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<document id="23D46B7026204F6CC1AE3EC1BB043ABB" ID-DOI="10.5852/ejt.2019.526" ID-GBIF-Dataset="2da65c0a-9c48-43f5-ada3-e088de28dec1" ID-ISSN="2118-9773" ID-Zenodo-Dep="3152217" ID-ZooBank="3FFC527D-E9CA-4D9F-BF59-076FA60171AC" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_requiresApprovalFor="existingObjects,operationResults,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1558535825802" checkinUser="plazi" docAuthor="Nurinsiyah, Ayu Savitri, Neiber, Marco T. &amp; Hausdorf, Bernhard" docDate="2019" docId="03B1DB4C8B014E374C6FFBCE3DD8FB17" docLanguage="en" docName="ej-526_nurinsiyah_neiber_hausdorf.pdf.imf" docOrigin="European Journal of Taxonomy 526" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Landouria tonywhitteni Nurinsiyah &amp; Neiber &amp; Hausdorf 2019, sp. nov." docType="treatment" docUuid="59260748-3438-411F-82B4-14A7847AFA58" docUuidSource="ZooBank" docVersion="16" lastPageNumber="47" masterDocId="FF88A3348B2D4E194E35FFFC3C0DFFDB" masterDocTitle="Revision of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from Java" masterLastPageNumber="73" masterPageNumber="1" pageNumber="45" updateTime="1716320655398" updateUser="carolina">
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<mods:title id="1CE263811D436A7D8334D0389AEA5E8E">Revision of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from Java</mods:title>
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<mods:namePart id="56D4BD0EADB0984225D3251B3BA447DF">Nurinsiyah, Ayu Savitri</mods:namePart>
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<mods:affiliation id="2D5A48B2ACFAAC754C583DDC5254B263">Museum Zoologicum Bogoriense, Research Center for Biology, Indonesian Institute of Science, Cibinong, West Java, Indonesia.</mods:affiliation>
<mods:nameIdentifier id="FE69D8ABCDA2730014D31BEF4F99A0C6" type="email">ayu_nurinsiyah@yahoo.com &amp; ayus002@lipi.go.id</mods:nameIdentifier>
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<mods:namePart id="F47ED1FA79DF6B28BDD0F9D778C47A31">Neiber, Marco T.</mods:namePart>
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<mods:affiliation id="DB7B7B2284D0420C45D8ED47BE0715FA">Zoological Museum, Center of Natural History, University of Hamburg, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany</mods:affiliation>
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<mods:namePart id="99ADDDEFEC86FE56996DB93CAC5B6E20">Hausdorf, Bernhard</mods:namePart>
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<mods:affiliation id="86761A175637C531609E5B4F12707593">Zoological Museum, Center of Natural History, University of Hamburg, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany.</mods:affiliation>
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<mods:date id="66E99947AAD095E94276DC56983AC148">2019</mods:date>
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<subSubSection id="C30239D18B014E354C6FFBCE3FD7FB96" box="[602,986,1074,1101]" pageId="44" pageNumber="45" type="nomenclature">
<paragraph id="8BA76A5A8B014E354C6FFBCE3FD7FB96" blockId="44.[380,1207,1074,1172]" box="[602,986,1074,1101]" pageId="44" pageNumber="45">
<heading id="D0EFDD368B014E354C6FFBCE3FD7FB96" box="[602,986,1074,1101]" centered="true" fontSize="11" level="2" pageId="44" pageNumber="45" reason="2">
<taxonomicName id="4C1811D98B014E354C6FFBCE3F75FB96" ID-CoL="7RTJF" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[602,888,1074,1101]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="44" pageNumber="45" phylum="Mollusca" rank="species" species="tonywhitteni" status="sp. nov.">
<emphasis id="B96CB6488B014E354C6FFBCE3F75FB96" bold="true" box="[602,888,1074,1101]" italics="true" pageId="44" pageNumber="45">Landouria tonywhitteni</emphasis>
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<taxonomicNameLabel id="A25F0B338B014E354D4AFBCF3FD7FB96" box="[895,986,1075,1101]" pageId="44" pageNumber="45" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C30239D18B014E354F49FBAA3FC2FB48" pageId="44" pageNumber="45" type="description">
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urn:lsid:zoobank.org:act: 
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<paragraph id="8BA76A5A8B014E354C50FB853FC2FB48" blockId="44.[380,1207,1074,1172]" box="[613,975,1145,1172]" pageId="44" pageNumber="45">
<figureCitation id="132376DF8B014E354C50FB853EB2FB48" box="[613,703,1145,1171]" captionStart="Figs 29–47" captionStartId="29.[189,241,1366,1392]" captionTargetBox="[189,1399,649,1337]" captionTargetId="figure@29.[189,1399,649,1337]" captionTargetPageId="29" captionText="Figs 29–47. Flagella of species of Landouria from Java. 29. L. winteriana (Pfeiffer, 1842), West Java, Padalarang, Gunung Masigit (neotype, ZMA 376193a). 30. L. rotatoria (Pfeiffer, 1842), West Java, Gunung Salak, 1166 m a.s.l. (neotype, MZB 19205). 31. L. epiplatia (Möllendorff, 1897)?, West Java, Bogor, Gunung Gede (ZMH 79946). 32. L. nusakambangensis sp. nov., Central Java, Nusa Kambangan Island, nature monument Gligir (holotype, ZMA 412298a). 33. L. petrukensis sp. nov., Central Java, Karangbolong, way to Petruk Cave (holotype, MZB 19203). 34. L. tholiformis sp. nov., Yogyakarta, Gunung Kidul, Tepus, near Poktunggal Beach (holotype, MZB 17999). 35. L. abdidalem sp. nov., Yogyakarta, Kulonprogo, Sabrang Kidul (holotype, MZB 17664). 36. L. madurensis sp. nov., East Java, Madura Island, Sampang, Nepa Monkey lowland rainforest (paratype, MZB 19209). 37. L. sewuensis sp. nov., Yogyakarta, Gunung Kidul, Tepus, Mbajo Hill (holotype, MZB 19255). 38. L. sukoliloensis sp. nov., Central Java, Kayen, near Watu Payung (holotype, MZB 19219). 39. L. tonywhitteni sp. nov., Central Java, Taruman Village, near Sendang Sedayu (holotype, MZB 19214). 40. L. nodifera sp. nov., Central Java, Larangan Village, Lanang Spring (holotype, MZB 19227). 41. L. pacitanensis sp. nov., East Java, Pacitan, near Srau Beach (MZB 19231). 42. L. intumescens (Martens, 1867), East Java, Guwoterus (MZB 19207). 43. L. smimensis (Mousson, 1848), East Java, Tosari (ZMA 390881). 44. L. zonifera sp. nov., Yogyakarta, Kulonprogo, Sibolong (holotype, MZB 17727). 45. L. ciliocincta (Möllendorff, 1897), East Java, Malang, Kondangmerak (MZB 19211). 46. L. pakidulan sp. nov., Central Java, Nusa Kambangan Island, nature monument Gligir (holotype, ZMA 417011a). 47. L. menorehensis sp. nov., Yogyakarta, Kulonprogo, Sibolong (holotype, MZB 19201). Scale bar: 1 mm." httpUri="https://zenodo.org/record/3152255/files/figure.png" pageId="44" pageNumber="45">Figs 39</figureCitation>
, 
<figureCitation id="132376DF8B014E354CFEFB853EE6FB4F" box="[715,747,1145,1172]" captionStart="Figs 52–55" captionStartId="33.[189,241,1717,1743]" captionTargetBox="[189,1398,265,1993]" captionTargetId="figure@34.[189,1398,265,1993]" captionTargetPageId="34" captionText="Figs 52–55 (opposite page). Distribution of species of Landouria in Java (10′ grid). 52. L. winteriana (Pfeiffer, 1842), L. naggsi sp. nov., L. nusakambangensis sp. nov., L. petrukensis sp. nov., L. tholiformis sp. nov., L. tonywhitteni sp. nov., L. smimensis (Mousson, 1848). 53. L. conoidea (Leschke, 1914), L. abdidalem sp. nov., L. sewuensis sp. nov., L. sukoliloensis sp. nov., L. pacitanensis sp. nov., L. intumescens (Martens, 1867), L. madurensis sp. nov. 54. L. rotatoria (Pfeiffer, 1842), L. nodifera sp. nov., L. parahyangensis sp. nov., L. schepmani (Möllendorff, 1897), L. zonifera sp. nov., L. ciliocincta (Möllendorff, 1897), L. pakidulan sp. nov. 55. L. epiplatia (Möllendorff, 1897), L. monticola van Benthem Jutting, 1950, L. davini Dharma, 2015, L. menorehensis sp. nov., L. dharmai sp. nov., L. moussoniana (Martens, 1867)." httpUri="https://zenodo.org/record/3152265/files/figure.png" pageId="44" pageNumber="45">52</figureCitation>
, 
<figureCitation id="132376DF8B014E354CC2FB853F1AFB48" box="[759,791,1145,1171]" captionStart="Figs 60–62" captionStartId="45.[189,241,1893,1919]" captionTargetBox="[210,1377,263,1867]" captionTargetId="figure@45.[210,1377,263,1867]" captionTargetPageId="45" captionText="Figs 60–62. Shells of species of Landouria from Java. 60. L. tonywhitteni sp. nov., Central Java, Taruman Village, near Sendang Sedayu (holotype, MZB 19214). 61. L. nodifera sp. nov., Central Java, Larangan Village, Lanang Spring (holotype, MZB 19227). 62. L. pacitanensis sp. nov., East Java, Pacitan, Srau Beach (holotype, MZB 21191). Scale bar: 5 mm." httpUri="https://zenodo.org/record/3152275/files/figure.png" pageId="44" pageNumber="45">60</figureCitation>
, 
<figureCitation id="132376DF8B014E354D16FB853F4EFB48" box="[803,835,1145,1171]" captionStart="Figs 63–71" captionStartId="48.[189,241,1668,1694]" captionTargetBox="[189,1394,515,1612]" captionTargetId="figure@48.[189,1398,510,1629]" captionTargetPageId="48" captionText="Figs 63–71. Genitalia of species of Landouria from Java. 63. L. tonywhitteni sp. nov., Central Java, Taruman Village, near Sendang Sedayu (holotype, MZB 19214). 64. L. nodifera sp. nov., Central Java, Larangan Village, Lanang Spring (holotype, MZB 19227). 65. L. pacitanensis sp. nov., East Java, Pacitan, near Srau Beach (MZB 19231). 66. L. intumescens (Martens, 1867), East Java, Guwoterus (MZB 19207). 67. L. smimensis (Mousson, 1848), East Java, Tosari (ZMA 390881). 68. L. zonifera sp. nov., Yogyakarta, Kulonprogo, Sibolong (holotype, MZB 17727). 69. L. ciliocincta (Möllendorff, 1897), East Java, Malang, Kondangmerak (MZB 19211). 70. L. pakidulan sp. nov., Central Java, Nusa Kambangan Island, nature monument Gligir (holotype, ZMA 417011a). 71. L. menorehensis sp. nov., Yogyakarta, Kulonprogo, Sibolong (holotype, MZB 19201). Scale bars: 1 mm." httpUri="https://zenodo.org/record/3152282/files/figure.png" pageId="44" pageNumber="45">63</figureCitation>
; 
<tableCitation id="C69A5FE18B014E354D7AFB853FC2FB48" box="[847,975,1145,1171]" captionStart-0="Table 3" captionStart-1="Table 4" captionStartId-0="13.[189,258,268,294]" captionStartId-1="22.[188,257,551,579]" captionTargetBox-0="[201,1370,481,1908]" captionTargetBox-1="[205,1929,661,1732]" captionTargetPageId-0="13" captionTargetPageId-1="22" captionText-0="Table 3 (continued on next 3 pages). Shell measurements for species of Landouria Godwin-Austen, 1918 (in mm). Abbreviations: D = shell diameter; D3W = diameter of the third whorl; da = diameter of the aperture; H = shell height; ha = height of the aperture; max = maximum; min = minimum; n = number of measured spec.; std = standard deviation; U = diameter of the umbilicus; W = number of whorls." captionText-1="Table 4 (continued on next 3 pages). Diagnostic characteristics of the species of Landouria Godwin-Austen, 1918 from Java. Characteristics most important for distinguishing similar taxa are printed in bold." httpUri-0="http://table.plazi.org/id/DF673AD28B204E144E88FEF03D19FE68" httpUri-1="http://table.plazi.org/id/DF673AD28B3B4E0F4E89FDDB3F73FDBC" pageId="44" pageNumber="45" tableUuid-0="DF673AD28B204E144E88FEF03D19FE68" tableUuid-1="DF673AD28B3B4E0F4E89FDDB3F73FDBC">Tables 3–4</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30239D18B014E354E88FB463D39FAB6" pageId="44" pageNumber="45" type="diagnosis">
<paragraph id="8BA76A5A8B014E354E88FB463D3BFB0F" blockId="44.[189,310,1210,1236]" box="[189,310,1210,1236]" pageId="44" pageNumber="45">
<heading id="D0EFDD368B014E354E88FB463D3BFB0F" bold="true" box="[189,310,1210,1236]" fontSize="11" level="3" pageId="44" pageNumber="45" reason="3">
<emphasis id="B96CB6488B014E354E88FB463D3BFB0F" bold="true" box="[189,310,1210,1236]" pageId="44" pageNumber="45">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BA76A5A8B014E354E88FB153D39FAB6" blockId="44.[189,1399,1257,1389]" pageId="44" pageNumber="45">
<taxonomicName id="4C1811D98B014E354E88FB153DD8FAD8" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[189,469,1257,1283]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="44" pageNumber="45" phylum="Mollusca" rank="species" species="tonywhitteni" status="sp. nov.">
<emphasis id="B96CB6488B014E354E88FB153DD8FAD8" box="[189,469,1257,1283]" italics="true" pageId="44" pageNumber="45">Landouria tonywhitteni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B014E354FD4FB163E32FADF" box="[481,575,1258,1284]" pageId="44" pageNumber="45" rank="species">sp. nov.</taxonomicNameLabel>
is characterized by having a strongly depressed conical, brownishcorneous, sharply keeled shell and a penis that does not become distinctly broader towards is proximal end, the lack of a coecum at the proximal end of the penis and the presence of a node at the base of the flagellum.
</paragraph>
</subSubSection>
<subSubSection id="C30239D18B014E354E88FA603D69F9F0" pageId="44" pageNumber="45" type="etymology">
<paragraph id="8BA76A5A8B014E354E88FA603D49FA6D" blockId="44.[189,324,1436,1462]" box="[189,324,1436,1462]" pageId="44" pageNumber="45">
<heading id="D0EFDD368B014E354E88FA603D49FA6D" bold="true" box="[189,324,1436,1462]" fontSize="11" level="3" pageId="44" pageNumber="45" reason="3">
<emphasis id="B96CB6488B014E354E88FA603D49FA6D" bold="true" box="[189,324,1436,1462]" pageId="44" pageNumber="45">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA76A5A8B014E354E88FA363D69F9F0" blockId="44.[189,1398,1482,1579]" pageId="44" pageNumber="45">
<taxonomicName id="4C1811D98B014E354E88FA363DDFFA3F" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[189,466,1482,1508]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="44" pageNumber="45" phylum="Mollusca" rank="species" species="tonywhitteni" status="sp. nov.">
<emphasis id="B96CB6488B014E354E88FA363DDFFA3F" box="[189,466,1482,1508]" italics="true" pageId="44" pageNumber="45">Landouria tonywhitteni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B014E354FEEFA373E3BFA3E" box="[475,566,1483,1509]" pageId="44" pageNumber="45" rank="species">sp. nov.</taxonomicNameLabel>
is named in honour of the late British naturalist Tony Whitten (
<date id="FFA64C9A8B014E354B19FA363CF1F9D3" pageId="44" pageNumber="45" value="1953" valueMax="2017">1953– 2017</date>
), who dedicated his life to the conservation of karst habitats and especially the snails that inhabit these habitats.
</paragraph>
</subSubSection>
<subSubSection id="C30239D18B014E354E88F9A53E67F83A" pageId="44" pageNumber="45" type="materials_examined">
<paragraph id="8BA76A5A8B014E354E88F9A53DA7F9A8" blockId="44.[189,426,1625,1651]" box="[189,426,1625,1651]" pageId="44" pageNumber="45">
<heading id="D0EFDD368B014E354E88F9A53DA7F9A8" bold="true" box="[189,426,1625,1651]" fontSize="11" level="3" pageId="44" pageNumber="45" reason="3">
<emphasis id="B96CB6488B014E354E88F9A53DA7F9A8" bold="true" box="[189,426,1625,1651]" pageId="44" pageNumber="45">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B7060078B014E354EDFF9743810F8D7" ID-GBIF-Occurrence="2243137545" collectingDate="2014-08-16" collectionCode="MZB" collectorName="A. S. Nurinsiyah &amp; F. L. H. Irsyad &amp; F. J. Sari &amp; E. Nurlaela" country="Indonesia" county="Grobogan" elevation="20" latitude="-7.0144444" location="near Sendang Sedayu" longLatPrecision="21" longitude="110.858055" municipality="Taruman" pageId="44" pageNumber="45" specimenCode="19214" specimenCount="1" stateProvince="Central Java" typeStatus="holotype">
<paragraph id="8BA76A5A8B014E354EDFF9743D51F979" blockId="44.[189,1398,1672,1804]" box="[234,348,1672,1698]" pageId="44" pageNumber="45">
<typeStatus id="54A3D4F88B014E354EDFF9743D51F979" box="[234,348,1672,1698]" pageId="44" pageNumber="45" type="holotype">
<emphasis id="B96CB6488B014E354EDFF9743D51F979" bold="true" box="[234,348,1672,1698]" pageId="44" pageNumber="45">Holotype</emphasis>
</typeStatus>
</paragraph>
<paragraph id="8BA76A5A8B014E354E88F9503810F8D7" blockId="44.[189,1398,1672,1804]" pageId="44" pageNumber="45">
<collectingCountry id="F30F2ACA8B014E354E88F9503D56F91D" box="[189,347,1708,1734]" name="Indonesia" pageId="44" pageNumber="45">INDONESIA</collectingCountry>
• holotype (det. anat.); 
<collectingRegion id="49DCA4B88B014E354CAAF9573F30F91D" box="[671,829,1707,1734]" country="Indonesia" name="Jawa Tengah" pageId="44" pageNumber="45">Central Java</collectingRegion>
, 
<collectingCounty id="62C612D68B014E354D67F9573FC7F91E" box="[850,970,1707,1733]" pageId="44" pageNumber="45">Grobogan</collectingCounty>
, 
<collectingMunicipality id="6BC3F0208B014E354DEAF9503847F91D" box="[991,1098,1708,1734]" pageId="44" pageNumber="45">Taruman</collectingMunicipality>
, 
<location id="8EC73C818B014E354A6AF950397FF91E" LSID="urn:lsid:plazi:treatment:03B1DB4C8B014E374C6FFBCE3DD8FB17:8EC73C818B014E354A6AF950397FF91E" box="[1119,1394,1707,1734]" country="Indonesia" county="Grobogan" latitude="-7.0144444" longLatPrecision="21" longitude="110.858055" municipality="Taruman" name="near Sendang Sedayu" pageId="44" pageNumber="45" stateProvince="Central Java">near Sendang Sedayu</location>
; 
<geoCoordinate id="EE2C0C9D8B014E354E88F9313D4FF932" box="[189,322,1741,1769]" degrees="7" direction="south" minutes="00" orientation="latitude" pageId="44" pageNumber="45" precision="15" seconds="52" value="-7.0144444">7°00′52″ S</geoCoordinate>
, 
<geoCoordinate id="EE2C0C9D8B014E354F66F9313DF5F932" box="[339,504,1741,1769]" degrees="110" direction="east" minutes="51" orientation="longitude" pageId="44" pageNumber="45" precision="15" seconds="29" value="110.858055">110°51′29″ E</geoCoordinate>
; 
<elevation id="00358D698B014E354C3DF9333E81F932" box="[520,652,1743,1769]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="44" pageNumber="45" unit="m" value="20.0">
<quantity id="4CE0C7BF8B014E354C3DF9333E45F932" box="[520,584,1743,1769]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="44" pageNumber="45" unit="m" value="20.0">20 m</quantity>
a.s.l.
</elevation>
; 
<date id="FFA64C9A8B014E354CA8F9323F45F932" box="[669,840,1742,1769]" pageId="44" pageNumber="45" value="2014-08-16">
<collectingDate id="EFE2B5728B014E354CA8F9323F45F932" box="[669,840,1742,1769]" pageId="44" pageNumber="45" value="2014-08-16">16 Aug. 2014</collectingDate>
</date>
; 
<collectorName id="26ED0F8C8B014E354D62F9333816F932" box="[855,1051,1743,1769]" pageId="44" pageNumber="45">A.S. Nurinsiyah</collectorName>
, 
<collectorName id="26ED0F8C8B014E354A1EF93338C3F932" box="[1067,1230,1743,1769]" pageId="44" pageNumber="45">F.L.H. Irsyad</collectorName>
, 
<collectorName id="26ED0F8C8B014E354AEBF933394DF932" box="[1246,1344,1743,1769]" pageId="44" pageNumber="45">F.J. Sari</collectorName>
and 
<collectorName id="26ED0F8C8B014E354E88F90E3D4EF8D7" box="[189,323,1778,1804]" pageId="44" pageNumber="45">E. Nurlaela</collectorName>
leg.; measurements: D = 
<quantity id="4CE0C7BF8B014E354C45F90E3ED1F8D7" box="[624,732,1778,1804]" metricMagnitude="-2" metricUnit="m" metricValue="1.76" pageId="44" pageNumber="45" unit="mm" value="17.6">17.6 mm</quantity>
, H = 
<quantity id="4CE0C7BF8B014E354D29F90E3F74F8D7" box="[796,889,1778,1804]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" pageId="44" pageNumber="45" unit="mm" value="7.6">7.6 mm</quantity>
; 
<collectionCode id="ED09F29F8B014E354DB0F90E3FC8F8D7" box="[901,965,1778,1804]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="44" pageNumber="45" type="Museum">MZB</collectionCode>
<specimenCode id="DBBEC2218B014E354DF9F90E3814F8D7" box="[972,1049,1778,1804]" pageId="44" pageNumber="45">19214</specimenCode>
.
</paragraph>
</materialsCitation>
<materialsCitation id="3B7060078B014E354EDFF8C73868F8A3" ID-GBIF-Occurrence="2243137468" collectingDate="2014-08-16" collectionCode="MZB" collectorName="A. S. Nurinsiyah &amp; F. L. H. Irsyad &amp; F. J. Sari &amp; E. Nurlaela" country="Indonesia" county="Grobogan" elevation="20" latitude="-7.0144444" location="Central Java" longLatPrecision="21" longitude="110.858055" municipality="Taruman" pageId="44" pageNumber="45" specimenCode="19217" specimenCount="9" stateProvince="Central Java" typeStatus="paratype">
<paragraph id="8BA76A5A8B014E354EDFF8C73D6AF88E" blockId="44.[189,1399,1851,2018]" box="[234,359,1851,1877]" pageId="44" pageNumber="45">
<typeStatus id="54A3D4F88B014E354EDFF8C73D6AF88E" box="[234,359,1851,1877]" pageId="44" pageNumber="45" type="paratype">
<emphasis id="B96CB6488B014E354EDFF8C73D6AF88E" bold="true" box="[234,359,1851,1877]" pageId="44" pageNumber="45">Paratypes</emphasis>
</typeStatus>
</paragraph>
<paragraph id="8BA76A5A8B014E354E88F8A23E67F83A" blockId="44.[189,1399,1851,2018]" pageId="44" pageNumber="45">
<collectingCountry id="F30F2ACA8B014E354E88F8A23D56F8A3" box="[189,347,1886,1912]" name="Indonesia" pageId="44" pageNumber="45">INDONESIA</collectingCountry>
– 
<collectingRegion id="49DCA4B88B014E354F4CF8A23E11F8A3" box="[377,540,1886,1912]" country="Indonesia" name="Jawa Tengah" pageId="44" pageNumber="45">
<emphasis id="B96CB6488B014E354F4CF8A23E11F8A3" bold="true" box="[377,540,1886,1912]" pageId="44" pageNumber="45">Central Java</emphasis>
</collectingRegion>
• 
<specimenCount id="9D1EA1D38B014E354C03F8A23E86F8A2" box="[566,651,1886,1913]" count="9" pageId="44" pageNumber="45" type="generic">9 spec.</specimenCount>
; same data as for holotype; 
<collectionCode id="ED09F29F8B014E354DE0F8A33818F8A2" box="[981,1045,1887,1913]" collectionName="MZB" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="44" pageNumber="45" type="Museum">MZB</collectionCode>
<specimenCode id="DBBEC2218B014E354A29F8A23868F8A3" box="[1052,1125,1886,1912]" pageId="44" pageNumber="45">19217</specimenCode>
• 
<materialsCitation id="3B7060078B014E354A4AF8A23DF4F840" ID-GBIF-Occurrence="2243137345" collectingDate="2014-08-16" collectionCode="ZMH" collectorName="A. S. Nurinsiyah &amp; F. L. H. Irsyad &amp; F. J. Sari &amp; E. Nurlaela" country="Indonesia" county="Grobogan" elevation="20" latitude="-7.0144444" location="Sendang Sedayu" longLatPrecision="21" longitude="110.858055" municipality="Taruman" pageId="44" pageNumber="45" specimenCode="ZMH 133434" specimenCount="5" stateProvince="Jawa Tengah" typeStatus="paratype">
<specimenCount id="9D1EA1D38B014E354A4AF8A238D8F8A2" box="[1151,1237,1886,1913]" count="5" pageId="44" pageNumber="45" type="generic">5 spec.</specimenCount>
; same data as for holotype; 
<specimenCode id="DBBEC2218B014E354F6FF87E3DF4F840" box="[346,505,1921,1948]" collectionCode="ZMH" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="44" pageNumber="45" type="Museum">ZMH 133434</specimenCode>
</materialsCitation>
• 
<materialsCitation id="3B7060078B014E354C25F87D3E71F864" collectionCode="MZB" elevation="23" latitude="-7.0144444" location="Grobogan" longLatPrecision="21" longitude="110.858055" municipality="Taruman" pageId="44" pageNumber="45" specimenCode="MZB 19215" specimenCount="9">
<specimenCount id="9D1EA1D38B014E354C25F87D3E6DF847" box="[528,608,1921,1948]" count="9" pageId="44" pageNumber="45" type="generic">9 spec.</specimenCount>
(det. anat.); 
<location id="8EC73C818B014E354CC4F87D3F64F840" LSID="urn:lsid:plazi:treatment:03B1DB4C8B014E374C6FFBCE3DD8FB17:8EC73C818B014E354CC4F87D3F64F840" box="[753,873,1921,1947]" latitude="-7.0144444" longLatPrecision="21" longitude="110.858055" municipality="Taruman" name="Grobogan" pageId="44" pageNumber="45">Grobogan</location>
, 
<collectingMunicipality id="6BC3F0208B014E354D41F87E3FD2F847" box="[884,991,1922,1948]" pageId="44" pageNumber="45">Taruman</collectingMunicipality>
, 
<location id="8EC73C818B014E354DDCF87E38EBF840" LSID="urn:lsid:plazi:treatment:03B1DB4C8B014E374C6FFBCE3DD8FB17:8EC73C818B014E354DDCF87E38EBF840" box="[1001,1254,1921,1948]" latitude="-7.0144444" longLatPrecision="21" longitude="110.858055" municipality="Taruman" name="near Sendang Sedayu" pageId="44" pageNumber="45">near Sendang Sedayu</location>
; 
<geoCoordinate id="EE2C0C9D8B014E354AC4F883397CF840" box="[1265,1393,1919,1947]" degrees="7" direction="south" minutes="00" orientation="latitude" pageId="44" pageNumber="45" precision="15" seconds="52" value="-7.0144444">7°00′52″ S</geoCoordinate>
, 
<geoCoordinate id="EE2C0C9D8B014E354E88F85F3D51F864" box="[189,348,1955,1983]" degrees="110" direction="east" minutes="51" orientation="longitude" pageId="44" pageNumber="45" precision="15" seconds="29" value="110.858055">110°51′29″ E</geoCoordinate>
; 
<quantity id="4CE0C7BF8B014E354F5DF8593DAFF864" box="[360,418,1957,1983]" metricMagnitude="1" metricUnit="m" metricValue="2.3" pageId="44" pageNumber="45" unit="m" value="23.0">
<elevation id="00358D698B014E354F5DF8593DAFF864" box="[360,418,1957,1983]" metricMagnitude="1" metricUnit="m" metricValue="2.3" pageId="44" pageNumber="45" unit="m" value="23.0">23 m</elevation>
</quantity>
a.s.l.; 
<specimenCode id="DBBEC2218B014E354FD8F8593E71F864" box="[493,636,1956,1983]" pageId="44" pageNumber="45">MZB 19215</specimenCode>
</materialsCitation>
• 
<materialsCitation id="3B7060078B014E354CA6F85938C9F865" ID-GBIF-Occurrence="2243137357" box="[659,1220,1956,1983]" collectingDate="2014-08-16" collectionCode="MZB" country="Indonesia" county="Grobogan" elevation="20" latitude="-7.0144444" location="Sendang Sedayu" longLatPrecision="21" longitude="110.858055" municipality="Taruman" pageId="44" pageNumber="45" specimenCode="21268" specimenCount="1" stateProvince="Jawa Tengah" typeStatus="paratype">
<specimenCount id="9D1EA1D38B014E354CA6F8593EEAF864" box="[659,743,1957,1983]" count="1" pageId="44" pageNumber="45" type="generic">1 spec.</specimenCount>
; same data as for preceding; 
<collectionCode id="ED09F29F8B014E354A00F8593878F864" box="[1077,1141,1957,1983]" collectionName="MZB" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="44" pageNumber="45" type="Museum">MZB</collectionCode>
<specimenCode id="DBBEC2218B014E354A4EF85838C9F865" box="[1147,1220,1956,1982]" pageId="44" pageNumber="45">21268</specimenCode>
</materialsCitation>
• 
<materialsCitation id="3B7060078B014E354AEEF8583E6BF83A" ID-GBIF-Occurrence="2243137319" collectingDate="2014-08-16" collectionCode="ZMH" country="Indonesia" county="Grobogan" elevation="20" latitude="-7.0144444" location="Sendang Sedayu" longLatPrecision="21" longitude="110.858055" municipality="Taruman" pageId="44" pageNumber="45" specimenCode="ZMH 133436" specimenCount="5" stateProvince="Jawa Tengah" typeStatus="paratype">
<specimenCount id="9D1EA1D38B014E354AEEF8583922F864" box="[1243,1327,1956,1983]" count="5" pageId="44" pageNumber="45" type="generic">5 spec.</specimenCount>
; same data as for preceding; 
<specimenCode id="DBBEC2218B014E354FF7F8343E6BF83A" box="[450,614,1991,2018]" collectionCode="ZMH" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="44" pageNumber="45" type="Museum">ZMH 133436</specimenCode>
</materialsCitation>
.
</paragraph>
</materialsCitation>
</subSubSection>
<caption id="DF673AD28B004E344E88F8993EF6F831" httpUri="https://zenodo.org/record/3152275/files/figure.png" pageId="45" pageNumber="46" startId="45.[189,241,1893,1919]" targetBox="[210,1377,263,1867]" targetPageId="45">
<paragraph id="8BA76A5A8B004E344E88F8993EF6F831" blockId="45.[189,1399,1893,2026]" pageId="45" pageNumber="46">
<emphasis id="B96CB6488B004E344E88F8993D4AF85B" bold="true" box="[189,327,1893,1920]" pageId="45" pageNumber="46">Figs 60–62.</emphasis>
Shells of species of 
<taxonomicName id="4C1811D98B004E344C1AF89A3EA6F85B" authorityName="Godwin-Austen" authorityYear="1918" box="[559,683,1894,1920]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="45" pageNumber="46" phylum="Mollusca" rank="genus">
<emphasis id="B96CB6488B004E344C1AF89A3EA6F85B" box="[559,683,1894,1920]" italics="true" pageId="45" pageNumber="46">Landouria</emphasis>
</taxonomicName>
from 
<collectingRegion id="49DCA4B88B004E344CDAF89A3F2AF85B" box="[751,807,1894,1920]" country="Indonesia" name="Jawa Timur" pageId="45" pageNumber="46">Java</collectingRegion>
. 
<emphasis id="B96CB6488B004E344D05F89A3F40F85B" bold="true" box="[816,845,1894,1920]" pageId="45" pageNumber="46">60</emphasis>
. 
<taxonomicName id="4C1811D98B004E344D6FF89B3805F85B" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[858,1032,1894,1920]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="45" pageNumber="46" phylum="Mollusca" rank="species" species="tonywhitteni" status="sp. nov.">
<emphasis id="B96CB6488B004E344D6FF89B3805F85B" box="[858,1032,1894,1920]" italics="true" pageId="45" pageNumber="46">L. tonywhitteni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B004E344A3BF89A3864F85B" box="[1038,1129,1894,1920]" pageId="45" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
, 
<collectingRegion id="49DCA4B88B004E344A46F89A3908F85B" box="[1139,1285,1894,1920]" country="Indonesia" name="Jawa Tengah" pageId="45" pageNumber="46">Central Java</collectingRegion>
, Taruman Village, near Sendang Sedayu (
<typeStatus id="54A3D4F88B004E344C1EF8753E9BF878" box="[555,662,1929,1955]" pageId="45" pageNumber="46" type="holotype">holotype</typeStatus>
, MZB 19214). 
<emphasis id="B96CB6488B004E344D7FF8753F6AF878" bold="true" box="[842,871,1929,1955]" pageId="45" pageNumber="46">61</emphasis>
. 
<taxonomicName id="4C1811D98B004E344D43F8763FF5F878" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[886,1016,1929,1955]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="45" pageNumber="46" phylum="Mollusca" rank="species" species="nodifera" status="sp. nov.">
<emphasis id="B96CB6488B004E344D43F8763FF5F878" box="[886,1016,1929,1955]" italics="true" pageId="45" pageNumber="46">L. nodifera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B004E344DCAF8753851F878" box="[1023,1116,1929,1955]" pageId="45" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
, 
<collectingRegion id="49DCA4B88B004E344A5DF87538F1F878" box="[1128,1276,1929,1955]" country="Indonesia" name="Jawa Tengah" pageId="45" pageNumber="46">Central Java</collectingRegion>
, Larangan Village, Lanang Spring (
<typeStatus id="54A3D4F88B004E344FD7F8503E40F81D" box="[482,589,1964,1990]" pageId="45" pageNumber="46" type="holotype">holotype</typeStatus>
, MZB 19227). 
<emphasis id="B96CB6488B004E344D33F8503F2EF81D" bold="true" box="[774,803,1964,1990]" pageId="45" pageNumber="46">62</emphasis>
. 
<taxonomicName id="4C1811D98B004E344D06F8513FEBF81D" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[819,998,1965,1990]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="45" pageNumber="46" phylum="Mollusca" rank="species" species="pacitanensis" status="sp. nov.">
<emphasis id="B96CB6488B004E344D06F8513FEBF81D" box="[819,998,1965,1990]" italics="true" pageId="45" pageNumber="46">L. pacitanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B004E344DDAF8513840F81C" box="[1007,1101,1965,1991]" pageId="45" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
, 
<collectingRegion id="49DCA4B88B004E344A6EF85138C0F81C" box="[1115,1229,1965,1991]" country="Indonesia" name="Jawa Timur" pageId="45" pageNumber="46">East Java</collectingRegion>
, Pacitan, Srau Beach (
<typeStatus id="54A3D4F88B004E344F23F8333D8CF832" box="[278,385,1999,2025]" pageId="45" pageNumber="46" type="holotype">holotype</typeStatus>
, MZB 21191). Scale bar: 
<quantity id="4CE0C7BF8B004E344C85F8333EFAF831" box="[688,759,1999,2026]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="45" pageNumber="46" unit="mm" value="5.0">5 mm</quantity>
.
</paragraph>
</caption>
<subSubSection id="C30239D18B034E374E88FEF73842FCE0" pageId="46" pageNumber="47" type="description">
<paragraph id="8BA76A5A8B034E374E88FEF73D43FEFE" blockId="46.[189,334,267,293]" box="[189,334,267,293]" pageId="46" pageNumber="47">
<heading id="D0EFDD368B034E374E88FEF73D43FEFE" bold="true" box="[189,334,267,293]" fontSize="11" level="3" pageId="46" pageNumber="47" reason="3">
<emphasis id="B96CB6488B034E374E88FEF73D43FEFE" bold="true" box="[189,334,267,293]" pageId="46" pageNumber="47">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BA76A5A8B034E374E88FEC63FA8FDD8" blockId="46.[189,1399,313,516]" pageId="46" pageNumber="47">
SHELL (
<figureCitation id="132376DF8B034E374F26FEC53D66FE88" box="[275,363,313,339]" captionStart="Figs 60–62" captionStartId="45.[189,241,1893,1919]" captionTargetBox="[210,1377,263,1867]" captionTargetId="figure@45.[210,1377,263,1867]" captionTargetPageId="45" captionText="Figs 60–62. Shells of species of Landouria from Java. 60. L. tonywhitteni sp. nov., Central Java, Taruman Village, near Sendang Sedayu (holotype, MZB 19214). 61. L. nodifera sp. nov., Central Java, Larangan Village, Lanang Spring (holotype, MZB 19227). 62. L. pacitanensis sp. nov., East Java, Pacitan, Srau Beach (holotype, MZB 21191). Scale bar: 5 mm." httpUri="https://zenodo.org/record/3152275/files/figure.png" pageId="46" pageNumber="47">Fig. 60</figureCitation>
; 
<tableCitation id="C69A5FE18B034E374F42FEC53DF6FE8F" box="[375,507,313,340]" captionStart-0="Table 3" captionStart-1="Table 4" captionStartId-0="13.[189,258,268,294]" captionStartId-1="22.[188,257,551,579]" captionTargetBox-0="[201,1370,481,1908]" captionTargetBox-1="[205,1929,661,1732]" captionTargetPageId-0="13" captionTargetPageId-1="22" captionText-0="Table 3 (continued on next 3 pages). Shell measurements for species of Landouria Godwin-Austen, 1918 (in mm). Abbreviations: D = shell diameter; D3W = diameter of the third whorl; da = diameter of the aperture; H = shell height; ha = height of the aperture; max = maximum; min = minimum; n = number of measured spec.; std = standard deviation; U = diameter of the umbilicus; W = number of whorls." captionText-1="Table 4 (continued on next 3 pages). Diagnostic characteristics of the species of Landouria Godwin-Austen, 1918 from Java. Characteristics most important for distinguishing similar taxa are printed in bold." httpUri-0="http://table.plazi.org/id/DF673AD28B204E144E88FEF03D19FE68" httpUri-1="http://table.plazi.org/id/DF673AD28B3B4E0F4E89FDDB3F73FDBC" pageId="46" pageNumber="47" tableUuid-0="DF673AD28B204E144E88FEF03D19FE68" tableUuid-1="DF673AD28B3B4E0F4E89FDDB3F73FDBC">Tables 3–4</tableCitation>
). Depressed conical, with 5.75–6 hardly convex whorls; protoconch almost smooth; teleoconch with fine, irregular wrinkles; apical side with indistinct spiral lines, umbilical side with indistinct incised spiral lines; with radial scaly processes; tuberculate around umbilicus; brownishcorneous; body whorl sharply keeled; aperture rhombic; upper insertion of peristome slightly descending; peristome expanded, reflexed and slightly thickened; umbilicus slightly eccentric, wide, comprising 32–41% of shell diameter, hardly obscured by columellar edge.
</paragraph>
<paragraph id="8BA76A5A8B034E374E88FDD73842FCE0" blockId="46.[189,1399,555,828]" pageId="46" pageNumber="47">
GENITALIA (
<figureCitation id="132376DF8B034E374F74FDD73D93FD9E" box="[321,414,555,581]" captionStart="Figs 29–47" captionStartId="29.[189,241,1366,1392]" captionTargetBox="[189,1399,649,1337]" captionTargetId="figure@29.[189,1399,649,1337]" captionTargetPageId="29" captionText="Figs 29–47. Flagella of species of Landouria from Java. 29. L. winteriana (Pfeiffer, 1842), West Java, Padalarang, Gunung Masigit (neotype, ZMA 376193a). 30. L. rotatoria (Pfeiffer, 1842), West Java, Gunung Salak, 1166 m a.s.l. (neotype, MZB 19205). 31. L. epiplatia (Möllendorff, 1897)?, West Java, Bogor, Gunung Gede (ZMH 79946). 32. L. nusakambangensis sp. nov., Central Java, Nusa Kambangan Island, nature monument Gligir (holotype, ZMA 412298a). 33. L. petrukensis sp. nov., Central Java, Karangbolong, way to Petruk Cave (holotype, MZB 19203). 34. L. tholiformis sp. nov., Yogyakarta, Gunung Kidul, Tepus, near Poktunggal Beach (holotype, MZB 17999). 35. L. abdidalem sp. nov., Yogyakarta, Kulonprogo, Sabrang Kidul (holotype, MZB 17664). 36. L. madurensis sp. nov., East Java, Madura Island, Sampang, Nepa Monkey lowland rainforest (paratype, MZB 19209). 37. L. sewuensis sp. nov., Yogyakarta, Gunung Kidul, Tepus, Mbajo Hill (holotype, MZB 19255). 38. L. sukoliloensis sp. nov., Central Java, Kayen, near Watu Payung (holotype, MZB 19219). 39. L. tonywhitteni sp. nov., Central Java, Taruman Village, near Sendang Sedayu (holotype, MZB 19214). 40. L. nodifera sp. nov., Central Java, Larangan Village, Lanang Spring (holotype, MZB 19227). 41. L. pacitanensis sp. nov., East Java, Pacitan, near Srau Beach (MZB 19231). 42. L. intumescens (Martens, 1867), East Java, Guwoterus (MZB 19207). 43. L. smimensis (Mousson, 1848), East Java, Tosari (ZMA 390881). 44. L. zonifera sp. nov., Yogyakarta, Kulonprogo, Sibolong (holotype, MZB 17727). 45. L. ciliocincta (Möllendorff, 1897), East Java, Malang, Kondangmerak (MZB 19211). 46. L. pakidulan sp. nov., Central Java, Nusa Kambangan Island, nature monument Gligir (holotype, ZMA 417011a). 47. L. menorehensis sp. nov., Yogyakarta, Kulonprogo, Sibolong (holotype, MZB 19201). Scale bar: 1 mm." httpUri="https://zenodo.org/record/3152255/files/figure.png" pageId="46" pageNumber="47">Figs 39</figureCitation>
, 
<figureCitation id="132376DF8B034E374F99FDD73DC1FD9E" box="[428,460,555,581]" captionStart="Figs 63–71" captionStartId="48.[189,241,1668,1694]" captionTargetBox="[189,1394,515,1612]" captionTargetId="figure@48.[189,1398,510,1629]" captionTargetPageId="48" captionText="Figs 63–71. Genitalia of species of Landouria from Java. 63. L. tonywhitteni sp. nov., Central Java, Taruman Village, near Sendang Sedayu (holotype, MZB 19214). 64. L. nodifera sp. nov., Central Java, Larangan Village, Lanang Spring (holotype, MZB 19227). 65. L. pacitanensis sp. nov., East Java, Pacitan, near Srau Beach (MZB 19231). 66. L. intumescens (Martens, 1867), East Java, Guwoterus (MZB 19207). 67. L. smimensis (Mousson, 1848), East Java, Tosari (ZMA 390881). 68. L. zonifera sp. nov., Yogyakarta, Kulonprogo, Sibolong (holotype, MZB 17727). 69. L. ciliocincta (Möllendorff, 1897), East Java, Malang, Kondangmerak (MZB 19211). 70. L. pakidulan sp. nov., Central Java, Nusa Kambangan Island, nature monument Gligir (holotype, ZMA 417011a). 71. L. menorehensis sp. nov., Yogyakarta, Kulonprogo, Sibolong (holotype, MZB 19201). Scale bars: 1 mm." httpUri="https://zenodo.org/record/3152282/files/figure.png" pageId="46" pageNumber="47">63</figureCitation>
; 
<tableCitation id="C69A5FE18B034E374FEFFDD73E38FD9E" box="[474,565,555,581]" captionStart="Table 4" captionStartId="22.[188,257,551,579]" captionTargetBox="[205,1929,661,1732]" captionTargetPageId="22" captionText="Table 4 (continued on next 3 pages). Diagnostic characteristics of the species of Landouria Godwin-Austen, 1918 from Java. Characteristics most important for distinguishing similar taxa are printed in bold." httpUri="http://table.plazi.org/id/DF673AD28B3B4E0F4E89FDDB3F73FDBC" pageId="46" pageNumber="47" tableUuid="DF673AD28B3B4E0F4E89FDDB3F73FDBC">Table 4</tableCitation>
). Atrium short; penis long, consisting of a long, cylindrical distal part covered by an additional sheath, which is connected by muscular tissue to proximal part of epiphallus and a dilated proximal part containing a short penial papilla; without penis coecum; penial retractor runs from diaphragm to distal part of long, narrow epiphallus; flagellum short, broadest at its base, where it is equipped with a globular node, tapering towards its proximal end, not distinctly crenated; vas deferens narrow and long; vagina long; peduncle of bursa copulatrix subdivided into a slightly broader, cylindrical distal part and a narrower, longer proximal part, ending in an elongate oval bursa beside albumen gland. Right ommatophoral retractor runs between penis and vagina.
</paragraph>
</subSubSection>
<subSubSection id="C30239D18B034E374E88FC9F397AFBE3" pageId="46" pageNumber="47" type="discussion">
<paragraph id="8BA76A5A8B034E374E88FC9F3D22FCA6" blockId="46.[189,303,867,893]" box="[189,303,867,893]" pageId="46" pageNumber="47">
<heading id="D0EFDD368B034E374E88FC9F3D22FCA6" bold="true" box="[189,303,867,893]" fontSize="11" level="3" pageId="46" pageNumber="47" reason="3">
<emphasis id="B96CB6488B034E374E88FC9F3D22FCA6" bold="true" box="[189,303,867,893]" pageId="46" pageNumber="47">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BA76A5A8B034E374E88FC6D397AFBE3" blockId="46.[189,1399,913,1081]" pageId="46" pageNumber="47">
<taxonomicName id="4C1811D98B034E374E88FC6D3DABFC70" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[189,422,913,939]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="nodifera" status="sp. nov.">
<emphasis id="B96CB6488B034E374E88FC6D3DABFC70" box="[189,422,913,939]" italics="true" pageId="46" pageNumber="47">Landouria nodifera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B034E374F84FC6E3E1CFC77" box="[433,529,914,940]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
, 
<taxonomicName id="4C1811D98B034E374C2AFC6E3EDFFC70" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[543,722,913,939]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="tonywhitteni" status="sp. nov.">
<emphasis id="B96CB6488B034E374C2AFC6E3EDFFC70" box="[543,722,913,939]" italics="true" pageId="46" pageNumber="47">L. tonywhitteni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B034E374CE9FC6E3F31FC77" box="[732,828,914,940]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
, 
<taxonomicName id="4C1811D98B034E374D7EFC6E380DFC70" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[843,1024,914,939]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="pacitanensis" status="sp. nov.">
<emphasis id="B96CB6488B034E374D7EFC6E380DFC70" box="[843,1024,914,939]" italics="true" pageId="46" pageNumber="47">L. pacitanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B034E374A3CFC6E3864FC77" box="[1033,1129,914,940]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
, 
<taxonomicName id="4C1811D98B034E374A4DFC6E3902FC70" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[1144,1295,914,939]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="sewuensis" status="sp. nov.">
<emphasis id="B96CB6488B034E374A4DFC6E3902FC70" box="[1144,1295,914,939]" italics="true" pageId="46" pageNumber="47">L. sewuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B034E374B2CFC6E397AFC77" box="[1305,1399,914,940]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
and 
<taxonomicName id="4C1811D98B034E374EC5FC493D92FC15" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" baseAuthorityName="Martens" baseAuthorityYear="1867" box="[240,415,949,974]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="intumescens">
<emphasis id="B96CB6488B034E374EC5FC493D92FC15" box="[240,415,949,974]" italics="true" pageId="46" pageNumber="47">L. intumescens</emphasis>
</taxonomicName>
share strongly depressed, brownish-corneous, sharply keeled shells. These species cannot be reliably distinguished by shell characters. 
<taxonomicName id="4C1811D98B034E374D04FC24384AFC29" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[817,1095,984,1010]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="tonywhitteni" status="sp. nov.">
<emphasis id="B96CB6488B034E374D04FC24384AFC29" box="[817,1095,984,1010]" italics="true" pageId="46" pageNumber="47">Landouria tonywhitteni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B034E374A64FC2438A0FC29" box="[1105,1197,984,1010]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
shares a node at the base of the flagellum with 
<taxonomicName id="4C1811D98B034E374C10FC003EA5FBCE" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[549,680,1019,1045]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="nodifera" status="sp. nov.">
<emphasis id="B96CB6488B034E374C10FC003EA5FBCE" box="[549,680,1019,1045]" italics="true" pageId="46" pageNumber="47">L. nodifera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B034E374C85FC073F03FBCE" box="[688,782,1019,1045]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
, from which it differs in the lack of a coecum at the proximal end of the penis and in having a penis that does not become broader towards its proximal end.
</paragraph>
</subSubSection>
<subSubSection id="C30239D18B034E374E88FBA33DD8FB17" pageId="46" pageNumber="47" type="distribution">
<paragraph id="8BA76A5A8B034E374E88FBA33D5BFBA2" blockId="46.[189,342,1119,1145]" box="[189,342,1119,1145]" pageId="46" pageNumber="47">
<heading id="D0EFDD368B034E374E88FBA33D5BFBA2" bold="true" box="[189,342,1119,1145]" fontSize="11" level="3" pageId="46" pageNumber="47" reason="3">
<emphasis id="B96CB6488B034E374E88FBA33D5BFBA2" bold="true" box="[189,342,1119,1145]" pageId="46" pageNumber="47">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BA76A5A8B034E374E88FB723DD8FB17" blockId="46.[189,1398,1166,1228]" pageId="46" pageNumber="47">
<taxonomicName id="4C1811D98B034E374E88FB723DC2FB73" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[189,463,1166,1192]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="tonywhitteni" status="sp. nov.">
<emphasis id="B96CB6488B034E374E88FB723DC2FB73" box="[189,463,1166,1192]" italics="true" pageId="46" pageNumber="47">Landouria tonywhitteni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B034E374FE1FB733E21FB72" box="[468,556,1167,1193]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
is endemic to Sukolilo Karst (Fig. 52), where it occurs sympatrically with 
<taxonomicName id="4C1811D98B034E374E88FB4E3D79FB10" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[189,372,1201,1227]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="sukoliloensis" status="sp. nov.">
<emphasis id="B96CB6488B034E374E88FB4E3D79FB10" box="[189,372,1201,1227]" italics="true" pageId="46" pageNumber="47">L. sukoliloensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B034E374F4EFB4E3DD8FB17" box="[379,469,1202,1228]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>