); 117 – Euspilotus (Neosaprinus) perrisi (Marseul, 1872); 118 – Exaesiopus grossipes grossipes (Marseul, 1855); 119 – Gnathoncus rotundatus (Kugelann, 1792); 120 – Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805); 121 – Hypocaccus (Baeckmanniolus) dimidiatus dimidiatus (Illiger, 1807); 122 – Hypocacculus (Hypocacculus) metallescens (Erichson, 1834); 123 – Hypocacculus (Colpellus) praecox (Erichson, 1834); 124 – Hypocacculus (Nessus) rubripes (Erichson, 1834); 125 – Microsaprinus therondianus (Dahlgren, 1973); 126 – Myrmetes paykulli Kanaar, 1979; 127 – Paravolvulus ovillum (Solskij, 1876); 128 – Philothis (Philothis) arcanus Reichardt, 1930; 129 – Philothis (Atavinus) atavus (Reichardt, 1931); 130 – Philothis (Farabius) hexeris Reichardt, 1930; 131 – Pholioxenus phoenix (Reichardt, 1930); 132 – Reichardtiolus duriculus (Reitter, 1904); 133 – Saprinillus paromaloides Kryzhanovskij, 1974; 134 – Saprinus (Saprinus) semistriatus (Scriba, 1790); 135 – Saprinus (Hemisaprinus) subvirescens (Ménétries, 1832); 136 – Saprinus (Phaonius) pharao (Marseul, 1855); 137 – Styphrus corpulentus Motschulsky, 1845; 138 – Turanostyphrus ignoratus Tishechkin, 2005; 139 – Xenonychus tridens (Jacquelin-Duval, 1852); 140 – Xenophilotis choumovitchi (Thérond, 1965); 141 – Zorius funereus (Schmidt, 1890)." figureDoi="http://doi.org/10.5281/zenodo.4272160" httpUri="https://zenodo.org/record/4272160/files/figure.png" pageId="144" pageNumber="141">Fig. 135</figureCitation>
; 
<treatmentCitation id="0A8D0659FF0709B9657CFB73CF38FB3F" author="MAZUR S." page="96" pageId="144" pageNumber="141" year="1984">
<bibRefCitation id="EFBD5DB9FF0709B9657CFB73CEDBFB3F" author="MAZUR S." pageId="144" pageNumber="141" pagination="1 - 376" refId="ref118519" refString="MAZUR S. 1984: A world catalogue of Histeridae. Polskie Pismo Entomologiczne 54 (3 - 4): 1 - 376." type="journal article" year="1984">MAZUR (1984)</bibRefCitation>
: 96
</treatmentCitation>
; 
<treatmentCitation id="0A8D0659FF0709B9604DFB49CFE8FB3F" author="MAZUR S." box="[282,450,1198,1217]" page="260" pageId="144" pageNumber="141" year="1997">
<bibRefCitation id="EFBD5DB9FF0709B9604DFB49CFB2FB3F" author="MAZUR S." box="[282,408,1198,1217]" pageId="144" pageNumber="141" pagination="1 - 373" refId="ref118633" refString="MAZUR S. 1997: A world catalogue of the Histeridae (Coleoptera: Histeridae). Genus, Supplement, pp. 1 - 373." type="book chapter" year="1997">MAZUR (1997)</bibRefCitation>
: 260
</treatmentCitation>
; 
<treatmentCitation id="0A8D0659FF0709B9609CFB49CCA9FB3F" author="YELAMOS T." box="[459,643,1198,1217]" page="328" pageId="144" pageNumber="141" year="2002">
<bibRefCitation id="EFBD5DB9FF0709B9609CFB49CC73FB3F" author="YELAMOS T." box="[459,601,1198,1217]" pageId="144" pageNumber="141" refId="ref122718" refString="YELAMOS T. 2002: Coleoptera, Histeridae. In: RAMOS M. A., TERCEDOR J. A, BELLES-ROS X., GOSAL- BEZ-NOGUERA J., SIERRA A. G., MAYOL E. M., PIERA F. M., MARINO J. S. &amp; GONZALEZ J. T. (eds.): Fauna Iberica. Vol. 17. Museo Nacional de Ciencias Naturales, CSCI, Madrid, 411 pp." type="book" year="2002">YÉLAMOS (2002)</bibRefCitation>
: 328
</treatmentCitation>
, 330, 
<figureCitation id="13173CCDFF0709B963E1FB49CDE5FB3E" box="[694,975,1198,1217]" captionStart="Figs" captionStartId="56.[152,191,1557,1576]" captionTargetBox="[227,1045,933,1531]" captionTargetId="figure-254@56.[220,1054,928,1534]" captionTargetPageId="56" captionText="Figs. 162–167.Euspilotus (Neosaprinus) perrisi (Marseul, 1872), variability of female pygidial sulci. Figs. 162–165 after OLEXA (1975)." figureDoi="http://doi.org/10.5281/zenodo.4272181" httpUri="https://zenodo.org/record/4272181/files/figure.png" pageId="144" pageNumber="141">
Figs. 163A, 164A, 166B; 
<treatmentCitation id="0A8D0659FF0709B962C4FB49CDE5FB3E" author="MAZUR S." box="[915,975,1198,1217]" page="94" pageId="144" pageNumber="141" year="2004">
<bibRefCitation id="EFBD5DB9FF0709B962C4FB49CDE5FB3E" author="MAZUR S." box="[915,975,1198,1217]" pageId="144" pageNumber="141" pagination="68 - 102" refId="ref118682" refString="MAZUR S. 2004: Family Histeridae. Pp. 68 - 102. In: LOBL I. &amp; SMETANA A. (eds.): Catalogue of Palaearctic Coleoptera, Vol. 2, Hydrophiloidea-Histeroidea-Staphylinidea. Apollo Books, Stenstrup, 942 pp." type="book chapter" year="2004">MAZUR</bibRefCitation>
</treatmentCitation>
</figureCitation>
(2004): 94.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C33673C3FF0709B961C0FB38CF82FAA3" pageId="144" pageNumber="141" type="materials_examined">
<paragraph id="8B932048FF0709B961C0FB38CC3EFB0C" blockId="144.[151,1128,1246,1374]" box="[151,532,1246,1267]" pageId="144" pageNumber="141">
<emphasis id="B958FC5AFF0709B961C0FB38CF39FB0C" bold="true" box="[151,275,1246,1266]" pageId="144" pageNumber="141">
<typeStatus id="54979EEAFF0709B961C0FB38CEEFFB0C" box="[151,197,1247,1266]" pageId="144" pageNumber="141">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F33B60D8FF0709B9604FFB38CF73FB0C" box="[280,345,1247,1266]" name="Austria" pageId="144" pageNumber="141">Austria</collectingCountry>
, Oerlander near Linz.
</paragraph>
<paragraph id="8B932048FF0709B961C0FB1ECF82FAA3" blockId="144.[151,1128,1246,1374]" pageId="144" pageNumber="141">
<emphasis id="B958FC5AFF0709B961C0FB1ECF61FAF3" bold="true" box="[151,331,1273,1293]" pageId="144" pageNumber="141">Material examined.</emphasis>
<materialsCitation id="3B442A15FF0709B96005FB1ECA0FFAF3" ID-GBIF-Occurrence="2981920370" box="[338,1061,1273,1293]" collectingDate="1948-07" collectorName="L. Krejcarek" country="Czech Republic" location="Bzenec" pageId="144" pageNumber="141" specimenCount="1" stateProvince="Jihomoravsky">
<emphasis id="B958FC5AFF0709B96005FB1ECC50FAF3" bold="true" box="[338,634,1273,1293]" pageId="144" pageNumber="141">
<collectingCountry id="F33B60D8FF0709B96005FB1ECC3DFAF3" box="[338,535,1273,1293]" name="Czech Republic" pageId="144" pageNumber="141">CZECH REPUBLIC</collectingCountry>
: 
<collectingRegion id="49E8EEAAFF0709B96375FB1DCC59FAF2" box="[546,627,1274,1293]" country="Czech Republic" name="Jihomoravsky" pageId="144" pageNumber="141">MORAVIA</collectingRegion>
:
</emphasis>
<location id="8EF37693FF0709B963D6FB1DCCEAFAF3" LSID="urn:lsid:plazi:treatment:0385915EFF0709BD601CFC84CA71FBED:8EF37693FF0709B963D6FB1DCCEAFAF3" box="[641,704,1274,1293]" country="Czech Republic" name="Bzenec" pageId="144" pageNumber="141" stateProvince="Jihomoravsky">Bzenec</location>
env., 
<date id="FF920688FF0709B963AFFB1DCD6AFAF3" box="[760,832,1274,1293]" pageId="144" pageNumber="141" value="1948-07">
<collectingDate id="EFD6FF60FF0709B963AFFB1DCD6AFAF3" box="[760,832,1274,1293]" pageId="144" pageNumber="141" value="1948-07">vii.1948</collectingDate>
</date>
, 
<specimenCount id="9D2AEBC1FF0709B9621CFB1DCDA6FAF3" box="[843,908,1274,1293]" pageId="144" pageNumber="141" type="generic">1 spec.</specimenCount>
, 
<collectorName id="26D9459EFF0709B962C2FB1DCA29FAF3" box="[917,1027,1274,1293]" pageId="144" pageNumber="141">L. Krejcárek</collectorName>
lgt
</materialsCitation>
.; 
<materialsCitation id="3B442A15FF0709B96563FB1DCC25FAD6" ID-GBIF-Occurrence="2981920344" collectingDate="1947-07" country="Czech Republic" location="Velke Pavlovice" pageId="144" pageNumber="141" specimenCount="1">
<location id="8EF37693FF0709B96563FB1DCEC7FAD6" LSID="urn:lsid:plazi:treatment:0385915EFF0709BD601CFC84CA71FBED:8EF37693FF0709B96563FB1DCEC7FAD6" country="Czech Republic" name="Velke Pavlovice" pageId="144" pageNumber="141">Velké Pavlovice</location>
, 
<date id="FF920688FF0709B961A2FAF2CF17FAD6" box="[245,317,1301,1320]" pageId="144" pageNumber="141" value="1947-07">
<collectingDate id="EFD6FF60FF0709B961A2FAF2CF17FAD6" box="[245,317,1301,1320]" pageId="144" pageNumber="141" value="1947-07">vii.1947</collectingDate>
</date>
, 1 ♁, collector unknown
</materialsCitation>
. 
<materialsCitation id="3B442A15FF0709B96341FAF3CA4EFAD6" ID-GBIF-Occurrence="2981920307" box="[534,1124,1300,1320]" collectingDate="1914-05-16" country="Italy" location="Millan" municipality="Les Vignes" pageId="144" pageNumber="141" specimenCount="1">
<emphasis id="B958FC5AFF0709B96341FAF3CC4BFAD6" bold="true" box="[534,609,1300,1320]" pageId="144" pageNumber="141">
<collectingCountry id="F33B60D8FF0709B96341FAF3CC77FAD6" box="[534,605,1300,1320]" name="Italy" pageId="144" pageNumber="141">ITALY</collectingCountry>
:
</emphasis>
<collectingMunicipality id="6BF7BA32FF0709B96332FAF2CCE2FAD6" box="[613,712,1301,1320]" pageId="144" pageNumber="141">Les Vignes</collectingMunicipality>
, a 
<location id="8EF37693FF0709B9638AFAF2CD32FAD6" LSID="urn:lsid:plazi:treatment:0385915EFF0709BD601CFC84CA71FBED:8EF37693FF0709B9638AFAF2CD32FAD6" box="[733,792,1301,1320]" country="Italy" municipality="Les Vignes" name="Millan" pageId="144" pageNumber="141">Millan</location>
, 
<date id="FF920688FF0709B96248FAF2CD5DFAD6" box="[799,887,1301,1320]" pageId="144" pageNumber="141" value="1914-05-16">
<collectingDate id="EFD6FF60FF0709B96248FAF2CD5DFAD6" box="[799,887,1301,1320]" pageId="144" pageNumber="141" value="1914-05-16">16.v.1914</collectingDate>
</date>
, 
<specimenCount id="9D2AEBC1FF0709B96229FAF2CD96FAD6" box="[894,956,1301,1320]" pageId="144" pageNumber="141" type="generic">1 spec.</specimenCount>
, collector unknown
</materialsCitation>
. 
<materialsCitation id="3B442A15FF0709B961C0FAC9CD51FABC" ID-GBIF-Occurrence="2981920321" box="[151,891,1326,1347]" collectingDate="1976-07-12" collectorName="J. Strejcek" country="Germany" location="Warnemunde" municipality="Markgratenheide" pageId="144" pageNumber="141" specimenCount="1">
<emphasis id="B958FC5AFF0709B961C0FAC9CF25FABC" bold="true" box="[151,271,1326,1346]" pageId="144" pageNumber="141">
<collectingCountry id="F33B60D8FF0709B961C0FAC9CF21FABC" box="[151,267,1326,1346]" name="Germany" pageId="144" pageNumber="141">GERMANY</collectingCountry>
:
</emphasis>
<collectingMunicipality id="6BF7BA32FF0709B96043FAC8CF82FABC" box="[276,424,1327,1346]" pageId="144" pageNumber="141">Markgratenheide</collectingMunicipality>
, near 
<location id="8EF37693FF0709B9608EFAC8CC64FABC" LSID="urn:lsid:plazi:treatment:0385915EFF0709BD601CFC84CA71FBED:8EF37693FF0709B9608EFAC8CC64FABC" box="[473,590,1327,1346]" country="Germany" municipality="Markgratenheide" name="Warnemunde" pageId="144" pageNumber="141">Warnemünde</location>
, 
<date id="FF920688FF0709B96302FAC8CC93FABC" box="[597,697,1327,1346]" pageId="144" pageNumber="141" value="1976-07-12">
<collectingDate id="EFD6FF60FF0709B96302FAC8CC93FABC" box="[597,697,1327,1346]" pageId="144" pageNumber="141" value="1976-07-12">12.vii.1976</collectingDate>
</date>
, 
<specimenCount id="9D2AEBC1FF0709B96396FAC8CCD4FABD" box="[705,766,1327,1347]" pageId="144" pageNumber="141" type="generic">1 spec.</specimenCount>
, 
<collectorName id="26D9459EFF0709B96252FAD7CD77FABC" box="[773,861,1327,1347]" pageId="144" pageNumber="141">J. Strejček</collectorName>
lgt
</materialsCitation>
. 
<materialsCitation id="3B442A15FF0709B962D5FAC9CF8EFAA3" ID-GBIF-Occurrence="2981920361" collectingDate="1975-05" collectionCode="TLAN" collectorName="A. Olexa" country="Ukraine" location="Kiev" pageId="144" pageNumber="141" specimenCount="1" specimenCount-female="1" stateProvince="Kyiv">
<emphasis id="B958FC5AFF0709B962D5FAC9CDC5FABC" bold="true" box="[898,1007,1326,1346]" pageId="144" pageNumber="141">
<collectingCountry id="F33B60D8FF0709B962D5FAC9CDC0FABC" box="[898,1002,1326,1346]" name="Ukraine" pageId="144" pageNumber="141">UKRAINE</collectingCountry>
:
</emphasis>
<collectingRegion id="49E8EEAAFF0709B962A4FAC8CA35FABC" box="[1011,1055,1327,1346]" country="Ukraine" name="Kyiv" pageId="144" pageNumber="141">Kiev</collectingRegion>
, 
<date id="FF920688FF0709B96571FAC8CA49FABC" box="[1062,1123,1327,1346]" pageId="144" pageNumber="141" value="1975-05">
<collectingDate id="EFD6FF60FF0709B96571FAC8CA49FABC" box="[1062,1123,1327,1346]" pageId="144" pageNumber="141" value="1975-05">v.1975</collectingDate>
</date>
, 1 ♁ 
<specimenCount id="9D2AEBC1FF0709B961EBFAADCEF6FAA0" box="[188,220,1351,1374]" pageId="144" pageNumber="141" type="female">1 ♀</specimenCount>
, 
<collectorName id="26D9459EFF0709B961B1FAADCF1EFAA3" box="[230,308,1354,1373]" pageId="144" pageNumber="141">A. Olexa</collectorName>
lgt. (
<collectionCode id="ED3DB88DFF0709B96037FAADCFB4FAA3" box="[352,414,1354,1373]" pageId="144" pageNumber="141">TLAN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33673C3FF0709BD61C0FA92CA71FBED" lastPageId="148" lastPageNumber="145" pageId="144" pageNumber="141" type="description">
<paragraph id="8B932048FF0709B961C0FA92CCAAFA53" blockId="144.[151,1127,1397,1613]" pageId="144" pageNumber="141">
<emphasis id="B958FC5AFF0709B961C0FA92CF17FA73" bold="true" box="[151,317,1397,1421]" pageId="144" pageNumber="141">Redescription.</emphasis>
Body length: PEL: 
<quantity id="4CD48DADFF0709B9635CFA92CC82FA73" box="[523,680,1397,1421]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" metricValueMax="3.5" metricValueMin="3.1" pageId="144" pageNumber="141" unit="mm" value="3.3" valueMax="3.5" valueMin="3.1">3.10–3.50 mm</quantity>
; APW: 
<quantity id="4CD48DADFF0709B963AFFA92CD88FA73" box="[760,930,1397,1421]" metricMagnitude="-3" metricUnit="m" metricValue="1.1875" metricValueMax="1.25" metricValueMin="1.125" pageId="144" pageNumber="141" unit="mm" value="1.1875" valueMax="1.25" valueMin="1.125">1.125–1.25 mm</quantity>
; PPW: 
<quantity id="4CD48DADFF0709B962B9FA92CEEEFA53" metricMagnitude="-3" metricUnit="m" metricValue="2.4375" metricValueMax="2.625" metricValueMin="2.25" pageId="144" pageNumber="141" unit="mm" value="2.4375" valueMax="2.625" valueMin="2.25">2.25–2.625 mm</quantity>
; EL: 2.00– 
<quantity id="4CD48DADFF0709B9606DFA72CFB6FA53" box="[314,412,1429,1453]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" pageId="144" pageNumber="141" unit="mm" value="2.25">2.25 mm</quantity>
; EW: 2.60–3.00 mm.
</paragraph>
<paragraph id="8B932048FF0709B961E2FA52CC2DF9F3" blockId="144.[151,1127,1397,1613]" pageId="144" pageNumber="141">
Body (
<figureCitation id="13173CCDFF0709B961ABFA52CFBFFA33" box="[252,405,1461,1485]" captionStart="Figs" captionStartId="146.[151,190,1489,1508]" captionTargetBox="[247,1029,217,1469]" captionTargetId="figure-11@146.[246,1029,217,1470]" captionTargetPageId="146" captionText="Figs. 433–443. Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805), SEM micrographs: 433 – habitus, dorsal view; 434 – ditto, ventral view; 435 – propygidium and pygidium; 436 – head, dorsal view; 437 – clypeus + labrum, dorsal view; 438 – head, ventral view; 439 – prosternum; 440 – antennal club, ventral view; 441 – lateral disc of metaventrite, metepisternum and fused metepimeron; 442 – protibia, dorsal view; 443 – ditto, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4272244" httpUri="https://zenodo.org/record/4272244/files/figure.png" pageId="144" pageNumber="141">Figs. 433–434</figureCitation>
) shortly oval, convex, cuticle dark brown to blackish-blue; pronotum occasionally with feeble bronze luster, elytra with greenish metallic tinge; legs, mouthparts and antennae dark brown to black.
</paragraph>
<paragraph id="8B932048FF0709B861E2F9F2CFE4FECD" blockId="144.[151,1127,1397,1613]" lastBlockId="145.[113,1089,219,1587]" lastPageId="145" lastPageNumber="138" pageId="144" pageNumber="141">
Antennal scape (
<figureCitation id="13173CCDFF0709B96030F9F2CFE8F9D3" box="[359,450,1557,1581]" captionStart="Figs" captionStartId="146.[151,190,1489,1508]" captionTargetBox="[247,1029,217,1469]" captionTargetId="figure-11@146.[246,1029,217,1470]" captionTargetPageId="146" captionText="Figs. 433–443. Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805), SEM micrographs: 433 – habitus, dorsal view; 434 – ditto, ventral view; 435 – propygidium and pygidium; 436 – head, dorsal view; 437 – clypeus + labrum, dorsal view; 438 – head, ventral view; 439 – prosternum; 440 – antennal club, ventral view; 441 – lateral disc of metaventrite, metepisternum and fused metepimeron; 442 – protibia, dorsal view; 443 – ditto, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4272244" httpUri="https://zenodo.org/record/4272244/files/figure.png" pageId="144" pageNumber="141">Fig. 436</figureCitation>
) slightly thickened, with few short setae; club (
<figureCitation id="13173CCDFF0709B962EEF9F2CA3EF9D3" box="[953,1044,1557,1581]" captionStart="Figs" captionStartId="146.[151,190,1489,1508]" captionTargetBox="[247,1029,217,1469]" captionTargetId="figure-11@146.[246,1029,217,1470]" captionTargetPageId="146" captionText="Figs. 433–443. Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805), SEM micrographs: 433 – habitus, dorsal view; 434 – ditto, ventral view; 435 – propygidium and pygidium; 436 – head, dorsal view; 437 – clypeus + labrum, dorsal view; 438 – head, ventral view; 439 – prosternum; 440 – antennal club, ventral view; 441 – lateral disc of metaventrite, metepisternum and fused metepimeron; 442 – protibia, dorsal view; 443 – ditto, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4272244" httpUri="https://zenodo.org/record/4272244/files/figure.png" pageId="144" pageNumber="141">Fig. 440</figureCitation>
) round, without visible articulation, entire surface covered with thick short yellow sensilla intermingled with sparse somewhat longer erect sensilla; sensory structures of antennal club (
<figureCitation id="13173CCDFF0609B86298FF3CCA37FF0A" box="[975,1053,219,244]" captionStart="Figs" captionStartId="25.[113,152,1384,1403]" captionTargetBox="[140,1062,219,1365]" captionTargetId="figure-12@25.[140,1062,219,1366]" captionTargetPageId="25" captionText="Figs. 9–28. Saprininae, sensory structures of the antennal club (vesicles marked with grey; sensory areas shaded): 9 – Saprinus (Saprinus) semistriatus (Scriba, 1790); 10 – Saprinus (Hemisaprinus) subvirescens (Ménétries, 1832); 11 – Myrmetes paykulli Kanaar, 1979; 12 – Styphrus corpulentus Motschulsky, 1845; 13 – Saprinus (Phaonius) pharao (Marseul, 1855); 14 – Ammostyphrus cerberus Reichardt, 1924; 15 – Chalcionellus amoenus (Erichson, 1834); 16 – Chivaenius kryzhanovskii Olexa,1980; 17 – Ctenophilothis chobauti (Théry,1900); 18 – Eopachylopus ripae (Lewis, 1885); 19 – Exaesiopus grossipes grossipes (Marseul, 1855); 20 – Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805); 21 – Hypocaccus (Baeckmanniolus) dimidiatus dimidiatus (Illiger, 1807); 22 – Hypocacculus (Hypocacculus) metallescens (Erichson, 1834); 23 – Hypocacculus (Colpellus) praecox (Erichson, 1834); 24 – Hypocacculus (Nessus) rubripes (Erichson, 1834); 25 – Paravolvulus lateristrius (Solskij, 1876); 26 – Pholioxenus phoenix (Reichardt, 1930); 27 – Reichardtiolus duriculus (Reitter, 1904); 28 – Xenonychus tridens (Jacquelin-Duval, 1852)." figureDoi="http://doi.org/10.5281/zenodo.4272139" httpUri="https://zenodo.org/record/4272139/files/figure.png" pageId="145" pageNumber="138">Fig. 20</figureCitation>
) in form of stipe-shaped vesicle situated under circular sensory area on internal distal margin of the ventral side of antennal club.
</paragraph>
<paragraph id="8B932048FF0609B861C7FEDCCEDDFD8D" blockId="145.[113,1089,219,1587]" pageId="145" pageNumber="138">
Mouthparts. Mandibles (
<figureCitation id="13173CCDFF0609B860CCFEDCCFC1FEAA" box="[411,491,315,340]" captionStart="Figs" captionStartId="33.[113,152,1542,1561]" captionTargetBox="[129,1076,1125,1526]" captionTargetId="figure-197@33.[129,1081,1125,1529]" captionTargetPageId="33" captionText="Figs.79–80. Saprininae, mandibular structures, ventral view: 79 – Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805); 80 – Microsaprinus therondianus (Dahlgren, 1973)." figureDoi="http://doi.org/10.5281/zenodo.4272149" httpUri="https://zenodo.org/record/4272149/files/figure.png" pageId="145" pageNumber="138">Fig. 79</figureCitation>
) with almost straight outer margin, moderately curved inwardly, acutely pointed; sub-apical tooth on inner margin of left mandible large, perpendicular; labrum (
<figureCitation id="13173CCDFF0609B86072FE9CCF54FE6D" box="[293,382,379,403]" captionStart="Figs" captionStartId="31.[122,161,1196,1215]" captionTargetId="figure-11@32.[151,1129,219,1531]" captionTargetPageId="32" captionText="Figs. 47–73. Saprininae, labral structures, dorsal view (left half showing external and right half showing internal structures): 47 – Alienocacculus neftensis (Olexa, 1984); 48 – Ammostyphrus cerberus Reichardt, 1924; 49 – Chalcionellus amoenus (Erichson, 1834); 50 – Chivaenius kryzhanovskii Olexa, 1980; 51 – Eopachylopus ripae (Lewis, 1885); 52 – Eremosaprinus vlasovi (Reichardt, 1941); 53 – Euspilotus (Neosaprinus) perrisi (Marseul, 1872); 54 – Exaesiopus grossipes grossipes (Marseul, 55); 55 – Gnathoncus rotundatus (Kugelann, 1792); 56 – Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805); 57 – Hypocacculus (Hypocacculus) metallescens (Erichson, 1834); 58 – Hypocacculus (Colpellus) praecox (Erichson, 1834); 59 – Hypocacculus (Nessus) rubripes (Erichson, 1834); 60 – Microsaprinus therondianus (Dahlgren, 1973); 61 – Myrmetes paykulli Kanaar, 1979; 62 – Paravolvulus lateristriatus (Solskij, 1876); 63 – Philothis (Philothis) arcanus Reichardt, 1930; 64 – Philothis (Atavinus) atavus (Reichardt, 1931); 65 – Philothis (Farabius) hexeris Reichardt, 1930; 66 – Pholioxenus phoenix (Reichardt, 1930); 67 – Reichardtiolus duriculus (Reitter, 1904); 68 – Saprinus (Saprinus) semistriatus (Scriba, 1790); 69 – Saprinus (Hemisaprinus) subvirescens (Ménétries, 1832); 70 – Saprinus (Phaonius) pharao (Marseul, 1855); 71 – Styphrus corpulentus Motschulsky, 1845; 72 – Xenonychus tridens (Jacquelin-Duval, 1852); 73 – Xenophilothis choumovitchi (Thérond, 1965)." figureDoi="http://doi.org/10.5281/zenodo.4288743" httpUri="https://zenodo.org/record/4288743/files/figure.png" pageId="145" pageNumber="138">Figs. 56</figureCitation>
, 
<figureCitation id="13173CCDFF0609B860DDFE9BCF99FE6A" box="[394,435,380,404]" captionStart="Figs" captionStartId="146.[151,190,1489,1508]" captionTargetBox="[247,1029,217,1469]" captionTargetId="figure-11@146.[246,1029,217,1470]" captionTargetPageId="146" captionText="Figs. 433–443. Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805), SEM micrographs: 433 – habitus, dorsal view; 434 – ditto, ventral view; 435 – propygidium and pygidium; 436 – head, dorsal view; 437 – clypeus + labrum, dorsal view; 438 – head, ventral view; 439 – prosternum; 440 – antennal club, ventral view; 441 – lateral disc of metaventrite, metepisternum and fused metepimeron; 442 – protibia, dorsal view; 443 – ditto, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4272244" httpUri="https://zenodo.org/record/4272244/files/figure.png" pageId="145" pageNumber="138">437</figureCitation>
) microscopically punctate, medially convex, anterior margin with slight median projection; with two labral pits, with two setae arising from each; terminal labial palpomere elongated, its width about one-third its length; mentum sub-trapezoid, anterior margin medially with shallow notch (
<figureCitation id="13173CCDFF0609B86355FE3CCC77FE0A" box="[514,605,475,500]" captionStart="Figs" captionStartId="38.[151,190,1224,1243]" captionTargetBox="[229,1046,222,1195]" captionTargetId="figure-11@38.[229,1046,219,1196]" captionTargetPageId="38" captionText="Figs. 109–141. Saprininae, anterior margin of mentum, ventral view: 109 – Alienocacculus neftensis (Olexa, 1984); 110 – Ammostyphrus cerberus Reichardt, 1924; 111 – Axelinus ghilarovi Kryzhanovskij, 1976; 112 – Chalcionellus amoenus (Erichson, 1834); 113 – Chivaenius kryzhanovskii Olexa, 1980; 114 – Ctenophilotis chobauti (Théry, 1900); 115 – Eopachylopus ripae (Lewis, 1885); 116 – Eremosaprinus vlasovi (Reichardt, 1941); 117 – Euspilotus (Neosaprinus) perrisi (Marseul, 1872); 118 – Exaesiopus grossipes grossipes (Marseul, 1855); 119 – Gnathoncus rotundatus (Kugelann, 1792); 120 – Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805); 121 – Hypocaccus (Baeckmanniolus) dimidiatus dimidiatus (Illiger, 1807); 122 – Hypocacculus (Hypocacculus) metallescens (Erichson, 1834); 123 – Hypocacculus (Colpellus) praecox (Erichson, 1834); 124 – Hypocacculus (Nessus) rubripes (Erichson, 1834); 125 – Microsaprinus therondianus (Dahlgren, 1973); 126 – Myrmetes paykulli Kanaar, 1979; 127 – Paravolvulus ovillum (Solskij, 1876); 128 – Philothis (Philothis) arcanus Reichardt, 1930; 129 – Philothis (Atavinus) atavus (Reichardt, 1931); 130 – Philothis (Farabius) hexeris Reichardt, 1930; 131 – Pholioxenus phoenix (Reichardt, 1930); 132 – Reichardtiolus duriculus (Reitter, 1904); 133 – Saprinillus paromaloides Kryzhanovskij, 1974; 134 – Saprinus (Saprinus) semistriatus (Scriba, 1790); 135 – Saprinus (Hemisaprinus) subvirescens (Ménétries, 1832); 136 – Saprinus (Phaonius) pharao (Marseul, 1855); 137 – Styphrus corpulentus Motschulsky, 1845; 138 – Turanostyphrus ignoratus Tishechkin, 2005; 139 – Xenonychus tridens (Jacquelin-Duval, 1852); 140 – Xenophilotis choumovitchi (Thérond, 1965); 141 – Zorius funereus (Schmidt, 1890)." figureDoi="http://doi.org/10.5281/zenodo.4272160" httpUri="https://zenodo.org/record/4272160/files/figure.png" pageId="145" pageNumber="138">Fig. 120</figureCitation>
) surrounded with few moderately long setae; lateral margins with a single row of sparse much shorter ramose setae; cardo of maxilla with few short setae on lateral margin; stipes triangular, with three much longer setae; terminal maxillary palpomere elongated, its width about one-fourth its length, about twice as long as penultimate.
</paragraph>
<paragraph id="8B932048FF0609B861C7FD9CCDA5FD0D" blockId="145.[113,1089,219,1587]" pageId="145" pageNumber="138">
Clypeus (
<figureCitation id="13173CCDFF0609B861AFFD9CCF7FFD6A" box="[248,341,635,660]" captionStart="Figs" captionStartId="146.[151,190,1489,1508]" captionTargetBox="[247,1029,217,1469]" captionTargetId="figure-11@146.[246,1029,217,1470]" captionTargetPageId="146" captionText="Figs. 433–443. Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805), SEM micrographs: 433 – habitus, dorsal view; 434 – ditto, ventral view; 435 – propygidium and pygidium; 436 – head, dorsal view; 437 – clypeus + labrum, dorsal view; 438 – head, ventral view; 439 – prosternum; 440 – antennal club, ventral view; 441 – lateral disc of metaventrite, metepisternum and fused metepimeron; 442 – protibia, dorsal view; 443 – ditto, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4272244" httpUri="https://zenodo.org/record/4272244/files/figure.png" pageId="145" pageNumber="138">Fig. 437</figureCitation>
) rectangular, rounded laterally, rugulose-lacunose; frontal stria (
<figureCitation id="13173CCDFF0609B86542FD9CCEB1FD4D" captionStart="Figs" captionStartId="146.[151,190,1489,1508]" captionTargetBox="[247,1029,217,1469]" captionTargetId="figure-11@146.[246,1029,217,1470]" captionTargetPageId="146" captionText="Figs. 433–443. Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805), SEM micrographs: 433 – habitus, dorsal view; 434 – ditto, ventral view; 435 – propygidium and pygidium; 436 – head, dorsal view; 437 – clypeus + labrum, dorsal view; 438 – head, ventral view; 439 – prosternum; 440 – antennal club, ventral view; 441 – lateral disc of metaventrite, metepisternum and fused metepimeron; 442 – protibia, dorsal view; 443 – ditto, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4272244" httpUri="https://zenodo.org/record/4272244/files/figure.png" pageId="145" pageNumber="138">Fig. 436</figureCitation>
) well impressed, almost straight (occasionally somewhat curved outwardly), carinate, continued as carinate supraorbital stria; frontal disc (
<figureCitation id="13173CCDFF0609B863C6FD5CCCC0FD2D" box="[657,746,699,723]" captionStart="Figs" captionStartId="146.[151,190,1489,1508]" captionTargetBox="[247,1029,217,1469]" captionTargetId="figure-11@146.[246,1029,217,1470]" captionTargetPageId="146" captionText="Figs. 433–443. Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805), SEM micrographs: 433 – habitus, dorsal view; 434 – ditto, ventral view; 435 – propygidium and pygidium; 436 – head, dorsal view; 437 – clypeus + labrum, dorsal view; 438 – head, ventral view; 439 – prosternum; 440 – antennal club, ventral view; 441 – lateral disc of metaventrite, metepisternum and fused metepimeron; 442 – protibia, dorsal view; 443 – ditto, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4272244" httpUri="https://zenodo.org/record/4272244/files/figure.png" pageId="145" pageNumber="138">Fig. 436</figureCitation>
) rugulose-lacunose, with several deeply impressed longitudinal rugae; eyes convex, observable from above.
</paragraph>
<paragraph id="8B932048FF0609B861C7FD1CCF80FC4D" blockId="145.[113,1089,219,1587]" pageId="145" pageNumber="138">
Pronotal sides (
<figureCitation id="13173CCDFF0609B8606EFD1CCFBCFCEA" box="[313,406,763,788]" captionStart="Figs" captionStartId="146.[151,190,1489,1508]" captionTargetBox="[247,1029,217,1469]" captionTargetId="figure-11@146.[246,1029,217,1470]" captionTargetPageId="146" captionText="Figs. 433–443. Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805), SEM micrographs: 433 – habitus, dorsal view; 434 – ditto, ventral view; 435 – propygidium and pygidium; 436 – head, dorsal view; 437 – clypeus + labrum, dorsal view; 438 – head, ventral view; 439 – prosternum; 440 – antennal club, ventral view; 441 – lateral disc of metaventrite, metepisternum and fused metepimeron; 442 – protibia, dorsal view; 443 – ditto, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4272244" httpUri="https://zenodo.org/record/4272244/files/figure.png" pageId="145" pageNumber="138">Fig. 433</figureCitation>
) almost parallel on apical two-thirds, thence curved inwardly; apical angles obtuse; marginal pronotal stria complete; pronotal disc convex, except for postero-median part entirely covered with dense punctation, punctures laterally form longitudinal rugae; postero-median part forming a well-defined triangular impunctate part; pronotal base with a double row of round dense punctures, with slight antero-scutellar depression; scutellum small, well visible.
</paragraph>
<paragraph id="8B932048FF0609B861C7FC5CCDF3FAED" blockId="145.[113,1089,219,1587]" pageId="145" pageNumber="138">Elytral humeri slightly prominent, epipleura with microscopic punctures; marginal epipleural stria complete; marginal elytral stria deeply impressed, continued as weakly impressed shortened apical elytral stria; along elytral marginal stria a regular row of round punctures present. Humeral elytral stria vaguely impressed on basal fifth; inner subhumeral stria present medially, deep and rather long, rarely joining humeral elytral stria; dorsal elytral striae almost unrecognizable beneath punctation; first dorsal elytral stria usually conspicuous, stopping short of elytral apex, apically slightly curved inwardly, second dorsal stria reaching approximately elytral half apically, third and fourth dorsal elytral striae do not reach elytral half apically, often unrecognizable beneath coarse elytral punctation; fourth dorsal elytral stria basally connected with sutural elytral stria; sutural elytral stria well impressed, in punctures, almost reaching elytral apex; between it and elytral suture a row of small round punctures present.</paragraph>
<paragraph id="8B932048FF0609B861C7FAFCCDCCFA6D" blockId="145.[113,1089,219,1587]" pageId="145" pageNumber="138">Elytral disc (with exception of elytral epipleura, humeri, smooth apical elytral band and a small ‘mirror’ between fourth dorsal elytral and sutural striae) completely covered with deep regular and very dense and coarse punctation; punctures separated by about half their diameter; elytral ‘mirrors’ between each other not divided by a band of punctation.</paragraph>
<paragraph id="8B932048FF0609B861C7FA7CCDE8FA0D" blockId="145.[113,1089,219,1587]" pageId="145" pageNumber="138">
Propygidium (
<figureCitation id="13173CCDFF0609B8607CFA7CCFADFA4D" box="[299,391,1435,1459]" captionStart="Figs" captionStartId="146.[151,190,1489,1508]" captionTargetBox="[247,1029,217,1469]" captionTargetId="figure-11@146.[246,1029,217,1470]" captionTargetPageId="146" captionText="Figs. 433–443. Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805), SEM micrographs: 433 – habitus, dorsal view; 434 – ditto, ventral view; 435 – propygidium and pygidium; 436 – head, dorsal view; 437 – clypeus + labrum, dorsal view; 438 – head, ventral view; 439 – prosternum; 440 – antennal club, ventral view; 441 – lateral disc of metaventrite, metepisternum and fused metepimeron; 442 – protibia, dorsal view; 443 – ditto, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4272244" httpUri="https://zenodo.org/record/4272244/files/figure.png" pageId="145" pageNumber="138">Fig. 435</figureCitation>
) almost completely exposed, about twice as broad as long, with dense and coarse punctation, punctures smaller than those of elytra; punctation of pygidium (
<figureCitation id="13173CCDFF0609B8612EFA3CCEFEFA0D" box="[121,212,1499,1523]" captionStart="Figs" captionStartId="146.[151,190,1489,1508]" captionTargetBox="[247,1029,217,1469]" captionTargetId="figure-11@146.[246,1029,217,1470]" captionTargetPageId="146" captionText="Figs. 433–443. Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805), SEM micrographs: 433 – habitus, dorsal view; 434 – ditto, ventral view; 435 – propygidium and pygidium; 436 – head, dorsal view; 437 – clypeus + labrum, dorsal view; 438 – head, ventral view; 439 – prosternum; 440 – antennal club, ventral view; 441 – lateral disc of metaventrite, metepisternum and fused metepimeron; 442 – protibia, dorsal view; 443 – ditto, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4272244" httpUri="https://zenodo.org/record/4272244/files/figure.png" pageId="145" pageNumber="138">Fig. 435</figureCitation>
) similar to that of propygidium, becoming somewhat sparser apically.
</paragraph>
<paragraph id="8B932048FF0609BA61C7FA1CCCBFFAB0" blockId="145.[113,1089,219,1587]" lastBlockId="147.[113,1089,1238,1614]" lastPageId="147" lastPageNumber="140" pageId="145" pageNumber="138">
Anterior margin of median portion of prosternum (
<figureCitation id="13173CCDFF0609B863EBFA1CCD33F9EA" box="[700,793,1531,1556]" captionStart="Figs" captionStartId="146.[151,190,1489,1508]" captionTargetBox="[247,1029,217,1469]" captionTargetId="figure-11@146.[246,1029,217,1470]" captionTargetPageId="146" captionText="Figs. 433–443. Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805), SEM micrographs: 433 – habitus, dorsal view; 434 – ditto, ventral view; 435 – propygidium and pygidium; 436 – head, dorsal view; 437 – clypeus + labrum, dorsal view; 438 – head, ventral view; 439 – prosternum; 440 – antennal club, ventral view; 441 – lateral disc of metaventrite, metepisternum and fused metepimeron; 442 – protibia, dorsal view; 443 – ditto, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4272244" httpUri="https://zenodo.org/record/4272244/files/figure.png" pageId="145" pageNumber="138">Fig. 439</figureCitation>
) obtuse-angulate; marginal prosternal stria present laterally; pre-apical foveae comparatively small; prosternal process slightly concave, surface between carinal striae smooth, laterally substrigulate with sparse deep punctures; carinal prosternal striae slightly divergent on prosternal apophysis, sub-parallel, usually not united apically; lateral prosternal striae strongly carinate, convergent anteriorly, united in front of apices of carinal prosternal striae.
</paragraph>
<caption id="DF5370C0FF0509BB61C0FA36CA07F9CA" ID-DOI="http://doi.org/10.5281/zenodo.4272244" ID-Zenodo-Dep="4272244" httpUri="https://zenodo.org/record/4272244/files/figure.png" pageId="146" pageNumber="143" startId="146.[151,190,1489,1508]" targetBox="[247,1029,217,1469]" targetPageId="146">
<paragraph id="8B932048FF0509BB61C0FA36CA07F9CA" blockId="146.[151,1127,1489,1589]" pageId="146" pageNumber="143">
Figs. 433–443. 
<taxonomicName id="4C2C5BCBFF0509BB604AFA35CD2BFA1A" authority="(Duftschmid, 1805)" baseAuthorityName="Duftschmid" baseAuthorityYear="1805" box="[285,769,1489,1509]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="146" pageNumber="143" phylum="Arthropoda" rank="species" species="rugiceps" subGenus="Hypocaccus">
<emphasis id="B958FC5AFF0509BB604AFA35CFAFFA1A" box="[285,389,1490,1508]" italics="true" pageId="146" pageNumber="143">Hypocaccus</emphasis>
(
<emphasis id="B958FC5AFF0509BB60C5FA35CFD0FA1A" box="[402,506,1490,1508]" italics="true" pageId="146" pageNumber="143">Hypocaccus</emphasis>
) 
<emphasis id="B958FC5AFF0509BB6350FA35CC7AFA1A" box="[519,592,1490,1508]" italics="true" pageId="146" pageNumber="143">rugiceps</emphasis>
(Duftschmid, 1805)
</taxonomicName>
, SEM micrographs: 433 – habitus, dorsal view; 434 – ditto, ventral view; 435 – propygidium and pygidium; 436 – head, dorsal view; 437 – clypeus + labrum, dorsal view; 438 – head, ventral view; 439 – prosternum; 440 – antennal club, ventral view; 441 – lateral disc of metaventrite, metepisternum and fused metepimeron; 442 – protibia, dorsal view; 443 – ditto, ventral view.
</paragraph>
</caption>
<caption id="DF5370C0FF0409BA6126FBC9CF52FB6F" ID-DOI="http://doi.org/10.5281/zenodo.4272246" ID-Zenodo-Dep="4272246" httpUri="https://zenodo.org/record/4272246/files/figure.png" pageId="147" pageNumber="140" startId="147.[113,152,1070,1089]" targetBox="[181,996,220,1036]" targetPageId="147">
<paragraph id="8B932048FF0409BA6126FBC9CF52FB6F" blockId="147.[113,1087,1068,1169]" pageId="147" pageNumber="140">
Figs. 444–452. 
<taxonomicName id="4C2C5BCBFF0409BA61ADFBC8CCCFFBBF" authority="(Duftschmid, 1805)" baseAuthorityName="Duftschmid" baseAuthorityYear="1805" box="[250,741,1070,1089]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="147" pageNumber="140" phylum="Arthropoda" rank="species" species="rugiceps" subGenus="Hypocaccus">
<emphasis id="B958FC5AFF0409BA61ADFBC8CF48FBBF" box="[250,354,1071,1089]" italics="true" pageId="147" pageNumber="140">Hypocaccus</emphasis>
(
<emphasis id="B958FC5AFF0409BA6026FBC8CFF3FBBF" box="[369,473,1071,1089]" italics="true" pageId="147" pageNumber="140">Hypocaccus</emphasis>
) 
<emphasis id="B958FC5AFF0409BA60BFFBC8CC1BFBBF" box="[488,561,1071,1089]" italics="true" pageId="147" pageNumber="140">rugiceps</emphasis>
(Duftschmid, 1805)
</taxonomicName>
, male terminalia: 444 – 8 
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sternite and tergite, dorsal view; 445 – ditto, ventral view; 446 – ditto, lateral view; 447 – 9 
<superScript id="7C598D00FF0409BA624AFBA0CD0DFBAD" attach="left" box="[797,807,1095,1107]" fontSize="5" pageId="147" pageNumber="140">th</superScript>
tergite, 10 
<superScript id="7C598D00FF0409BA62D3FBA0CDA4FBAD" attach="left" box="[900,910,1095,1107]" fontSize="5" pageId="147" pageNumber="140">th</superScript>
tergite, dorsal view; 448 – ditto, lateral view; 449 – aedeagus, dorsal view; 450 – ditto, lateral view; 451– spiculum gastrale, ventral view; 452 – ditto, lateral view.
</paragraph>
</caption>
<paragraph id="8B932048FF0409BA61C7FAB1CFD8F9B0" blockId="147.[113,1089,1238,1614]" pageId="147" pageNumber="140">
Anterior margin of mesoventrite slightly emarginate medially; discal marginal mesoventral stria well impressed, carinate; disc of mesoventrite flat, smooth (at times sparsely punctate); meso-metaventral sutural stria well impressed, undulate; intercoxal disc of metaventrite smooth, with a longitudinal depression in male, area behind hind coxa with sparse punctures; lateral metaventral stria well impressed, curved outwardly, stopping short of metacoxa; lateral disc of metaventrite (
<figureCitation id="13173CCDFF0409BA60F0FA11CC2EF9F0" box="[423,516,1526,1550]" captionStart="Figs" captionStartId="146.[151,190,1489,1508]" captionTargetBox="[247,1029,217,1469]" captionTargetId="figure-11@146.[246,1029,217,1470]" captionTargetPageId="146" captionText="Figs. 433–443. Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805), SEM micrographs: 433 – habitus, dorsal view; 434 – ditto, ventral view; 435 – propygidium and pygidium; 436 – head, dorsal view; 437 – clypeus + labrum, dorsal view; 438 – head, ventral view; 439 – prosternum; 440 – antennal club, ventral view; 441 – lateral disc of metaventrite, metepisternum and fused metepimeron; 442 – protibia, dorsal view; 443 – ditto, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4272244" httpUri="https://zenodo.org/record/4272244/files/figure.png" pageId="147" pageNumber="140">Fig. 441</figureCitation>
) slightly concave, with deep setiferous punctures of various sizes; metepisternum + fused metepimeron (
<figureCitation id="13173CCDFF0409BA63CBF9F1CCDCF9D0" box="[668,758,1558,1582]" captionStart="Figs" captionStartId="146.[151,190,1489,1508]" captionTargetBox="[247,1029,217,1469]" captionTargetId="figure-11@146.[246,1029,217,1470]" captionTargetPageId="146" captionText="Figs. 433–443. Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805), SEM micrographs: 433 – habitus, dorsal view; 434 – ditto, ventral view; 435 – propygidium and pygidium; 436 – head, dorsal view; 437 – clypeus + labrum, dorsal view; 438 – head, ventral view; 439 – prosternum; 440 – antennal club, ventral view; 441 – lateral disc of metaventrite, metepisternum and fused metepimeron; 442 – protibia, dorsal view; 443 – ditto, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4272244" httpUri="https://zenodo.org/record/4272244/files/figure.png" pageId="147" pageNumber="140">Fig. 441</figureCitation>
) with much coarser and denser punctation, punctures without setae.
</paragraph>
<paragraph id="8B932048FF0309BD61E2FF3CCD4AFEED" blockId="148.[151,1127,219,1043]" pageId="148" pageNumber="145">Intercoxal disc of the first abdominal sternite completely striate laterally; almost smooth, with sparse punctures along lateral, anterior and posterior margins.</paragraph>
<paragraph id="8B932048FF0309BD61E2FEFCCC58FD8D" blockId="148.[151,1127,219,1043]" pageId="148" pageNumber="145">
Protibia (
<figureCitation id="13173CCDFF0309BD6042FEFCCF86FECA" box="[277,428,283,308]" captionStart="Figs" captionStartId="146.[151,190,1489,1508]" captionTargetBox="[247,1029,217,1469]" captionTargetId="figure-11@146.[246,1029,217,1470]" captionTargetPageId="146" captionText="Figs. 433–443. Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805), SEM micrographs: 433 – habitus, dorsal view; 434 – ditto, ventral view; 435 – propygidium and pygidium; 436 – head, dorsal view; 437 – clypeus + labrum, dorsal view; 438 – head, ventral view; 439 – prosternum; 440 – antennal club, ventral view; 441 – lateral disc of metaventrite, metepisternum and fused metepimeron; 442 – protibia, dorsal view; 443 – ditto, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4272244" httpUri="https://zenodo.org/record/4272244/files/figure.png" pageId="148" pageNumber="145">Figs. 442–443</figureCitation>
) flattened and somewhat dilated, outer margin with five moderately large triangular teeth topped with short rounded denticle followed by two minuscule denticles; setae of outer row moderately dense, regular and short; setae of median row similarly dense and regular, somewhat shorter than those of outer row; protarsal groove moderately deep; anterior protibial stria carinate, almost complete, shortened apically; one rather long and thin + one shorter but stouter tarsal denticle present apically; protibial spur tiny, straight, growing out from apical protibial margin; apical margin of protibia posteriorly with two tiny rounded apical denticles; outer part of posterior surface of protibia (
<figureCitation id="13173CCDFF0309BD625FFE1CCD48FDEA" box="[776,866,507,532]" captionStart="Figs" captionStartId="146.[151,190,1489,1508]" captionTargetBox="[247,1029,217,1469]" captionTargetId="figure-11@146.[246,1029,217,1470]" captionTargetPageId="146" captionText="Figs. 433–443. Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805), SEM micrographs: 433 – habitus, dorsal view; 434 – ditto, ventral view; 435 – propygidium and pygidium; 436 – head, dorsal view; 437 – clypeus + labrum, dorsal view; 438 – head, ventral view; 439 – prosternum; 440 – antennal club, ventral view; 441 – lateral disc of metaventrite, metepisternum and fused metepimeron; 442 – protibia, dorsal view; 443 – ditto, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4272244" httpUri="https://zenodo.org/record/4272244/files/figure.png" pageId="148" pageNumber="145">Fig. 443</figureCitation>
) areolate-rugose; distinctly separated from glabrous median part of posterior surface, basally with two minuscule denticles; posterior protibial stria complete, terminating in three tiny inner-posterior denticles; inner margin with double row of short setae.
</paragraph>
<paragraph id="8B932048FF0309BD61E2FD9CCF48FCAD" blockId="148.[151,1127,219,1043]" pageId="148" pageNumber="145">Mesotibia slender, outer margin with two sparse rows of thin denticles growing in size apically; setae of outer row sparse, strongly sclerotized; setae of median row irregular, microscopic; posterior mesotibial stria shortened apically; anterior surface of mesotibia glabrous; anterior mesotibial stria shortened apically; mesotibial spur stout, rather long; apical margin with several tiny denticles; claws of apical tarsomere shorter than half its length; metatibia basically similar to mesotibia, but slenderer and denticles of outer margin much sparser than those of mesotibia.</paragraph>
<paragraph id="8B932048FF0309BD61E2FCBCCA71FBED" blockId="148.[151,1127,219,1043]" pageId="148" pageNumber="145">
Male genitalia. Eighth sternite (
<figureCitation id="13173CCDFF0309BD6343FCBCCC99FC8D" box="[532,691,859,883]" captionStart="Figs" captionStartId="147.[113,152,1070,1089]" captionTargetBox="[181,996,220,1036]" captionTargetId="figure-143@147.[167,1035,208,1066]" captionTargetPageId="147" captionText="Figs. 444–452. Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805), male terminalia: 444 – 8th sternite and tergite, dorsal view; 445 – ditto, ventral view; 446 – ditto, lateral view; 447 – 9th tergite, 10th tergite, dorsal view; 448 – ditto, lateral view; 449 – aedeagus, dorsal view; 450 – ditto, lateral view; 451– spiculum gastrale, ventral view; 452 – ditto, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4272246" httpUri="https://zenodo.org/record/4272246/files/figure.png" pageId="148" pageNumber="145">Figs. 444–445</figureCitation>
) longitudinally entirely fused medially, apically with large inflatable membrane (velum); furnished with sparse microscopic setae; eighth tergite and eighth sternite fused laterally (
<figureCitation id="13173CCDFF0309BD63F5FC7CCD2AFC4D" box="[674,768,923,947]" captionStart="Figs" captionStartId="147.[113,152,1070,1089]" captionTargetBox="[181,996,220,1036]" captionTargetId="figure-143@147.[167,1035,208,1066]" captionTargetPageId="147" captionText="Figs. 444–452. Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805), male terminalia: 444 – 8th sternite and tergite, dorsal view; 445 – ditto, ventral view; 446 – ditto, lateral view; 447 – 9th tergite, 10th tergite, dorsal view; 448 – ditto, lateral view; 449 – aedeagus, dorsal view; 450 – ditto, lateral view; 451– spiculum gastrale, ventral view; 452 – ditto, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4272246" httpUri="https://zenodo.org/record/4272246/files/figure.png" pageId="148" pageNumber="145">Fig. 446</figureCitation>
). Morphology of 9 
<superScript id="7C598D00FF0309BD629DFC7DCDFCFC57" attach="left" box="[970,982,922,937]" fontSize="6" pageId="148" pageNumber="145">th</superScript>
tergite (
<figureCitation id="13173CCDFF0309BD6566FC7CCEDFFC2A" captionStart="Figs" captionStartId="147.[113,152,1070,1089]" captionTargetBox="[181,996,220,1036]" captionTargetId="figure-143@147.[167,1035,208,1066]" captionTargetPageId="147" captionText="Figs. 444–452. Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805), male terminalia: 444 – 8th sternite and tergite, dorsal view; 445 – ditto, ventral view; 446 – ditto, lateral view; 447 – 9th tergite, 10th tergite, dorsal view; 448 – ditto, lateral view; 449 – aedeagus, dorsal view; 450 – ditto, lateral view; 451– spiculum gastrale, ventral view; 452 – ditto, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4272246" httpUri="https://zenodo.org/record/4272246/files/figure.png" pageId="148" pageNumber="145">Figs. 447–448</figureCitation>
) typical for the subfamily; spiculum gastrale (
<figureCitation id="13173CCDFF0309BD6398FC5CCD0DFC2D" box="[719,807,955,979]" captionStart="Figs" captionStartId="147.[113,152,1070,1089]" captionTargetBox="[181,996,220,1036]" captionTargetId="figure-143@147.[167,1035,208,1066]" captionTargetPageId="147" captionText="Figs. 444–452. Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805), male terminalia: 444 – 8th sternite and tergite, dorsal view; 445 – ditto, ventral view; 446 – ditto, lateral view; 447 – 9th tergite, 10th tergite, dorsal view; 448 – ditto, lateral view; 449 – aedeagus, dorsal view; 450 – ditto, lateral view; 451– spiculum gastrale, ventral view; 452 – ditto, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4272246" httpUri="https://zenodo.org/record/4272246/files/figure.png" pageId="148" pageNumber="145">Fig. 451</figureCitation>
) expanded on both ends. Basal piece of aedeagus (
<figureCitation id="13173CCDFF0309BD6008FC3CCFD3FC0D" box="[351,505,987,1011]" captionStart="Figs" captionStartId="147.[113,152,1070,1089]" captionTargetBox="[181,996,220,1036]" captionTargetId="figure-143@147.[167,1035,208,1066]" captionTargetPageId="147" captionText="Figs. 444–452. Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805), male terminalia: 444 – 8th sternite and tergite, dorsal view; 445 – ditto, ventral view; 446 – ditto, lateral view; 447 – 9th tergite, 10th tergite, dorsal view; 448 – ditto, lateral view; 449 – aedeagus, dorsal view; 450 – ditto, lateral view; 451– spiculum gastrale, ventral view; 452 – ditto, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4272246" httpUri="https://zenodo.org/record/4272246/files/figure.png" pageId="148" pageNumber="145">Figs. 449–450</figureCitation>
) rather long, ratio of its length: length of parameres 1: 2.5; parameres fused approximately along their basal half; aedeagus curved ventrad (
<figureCitation id="13173CCDFF0309BD62A6FC1CCA65FBED" box="[1009,1103,1019,1043]" captionStart="Figs" captionStartId="147.[113,152,1070,1089]" captionTargetBox="[181,996,220,1036]" captionTargetId="figure-143@147.[167,1035,208,1066]" captionTargetPageId="147" captionText="Figs. 444–452. Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805), male terminalia: 444 – 8th sternite and tergite, dorsal view; 445 – ditto, ventral view; 446 – ditto, lateral view; 447 – 9th tergite, 10th tergite, dorsal view; 448 – ditto, lateral view; 449 – aedeagus, dorsal view; 450 – ditto, lateral view; 451– spiculum gastrale, ventral view; 452 – ditto, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4272246" httpUri="https://zenodo.org/record/4272246/files/figure.png" pageId="148" pageNumber="145">Fig. 450</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>