treatments-xml/data/03/A1/11/03A1112CFFB7FFA116AFD29E89C6D7A0.xml
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<document id="26FC276C1FA4869A1044187C69AC6DF7" ID-CLB-Dataset="45929" ID-DOI="10.5281/zenodo.205547" ID-GBIF-Dataset="ca67f34e-62ea-43e7-a445-8514237a2e8a" ID-ISSN="1175-5326" ID-Zenodo-Dep="205547" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460412808958" checkinUser="plazi" docAuthor="Yamashita, Masahiro, Golani, Daniel &amp; Motomura, Hiroyuki" docDate="2011" docId="03A1112CFFB7FFA116AFD29E89C6D7A0" docLanguage="en" docName="zt03107p058.pdf" docOrigin="Zootaxa 3107" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Upeneus itoui Yamashita, Golani &amp; Motomura, 2011, n. sp." docType="treatment" docVersion="10" lastPageNumber="56" masterDocId="FF986954FFB6FFA81638D2098D6FD343" masterDocTitle="A new species of Upeneus (Perciformes: Mullidae) from southern Japan" masterLastPageNumber="58" masterPageNumber="47" pageNumber="48" updateTime="1698301161424" updateUser="ExternalLinkService">
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<mods:title id="4B0B20E323A9613BE49B075226745405">A new species of Upeneus (Perciformes: Mullidae) from southern Japan</mods:title>
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<mods:name id="6F3D244819E5D54B04DBD88B4A60FA9F" type="personal">
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<mods:namePart id="287E6580A1E00EA8F719F20437E6A3B1">Yamashita, Masahiro</mods:namePart>
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<mods:namePart id="BE38CDABE03370E6AB7CF5910602E8F5">Golani, Daniel</mods:namePart>
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<mods:namePart id="736D68BAD1CC6BA14A940AF288D4F733">Motomura, Hiroyuki</mods:namePart>
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<mods:title id="9EC5F465A939D6ED8750BCC90D5E77AF">Zootaxa</mods:title>
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<mods:date id="40159800AF088C1614A969D7B1053251">2011</mods:date>
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<taxonomicName id="4C08DBB9FFB7FFA916AFD29E8C50D3F2" ID-CoL="7DM3Y" box="[151,319,151,177]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="48" phylum="Chordata" rank="species" species="itoui" status="sp. nov.">
<emphasis id="B97C7C28FFB7FFA916AFD29E8C50D3F2" bold="true" box="[151,319,151,177]" italics="true" pageId="1" pageNumber="48">Upeneus itoui</emphasis>
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<paragraph id="8BB7A03AFFB7FFA916AFD2B38CF5D391" blockId="1.[151,410,151,210]" box="[151,410,186,210]" pageId="1" pageNumber="48">(Figs. 165; Tables 12)</paragraph>
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.
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NSMT-P
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(ex.
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I. 21091),
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, east of Sakinoyama, Kataura, Kasasa, Minamisatsuma, Kagoshima,
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, 31°25ʹ44ʺN, 130°11ʹ49ʺE,
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, set net,
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, M. Itou.
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.
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(
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<collectionCode id="ED1938FFFFB7FFA9148CD3408FB9D222" box="[692,726,329,353]" pageId="1" pageNumber="48">SL</collectionCode>
) from southern
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.
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West coast of Kagoshima Pref. (East
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Sea)
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: the following specimens have the same data as the
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, except for collection dates
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I.
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, 111.7 mm
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,
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;
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I.
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,
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I.
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, date unknown;
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I.
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; AMS I. 45460-001 (ex.
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I. 10251),
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,
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H 7133-01 (ex.
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I. 10252),
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I.
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I.
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I.
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2010-1050 (ex.
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I. 10253),
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399269 (ex.
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I. 10254),
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,
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;
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I.
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;
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I. 10384,
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,
<date id="FFB686FAFFB7FFA914F4D0488E04D11A" box="[716,875,577,602]" pageId="1" pageNumber="48" value="2008-04-14">14 Apr. 2008</date>
;
<collectionCode id="ED1938FFFFB7FFA91544D04B8EBAD119" box="[892,981,578,602]" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="1" pageNumber="48">KAUM</collectionCode>
I.
<quantity id="4CF00DDFFFB7FFA915C3D04889A5D11A" box="[1019,1226,577,601]" metricMagnitude="4" metricUnit="m" metricValue="1.1891106499999997" pageId="1" pageNumber="48" unit="mm" value="1.18911065E7">11891, 106.5 mm</quantity>
<collectionCode id="ED1938FFFFB7FFA912EDD0488997D11A" box="[1237,1272,577,601]" pageId="1" pageNumber="48">SL</collectionCode>
,
<date id="FFB686FAFFB7FFA91331D04888F7D11A" box="[1289,1432,577,601]" pageId="1" pageNumber="48" value="2008-07-05">5 July 2008</date>
,
<collectionCode id="ED1938FFFFB7FFA916AFD06F8D9FD13D" box="[151,240,614,638]" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="1" pageNumber="48">KAUM</collectionCode>
I.
<quantity id="4CF00DDFFFB7FFA9172AD06C8CB7D13E" box="[274,472,613,637]" metricMagnitude="4" metricUnit="m" metricValue="1.32651168" pageId="1" pageNumber="48" unit="mm" value="1.32651168E7">13265, 116.8 mm</quantity>
<collectionCode id="ED1938FFFFB7FFA917E7D06C8F6DD13E" box="[479,514,613,637]" pageId="1" pageNumber="48">SL</collectionCode>
,
<date id="FFB686FAFFB7FFA91436D06C8FC7D13E" box="[526,680,613,637]" pageId="1" pageNumber="48" value="2008-12-20">20 Dec. 2008</date>
;
<collectionCode id="ED1938FFFFB7FFA9148DD06F8E61D13D" box="[693,782,614,638]" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="1" pageNumber="48">KAUM</collectionCode>
I.
<quantity id="4CF00DDFFFB7FFA91508D06C8E99D13E" box="[816,1014,613,637]" metricMagnitude="4" metricUnit="m" metricValue="1.35951149" pageId="1" pageNumber="48" unit="mm" value="1.35951149E7">13595, 114.9 mm</quantity>
<collectionCode id="ED1938FFFFB7FFA915C5D06C894FD13E" box="[1021,1056,613,637]" pageId="1" pageNumber="48">SL</collectionCode>
,
<date id="FFB686FAFFB7FFA91214D06C89A8D13E" box="[1068,1223,613,637]" pageId="1" pageNumber="48" value="2008-11-27">27 Nov. 2008</date>
;
<collectionCode id="ED1938FFFFB7FFA912ECD06F8842D13D" box="[1236,1325,614,638]" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="1" pageNumber="48">KAUM</collectionCode>
I.
<quantity id="4CF00DDFFFB7FFA91377D06C8C62D1E3" metricMagnitude="4" metricUnit="m" metricValue="1.7596124499999999" pageId="1" pageNumber="48" unit="mm" value="1.75961245E7">17596, 124.5 mm</quantity>
<collectionCode id="ED1938FFFFB7FFA91720D0818C54D1E3" box="[280,315,648,672]" pageId="1" pageNumber="48">SL</collectionCode>
,
<collectionCode id="ED1938FFFFB7FFA91773D0808CCBD1E2" box="[331,420,649,673]" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="1" pageNumber="48">KAUM</collectionCode>
I.
<quantity id="4CF00DDFFFB7FFA917F2D0818FF6D1E3" box="[458,665,648,672]" metricMagnitude="4" metricUnit="m" metricValue="1.7597105699999998" pageId="1" pageNumber="48" unit="mm" value="1.75971057E7">17597, 105.7 mm</quantity>
<collectionCode id="ED1938FFFFB7FFA9149CD0818FA8D1E3" box="[676,711,648,672]" pageId="1" pageNumber="48">SL</collectionCode>
,
<date id="FFB686FAFFB7FFA914EFD0818E16D1E3" box="[727,889,648,672]" pageId="1" pageNumber="48" value="2009-03-16">16 Mar. 2009</date>
;
<collectionCode id="ED1938FFFFB7FFA915B0D0808E8ED1E2" box="[904,993,649,673]" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="1" pageNumber="48">KAUM</collectionCode>
I. 17747,
<quantity id="4CF00DDFFFB7FFA91258D08189A7D1E3" box="[1120,1224,648,672]" metricMagnitude="-2" metricUnit="m" metricValue="9.29" pageId="1" pageNumber="48" unit="mm" value="92.9">92.9 mm</quantity>
<collectionCode id="ED1938FFFFB7FFA912ECD0818998D1E3" box="[1236,1271,648,672]" pageId="1" pageNumber="48">SL</collectionCode>
,
<date id="FFB686FAFFB7FFA9133FD08188F8D1E3" box="[1287,1431,648,673]" pageId="1" pageNumber="48" value="2008-04-03">3 Apr. 2008</date>
;
<collectionCode id="ED1938FFFFB7FFA916AFD0A48D9FD186" box="[151,240,685,709]" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="1" pageNumber="48">KAUM</collectionCode>
I.
<quantity id="4CF00DDFFFB7FFA9172CD0A58CB1D187" box="[276,478,684,708]" metricMagnitude="4" metricUnit="m" metricValue="2.03841226" pageId="1" pageNumber="48" unit="mm" value="2.03841226E7">20384, 122.6 mm</quantity>
<collectionCode id="ED1938FFFFB7FFA917DFD0A58F65D187" box="[487,522,684,708]" pageId="1" pageNumber="48">SL</collectionCode>
,
<date id="FFB686FAFFB7FFA9142FD0A58FC6D187" box="[535,681,684,708]" pageId="1" pageNumber="48" value="2008-11-03">3 Nov. 2008</date>
;
<collectionCode id="ED1938FFFFB7FFA9148DD0A48E61D186" box="[693,782,685,709]" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="1" pageNumber="48">KAUM</collectionCode>
I. 25859, 92.0 mm
<collectionCode id="ED1938FFFFB7FFA915CFD0A58975D187" box="[1015,1050,684,708]" pageId="1" pageNumber="48">SL</collectionCode>
,
<date id="FFB686FAFFB7FFA91210D0A589ABD187" box="[1064,1220,684,709]" pageId="1" pageNumber="48" value="2009-05-16">16 May 2009</date>
;
<collectionCode id="ED1938FFFFB7FFA912EAD0A48844D186" box="[1234,1323,685,709]" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="1" pageNumber="48">KAUM</collectionCode>
I.
<quantity id="4CF00DDFFFB7FFA91377D0A58C65D1AB" metricMagnitude="4" metricUnit="m" metricValue="2.87901213" pageId="1" pageNumber="48" unit="mm" value="2.87901213E7">28790, 121.3 mm</quantity>
<collectionCode id="ED1938FFFFB7FFA91729D0D98C5BD1AB" box="[273,308,720,744]" pageId="1" pageNumber="48">SL</collectionCode>
,
<date id="FFB686FAFFB7FFA91779D0D98CB4D1AB" box="[321,475,720,745]" pageId="1" pageNumber="48" value="2010-05-16">16 May 2010</date>
.
<collectionCode id="ED1938FFFFB7FFA917DED0D98F50D1AB" box="[486,575,720,744]" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="1" pageNumber="48">KAUM</collectionCode>
I.
<quantity id="4CF00DDFFFB7FFA9145AD0D98E73D1AB" box="[610,796,720,744]" metricMagnitude="3" metricUnit="m" metricValue="7.2821334" pageId="1" pageNumber="48" unit="mm" value="7282133.4">7282, 133.4 mm</quantity>
<collectionCode id="ED1938FFFFB7FFA9151BD0D98E29D1AB" box="[803,838,720,744]" pageId="1" pageNumber="48">SL</collectionCode>
,
<collectionCode id="ED1938FFFFB7FFA9156BD0D98EC3D1AB" box="[851,940,720,744]" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="1" pageNumber="48">KAUM</collectionCode>
I.
<quantity id="4CF00DDFFFB7FFA915F6D0D989E7D1AB" box="[974,1160,720,744]" metricMagnitude="3" metricUnit="m" metricValue="7.2831217" pageId="1" pageNumber="48" unit="mm" value="7283121.7">7283, 121.7 mm</quantity>
<collectionCode id="ED1938FFFFB7FFA912B7D0D989DDD1AB" box="[1167,1202,720,744]" pageId="1" pageNumber="48">SL</collectionCode>
, northeast of Matsushima Island, Kasasa, Minamisatsuma, 31°25ʹ0 6ʺN, 130°12ʹ32ʺE,
<date id="FFB686FAFFB7FFA915BBD0FA8972D048" box="[899,1053,755,779]" pageId="1" pageNumber="48" value="2008-11-10">10 Nov. 2008</date>
, set net,
<quantity id="4CF00DDFFFB7FFA91246D0FA89D6D048" box="[1150,1209,755,779]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="48" unit="m" value="20.0">20 m</quantity>
, M. Itou;
<collectionCode id="ED1938FFFFB7FFA91310D0FA88EED048" box="[1320,1409,755,779]" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="1" pageNumber="48">KAUM</collectionCode>
I.
<quantity id="4CF00DDFFFB7FFA916AFD11F8C35D06D" box="[151,346,790,814]" metricMagnitude="4" metricUnit="m" metricValue="1.28431148" pageId="1" pageNumber="48" unit="mm" value="1.28431148E7">12843, 114.8 mm</quantity>
<collectionCode id="ED1938FFFFB7FFA91758D11F8CECD06D" box="[352,387,790,814]" pageId="1" pageNumber="48">SL</collectionCode>
, same data as
<collectionCode id="ED1938FFFFB7FFA91419D11E8F15D06C" box="[545,634,791,815]" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="1" pageNumber="48">KAUM</collectionCode>
I. 7282, except date,
<date id="FFB686FAFFB7FFA91554D11F8E90D06D" box="[876,1023,790,814]" pageId="1" pageNumber="48" value="2008-10-13">13 Oct. 2008</date>
;
<collectionCode id="ED1938FFFFB7FFA91232D11E890CD06C" box="[1034,1123,791,815]" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="1" pageNumber="48">KAUM</collectionCode>
I.
<quantity id="4CF00DDFFFB7FFA912BCD11F8827D06D" box="[1156,1352,790,814]" metricMagnitude="4" metricUnit="m" metricValue="1.35941148" pageId="1" pageNumber="48" unit="mm" value="1.35941148E7">13594, 114.8 mm</quantity>
<collectionCode id="ED1938FFFFB7FFA91376D11F881ED06D" box="[1358,1393,790,814]" pageId="1" pageNumber="48">SL</collectionCode>
, off Takasaki-yama, Kataura, Kasasa, Minamisatsuma, 31°26ʹ0 0ʺN, 130°10ʹ0 5ʺE,
<date id="FFB686FAFFB7FFA9123BD13389F5D011" box="[1027,1178,826,850]" pageId="1" pageNumber="48" value="2008-12-11">11 Dec. 2008</date>
, set net,
<quantity id="4CF00DDFFFB7FFA912C3D133885AD011" box="[1275,1333,826,850]" metricMagnitude="1" metricUnit="m" metricValue="3.6" pageId="1" pageNumber="48" unit="m" value="36.0">36 m</quantity>
, M. Itou.
<emphasis id="B97C7C28FFB7FFA916AFD1548C09D035" bold="true" box="[151,358,861,886]" pageId="1" pageNumber="48">Kagoshima Bay:</emphasis>
<collectionCode id="ED1938FFFFB7FFA9174ED1578CA0D035" box="[374,463,862,886]" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="1" pageNumber="48">KAUM</collectionCode>
I.
<quantity id="4CF00DDFFFB7FFA917C2D1548FBDD036" box="[506,722,861,885]" metricMagnitude="4" metricUnit="m" metricValue="1.77141196" pageId="1" pageNumber="48" unit="mm" value="1.77141196E7">17714, 119.6 mm</quantity>
<collectionCode id="ED1938FFFFB7FFA914D9D1548E6BD036" box="[737,772,861,885]" pageId="1" pageNumber="48">SL</collectionCode>
, off Chiringa-jima Island, Ibusuki, Kagoshima,
<collectingCountry id="F31FE0AAFFB7FFA9136DD15788F7D036" box="[1365,1432,862,885]" name="Japan" pageId="1" pageNumber="48">Japan</collectingCountry>
, 31°16ʹ38ʺN, 130°40ʹ18ʺE, set net,
<quantity id="4CF00DDFFFB7FFA91412D1888F07D0DA" box="[554,616,897,921]" metricMagnitude="1" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="48" unit="m" value="25.0">25 m</quantity>
,
<date id="FFB686FAFFB7FFA9144FD1888E6BD0DA" box="[631,772,897,921]" pageId="1" pageNumber="48" value="2009-04-01">1 Apr. 2009</date>
, Orita Fishery; data for the following specimens as
<quantity id="4CF00DDFFFB7FFA91358D18888F3D0DA" box="[1376,1436,897,921]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="48" unit="km" value="1.0">1 km</quantity>
southwest off Kawajiri Fishing Port, Kaimonkawajiri, Ibusuki, 31°10ʹN, 130°32ʹE, set net,
<quantity id="4CF00DDFFFB7FFA912A9D1AD89A3D0FF" box="[1169,1228,932,956]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="48" unit="m" value="40.0">40 m</quantity>
, G. Ogihara and T. Yoshida —
<collectionCode id="ED1938FFFFB7FFA91724D1C18C1AD0A3" box="[284,373,968,992]" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="1" pageNumber="48">KAUM</collectionCode>
I.
<quantity id="4CF00DDFFFB7FFA917AFD1CE8F21D09C" box="[407,590,967,991]" metricMagnitude="3" metricUnit="m" metricValue="9.2111204" pageId="1" pageNumber="48" unit="mm" value="9211120.4">9211, 120.4 mm</quantity>
<collectionCode id="ED1938FFFFB7FFA9146DD1CE8F17D09C" box="[597,632,967,991]" pageId="1" pageNumber="48">SL</collectionCode>
,
<date id="FFB686FAFFB7FFA914BCD1CE8E63D09C" box="[644,780,967,992]" pageId="1" pageNumber="48" value="2008-04-09">9 Apr. 2008</date>
,
<collectionCode id="ED1938FFFFB7FFA9152FD1C18E1FD0A3" box="[791,880,968,992]" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="1" pageNumber="48">KAUM</collectionCode>
I.
<quantity id="4CF00DDFFFB7FFA915ABD1CE8936D09C" box="[915,1113,967,991]" metricMagnitude="4" metricUnit="m" metricValue="1.3715133599999996" pageId="1" pageNumber="48" unit="mm" value="1.37151336E7">13715, 133.6 mm</quantity>
<collectionCode id="ED1938FFFFB7FFA91258D1CE89ECD09C" box="[1120,1155,967,991]" pageId="1" pageNumber="48">SL</collectionCode>
,
<collectionCode id="ED1938FFFFB7FFA912B7D1C18987D0A3" box="[1167,1256,968,992]" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="1" pageNumber="48">KAUM</collectionCode>
I.
<quantity id="4CF00DDFFFB7FFA91332D1CE8DACD740" metricMagnitude="4" metricUnit="m" metricValue="1.3716114799999999" pageId="1" pageNumber="48" unit="mm" value="1.37161148E7">13716, 114.8 mm</quantity>
<collectionCode id="ED1938FFFFB7FFA916F3D1E28D81D740" box="[203,238,1003,1027]" pageId="1" pageNumber="48">SL</collectionCode>
,
<collectionCode id="ED1938FFFFB7FFA916C3D1E58C3BD747" box="[251,340,1004,1028]" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="1" pageNumber="48">KAUM</collectionCode>
I.
<quantity id="4CF00DDFFFB7FFA91740D1E28F2ED740" box="[376,577,1003,1027]" metricMagnitude="4" metricUnit="m" metricValue="1.3717108599999999" pageId="1" pageNumber="48" unit="mm" value="1.37171086E7">13717, 108.6 mm</quantity>
<collectionCode id="ED1938FFFFB7FFA91471D1E28F03D740" box="[585,620,1003,1027]" pageId="1" pageNumber="48">SL</collectionCode>
,
<date id="FFB686FAFFB7FFA91441D1E28E61D740" box="[633,782,1003,1027]" pageId="1" pageNumber="48" value="2009-01-21">21 Jan. 2009</date>
;
<collectionCode id="ED1938FFFFB7FFA91523D1E58E1BD747" box="[795,884,1004,1028]" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="1" pageNumber="48">KAUM</collectionCode>
I.
<quantity id="4CF00DDFFFB7FFA915A0D1E2890FD740" box="[920,1120,1003,1027]" metricMagnitude="4" metricUnit="m" metricValue="1.46621075" pageId="1" pageNumber="48" unit="mm" value="1.46621075E7">14662, 107.5 mm</quantity>
<collectionCode id="ED1938FFFFB7FFA91251D1E289E3D740" box="[1129,1164,1003,1027]" pageId="1" pageNumber="48">SL</collectionCode>
,
<collectionCode id="ED1938FFFFB7FFA912A1D1E5899DD747" box="[1177,1266,1004,1028]" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="1" pageNumber="48">KAUM</collectionCode>
I. 14663,
<quantity id="4CF00DDFFFB7FFA91353D1E28DACD764" metricMagnitude="-2" metricUnit="m" metricValue="8.62" pageId="1" pageNumber="48" unit="mm" value="86.2">86.2 mm</quantity>
<collectionCode id="ED1938FFFFB7FFA916F3D6068D81D764" box="[203,238,1039,1063]" pageId="1" pageNumber="48">SL</collectionCode>
,
<collectionCode id="ED1938FFFFB7FFA916C3D6068C3BD764" box="[251,340,1039,1063]" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="1" pageNumber="48">KAUM</collectionCode>
I.
<quantity id="4CF00DDFFFB7FFA9174FD6068F50D764" box="[375,575,1039,1063]" metricMagnitude="4" metricUnit="m" metricValue="1.46641179" pageId="1" pageNumber="48" unit="mm" value="1.46641179E7">14664, 117.9 mm</quantity>
<collectionCode id="ED1938FFFFB7FFA9147FD6068F05D764" box="[583,618,1039,1063]" pageId="1" pageNumber="48">SL</collectionCode>
,
<collectionCode id="ED1938FFFFB7FFA9144FD6068FBFD764" box="[631,720,1039,1063]" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="1" pageNumber="48">KAUM</collectionCode>
I. 14665,
<quantity id="4CF00DDFFFB7FFA91570D6068EC1D764" box="[840,942,1039,1063]" metricMagnitude="-2" metricUnit="m" metricValue="8.959999999999999" pageId="1" pageNumber="48" unit="mm" value="89.6">89.6 mm</quantity>
<collectionCode id="ED1938FFFFB7FFA9158ED6068EB6D764" box="[950,985,1039,1063]" pageId="1" pageNumber="48">SL</collectionCode>
,
<date id="FFB686FAFFB7FFA915DED6068911D764" box="[998,1150,1039,1063]" pageId="1" pageNumber="48" value="2009-02-18">18 Feb. 2009</date>
;
<collectionCode id="ED1938FFFFB7FFA912B3D606898BD764" box="[1163,1252,1039,1063]" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="1" pageNumber="48">KAUM</collectionCode>
I.
<quantity id="4CF00DDFFFB7FFA91330D6068DACD709" metricMagnitude="4" metricUnit="m" metricValue="1.89551042" pageId="1" pageNumber="48" unit="mm" value="1.89551042E7">18955, 104.2 mm</quantity>
<collectionCode id="ED1938FFFFB7FFA916F3D63B8D81D709" box="[203,238,1074,1098]" pageId="1" pageNumber="48">SL</collectionCode>
,
<date id="FFB686FAFFB7FFA916C3D63B8CFBD709" box="[251,404,1074,1099]" pageId="1" pageNumber="48" value="2009-04-15">15 Apr. 2009</date>
;
<collectionCode id="ED1938FFFFB7FFA91799D63A8C95D708" box="[417,506,1075,1099]" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="1" pageNumber="48">KAUM</collectionCode>
I.
<quantity id="4CF00DDFFFB7FFA91425D63B8F8AD709" box="[541,741,1074,1098]" metricMagnitude="4" metricUnit="m" metricValue="2.26511177" pageId="1" pageNumber="48" unit="mm" value="2.26511177E7">22651, 117.7 mm</quantity>
<collectionCode id="ED1938FFFFB7FFA914D5D63B8E7FD709" box="[749,784,1074,1098]" pageId="1" pageNumber="48">SL</collectionCode>
,
<date id="FFB686FAFFB7FFA91525D63B8ED6D709" box="[797,953,1074,1098]" pageId="1" pageNumber="48" value="2009-11-11">11 Nov. 2009</date>
;
<collectionCode id="ED1938FFFFB7FFA915FFD63A894FD708" box="[967,1056,1075,1099]" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="1" pageNumber="48">KAUM</collectionCode>
I.
<quantity id="4CF00DDFFFB7FFA9127BD63B8863D709" box="[1091,1292,1074,1098]" metricMagnitude="4" metricUnit="m" metricValue="2.75501068" pageId="1" pageNumber="48" unit="mm" value="2.75501068E7">27550, 106.8 mm</quantity>
<collectionCode id="ED1938FFFFB7FFA9132BD63B8859D709" box="[1299,1334,1074,1098]" pageId="1" pageNumber="48">SL</collectionCode>
,
<date id="FFB686FAFFB7FFA9137CD63B8DBCD72D" pageId="1" pageNumber="48" value="2010-03-10">10 Mar. 2010</date>
;
<collectionCode id="ED1938FFFFB7FFA916D8D65F8C56D72D" box="[224,313,1110,1134]" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="1" pageNumber="48">KAUM</collectionCode>
I.
<quantity id="4CF00DDFFFB7FFA91765D65F8F49D72D" box="[349,550,1110,1134]" metricMagnitude="4" metricUnit="m" metricValue="2.7634114199999997" pageId="1" pageNumber="48" unit="mm" value="2.76341142E7">27634, 114.2 mm</quantity>
<collectionCode id="ED1938FFFFB7FFA91416D65F8F3ED72D" box="[558,593,1110,1134]" pageId="1" pageNumber="48">SL</collectionCode>
,
<collectionCode id="ED1938FFFFB7FFA91467D65F8FD7D72D" box="[607,696,1110,1134]" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="1" pageNumber="48">KAUM</collectionCode>
I.
<quantity id="4CF00DDFFFB7FFA914E4D65F8EC9D72D" box="[732,934,1110,1134]" metricMagnitude="4" metricUnit="m" metricValue="2.76351353" pageId="1" pageNumber="48" unit="mm" value="2.76351353E7">27635, 135.3 mm</quantity>
<collectionCode id="ED1938FFFFB7FFA91597D65F8EBDD72D" box="[943,978,1110,1134]" pageId="1" pageNumber="48">SL</collectionCode>
,
<date id="FFB686FAFFB7FFA915D8D65F8911D72D" box="[992,1150,1110,1134]" pageId="1" pageNumber="48" value="2010-03-17">17 Mar. 2010</date>
;
<collectionCode id="ED1938FFFFB7FFA912B3D65F898BD72D" box="[1163,1252,1110,1134]" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="1" pageNumber="48">KAUM</collectionCode>
I.
<quantity id="4CF00DDFFFB7FFA91330D65F8DACD7D2" metricMagnitude="4" metricUnit="m" metricValue="1.35901189" pageId="1" pageNumber="48" unit="mm" value="1.35901189E7">13590, 118.9 mm</quantity>
<collectionCode id="ED1938FFFFB7FFA916F4D6708D80D7D2" box="[204,239,1145,1169]" pageId="1" pageNumber="48">SL</collectionCode>
,
<collectionCode id="ED1938FFFFB7FFA916C5D6738C39D7D1" box="[253,342,1146,1170]" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="1" pageNumber="48">KAUM</collectionCode>
I.
<quantity id="4CF00DDFFFB7FFA91742D6708F2AD7D2" box="[378,581,1145,1169]" metricMagnitude="4" metricUnit="m" metricValue="1.3591104499999997" pageId="1" pageNumber="48" unit="mm" value="1.35911045E7">13591, 104.5 mm</quantity>
<collectionCode id="ED1938FFFFB7FFA91476D6708F1ED7D2" box="[590,625,1145,1169]" pageId="1" pageNumber="48">SL</collectionCode>
,
<collectionCode id="ED1938FFFFB7FFA91446D6738FB8D7D1" box="[638,727,1146,1170]" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="1" pageNumber="48">KAUM</collectionCode>
I. 13592, 119.0 mm
<collectionCode id="ED1938FFFFB7FFA915F6D6708E9ED7D2" box="[974,1009,1145,1169]" pageId="1" pageNumber="48">SL</collectionCode>
,
<collectionCode id="ED1938FFFFB7FFA915C7D6738937D7D1" box="[1023,1112,1146,1170]" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="1" pageNumber="48">KAUM</collectionCode>
I.
<quantity id="4CF00DDFFFB7FFA91245D6708829D7D2" box="[1149,1350,1145,1169]" metricMagnitude="4" metricUnit="m" metricValue="1.3593118499999999" pageId="1" pageNumber="48" unit="mm" value="1.35931185E7">13593, 118.5 mm</quantity>
<collectionCode id="ED1938FFFFB7FFA91377D670881DD7D2" box="[1359,1394,1145,1169]" pageId="1" pageNumber="48">SL</collectionCode>
,
<date id="FFB686FAFFB7FFA913B8D6708C60D7F7" pageId="1" pageNumber="48" value="2008-11-21">21 Nov. 2008</date>
;
<collectionCode id="ED1938FFFFB7FFA91722D6948C1CD7F6" box="[282,371,1181,1205]" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="1" pageNumber="48">KAUM</collectionCode>
I.
<quantity id="4CF00DDFFFB7FFA917ADD6958F34D7F7" box="[405,603,1180,1204]" metricMagnitude="4" metricUnit="m" metricValue="1.98901438" pageId="1" pageNumber="48" unit="mm" value="1.98901438E7">19890, 143.8 mm</quantity>
<collectionCode id="ED1938FFFFB7FFA9145AD6958FEAD7F7" box="[610,645,1180,1204]" pageId="1" pageNumber="48">SL</collectionCode>
,
<date id="FFB686FAFFB7FFA914A9D6958E4AD7F7" box="[657,805,1180,1204]" pageId="1" pageNumber="48" value="2008-10-15">15 Oct. 2008</date>
, staff of Kagoshima Aquarium;
<collectionCode id="ED1938FFFFB7FFA912B6D6948988D7F6" box="[1166,1255,1181,1205]" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="1" pageNumber="48">KAUM</collectionCode>
I.
<quantity id="4CF00DDFFFB7FFA91332D6958DACD79B" metricMagnitude="4" metricUnit="m" metricValue="2.0577116699999998" pageId="1" pageNumber="48" unit="mm" value="2.05771167E7">20577, 116.7 mm</quantity>
<collectionCode id="ED1938FFFFB7FFA916F7D6C98D9DD79B" box="[207,242,1216,1240]" pageId="1" pageNumber="48">SL</collectionCode>
,
<collectionCode id="ED1938FFFFB7FFA9173CD6C88C32D79A" box="[260,349,1217,1241]" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="1" pageNumber="48">KAUM</collectionCode>
I.
<quantity id="4CF00DDFFFB7FFA917BCD6C98F3AD79B" box="[388,597,1216,1240]" metricMagnitude="4" metricUnit="m" metricValue="2.05781074" pageId="1" pageNumber="48" unit="mm" value="2.05781074E7">20578, 107.4 mm</quantity>
<collectionCode id="ED1938FFFFB7FFA9145AD6C98FEAD79B" box="[610,645,1216,1240]" pageId="1" pageNumber="48">SL</collectionCode>
,
<date id="FFB686FAFFB7FFA914AED6C98E52D79B" box="[662,829,1216,1240]" pageId="1" pageNumber="48" value="2008-11-20">20 Nov. 2008</date>
.
<emphasis id="B97C7C28FFB7FFA91574D6C988F3D79B" bold="true" box="[844,1436,1215,1240]" pageId="1" pageNumber="48">East coast of Kagoshima Pref. (Pacific Ocean):</emphasis>
<collectionCode id="ED1938FFFFB7FFA916AFD6ED8D9FD7BF" box="[151,240,1252,1276]" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="1" pageNumber="48">KAUM</collectionCode>
I.
<quantity id="4CF00DDFFFB7FFA9172BD6EA8CAED7B8" box="[275,449,1251,1275]" metricMagnitude="2" metricUnit="m" metricValue="4.771313" pageId="1" pageNumber="48" unit="mm" value="477131.3">477, 131.3 mm</quantity>
<collectionCode id="ED1938FFFFB7FFA917F1D6EA8C83D7B8" box="[457,492,1251,1275]" pageId="1" pageNumber="48">SL</collectionCode>
,
<collectionCode id="ED1938FFFFB7FFA917C1D6ED8F3DD7BF" box="[505,594,1252,1276]" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="1" pageNumber="48">KAUM</collectionCode>
I.
<quantity id="4CF00DDFFFB7FFA9144ED6EA8E4CD7B8" box="[630,803,1251,1275]" metricMagnitude="2" metricUnit="m" metricValue="4.781250999999999" pageId="1" pageNumber="48" unit="mm" value="478125.1">478, 125.1 mm</quantity>
<collectionCode id="ED1938FFFFB7FFA91513D6EA8E21D7B8" box="[811,846,1251,1275]" pageId="1" pageNumber="48">SL</collectionCode>
, Kimotsuki, Uchinoura Bay, 31°20ʹN, 130°04ʹE,
<date id="FFB686FAFFB7FFA913B8D6EA8C63D65C" pageId="1" pageNumber="48" value="2006-02-24">24 Feb. 2006</date>
, set net,
<quantity id="4CF00DDFFFB7FFA9174CD70E8CDED65C" box="[372,433,1287,1311]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="48" unit="m" value="40.0">40 m</quantity>
, K. Nakahata.
<emphasis id="B97C7C28FFB7FFA91466D70F8EB3D65C" bold="true" box="[606,988,1286,1311]" pageId="1" pageNumber="48">Miyazaki Pref. (Pacific Ocean):</emphasis>
<collectionCode id="ED1938FFFFB7FFA915DDD70E895ED65C" box="[997,1073,1287,1311]" httpUri="http://grbio.org/cool/ifay-a7s4" name="Department of Animal Science" pageId="1" pageNumber="48">MUFS</collectionCode>
26238,
<quantity id="4CF00DDFFFB7FFA912A9D70E8998D65C" box="[1169,1271,1287,1311]" metricMagnitude="-2" metricUnit="m" metricValue="9.14" pageId="1" pageNumber="48" unit="mm" value="91.4">91.4 mm</quantity>
<collectionCode id="ED1938FFFFB7FFA91338D70E884CD65C" box="[1280,1315,1287,1311]" pageId="1" pageNumber="48">SL</collectionCode>
, Iorigawa, Kadogawa, Higashiusu,
<date id="FFB686FAFFB7FFA91791D7238F2FD601" box="[425,576,1322,1346]" pageId="1" pageNumber="48" value="2008-09-29">29 Sep. 2008</date>
;
<collectionCode id="ED1938FFFFB7FFA91474D7238FF7D601" box="[588,664,1322,1346]" httpUri="http://grbio.org/cool/ifay-a7s4" name="Department of Animal Science" pageId="1" pageNumber="48">MUFS</collectionCode>
27037, 100.0 mm
<collectionCode id="ED1938FFFFB7FFA91555D7238EFFD601" box="[877,912,1322,1346]" pageId="1" pageNumber="48">SL</collectionCode>
, Kushima,
<date id="FFB686FAFFB7FFA91237D72389C5D601" box="[1039,1194,1322,1346]" pageId="1" pageNumber="48" value="2008-11-15">15 Nov. 2008</date>
;
<collectionCode id="ED1938FFFFB7FFA9128ED723886DD601" box="[1206,1282,1322,1346]" httpUri="http://grbio.org/cool/ifay-a7s4" name="Department of Animal Science" pageId="1" pageNumber="48">MUFS</collectionCode>
28227, 118.0 mm
<collectionCode id="ED1938FFFFB7FFA916F1D7478D83D625" box="[201,236,1358,1382]" pageId="1" pageNumber="48">SL</collectionCode>
, Meitsu, Nango,
<date id="FFB686FAFFB7FFA91795D7478F28D625" box="[429,583,1358,1382]" pageId="1" pageNumber="48" value="2009-03-21">21 Mar. 2009</date>
.
</paragraph>
<paragraph id="8BB7A03AFFB7FFA916FDD77B88EED554" blockId="1.[151,1437,258,2021]" pageId="1" pageNumber="48">
<emphasis id="B97C7C28FFB7FFA916FDD77B8CD1D6C9" bold="true" box="[197,446,1393,1418]" pageId="1" pageNumber="48">Non-type specimens.</emphasis>
<specimenCount id="9D0E6BB3FFB7FFA917F0D7788F0CD6CA" box="[456,611,1393,1417]" pageId="1" pageNumber="48" type="generic">13 specimens</specimenCount>
(93.3115.0 mm
<collectionCode id="ED1938FFFFB7FFA9150BD7788E3AD6CA" box="[819,853,1393,1417]" pageId="1" pageNumber="48">SL</collectionCode>
) from Okinawa-jima Island,
<collectingCountry id="F31FE0AAFFB7FFA91290D77B8983D6CA" box="[1192,1260,1394,1417]" name="Japan" pageId="1" pageNumber="48">Japan</collectingCountry>
: URM-P 8180, 103.0 mm
<collectionCode id="ED1938FFFFB7FFA91729D79C8C5BD6EE" box="[273,308,1429,1453]" pageId="1" pageNumber="48">SL</collectionCode>
, URM-P
<quantity id="4CF00DDFFFB7FFA91798D79C8F37D6EE" box="[416,600,1429,1453]" metricMagnitude="3" metricUnit="m" metricValue="8.1811139" pageId="1" pageNumber="48" unit="mm" value="8181113.9">8181, 113.9 mm</quantity>
<collectionCode id="ED1938FFFFB7FFA91458D79C8FECD6EE" box="[608,643,1429,1453]" pageId="1" pageNumber="48">SL</collectionCode>
, URM-P 8182,
<quantity id="4CF00DDFFFB7FFA9150ED79C8EF5D6EE" box="[822,922,1429,1453]" metricMagnitude="-2" metricUnit="m" metricValue="9.52" pageId="1" pageNumber="48" unit="mm" value="95.2">95.2 mm</quantity>
<collectionCode id="ED1938FFFFB7FFA9159AD79C8EAAD6EE" box="[930,965,1429,1453]" pageId="1" pageNumber="48">SL</collectionCode>
, URM-P 8183,
<quantity id="4CF00DDFFFB7FFA91240D79C89B2D6EE" box="[1144,1245,1429,1453]" metricMagnitude="-2" metricUnit="m" metricValue="9.33" pageId="1" pageNumber="48" unit="mm" value="93.3">93.3 mm</quantity>
<collectionCode id="ED1938FFFFB7FFA912DCD79C8868D6EE" box="[1252,1287,1429,1453]" pageId="1" pageNumber="48">SL</collectionCode>
, Chinen Fish Market, Chinen,
<date id="FFB686FAFFB7FFA9176CD7B18C8BD693" box="[340,484,1464,1488]" pageId="1" pageNumber="48" value="1983-09-09">9 Sept. 1983</date>
; URM-P
<quantity id="4CF00DDFFFB7FFA91477D7B18E69D693" box="[591,774,1464,1488]" metricMagnitude="3" metricUnit="m" metricValue="8.2641086" pageId="1" pageNumber="48" unit="mm" value="8264108.6">8264, 108.6 mm</quantity>
<collectionCode id="ED1938FFFFB7FFA91535D7B18E5FD693" box="[781,816,1464,1488]" pageId="1" pageNumber="48">SL</collectionCode>
, URM-P 8265, 115.0 mm
<collectionCode id="ED1938FFFFB7FFA91260D7B18914D693" box="[1112,1147,1464,1488]" pageId="1" pageNumber="48">SL</collectionCode>
, URM-P
<quantity id="4CF00DDFFFB7FFA912DED7B188F2D693" box="[1254,1437,1464,1488]" metricMagnitude="3" metricUnit="m" metricValue="8.2661104" pageId="1" pageNumber="48" unit="mm" value="8266110.4">8266, 110.4 mm</quantity>
<collectionCode id="ED1938FFFFB7FFA916AFD7D58DD5D6B7" box="[151,186,1500,1524]" pageId="1" pageNumber="48">SL</collectionCode>
, URM-P
<quantity id="4CF00DDFFFB7FFA91710D7D58C8CD6B7" box="[296,483,1500,1524]" metricMagnitude="3" metricUnit="m" metricValue="8.267104499999999" pageId="1" pageNumber="48" unit="mm" value="8267104.5">8267, 104.5 mm</quantity>
<collectionCode id="ED1938FFFFB7FFA917D3D7D58F61D6B7" box="[491,526,1500,1524]" pageId="1" pageNumber="48">SL</collectionCode>
, URM-P 8268,
<quantity id="4CF00DDFFFB7FFA914FCD7D58E46D6B7" box="[708,809,1500,1524]" metricMagnitude="-2" metricUnit="m" metricValue="9.629999999999999" pageId="1" pageNumber="48" unit="mm" value="96.3">96.3 mm</quantity>
<collectionCode id="ED1938FFFFB7FFA91509D7D58E3BD6B7" box="[817,852,1500,1524]" pageId="1" pageNumber="48">SL</collectionCode>
, URM-P
<quantity id="4CF00DDFFFB7FFA915FAD7D58913D6B7" box="[962,1148,1500,1524]" metricMagnitude="3" metricUnit="m" metricValue="8.2691104" pageId="1" pageNumber="48" unit="mm" value="8269110.4">8269, 110.4 mm</quantity>
<collectionCode id="ED1938FFFFB7FFA912BCD7D589C8D6B7" box="[1156,1191,1500,1524]" pageId="1" pageNumber="48">SL</collectionCode>
, URM-P
<quantity id="4CF00DDFFFB7FFA9132ED7D58DACD554" metricMagnitude="3" metricUnit="m" metricValue="8.2701083" pageId="1" pageNumber="48" unit="mm" value="8270108.3">8270, 108.3 mm</quantity>
<collectionCode id="ED1938FFFFB7FFA916F1D7F68D83D554" box="[201,236,1535,1559]" pageId="1" pageNumber="48">SL</collectionCode>
, URM-P
<quantity id="4CF00DDFFFB7FFA91760D7F68F7FD554" box="[344,528,1535,1559]" metricMagnitude="3" metricUnit="m" metricValue="8.271100399999998" pageId="1" pageNumber="48" unit="mm" value="8271100.4">8271, 100.4 mm</quantity>
<collectionCode id="ED1938FFFFB7FFA9142ED7F68F56D554" box="[534,569,1535,1559]" pageId="1" pageNumber="48">SL</collectionCode>
, URM-P
<quantity id="4CF00DDFFFB7FFA9149CD7F68E32D554" box="[676,861,1535,1559]" metricMagnitude="3" metricUnit="m" metricValue="8.2721084" pageId="1" pageNumber="48" unit="mm" value="8272108.4">8272, 108.4 mm</quantity>
<collectionCode id="ED1938FFFFB7FFA9155BD7F68EE9D554" box="[867,902,1535,1559]" pageId="1" pageNumber="48">SL</collectionCode>
, Chinen Fish Market, Chinen,
<date id="FFB686FAFFB7FFA912E7D7F68813D554" box="[1247,1404,1535,1559]" pageId="1" pageNumber="48" value="1983-09-14">14 Sept. 1983</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C312F3B1FFB7FFA916FFD42B8978D439" pageId="1" pageNumber="48" type="diagnosis">
<paragraph id="8BB7A03AFFB7FFA916FFD42B8978D439" blockId="1.[151,1437,258,2021]" pageId="1" pageNumber="48">
<emphasis id="B97C7C28FFB7FFA916FFD42B8C2ED578" bold="true" box="[199,321,1570,1595]" pageId="1" pageNumber="48">Diagnosis.</emphasis>
A species of
<taxonomicName id="4C08DBB9FFB7FFA917D8D42D8F2DD578" box="[480,578,1572,1595]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="48" phylum="Chordata" rank="genus">
<emphasis id="B97C7C28FFB7FFA917D8D42D8F2DD578" box="[480,578,1572,1595]" italics="true" pageId="1" pageNumber="48">Upeneus</emphasis>
</taxonomicName>
with the following combination of characters: dorsal-fin rays VII + 9, first dorsal-fin spine longest; pectoral-fin rays 1315 (usually 14); gill rakers 6 or 7 (mode 6) + 1618 (17) = 2225 (23); pored lateral-line scales 2830 (29); no teeth on ectopterygoids; maximum body depth 20.924.7% SL (mean 22.7%); maximum head depth 18.020.6% SL (19.2%); barbel width 6.07.8% HL (7.0%); barbels white; a distinct reddish brown stripe from tip of snout to caudal-fin base through eye in life and fresh specimens; first dorsal fin with 4 irregular white bands in life, 46 irregular reddish brown bands (retained as dark bands in preserved specimens) and 46 white bands in fresh specimens; upper caudal-fin lobe pale white with 57 reddish brown bands (retained as dark bands in preserved specimens) and 59 irregular pure white bands in life and fresh specimens; lower caudal-fin lobe reddish with 59 short red bands (retained as dark bands in preserved specimens) and 59 short white bands or spots along ventral margin in life and fresh specimens.
</paragraph>
</subSubSection>
<subSubSection id="C312F3B1FFB7FFAF16FFD58C8FF7D254" lastPageId="7" lastPageNumber="54" pageId="1" pageNumber="48" type="description">
<paragraph id="8BB7A03AFFB7FFA916FFD58C8EB6D4A6" blockId="1.[151,1437,258,2021]" pageId="1" pageNumber="48">
<emphasis id="B97C7C28FFB7FFA916FFD58C8C36D4DD" bold="true" box="[199,345,1925,1950]" pageId="1" pageNumber="48">Description.</emphasis>
Counts and measurements, expressed as percentages of SL, are given in Table 1. Frequency distribution of the gill-raker counts are given in
<tableCitation id="C68A9581FFB7FFA914A2D5A08F9ED482" box="[666,753,1961,1985]" captionStart="TABLE 2" captionStartId="4.[151,235,152,173]" captionText="TABLE 2. Counts of gill rakers of Upeneus itoui n. sp. and U. pori." pageId="1" pageNumber="48">Table 2</tableCitation>
. Data for the
<typeStatus id="54B31E98FFB7FFA915B6D5A08E9FD482" box="[910,1008,1961,1985]" pageId="1" pageNumber="48" type="holotype">holotype</typeStatus>
are presented first, followed by
<typeStatus id="54B31E98FFB7FFA9135AD5A38DA8D4A6" pageId="1" pageNumber="48" type="paratype">paratype</typeStatus>
data in parentheses. Osteological characters are based on 7
<typeStatus id="54B31E98FFB7FFA9155ED5C48EBAD4A6" box="[870,981,1997,2021]" pageId="1" pageNumber="48" type="paratype">paratypes</typeStatus>
.
</paragraph>
<caption id="DF77F0B2FFB2FFAC16AFD2918E27D3ED" box="[151,840,152,174]" pageId="4" pageNumber="51">
<paragraph id="8BB7A03AFFB2FFAC16AFD2918E27D3ED" blockId="4.[151,1411,152,350]" box="[151,840,152,174]" pageId="4" pageNumber="51">
<emphasis id="B97C7C28FFB2FFAC16AFD2918C6BD3EE" bold="true" box="[151,260,152,173]" pageId="4" pageNumber="51">TABLE 2.</emphasis>
Counts of gill rakers of
<taxonomicName id="4C08DBB9FFB2FFAC17C4D2908FE7D3ED" box="[508,648,153,174]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="51" phylum="Chordata" rank="species" species="itoui" status="sp. nov.">
<emphasis id="B97C7C28FFB2FFAC17C4D2908FE7D3ED" box="[508,648,153,174]" italics="true" pageId="4" pageNumber="51">Upeneus itoui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24FC153FFB2FFAC14B7D2918FA7D3EE" box="[655,712,152,173]" pageId="4" pageNumber="51" rank="species">
<emphasis id="B97C7C28FFB2FFAC14B7D2918FA7D3EE" bold="true" box="[655,712,152,173]" pageId="4" pageNumber="51">n. sp.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="4C08DBB9FFB2FFAC14C1D2908E2DD3ED" box="[761,834,153,174]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="51" phylum="Chordata" rank="species" species="pori">
<emphasis id="B97C7C28FFB2FFAC14C1D2908E2DD3ED" box="[761,834,153,174]" italics="true" pageId="4" pageNumber="51">U. pori</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BB7A03AFFB2FFAC175DD2C38949D39C" blockId="4.[151,1411,152,350]" box="[357,1062,201,223]" pageId="4" pageNumber="51">Upper limb Lower limb Total</paragraph>
<paragraph id="8BB7A03AFFB2FFAC175DD2FB88ECD24B" blockId="4.[151,1411,152,350]" box="[357,1411,242,264]" pageId="4" pageNumber="51">
6
<date id="FFB686FAFFB2FFAC1798D2FA8F04D24B" box="[416,619,242,264]" pageId="4" pageNumber="51" value="1916-08-07">7 8 16</date>
17 18 19 20 21 22 23 24 25 26 27 28
</paragraph>
<paragraph id="8BB7A03AFFB2FFAC16A7D3158818D21E" blockId="4.[151,1411,152,350]" pageId="4" pageNumber="51">
<taxonomicName id="4C08DBB9FFB2FFAC16A7D3158D87D272" box="[159,232,284,305]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="51" phylum="Chordata" rank="species" species="itoui" status="sp. nov.">
<emphasis id="B97C7C28FFB2FFAC16A7D3158DDFD272" box="[159,176,284,305]" italics="true" pageId="4" pageNumber="51">U</emphasis>
.
<emphasis id="B97C7C28FFB2FFAC1684D3158D87D272" box="[188,232,284,305]" italics="true" pageId="4" pageNumber="51">itoui</emphasis>
</taxonomicName>
<emphasis id="B97C7C28FFB2FFAC16D6D3158C48D272" bold="true" box="[238,295,284,305]" pageId="4" pageNumber="51">
<taxonomicNameLabel id="A24FC153FFB2FFAC16D6D3158C48D272" box="[238,295,284,305]" pageId="4" pageNumber="51" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
37 28H 12 48H
<date id="FFB686FAFFB2FFAC14E5D3128929D272" box="[733,1094,283,305]" pageId="4" pageNumber="51" value="1933-08-05">5 8 33</date>
22H 2
<taxonomicName id="4C08DBB9FFB2FFAC16A7D3408D87D21D" box="[159,232,328,350]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="51" phylum="Chordata" rank="species" species="pori">
<emphasis id="B97C7C28FFB2FFAC16A7D3408DDED21D" box="[159,177,329,350]" italics="true" pageId="4" pageNumber="51">U</emphasis>
.
<emphasis id="B97C7C28FFB2FFAC1686D3408D87D21D" box="[190,232,329,350]" italics="true" pageId="4" pageNumber="51">pori</emphasis>
</taxonomicName>
29 13
<date id="FFB686FAFFB2FFAC14E5D3418E0ED21E" box="[733,865,328,349]" pageId="4" pageNumber="51" value="1927-04-08">4 27 8</date>
<date id="FFB686FAFFB2FFAC15B7D3418862D21E" box="[911,1293,328,349]" pageId="4" pageNumber="51" value="1922-02-03">3 2 22</date>
11 7
</paragraph>
<paragraph id="8BB7A03AFFB2FFAC16AFD3778CD9D2D4" blockId="4.[151,438,382,407]" box="[151,438,382,407]" pageId="4" pageNumber="51">
H Includes count of
<typeStatus id="54B31E98FFB2FFAC176FD3888CDDD2D4" box="[343,434,385,407]" pageId="4" pageNumber="51" type="holotype">holotype</typeStatus>
.
</paragraph>
<caption id="DF77F0B2FFB2FFAC16AFD7A18C29D69E" httpUri="https://zenodo.org/record/205548/files/figure.png" pageId="4" pageNumber="51" targetBox="[191,1396,459,1436]" targetPageId="4">
<paragraph id="8BB7A03AFFB2FFAC16AFD7A18C29D69E" blockId="4.[151,1436,1448,1501]" pageId="4" pageNumber="51">
<emphasis id="B97C7C28FFB2FFAC16AFD7A18C7CD6FD" bold="true" box="[151,275,1448,1471]" pageId="4" pageNumber="51">FIGURE 1.</emphasis>
Holotype of
<taxonomicName id="4C08DBB9FFB2FFAC17A2D7A38F48D6FC" box="[410,551,1450,1471]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="51" phylum="Chordata" rank="species" species="itoui" status="sp. nov.">
<emphasis id="B97C7C28FFB2FFAC17A2D7A38F48D6FC" box="[410,551,1450,1471]" italics="true" pageId="4" pageNumber="51">Upeneus itoui</emphasis>
</taxonomicName>
<emphasis id="B97C7C28FFB2FFAC1416D7A08F07D6FD" bold="true" box="[558,616,1449,1470]" pageId="4" pageNumber="51">
<taxonomicNameLabel id="A24FC153FFB2FFAC1416D7A08F07D6FD" box="[558,616,1449,1470]" pageId="4" pageNumber="51" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(NSMT-P 102554, 122.9 mm SL, Kagoshima, Japan).
<emphasis id="B97C7C28FFB2FFAC12A1D7A089CAD6FD" bold="true" box="[1177,1189,1449,1470]" pageId="4" pageNumber="51">a</emphasis>
, fresh specimen.
<emphasis id="B97C7C28FFB2FFAC1360D7A08809D6FD" bold="true" box="[1368,1382,1449,1470]" pageId="4" pageNumber="51">b</emphasis>
, preserved specimen.
</paragraph>
</caption>
<paragraph id="8BB7A03AFFB2FFAD16FDD40C8EDCD3B4" blockId="4.[151,1437,1541,2033]" lastBlockId="5.[151,1437,151,535]" lastPageId="5" lastPageNumber="52" pageId="4" pageNumber="51">First dorsal-fin spine longest; first dorsal-fin soft ray unbranched, remaining soft rays branched. Anal-fin spine extremely short; first anal-fin soft ray unbranched and spinous, segmented in outer half. Mouth small, ventral and oblique; posterior margin of maxilla reaching (or just short of) a vertical through anterior margin of orbit; posterior margin of maxilla membranous and convex; upper-jaw length less than half head length. A band of small nodular teeth in each jaw, about 3 (13) rows of teeth at front, 5 (25) rows in middle, becoming a single row posteriorly. Vomer with a V-shaped band of villiform teeth, 2 (or 1) rows of teeth at front, 3 (or 2) rows in middle, becoming 1 (or 2) row(s) posteriorly. Palatines with a band of villiform teeth. No teeth on ectopterygoids. Tongue triangular and fused to floor of mouth. Barbels reaching a vertical through preopercular margin. Anterior nostril a short vertically oval opening, anterior to middle of eye; posterior nostril a curved vertical slit, anterior to eye at level of middle of pupil horizontally. Length of longest gill raker on first gill arch subequal to that of longest gill filaments. Opercular spine at level of middle of eye, spine tip reaching (or just short of) opercular margin. Scales finely ctenoid; body scales not extending onto bases of fins, except for second dorsal-fin, anal-fin and caudal-fin bases. First dorsal-fin origin above fourth pored lateral-line scale. Second dorsal-fin origin anterior to anal-fin origin. Posterior tip of depressed pelvic fin extending slightly beyond a vertical through pectoral-fin tip, but not reaching to anus. Formula for configuration of supraneural bones, anterior neural spines, and anterior dorsal pterygiophores 0/ 0/0+1/1+1/1/1/1/1/ or 0/0/0+1/1/1+1/1/1/1/. Vertebrae 10 + 14. Upper series of procurrent caudal-fin rays 8 or 9, lower series 69; principal caudal-fin rays 8 + 7. Swimbladder present.</paragraph>
<paragraph id="8BB7A03AFFB3FFAD16FDD30D8FACD154" blockId="5.[151,1437,151,535]" pageId="5" pageNumber="52">
Color in life [based on aquarium photographs of KAUMI. 28790 (
<figureCitation id="1333BCBFFFB3FFAD158ED30D8E97D25F" box="[950,1016,260,284]" captionStart="FIGURE 2" captionStartId="5.[151,250,1464,1487]" captionTargetBox="[151,1436,589,1442]" captionTargetId="figure@5.[151,1436,589,1443]" captionTargetPageId="5" captionText="FIGURE 2. Underwater photographs of Upeneus itoui n. sp. a b, Approximately 15 cm TL, Morode Beach, Ainan, Ehime, Japan, 33 ° 00 ˏ 15 ʺN, 132 ° 30 ˏ 22 ʺE, 4 5 m depth, 19 Oct. 1999 (photos by J. Shibata). c, Approximately 4 cm TL, Morode Beach, 6 m depth, 13 Nov. 1999 (photo by T. Hirata); U. itoui is the individual in center of the photo, other individuals are U. tragula. d, Aquarium photograph of KAUM I. 28790, paratype, 121.3 mm SL, Kagoshima, Japan, just after capture." httpUri="https://zenodo.org/record/205549/files/figure.png" pageId="5" pageNumber="52">Fig. 2</figureCitation>
d) and underwater photographs (
<figureCitation id="1333BCBFFFB3FFAD135BD30D8DC9D27C" captionStart="FIGURE 2" captionStartId="5.[151,250,1464,1487]" captionTargetBox="[151,1436,589,1442]" captionTargetId="figure@5.[151,1436,589,1443]" captionTargetPageId="5" captionText="FIGURE 2. Underwater photographs of Upeneus itoui n. sp. a b, Approximately 15 cm TL, Morode Beach, Ainan, Ehime, Japan, 33 ° 00 ˏ 15 ʺN, 132 ° 30 ˏ 22 ʺE, 4 5 m depth, 19 Oct. 1999 (photos by J. Shibata). c, Approximately 4 cm TL, Morode Beach, 6 m depth, 13 Nov. 1999 (photo by T. Hirata); U. itoui is the individual in center of the photo, other individuals are U. tragula. d, Aquarium photograph of KAUM I. 28790, paratype, 121.3 mm SL, Kagoshima, Japan, just after capture." httpUri="https://zenodo.org/record/205549/files/figure.png" pageId="5" pageNumber="52">Figs. 2</figureCitation>
ac)]: Head and body grayish dorsally, whitish ventrally; a distinct reddish brown stripe from tip of snout to caudal-fin base through eye. Each scale on dorsal body, above the reddish brown stripe, with blackish posterior margin. No vertical red bar below eye. Barbel white. Dorsal fins semitransparent with white spots, forming 4 irregular white bands on each fin; no blackish blotch on tip of the fins. Pectoral fin transparent, without spots or blotches. Pelvic and anal fins semitransparent with whitish margins. Upper caudal-fin lobe semitransparent, with 5 or 6 dark brown or reddish brown cross bands and irregular whitish bands. Lower caudal-fin lobe reddish brown medially with white bar or spots in a row along fin margin.
</paragraph>
<caption id="DF77F0B2FFB3FFAD16AFD7B1898DD569" httpUri="https://zenodo.org/record/205549/files/figure.png" pageId="5" pageNumber="52" targetBox="[151,1436,589,1442]" targetPageId="5">
<paragraph id="8BB7A03AFFB3FFAD16AFD7B1898DD569" blockId="5.[151,1436,1464,1578]" pageId="5" pageNumber="52">
<emphasis id="B97C7C28FFB3FFAD16AFD7B18C7BD68E" bold="true" box="[151,276,1464,1487]" pageId="5" pageNumber="52">FIGURE 2.</emphasis>
Underwater photographs of
<taxonomicName id="4C08DBB9FFB3FFAD1405D7B08FA3D68D" box="[573,716,1465,1486]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="52" phylum="Chordata" rank="species" species="itoui" status="sp. nov.">
<emphasis id="B97C7C28FFB3FFAD1405D7B08FA3D68D" box="[573,716,1465,1486]" italics="true" pageId="5" pageNumber="52">Upeneus itoui</emphasis>
</taxonomicName>
<emphasis id="B97C7C28FFB3FFAD14ECD7B08E51D68D" bold="true" box="[724,830,1465,1486]" pageId="5" pageNumber="52">
<taxonomicNameLabel id="A24FC153FFB3FFAD14ECD7B08E60D68D" box="[724,783,1465,1486]" pageId="5" pageNumber="52" rank="species">n. sp.</taxonomicNameLabel>
ab
</emphasis>
, Approximately 15 cm TL, Morode Beach, Ainan, Ehime, Japan, 33°00ˏ15ʺN, 132°30ˏ22ʺE, 45 m depth, 19 Oct. 1999 (photos by J. Shibata).
<emphasis id="B97C7C28FFB3FFAD15D0D7DE8E9CD6AF" bold="true" box="[1000,1011,1495,1516]" pageId="5" pageNumber="52">c</emphasis>
, Approximately 4 cm TL, Morode Beach, 6 m depth, 13 Nov. 1999 (photo by T. Hirata);
<taxonomicName id="4C08DBB9FFB3FFAD1449D7FE8FD1D54F" box="[625,702,1527,1548]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="52" phylum="Chordata" rank="species" species="itoui">
<emphasis id="B97C7C28FFB3FFAD1449D7FE8FD1D54F" box="[625,702,1527,1548]" italics="true" pageId="5" pageNumber="52">U. itoui</emphasis>
</taxonomicName>
is the individual in center of the photo, other individuals are
<taxonomicName id="4C08DBB9FFB3FFAD1314D7FE88F9D54F" box="[1324,1430,1526,1548]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="52" phylum="Chordata" rank="species" species="tragula">
<emphasis id="B97C7C28FFB3FFAD1314D7FE88F9D54F" box="[1324,1430,1526,1548]" italics="true" pageId="5" pageNumber="52">U. tragula</emphasis>
</taxonomicName>
.
<emphasis id="B97C7C28FFB3FFAD16AFD41C8DCAD569" bold="true" box="[151,165,1557,1578]" pageId="5" pageNumber="52">d</emphasis>
, Aquarium photograph of KAUMI. 28790, paratype, 121.3 mm SL, Kagoshima, Japan, just after capture.
</paragraph>
</caption>
<paragraph id="8BB7A03AFFB3FFAD16FDD45E8C8FD494" blockId="5.[151,1437,1623,2007]" pageId="5" pageNumber="52">
Color when fresh (based on color photographs of 44
<typeStatus id="54B31E98FFB3FFAD1524D4518E23D533" box="[796,844,1624,1648]" pageId="5" pageNumber="52">type</typeStatus>
specimens, 5 of which were reproduced in
<figureCitation id="1333BCBFFFB3FFAD1303D45E88E9D52C" box="[1339,1414,1623,1647]" captionStart="FIGURE 1" captionStartId="4.[151,250,1448,1471]" captionTargetBox="[191,1396,459,1436]" captionTargetId="figure@4.[191,1396,459,1436]" captionTargetPageId="4" captionText="FIGURE 1. Holotype of Upeneus itoui n. sp. (NSMT-P 102554, 122.9 mm SL, Kagoshima, Japan). a, fresh specimen. b, preserved specimen." httpUri="https://zenodo.org/record/205548/files/figure.png" pageId="5" pageNumber="52">Figs. 1</figureCitation>
a, 3): Head and body yellowish-brown to reddish brown dorsally; lower lateral surface of body, below the reddish longitudinal stripe, with poorly defined, large, reddish blotches (or lower lateral surface becoming uniformly reddish in some
<typeStatus id="54B31E98FFB3FFAD1708D4CD8CF0D59F" box="[304,415,1732,1756]" pageId="5" pageNumber="52" type="paratype">paratypes</typeStatus>
; such reddish coloration may be caused by stress when captured). A distinct longitudinal, reddish-brown stripe, its width subequal to pupil diameter, from tip of snout to caudal-fin base through eye. Barbels white. First dorsal fin with 5 (
<quantity id="4CF00DDFFFB3FFAD17C9D5028F53D460" box="[497,572,1803,1827]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" metricValueMax="1.524" metricValueMin="1.016" pageId="5" pageNumber="52" unit="in" value="5.0" valueMax="6.0" valueMin="4.0">46 in</quantity>
<typeStatus id="54B31E98FFB3FFAD147CD5028FDED460" box="[580,689,1803,1827]" pageId="5" pageNumber="52" type="paratype">paratypes</typeStatus>
) reddish brown bands and 5 (46) irregular white bands. Second dorsal fin with 4 (or 5) reddish brown bands and 4 (or 5) irregular white bands. Pectoral fin semitransparent. Pelvic fin yellowish white (or white). Anal fin white. Upper caudal-fin lobe pale white, with 6 (57) reddish brown bands and 5 (46) irregular pure white bands. Lower caudal-fin lobe reddish, with 8 (59) short, reddish bands, and 8 (5 9) short, white bands (or spots) along ventral margin; white spots along posterior margin of lower lobe. Fresh color variations are given in
<figureCitation id="1333BCBFFFB3FFAD17AFD5B68CB4D494" box="[407,475,1983,2007]" captionStart="FIGURE 3" captionStartId="6.[151,250,1953,1976]" captionTargetBox="[295,1302,193,1931]" captionTargetId="figure@6.[295,1302,193,1932]" captionTargetPageId="6" captionText="FIGURE 3. Variations of body and fin fresh coloration of Upeneus itoui n. sp. a, KAUM I. 17714, paratype, 119.6 mm SL. b, KAUM I. 477, paratype, 131.3 mm SL. c, KAUM I. 13595, paratype, 114.9 mm SL. d, KAUM I. 11891, paratype, 106.5 mm SL." httpUri="https://zenodo.org/record/205550/files/figure.png" pageId="5" pageNumber="52">Fig. 3</figureCitation>
.
</paragraph>
<caption id="DF77F0B2FFB0FFAE16AFD5A888F1D495" httpUri="https://zenodo.org/record/205550/files/figure.png" pageId="6" pageNumber="53" targetBox="[295,1302,193,1931]" targetPageId="6">
<paragraph id="8BB7A03AFFB0FFAE16AFD5A888F1D495" blockId="6.[151,1438,1953,2006]" pageId="6" pageNumber="53">
<emphasis id="B97C7C28FFB0FFAE16AFD5A88C7BD4F4" bold="true" box="[151,276,1953,1976]" pageId="6" pageNumber="53">FIGURE 3.</emphasis>
Variations of body and fin fresh coloration of
<taxonomicName id="4C08DBB9FFB0FFAE14D2D5AA8E1BD4F4" box="[746,884,1954,1976]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="53" phylum="Chordata" rank="species" species="itoui" status="sp. nov.">
<emphasis id="B97C7C28FFB0FFAE14D2D5AA8E1BD4F4" box="[746,884,1954,1976]" italics="true" pageId="6" pageNumber="53">Upeneus itoui</emphasis>
</taxonomicName>
<emphasis id="B97C7C28FFB0FFAE1544D5AB8EA7D4F4" bold="true" box="[892,968,1954,1975]" pageId="6" pageNumber="53">
<taxonomicNameLabel id="A24FC153FFB0FFAE1544D5AB8EDAD4F4" box="[892,949,1954,1975]" pageId="6" pageNumber="53" rank="species">n. sp.</taxonomicNameLabel>
a
</emphasis>
, KAUMI. 17714, paratype, 119.6 mm SL.
<emphasis id="B97C7C28FFB0FFAE13B0D5AB88FAD4F4" bold="true" box="[1416,1429,1954,1975]" pageId="6" pageNumber="53">b</emphasis>
, KAUMI. 477, paratype, 131.3 mm SL.
<emphasis id="B97C7C28FFB0FFAE141DD5C88F5FD495" bold="true" box="[549,560,1985,2006]" pageId="6" pageNumber="53">c</emphasis>
, KAUMI. 13595, paratype, 114.9 mm SL.
<emphasis id="B97C7C28FFB0FFAE15DBD5C88E9ED495" bold="true" box="[995,1009,1985,2006]" pageId="6" pageNumber="53">d</emphasis>
, KAUMI. 11891, paratype, 106.5 mm SL.
</paragraph>
</caption>
<paragraph id="8BB7A03AFFB1FFAF16FDD29E8FF7D254" blockId="7.[151,1436,151,2013]" pageId="7" pageNumber="54">
Color of preserved specimens (
<figureCitation id="1333BCBFFFB1FFAF141ED29E8F08D3EC" box="[550,615,151,175]" captionStart="FIGURE 1" captionStartId="4.[151,250,1448,1471]" captionTargetBox="[191,1396,459,1436]" captionTargetId="figure@4.[191,1396,459,1436]" captionTargetPageId="4" captionText="FIGURE 1. Holotype of Upeneus itoui n. sp. (NSMT-P 102554, 122.9 mm SL, Kagoshima, Japan). a, fresh specimen. b, preserved specimen." httpUri="https://zenodo.org/record/205548/files/figure.png" pageId="7" pageNumber="54">Fig. 1</figureCitation>
b): Head and body dark brown dorsally, pale yellow ventrally, without a longitudinal stripe. Barbels pale. First dorsal fin with 5 (46) grayish bands. Second dorsal fin with 4 (or 5) grayish bands. Pectoral, pelvic and anal fins whitish, without melanophores. Upper caudal-fin lobe with 6 (57) grayish bands; lower lobe with 7 (59) grayish bands.
</paragraph>
</subSubSection>
<subSubSection id="C312F3B1FFB1FFAF16FDD32B8945D2E1" pageId="7" pageNumber="54" type="distribution">
<paragraph id="8BB7A03AFFB1FFAF16FDD32B8945D2E1" blockId="7.[151,1436,151,2013]" pageId="7" pageNumber="54">
<emphasis id="B97C7C28FFB1FFAF16FDD32B8C31D278" bold="true" box="[197,350,290,315]" pageId="7" pageNumber="54">Distribution.</emphasis>
Currently known only from southern
<collectingCountry id="F31FE0AAFFB1FFAF1512D32A8E02D279" box="[810,877,291,314]" name="Japan" pageId="7" pageNumber="54">Japan</collectingCountry>
, including Okinawa-jima Island in the Ryukyu Islands, and Kagoshima, Miyazaki,
<collectingCountry id="F31FE0AAFFB1FFAF1412D34C8F1ED21E" box="[554,625,325,349]" name="India" pageId="7" pageNumber="54">Kochi</collectingCountry>
, and Ehime Prefectures. The last locality was based on underwater photographs reported as “
<taxonomicName id="4C08DBB9FFB1FFAF17A8D36E8C95D23C" box="[400,506,359,383]" class="Actinopterygii" family="Mullidae" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="54" phylum="Chordata" rank="family">Mullidae</taxonomicName>
, indet. Gen. and sp. 1” by
<bibRefCitation id="EF99DDCBFFB1FFAF1515D36E8966D23C" author="Takagi" box="[813,1033,359,384]" pageId="7" pageNumber="54" refString="Takagi, M., Hirata, T., Hirata, S. &amp; Nakata, C. (2010) Fishes of Ainan Ehime. Soufusha Press, Matsuyama, 250 pp." type="book" year="2010">
Takagi
<emphasis id="B97C7C28FFB1FFAF15B9D3608ED4D2C3" box="[897,955,360,384]" italics="true" pageId="7" pageNumber="54">et al.</emphasis>
(2010
</bibRefCitation>
: 65). It occurs on sandy bottoms in depths of
<quantity id="4CF00DDFFFB1FFAF173DD3838C33D2E1" box="[261,348,394,418]" metricMagnitude="1" metricUnit="m" metricValue="2.2" metricValueMax="4.0" metricValueMin="0.4" pageId="7" pageNumber="54" unit="m" value="22.0" valueMax="40.0" valueMin="4.0">440 m</quantity>
, based on underwater photographs and collected specimen data.
</paragraph>
</subSubSection>
<subSubSection id="C312F3B1FFB1FFAF16FDD3A48EEAD149" pageId="7" pageNumber="54" type="etymology">
<paragraph id="8BB7A03AFFB1FFAF16FDD3A48EEAD149" blockId="7.[151,1436,151,2013]" pageId="7" pageNumber="54">
<emphasis id="B97C7C28FFB1FFAF16FDD3A48C24D286" bold="true" box="[197,331,429,453]" pageId="7" pageNumber="54">Etymology.</emphasis>
Named for Mr. Masahide Itou who collected almost all Kagoshima specimens of the new species and kindly made them available to the authors. Sakuya-himeji and Oriental Goatfish are herein proposed as the standard Japanese and English names, respectively, for the species.
</paragraph>
</subSubSection>
<subSubSection id="C312F3B1FFB1FFAF16FDD01C8C46D063" pageId="7" pageNumber="54" type="discussion">
<paragraph id="8BB7A03AFFB1FFAF16FDD01C8C46D063" blockId="7.[151,1436,151,2013]" pageId="7" pageNumber="54">
<emphasis id="B97C7C28FFB1FFAF16FDD01C8C56D16E" bold="true" box="[197,313,533,557]" pageId="7" pageNumber="54">Remarks.</emphasis>
The genus
<taxonomicName id="4C08DBB9FFB1FFAF17F0D01F8F45D16E" box="[456,554,534,557]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="54" phylum="Chordata" rank="genus">
<emphasis id="B97C7C28FFB1FFAF17F0D01F8F45D16E" box="[456,554,534,557]" italics="true" pageId="7" pageNumber="54">Upeneus</emphasis>
</taxonomicName>
is divided into two species groups based on the number of first dorsal-fin spines, i.e., seven or eight spines (
<bibRefCitation id="EF99DDCBFFB1FFAF1429D03E8FDFD10C" author="Lachner" box="[529,688,567,591]" pageId="7" pageNumber="54" refString="Lachner, E. A. (1954) A revision of the goatfish genus Upeneus with descriptions of two new species. Proceeding of the United States National Museum, 103, 497 - 532." type="journal article" year="1954">Lachner 1954</bibRefCitation>
;
<bibRefCitation id="EF99DDCBFFB1FFAF1484D03E8EC5D10C" author="Kim" box="[700,938,567,591]" pageId="7" pageNumber="54" refString="Kim, B. - J. &amp; Nakaya, K. (2002) Upeneus australiae, a new goatfish (Mullidae: Perciformes) from Australia. Ichthyological Research, 49, 128 - 132." type="journal article" year="2002">Kim &amp; Nakaya 2002</bibRefCitation>
).
<bibRefCitation id="EF99DDCBFFB1FFAF1584D03E8980D10C" author="Uiblein" box="[956,1263,567,592]" pageId="7" pageNumber="54" refString="Uiblein, F. &amp; Heemstra, P. C. (2010) A taxonomic review of the western Indian Ocean goatfishes of the genus Upeneus (family Mullidae), with descriptions of four new species. Smithiana, 11, 35 - 71." type="journal article" year="2010">Uiblein &amp; Heemstra (2010)</bibRefCitation>
further divided the eight-spine group into three species groups.
<taxonomicName id="4C08DBB9FFB1FFAF148CD0558E3ED130" box="[692,849,604,627]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="54" phylum="Chordata" rank="species" species="itoui" status="sp. nov.">
<emphasis id="B97C7C28FFB1FFAF148CD0558E3ED130" box="[692,849,604,627]" italics="true" pageId="7" pageNumber="54">Upeneus itoui</emphasis>
</taxonomicName>
<emphasis id="B97C7C28FFB1FFAF1561D0528EF6D130" bold="true" box="[857,921,603,627]" pageId="7" pageNumber="54">
<taxonomicNameLabel id="A24FC153FFB1FFAF1561D0528EF6D130" box="[857,921,603,627]" pageId="7" pageNumber="54" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
belongs to the seven-spine group, which also contains
<taxonomicName id="4C08DBB9FFB1FFAF16C4D0778F0AD1D6" authority="Lachner, 1954" authorityName="Lachner" authorityYear="1954" box="[252,613,637,661]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="54" phylum="Chordata" rank="species" species="asymmetricus">
<emphasis id="B97C7C28FFB1FFAF16C4D0778CD6D1D6" box="[252,441,638,661]" italics="true" pageId="7" pageNumber="54">U. asymmetricus</emphasis>
<bibRefCitation id="EF99DDCBFFB1FFAF17F9D0748F0AD1D6" author="Lachner" box="[449,613,637,661]" pageId="7" pageNumber="54" refString="Lachner, E. A. (1954) A revision of the goatfish genus Upeneus with descriptions of two new species. Proceeding of the United States National Museum, 103, 497 - 532." type="journal article" year="1954">Lachner, 1954</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C08DBB9FFB1FFAF1449D0778E91D1D6" authority="Kim &amp; Nakaya 2002" authorityName="Kim &amp; Nakaya" authorityYear="2002" box="[625,1022,637,661]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="54" phylum="Chordata" rank="species" species="australiae">
<emphasis id="B97C7C28FFB1FFAF1449D0778E6AD1D6" box="[625,773,637,661]" italics="true" pageId="7" pageNumber="54">U. australiae</emphasis>
<bibRefCitation id="EF99DDCBFFB1FFAF1536D0748E91D1D6" author="Kim" box="[782,1022,637,661]" pageId="7" pageNumber="54" refString="Kim, B. - J. &amp; Nakaya, K. (2002) Upeneus australiae, a new goatfish (Mullidae: Perciformes) from Australia. Ichthyological Research, 49, 128 - 132." type="journal article" year="2002">Kim &amp; Nakaya 2002</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C08DBB9FFB1FFAF1231D07788F7D1D6" authority="Randall &amp; Gueze, 1992" authorityName="Randall &amp; Gueze" authorityYear="1992" box="[1033,1432,637,661]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="54" phylum="Chordata" rank="species" species="francisi">
<emphasis id="B97C7C28FFB1FFAF1231D07789EDD1D6" box="[1033,1154,637,661]" italics="true" pageId="7" pageNumber="54">U. francisi</emphasis>
<bibRefCitation id="EF99DDCBFFB1FFAF12B2D07488F7D1D6" author="Randall" box="[1162,1432,637,661]" pageId="7" pageNumber="54" refString="Randall, J. E. &amp; Gueze, P. (1992) Upeneus francisi, a new goat fish (Perciformes: Mullidae) from Norfolk Island and New Zealand. Cybium, 16, 21 - 29." type="journal article" year="1992">Randall &amp; Guézé, 1992</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C08DBB9FFB1FFAF16AFD0A88CCED1F4" authority="Day, 1868" authorityName="Day" authorityYear="1868" box="[151,417,671,696]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="54" phylum="Chordata" rank="species" species="guttatus">
<emphasis id="B97C7C28FFB1FFAF16AFD0A88C7CD1FB" box="[151,275,673,696]" italics="true" pageId="7" pageNumber="54">U. guttatus</emphasis>
(
<bibRefCitation id="EF99DDCBFFB1FFAF1719D0968CF7D1F4" author="Day" box="[289,408,671,695]" pageId="7" pageNumber="54" refString="Day, F. 1868. On some new or imperfectly known fishes of India. Proceedings of the General Meetings for Scientific Business of the Zoological Society of London, 1867, 935 - 942." type="proceedings paper" year="1868">Day, 1868</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C08DBB9FFB1FFAF1794D0A88E69D1F4" authority="Houttuyn, 1782" authorityName="Houttuyn" authorityYear="1782" box="[428,774,671,696]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="54" phylum="Chordata" rank="species" species="japonicus">
<emphasis id="B97C7C28FFB1FFAF1794D0A88F54D1FB" box="[428,571,673,696]" italics="true" pageId="7" pageNumber="54">U. japonicus</emphasis>
(
<bibRefCitation id="EF99DDCBFFB1FFAF1471D0968F91D1F4" author="Houttuyn" box="[585,766,671,695]" pageId="7" pageNumber="54" refString="Houttuyn, M. (1782) Beschryving van eenige Japanese visschen, en andere zee-schepzelen. Verhandelingen der Hollandsche Maatschappij der Wetenschappen, Haarlem, 20, 311 - 350." type="journal article" year="1782">Houttuyn, 1782</bibRefCitation>
)
</taxonomicName>
, and
<taxonomicName id="4C08DBB9FFB1FFAF1578D0A889AAD1F4" authority="Ben-Tuvia &amp; Golani, 1989" authorityName="Ben-Tuvia &amp; Golani" authorityYear="1989" box="[832,1221,671,696]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="54" phylum="Chordata" rank="species" species="pori">
<emphasis id="B97C7C28FFB1FFAF1578D0A88E3BD1FB" box="[832,852,673,696]" italics="true" pageId="7" pageNumber="54">U</emphasis>
.
<emphasis id="B97C7C28FFB1FFAF155AD0A88EFED1FB" box="[866,913,673,696]" italics="true" pageId="7" pageNumber="54">pori</emphasis>
<bibRefCitation id="EF99DDCBFFB1FFAF15AFD09689AAD1F4" author="Ben-Tuvia" box="[919,1221,671,695]" pageId="7" pageNumber="54" refString="Ben-Tuvia, A. &amp; Golani, D. (1989) A new species of the goatfish (Mullidae) of the genus Upeneus from the Red Sea and the eastern Mediterranean. Israel Journal of Zoology, 36, 103 - 112." type="journal article" year="1989">Ben-Tuvia &amp; Golani, 1989</bibRefCitation>
</taxonomicName>
. Although
<bibRefCitation id="EF99DDCBFFB1FFAF1378D0968D8BD199" author="Lachner" pageId="7" pageNumber="54" refString="Lachner, E. A. (1954) A revision of the goatfish genus Upeneus with descriptions of two new species. Proceeding of the United States National Museum, 103, 497 - 532." type="journal article" year="1954">Lachner (1954)</bibRefCitation>
,
<bibRefCitation id="EF99DDCBFFB1FFAF16D7D0CB8F6CD199" author="Randall" box="[239,515,706,730]" pageId="7" pageNumber="54" refString="Randall, J. E. &amp; Gueze, P. (1992) Upeneus francisi, a new goat fish (Perciformes: Mullidae) from Norfolk Island and New Zealand. Cybium, 16, 21 - 29." type="journal article" year="1992">Randall &amp; Guézé (1992)</bibRefCitation>
and
<bibRefCitation id="EF99DDCBFFB1FFAF1400D0CB8E02D199" author="Uiblein" box="[568,877,706,730]" pageId="7" pageNumber="54" refString="Uiblein, F. &amp; Heemstra, P. C. (2010) A taxonomic review of the western Indian Ocean goatfishes of the genus Upeneus (family Mullidae), with descriptions of four new species. Smithiana, 11, 35 - 71." type="journal article" year="2010">Uiblein &amp; Heemstra (2010)</bibRefCitation>
included
<taxonomicName id="4C08DBB9FFB1FFAF15E2D0CD89A4D199" authority="Poey, 1852" authorityName="Poey" authorityYear="1852" box="[986,1227,706,731]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="54" phylum="Chordata" rank="species" species="parvus">
<emphasis id="B97C7C28FFB1FFAF15E2D0CD8927D198" box="[986,1096,708,731]" italics="true" pageId="7" pageNumber="54">U. parvus</emphasis>
Poey,
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</taxonomicName>
in the seven-spine group,
<bibRefCitation id="EF99DDCBFFB1FFAF16D0D0EC8C82D1BE" author="Kim" box="[232,493,741,765]" pageId="7" pageNumber="54" refString="Kim, B. - J. &amp; Nakaya, K. (2002) Upeneus australiae, a new goatfish (Mullidae: Perciformes) from Australia. Ichthyological Research, 49, 128 - 132." type="journal article" year="2002">Kim &amp; Nakaya (2002)</bibRefCitation>
regarded the species as a member of the eight-spine group based on radiographic examination.
</paragraph>
</subSubSection>
<subSubSection id="C312F3B1FFB1FFA116FDD12589C6D7A0" lastPageId="9" lastPageNumber="56" pageId="7" pageNumber="54" type="reference_group">
<paragraph id="8BB7A03AFFB1FFAF16FDD1258FC0D7DE" blockId="7.[151,1436,151,2013]" pageId="7" pageNumber="54">
<taxonomicName id="4C08DBB9FFB1FFAF16FDD1258C0ED000" ID-CoL="7DM3Y" box="[197,353,812,835]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="54" phylum="Chordata" rank="species" species="itoui">
<emphasis id="B97C7C28FFB1FFAF16FDD1258C0ED000" box="[197,353,812,835]" italics="true" pageId="7" pageNumber="54">Upeneus itoui</emphasis>
</taxonomicName>
can be easily distinguished from
<taxonomicName id="4C08DBB9FFB1FFAF14E2D1258E39D000" ID-CoL="7DLTN" box="[730,854,812,835]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="54" phylum="Chordata" rank="species" species="guttatus">
<emphasis id="B97C7C28FFB1FFAF14E2D1258E39D000" box="[730,854,812,835]" italics="true" pageId="7" pageNumber="54">U. guttatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C08DBB9FFB1FFAF15B5D1258972D000" ID-CoL="7DLS9" box="[909,1053,812,835]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="54" phylum="Chordata" rank="species" species="japonicus">
<emphasis id="B97C7C28FFB1FFAF15B5D1258972D000" box="[909,1053,812,835]" italics="true" pageId="7" pageNumber="54">U. japonicus</emphasis>
</taxonomicName>
by lacking teeth on the ectopterygoids (
<bibRefCitation id="EF99DDCBFFB1FFAF16DAD1448F22D026" author="Kim" box="[226,589,845,869]" pageId="7" pageNumber="54" refString="Kim, B. - J. &amp; Nakaya, K. (2002) Upeneus australiae, a new goatfish (Mullidae: Perciformes) from Australia. Ichthyological Research, 49, 128 - 132." type="journal article" year="2002">Kim &amp; Nakaya 2002; this study</bibRefCitation>
). The new species differs from
<taxonomicName id="4C08DBB9FFB1FFAF1596D1478905D026" ID-CoL="7DLTB" box="[942,1130,846,869]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="54" phylum="Chordata" rank="species" species="asymmetricus">
<emphasis id="B97C7C28FFB1FFAF1596D1478905D026" box="[942,1130,846,869]" italics="true" pageId="7" pageNumber="54">U. asymmetricus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C08DBB9FFB1FFAF1298D1478877D026" ID-CoL="7DLTQ" box="[1184,1304,845,869]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="54" phylum="Chordata" rank="species" species="francisi">
<emphasis id="B97C7C28FFB1FFAF1298D1478877D026" box="[1184,1304,845,869]" italics="true" pageId="7" pageNumber="54">U. francisi</emphasis>
</taxonomicName>
in having a lower number of total gill rakers [2225 vs.
<quantity id="4CF00DDFFFB1FFAF14B0D1798F84D0CB" box="[648,747,880,904]" metricMagnitude="-1" metricUnit="m" metricValue="7.239" metricValueMax="7.366" metricValueMin="7.112" pageId="7" pageNumber="54" unit="in" value="28.5" valueMax="29.0" valueMin="28.0">2829 in</quantity>
<taxonomicName id="4C08DBB9FFB1FFAF14CAD1788EC1D0CB" ID-CoL="7DLTB" box="[754,942,881,904]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="54" phylum="Chordata" rank="species" species="asymmetricus">
<emphasis id="B97C7C28FFB1FFAF14CAD1788EC1D0CB" box="[754,942,881,904]" italics="true" pageId="7" pageNumber="54">U. asymmetricus</emphasis>
</taxonomicName>
and
<quantity id="4CF00DDFFFB1FFAF15DDD1798927D0CB" box="[997,1096,880,904]" metricMagnitude="-1" metricUnit="m" metricValue="8.001" metricValueMax="8.382" metricValueMin="7.62" pageId="7" pageNumber="54" unit="in" value="31.5" valueMax="33.0" valueMin="30.0">3033 in</quantity>
<taxonomicName id="4C08DBB9FFB1FFAF1277D17889A8D0CB" ID-CoL="7DLTQ" box="[1103,1223,880,904]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="54" phylum="Chordata" rank="species" species="francisi">
<emphasis id="B97C7C28FFB1FFAF1277D17889A8D0CB" box="[1103,1223,880,904]" italics="true" pageId="7" pageNumber="54">U. francisi</emphasis>
</taxonomicName>
; based on
<bibRefCitation id="EF99DDCBFFB1FFAF1378D1798D8ED0E9" author="Lachner" pageId="7" pageNumber="54" refString="Lachner, E. A. (1954) A revision of the goatfish genus Upeneus with descriptions of two new species. Proceeding of the United States National Museum, 103, 497 - 532." type="journal article" year="1954">Lachner (1954)</bibRefCitation>
and
<bibRefCitation id="EF99DDCBFFB1FFAF1723D19B8F5CD0E9" author="Randall" box="[283,563,914,938]" pageId="7" pageNumber="54" refString="Randall, J. E. &amp; Gueze, P. (1992) Upeneus francisi, a new goat fish (Perciformes: Mullidae) from Norfolk Island and New Zealand. Cybium, 16, 21 - 29." type="journal article" year="1992">Randall &amp; Guézé (1992)</bibRefCitation>
respectively]. In addition,
<taxonomicName id="4C08DBB9FFB1FFAF155ED19D8ED2D0E8" ID-CoL="7DM3Y" box="[870,957,916,939]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="54" phylum="Chordata" rank="species" species="itoui">
<emphasis id="B97C7C28FFB1FFAF155ED19D8ED2D0E8" box="[870,957,916,939]" italics="true" pageId="7" pageNumber="54">U. itoui</emphasis>
</taxonomicName>
and
<taxonomicName id="4C08DBB9FFB1FFAF15CED19D89DBD0E8" ID-CoL="7DLTB" box="[1014,1204,916,939]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="54" phylum="Chordata" rank="species" species="asymmetricus">
<emphasis id="B97C7C28FFB1FFAF15CED19D89DBD0E8" box="[1014,1204,916,939]" italics="true" pageId="7" pageNumber="54">U. asymmetricus</emphasis>
</taxonomicName>
can be separated by coloration. The latter has a dark brown saddle-like spot on the caudal peduncle just posterior to the second dorsal fin (vs. spot absent in
<taxonomicName id="4C08DBB9FFB1FFAF17A5D1D08C99D0B3" ID-CoL="7DM3Y" box="[413,502,985,1008]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="54" phylum="Chordata" rank="species" species="itoui">
<emphasis id="B97C7C28FFB1FFAF17A5D1D08C99D0B3" box="[413,502,985,1008]" italics="true" pageId="7" pageNumber="54">U. itoui</emphasis>
</taxonomicName>
) and three dark bands on the caudal-fin upper lobe (vs. 57 bands) (data for
<taxonomicName id="4C08DBB9FFB1FFAF13B9D1D08C5ED750" ID-CoL="7DLTB" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="54" phylum="Chordata" rank="species" species="asymmetricus">
<emphasis id="B97C7C28FFB1FFAF13B9D1D08C5ED750" italics="true" pageId="7" pageNumber="54">U. asymmetricus</emphasis>
</taxonomicName>
from
<bibRefCitation id="EF99DDCBFFB1FFAF174CD1F38F62D751" author="Randall" box="[372,525,1018,1042]" pageId="7" pageNumber="54" refString="Randall, J. E. (2001) Mullidae. Goatfishes (surmullets). In: Carpenter, K. &amp; Niem, V. (Eds.), FAO species identification guide for fishery purposes. The living marine resources of the western central Pacific, Vol. 5. Bony fishes part 3 (Menidae to Pomacentridae), FAO, Rome, pp. 3175 - 3200." type="journal article" year="2001">Randall 2001</bibRefCitation>
). Although most meristic characters of
<taxonomicName id="4C08DBB9FFB1FFAF15FBD1F58978D750" ID-CoL="7DM3Y" box="[963,1047,1020,1043]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="54" phylum="Chordata" rank="species" species="itoui">
<emphasis id="B97C7C28FFB1FFAF15FBD1F58978D750" box="[963,1047,1020,1043]" italics="true" pageId="7" pageNumber="54">U. itoui</emphasis>
</taxonomicName>
agreed with those of an Australian species,
<taxonomicName id="4C08DBB9FFB1FFAF16C4D6178CFAD776" ID-CoL="7DLTL" box="[252,405,1053,1077]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="54" phylum="Chordata" rank="species" species="australiae">
<emphasis id="B97C7C28FFB1FFAF16C4D6178CFAD776" box="[252,405,1053,1077]" italics="true" pageId="7" pageNumber="54">U. australiae</emphasis>
</taxonomicName>
, the former differs in having 59 narrow grayish bands on the lower caudal-fin lobe, narrower than bands on the upper lobe, in preserved specimens [
<figureCitation id="1333BCBFFFB1FFAF155DD6498EC8D71B" box="[869,935,1088,1112]" captionStart="FIGURE 1" captionStartId="4.[151,250,1448,1471]" captionTargetBox="[191,1396,459,1436]" captionTargetId="figure@4.[191,1396,459,1436]" captionTargetPageId="4" captionText="FIGURE 1. Holotype of Upeneus itoui n. sp. (NSMT-P 102554, 122.9 mm SL, Kagoshima, Japan). a, fresh specimen. b, preserved specimen." httpUri="https://zenodo.org/record/205548/files/figure.png" pageId="7" pageNumber="54">Fig. 1</figureCitation>
b: vs. 6 broad black bands, widths more than twice of those of bands on the upper lobe, and posterior bands (fifth and sixth bands) forming subcircular blotches in
<taxonomicName id="4C08DBB9FFB1FFAF168CD68F8C27D7DE" ID-CoL="7DLTL" box="[180,328,1157,1181]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="54" phylum="Chordata" rank="species" species="australiae">
<emphasis id="B97C7C28FFB1FFAF168CD68F8C27D7DE" box="[180,328,1157,1181]" italics="true" pageId="7" pageNumber="54">U. australiae</emphasis>
</taxonomicName>
;
<bibRefCitation id="EF99DDCBFFB1FFAF176ED68C8F2BD7DE" author="Kim" box="[342,580,1157,1181]" pageId="7" pageNumber="54" refString="Kim, B. - J. &amp; Nakaya, K. (2002) Upeneus australiae, a new goatfish (Mullidae: Perciformes) from Australia. Ichthyological Research, 49, 128 - 132." type="journal article" year="2002">Kim &amp; Nakaya 2002</bibRefCitation>
: fig. 3A].
</paragraph>
<paragraph id="8BB7A03AFFB1FFAF16FDD6A08E61D6F1" blockId="7.[151,1436,151,2013]" pageId="7" pageNumber="54">
<taxonomicName id="4C08DBB9FFB1FFAF16FDD6A08C0DD783" ID-CoL="7DM3Y" box="[197,354,1193,1216]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="54" phylum="Chordata" rank="species" species="itoui">
<emphasis id="B97C7C28FFB1FFAF16FDD6A08C0DD783" box="[197,354,1193,1216]" italics="true" pageId="7" pageNumber="54">Upeneus itoui</emphasis>
</taxonomicName>
is most similar to a western Indian Ocean species,
<taxonomicName id="4C08DBB9FFB1FFAF1596D6A0896ED783" ID-CoL="7DM4K" box="[942,1025,1193,1216]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="54" phylum="Chordata" rank="species" species="pori">
<emphasis id="B97C7C28FFB1FFAF1596D6A08EADD783" box="[942,962,1193,1216]" italics="true" pageId="7" pageNumber="54">U</emphasis>
.
<emphasis id="B97C7C28FFB1FFAF15EAD6A0896ED783" box="[978,1025,1193,1216]" italics="true" pageId="7" pageNumber="54">pori</emphasis>
</taxonomicName>
, in overall body appearance and fin coloration, and the two allopatric species are difficult to distinguish from each other at a glance. However, the body of
<taxonomicName id="4C08DBB9FFB1FFAF168DD6E78C64D646" ID-CoL="7DM3Y" box="[181,267,1262,1285]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="54" phylum="Chordata" rank="species" species="itoui">
<emphasis id="B97C7C28FFB1FFAF168DD6E78C64D646" box="[181,267,1262,1285]" italics="true" pageId="7" pageNumber="54">U. itoui</emphasis>
</taxonomicName>
tends to be shallower than that of
<taxonomicName id="4C08DBB9FFB1FFAF14B5D6E78FB0D646" ID-CoL="7DM4K" box="[653,735,1262,1285]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="54" phylum="Chordata" rank="species" species="pori">
<emphasis id="B97C7C28FFB1FFAF14B5D6E78FB0D646" box="[653,735,1262,1285]" italics="true" pageId="7" pageNumber="54">U. pori</emphasis>
</taxonomicName>
[body depth 20.924.7% of SL (mean 22.7%) vs. 21.125.2% (23.5%) in
<taxonomicName id="4C08DBB9FFB1FFAF172DD7188C09D66B" ID-CoL="7DM4K" box="[277,358,1297,1320]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="54" phylum="Chordata" rank="species" species="pori">
<emphasis id="B97C7C28FFB1FFAF172DD7188C09D66B" box="[277,358,1297,1320]" italics="true" pageId="7" pageNumber="54">U. pori</emphasis>
</taxonomicName>
,
<figureCitation id="1333BCBFFFB1FFAF174BD7068CD9D664" box="[371,438,1295,1319]" captionStart="FIGURE 4" captionStartId="8.[151,250,1209,1232]" captionTargetBox="[184,1394,211,1184]" captionTargetId="figure@8.[172,1416,193,1191]" captionTargetPageId="8" captionText="FIGURE 4. Relationships of maximum body depth to standard length in Upeneus itoui n. sp. [open symbols (star = holotype)] and U. pori (closed triangles)." httpUri="https://zenodo.org/record/205551/files/figure.png" pageId="7" pageNumber="54">Fig. 4</figureCitation>
, Table 1; head depth 18.020.6% of SL (mean 19.2%) vs. 18.421.7% (20.0%), Table 1]. The barbel width at its base of
<taxonomicName id="4C08DBB9FFB1FFAF17CDD73D8F25D608" ID-CoL="7DM3Y" box="[501,586,1332,1355]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="54" phylum="Chordata" rank="species" species="itoui">
<emphasis id="B97C7C28FFB1FFAF17CDD73D8F25D608" box="[501,586,1332,1355]" italics="true" pageId="7" pageNumber="54">U. itoui</emphasis>
</taxonomicName>
also tends to be narrower than that of
<taxonomicName id="4C08DBB9FFB1FFAF15C6D73D893FD608" ID-CoL="7DM4K" box="[1022,1104,1332,1355]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="54" phylum="Chordata" rank="species" species="pori">
<emphasis id="B97C7C28FFB1FFAF15C6D73D893FD608" box="[1022,1104,1332,1355]" italics="true" pageId="7" pageNumber="54">U. pori</emphasis>
</taxonomicName>
(
<figureCitation id="1333BCBFFFB1FFAF1258D73B89CAD609" box="[1120,1189,1330,1354]" captionStart="FIGURE 5" captionStartId="8.[151,250,1967,1990]" captionTargetBox="[153,1430,1405,1920]" captionTargetId="figure@8.[151,1436,1312,1946]" captionTargetPageId="8" captionText="FIGURE 5. Relationships of barbel width to standard length (a) and barbel length (b) in Upeneus itoui n. sp. [open symbols (star = holotype)] and U. pori (closed triangles)." httpUri="https://zenodo.org/record/205552/files/figure.png" pageId="7" pageNumber="54">Fig. 5</figureCitation>
). Although the barbel width of the two species when young mostly overlapped [6.07.8% of HL (mean 7.0%) in
<taxonomicName id="4C08DBB9FFB1FFAF1290D75F8990D62E" ID-CoL="7DM3Y" box="[1192,1279,1366,1389]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="54" phylum="Chordata" rank="species" species="itoui">
<emphasis id="B97C7C28FFB1FFAF1290D75F8990D62E" box="[1192,1279,1366,1389]" italics="true" pageId="7" pageNumber="54">U. itoui</emphasis>
</taxonomicName>
vs. 6.49.7% (7.7%) in
<taxonomicName id="4C08DBB9FFB1FFAF173FD7708C37D6D3" ID-CoL="7DM4K" box="[263,344,1401,1424]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="54" phylum="Chordata" rank="species" species="pori">
<emphasis id="B97C7C28FFB1FFAF173FD7708C37D6D3" box="[263,344,1401,1424]" italics="true" pageId="7" pageNumber="54">U. pori</emphasis>
</taxonomicName>
], it became distinct with growth [in over
<quantity id="4CF00DDFFFB1FFAF151FD77E8EECD6CC" box="[807,899,1399,1423]" metricMagnitude="-1" metricUnit="m" metricValue="1.1" pageId="7" pageNumber="54" unit="mm" value="110.0">110 mm</quantity>
SL specimens, 6.07.8% of HL (mean 7.0%) in
<taxonomicName id="4C08DBB9FFB1FFAF16AFD7958D83D6F0" ID-CoL="7DM3Y" box="[151,236,1436,1459]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="54" phylum="Chordata" rank="species" species="itoui">
<emphasis id="B97C7C28FFB1FFAF16AFD7958D83D6F0" box="[151,236,1436,1459]" italics="true" pageId="7" pageNumber="54">U. itoui</emphasis>
</taxonomicName>
vs. 7.49.7% (8.3%) in
<taxonomicName id="4C08DBB9FFB1FFAF17C5D7958F21D6F0" ID-CoL="7DM4K" box="[509,590,1436,1459]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="54" phylum="Chordata" rank="species" species="pori">
<emphasis id="B97C7C28FFB1FFAF17C5D7958F21D6F0" box="[509,590,1436,1459]" italics="true" pageId="7" pageNumber="54">U. pori</emphasis>
</taxonomicName>
; also see
<figureCitation id="1333BCBFFFB1FFAF1484D7938E6FD6F1" box="[700,768,1434,1458]" captionStart="FIGURE 5" captionStartId="8.[151,250,1967,1990]" captionTargetBox="[153,1430,1405,1920]" captionTargetId="figure@8.[151,1436,1312,1946]" captionTargetPageId="8" captionText="FIGURE 5. Relationships of barbel width to standard length (a) and barbel length (b) in Upeneus itoui n. sp. [open symbols (star = holotype)] and U. pori (closed triangles)." httpUri="https://zenodo.org/record/205552/files/figure.png" pageId="7" pageNumber="54">Fig. 5</figureCitation>
].
</paragraph>
<paragraph id="8BB7A03AFFB1FFAF16FDD7B48CCDD57E" blockId="7.[151,1436,151,2013]" pageId="7" pageNumber="54">
The number of gill rakers also distinguishes
<taxonomicName id="4C08DBB9FFB1FFAF1481D7B78E60D696" ID-CoL="7DM3Y" box="[697,783,1470,1493]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="54" phylum="Chordata" rank="species" species="itoui">
<emphasis id="B97C7C28FFB1FFAF1481D7B78E60D696" box="[697,783,1470,1493]" italics="true" pageId="7" pageNumber="54">U. itoui</emphasis>
</taxonomicName>
(67 on upper limb, 1618 on lower limb,
<quantity id="4CF00DDFFFB1FFAF12CFD7B48835D696" box="[1271,1370,1469,1493]" metricMagnitude="-1" metricUnit="m" metricValue="5.968999999999999" metricValueMax="6.35" metricValueMin="5.588" pageId="7" pageNumber="54" unit="in" value="23.5" valueMax="25.0" valueMin="22.0">2225 in</quantity>
total) from
<taxonomicName id="4C08DBB9FFB1FFAF16EDD7E88C49D6BB" ID-CoL="7DM4K" box="[213,294,1505,1528]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="54" phylum="Chordata" rank="species" species="pori">
<emphasis id="B97C7C28FFB1FFAF16EDD7E88C49D6BB" box="[213,294,1505,1528]" italics="true" pageId="7" pageNumber="54">U. pori</emphasis>
</taxonomicName>
(78, 1820, 2527, respectively; Tables 12). Although the number of gill rakers of the two species overlaps in 25, specimens with 25 rakers of the two species can be easily separated by the above mentioned morphometrics (
<figureCitation id="1333BCBFFFB1FFAF1768D42C8CFBD57E" box="[336,404,1573,1597]" captionStart="FIGURE 6" captionStartId="9.[151,250,1022,1045]" captionTargetBox="[317,1285,201,995]" captionTargetId="figure@9.[294,1293,193,1001]" captionTargetPageId="9" captionText="FIGURE 6. Relationships of maximum body depth plus barbel width (% of standard length) to number of gill rakers in Upeneus itoui n. sp. [open symbols (star = holotype)] and U. pori (closed triangles)." httpUri="https://zenodo.org/record/205553/files/figure.png" pageId="7" pageNumber="54">Fig. 6</figureCitation>
).
</paragraph>
<paragraph id="8BB7A03AFFB1FFAF16FDD4408C9BD46C" blockId="7.[151,1436,151,2013]" pageId="7" pageNumber="54">
In addition, underwater photographs show that
<taxonomicName id="4C08DBB9FFB1FFAF14E1D4408E45D523" ID-CoL="7DM4K" box="[729,810,1609,1632]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="54" phylum="Chordata" rank="species" species="pori">
<emphasis id="B97C7C28FFB1FFAF14E1D4408E45D523" box="[729,810,1609,1632]" italics="true" pageId="7" pageNumber="54">U. pori</emphasis>
</taxonomicName>
usually has a vertical, broad, dark brown or red bar (its width wider than pupil diameter) from the eye to the ventral margin of the head between the posterior margin of the upper jaw and preopercle (see
<bibRefCitation id="EF99DDCBFFB1FFAF17CDD4848E4CD5E6" author="Ben-Tuvia" box="[501,803,1677,1701]" pageId="7" pageNumber="54" refString="Ben-Tuvia, A. &amp; Golani, D. (1989) A new species of the goatfish (Mullidae) of the genus Upeneus from the Red Sea and the eastern Mediterranean. Israel Journal of Zoology, 36, 103 - 112." type="journal article" year="1989">Ben-Tuvia &amp; Golani 1989</bibRefCitation>
: fig. 2;
<bibRefCitation id="EF99DDCBFFB1FFAF1544D4848977D5E6" author="Randall" box="[892,1048,1677,1701]" pageId="7" pageNumber="54" refString="Randall, J. E. (1995) Coastal fishes of Oman. University of Hawai,i Press, Honolulu, xvi + 439 pp." type="book" year="1995">Randall 1995</bibRefCitation>
: fig. 634), whereas such a bar has not been observed in
<taxonomicName id="4C08DBB9FFB1FFAF17A5D4B88C96D58B" ID-CoL="7DM3Y" box="[413,505,1713,1736]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="54" phylum="Chordata" rank="species" species="itoui">
<emphasis id="B97C7C28FFB1FFAF17A5D4B88C96D58B" box="[413,505,1713,1736]" italics="true" pageId="7" pageNumber="54">U. itoui</emphasis>
</taxonomicName>
(
<figureCitation id="1333BCBFFFB1FFAF1436D4A68F36D584" box="[526,601,1711,1735]" captionStart="FIGURE 2" captionStartId="5.[151,250,1464,1487]" captionTargetBox="[151,1436,589,1442]" captionTargetId="figure@5.[151,1436,589,1443]" captionTargetPageId="5" captionText="FIGURE 2. Underwater photographs of Upeneus itoui n. sp. a b, Approximately 15 cm TL, Morode Beach, Ainan, Ehime, Japan, 33 ° 00 ˏ 15 ʺN, 132 ° 30 ˏ 22 ʺE, 4 5 m depth, 19 Oct. 1999 (photos by J. Shibata). c, Approximately 4 cm TL, Morode Beach, 6 m depth, 13 Nov. 1999 (photo by T. Hirata); U. itoui is the individual in center of the photo, other individuals are U. tragula. d, Aquarium photograph of KAUM I. 28790, paratype, 121.3 mm SL, Kagoshima, Japan, just after capture." httpUri="https://zenodo.org/record/205549/files/figure.png" pageId="7" pageNumber="54">Fig. 2</figureCitation>
). Preliminary results of mitochondrial DNA analysis of
<taxonomicName id="4C08DBB9FFB1FFAF12C5D4B88836D58B" ID-CoL="7DM3Y" box="[1277,1369,1713,1736]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="54" phylum="Chordata" rank="species" species="itoui">
<emphasis id="B97C7C28FFB1FFAF12C5D4B88836D58B" box="[1277,1369,1713,1736]" italics="true" pageId="7" pageNumber="54">U. itoui</emphasis>
</taxonomicName>
from Kagoshima and
<taxonomicName id="4C08DBB9FFB1FFAF176CD4DD8CC7D5A8" ID-CoL="7DM4K" box="[340,424,1748,1771]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="54" phylum="Chordata" rank="species" species="pori">
<emphasis id="B97C7C28FFB1FFAF176CD4DD8CC7D5A8" box="[340,424,1748,1771]" italics="true" pageId="7" pageNumber="54">U. pori</emphasis>
</taxonomicName>
from the Red Sea and the Mediterranean (the latter species is a Lessepsian migrant, a colonizer of the Mediterranean via the Suez Canal) reveal a small although constant divergence between the two species (Shirak
<emphasis id="B97C7C28FFB1FFAF1770D5108CEFD473" box="[328,384,1816,1840]" italics="true" pageId="7" pageNumber="54">et al.</emphasis>
, in prep.).
</paragraph>
<paragraph id="8BB7A03AFFB1FFAF16FDD5338E3AD49E" blockId="7.[151,1436,151,2013]" pageId="7" pageNumber="54">
Underwater photographs of
<taxonomicName id="4C08DBB9FFB1FFAF143AD5358F37D410" ID-CoL="7DM3Y" box="[514,600,1852,1875]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="54" phylum="Chordata" rank="species" species="itoui">
<emphasis id="B97C7C28FFB1FFAF143AD5358F37D410" box="[514,600,1852,1875]" italics="true" pageId="7" pageNumber="54">U. itoui</emphasis>
</taxonomicName>
reported by
<bibRefCitation id="EF99DDCBFFB1FFAF14D1D5338EADD411" author="Takagi" box="[745,962,1850,1875]" pageId="7" pageNumber="54" refString="Takagi, M., Hirata, T., Hirata, S. &amp; Nakata, C. (2010) Fishes of Ainan Ehime. Soufusha Press, Matsuyama, 250 pp." type="book" year="2010">
Takagi
<emphasis id="B97C7C28FFB1FFAF1504D5358E1AD410" box="[828,885,1851,1875]" italics="true" pageId="7" pageNumber="54">et al.</emphasis>
(2010
</bibRefCitation>
: 65; reproduced here as
<figureCitation id="1333BCBFFFB1FFAF12EED5338876D411" box="[1238,1305,1850,1874]" captionStart="FIGURE 2" captionStartId="5.[151,250,1464,1487]" captionTargetBox="[151,1436,589,1442]" captionTargetId="figure@5.[151,1436,589,1443]" captionTargetPageId="5" captionText="FIGURE 2. Underwater photographs of Upeneus itoui n. sp. a b, Approximately 15 cm TL, Morode Beach, Ainan, Ehime, Japan, 33 ° 00 ˏ 15 ʺN, 132 ° 30 ˏ 22 ʺE, 4 5 m depth, 19 Oct. 1999 (photos by J. Shibata). c, Approximately 4 cm TL, Morode Beach, 6 m depth, 13 Nov. 1999 (photo by T. Hirata); U. itoui is the individual in center of the photo, other individuals are U. tragula. d, Aquarium photograph of KAUM I. 28790, paratype, 121.3 mm SL, Kagoshima, Japan, just after capture." httpUri="https://zenodo.org/record/205549/files/figure.png" pageId="7" pageNumber="54">Fig. 2</figureCitation>
c) indicated that
<taxonomicName id="4C08DBB9FFB1FFAF16F0D5578C72D436" ID-CoL="7DM3Y" box="[200,285,1886,1909]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="54" phylum="Chordata" rank="species" species="itoui">
<emphasis id="B97C7C28FFB1FFAF16F0D5578C72D436" box="[200,285,1886,1909]" italics="true" pageId="7" pageNumber="54">U. itoui</emphasis>
</taxonomicName>
formed a school with
<taxonomicName id="4C08DBB9FFB1FFAF1422D5578E31D436" ID-CoL="7DM6Z" authority="Richardson, 1846" authorityName="Richardson" authorityYear="1846" box="[538,862,1885,1909]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="54" phylum="Chordata" rank="species" species="tragula">
<emphasis id="B97C7C28FFB1FFAF1422D5578FE1D436" box="[538,654,1885,1909]" italics="true" pageId="7" pageNumber="54">U. tragula</emphasis>
Richardson, 1846
</taxonomicName>
. Most specimens of
<taxonomicName id="4C08DBB9FFB1FFAF127CD55789F6D436" ID-CoL="7DM3Y" box="[1092,1177,1886,1909]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="54" phylum="Chordata" rank="species" species="itoui">
<emphasis id="B97C7C28FFB1FFAF127CD55789F6D436" box="[1092,1177,1886,1909]" italics="true" pageId="7" pageNumber="54">U. itoui</emphasis>
</taxonomicName>
from Kagoshima were also collected with
<taxonomicName id="4C08DBB9FFB1FFAF1757D5888C8CD4DB" ID-CoL="7DM6Z" box="[367,483,1920,1944]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="54" phylum="Chordata" rank="species" species="tragula">
<emphasis id="B97C7C28FFB1FFAF1757D5888C8CD4DB" box="[367,483,1920,1944]" italics="true" pageId="7" pageNumber="54">U. tragula</emphasis>
</taxonomicName>
. Although the two species are similar to each other when alive,
<taxonomicName id="4C08DBB9FFB1FFAF1297D588886BD4DB" ID-CoL="7DM3Y" box="[1199,1284,1921,1944]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="54" phylum="Chordata" rank="species" species="itoui">
<emphasis id="B97C7C28FFB1FFAF1297D588886BD4DB" box="[1199,1284,1921,1944]" italics="true" pageId="7" pageNumber="54">U. itoui</emphasis>
</taxonomicName>
can be easily distinguished from
<taxonomicName id="4C08DBB9FFB1FFAF174BD5AD8C86D4F8" ID-CoL="7DM6Z" box="[371,489,1955,1979]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="54" phylum="Chordata" rank="species" species="tragula">
<emphasis id="B97C7C28FFB1FFAF174BD5AD8C86D4F8" box="[371,489,1955,1979]" italics="true" pageId="7" pageNumber="54">U. tragula</emphasis>
</taxonomicName>
by having white barbels (vs. yellow in the latter) and lacking a blotch on the tip of the first dorsal fin (vs. large, dark red blotch present) (
<figureCitation id="1333BCBFFFB1FFAF14CDD5CC8E55D49E" box="[757,826,1989,2013]" captionStart="FIGURE 2" captionStartId="5.[151,250,1464,1487]" captionTargetBox="[151,1436,589,1442]" captionTargetId="figure@5.[151,1436,589,1443]" captionTargetPageId="5" captionText="FIGURE 2. Underwater photographs of Upeneus itoui n. sp. a b, Approximately 15 cm TL, Morode Beach, Ainan, Ehime, Japan, 33 ° 00 ˏ 15 ʺN, 132 ° 30 ˏ 22 ʺE, 4 5 m depth, 19 Oct. 1999 (photos by J. Shibata). c, Approximately 4 cm TL, Morode Beach, 6 m depth, 13 Nov. 1999 (photo by T. Hirata); U. itoui is the individual in center of the photo, other individuals are U. tragula. d, Aquarium photograph of KAUM I. 28790, paratype, 121.3 mm SL, Kagoshima, Japan, just after capture." httpUri="https://zenodo.org/record/205549/files/figure.png" pageId="7" pageNumber="54">Fig. 2</figureCitation>
c).
</paragraph>
<caption id="DF77F0B2FFBEFFA016AFD6B08CA8D7AC" httpUri="https://zenodo.org/record/205551/files/figure.png" pageId="8" pageNumber="55" targetBox="[184,1394,211,1184]" targetPageId="8">
<paragraph id="8BB7A03AFFBEFFA016AFD6B08CA8D7AC" blockId="8.[151,1436,1209,1263]" pageId="8" pageNumber="55">
<emphasis id="B97C7C28FFBEFFA016AFD6B08C7DD78C" bold="true" box="[151,274,1209,1232]" pageId="8" pageNumber="55">FIGURE 4.</emphasis>
Relationships of maximum body depth to standard length in
<taxonomicName id="4C08DBB9FFBEFFA01545D6B28965D793" ID-CoL="7DM3Y" box="[893,1034,1211,1232]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="55" phylum="Chordata" rank="species" species="itoui" status="sp. nov.">
<emphasis id="B97C7C28FFBEFFA01545D6B28965D793" box="[893,1034,1211,1232]" italics="true" pageId="8" pageNumber="55">Upeneus itoui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24FC153FFBEFFA01229D6B38925D78C" box="[1041,1098,1210,1231]" pageId="8" pageNumber="55" rank="species">
<emphasis id="B97C7C28FFBEFFA01229D6B38925D78C" bold="true" box="[1041,1098,1210,1231]" pageId="8" pageNumber="55">n. sp.</emphasis>
</taxonomicNameLabel>
[open symbols (star = holotype)] and
<taxonomicName id="4C08DBB9FFBEFFA016FAD6D38C64D7AC" ID-CoL="7DM4K" box="[194,267,1242,1263]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="55" phylum="Chordata" rank="species" species="pori">
<emphasis id="B97C7C28FFBEFFA016FAD6D38C64D7AC" box="[194,267,1242,1263]" italics="true" pageId="8" pageNumber="55">U. pori</emphasis>
</taxonomicName>
(closed triangles).
</paragraph>
</caption>
<caption id="DF77F0B2FFBEFFA016AFD5A68F10D4A7" httpUri="https://zenodo.org/record/205552/files/figure.png" pageId="8" pageNumber="55" targetBox="[153,1430,1405,1920]" targetPageId="8">
<paragraph id="8BB7A03AFFBEFFA016AFD5A68F10D4A7" blockId="8.[151,1435,1967,2021]" pageId="8" pageNumber="55">
<emphasis id="B97C7C28FFBEFFA016AFD5A68C7BD486" bold="true" box="[151,276,1967,1990]" pageId="8" pageNumber="55">FIGURE 5.</emphasis>
Relationships of barbel width to standard length (a) and barbel length (b) in
<taxonomicName id="4C08DBB9FFBEFFA01216D5B889D2D485" ID-CoL="7DM3Y" box="[1070,1213,1969,1990]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="55" phylum="Chordata" rank="species" species="itoui" status="sp. nov.">
<emphasis id="B97C7C28FFBEFFA01216D5B889D2D485" box="[1070,1213,1969,1990]" italics="true" pageId="8" pageNumber="55">Upeneus itoui</emphasis>
</taxonomicName>
<emphasis id="B97C7C28FFBEFFA012FCD5B98991D486" bold="true" box="[1220,1278,1968,1989]" pageId="8" pageNumber="55">
<taxonomicNameLabel id="A24FC153FFBEFFA012FCD5B98991D486" box="[1220,1278,1968,1989]" pageId="8" pageNumber="55" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
[open symbols (star = holotype)] and
<taxonomicName id="4C08DBB9FFBEFFA01741D5D98CADD4A6" ID-CoL="7DM4K" box="[377,450,2000,2021]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="55" phylum="Chordata" rank="species" species="pori">
<emphasis id="B97C7C28FFBEFFA01741D5D98CADD4A6" box="[377,450,2000,2021]" italics="true" pageId="8" pageNumber="55">U. pori</emphasis>
</taxonomicName>
(closed triangles).
</paragraph>
</caption>
<caption id="DF77F0B2FFBFFFA116AFD1F78EACD770" httpUri="https://zenodo.org/record/205553/files/figure.png" pageId="9" pageNumber="56" targetBox="[317,1285,201,995]" targetPageId="9">
<paragraph id="8BB7A03AFFBFFFA116AFD1F78EACD770" blockId="9.[151,1436,1022,1075]" pageId="9" pageNumber="56">
<emphasis id="B97C7C28FFBFFFA116AFD1F78C7CD757" bold="true" box="[151,275,1022,1045]" pageId="9" pageNumber="56">FIGURE 6.</emphasis>
Relationships of maximum body depth plus barbel width (% of standard length) to number of gill rakers in
<taxonomicName id="4C08DBB9FFBFFFA11351D6098D94D770" ID-CoL="7DM3Y" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="56" phylum="Chordata" rank="species" species="itoui" status="sp. nov.">
<emphasis id="B97C7C28FFBFFFA11351D6098D94D770" italics="true" pageId="9" pageNumber="56">Upeneus itoui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24FC153FFBFFFA11738D6178C56D770" box="[256,313,1054,1075]" pageId="9" pageNumber="56" rank="species">
<emphasis id="B97C7C28FFBFFFA11738D6178C56D770" bold="true" box="[256,313,1054,1075]" pageId="9" pageNumber="56">n. sp.</emphasis>
</taxonomicNameLabel>
[open symbols (star = holotype)] and
<taxonomicName id="4C08DBB9FFBFFFA11485D6178E69D770" ID-CoL="7DM4K" box="[701,774,1054,1075]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="56" phylum="Chordata" rank="species" species="pori">
<emphasis id="B97C7C28FFBFFFA11485D6178E69D770" box="[701,774,1054,1075]" italics="true" pageId="9" pageNumber="56">U. pori</emphasis>
</taxonomicName>
(closed triangles).
</paragraph>
</caption>
<paragraph id="8BB7A03AFFBFFFA116FDD65689C6D7A0" blockId="9.[151,1436,1119,1251]" pageId="9" pageNumber="56">
Specimens of
<taxonomicName id="4C08DBB9FFBFFFA1175DD6698CD4D734" ID-CoL="7DM3Y" box="[357,443,1120,1143]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="56" phylum="Chordata" rank="species" species="itoui">
<emphasis id="B97C7C28FFBFFFA1175DD6698CD4D734" box="[357,443,1120,1143]" italics="true" pageId="9" pageNumber="56">U. itoui</emphasis>
</taxonomicName>
from Okinawa-jima Island (
<specimenCount id="9D0E6BB3FFBFFFA114C7D6568EF4D734" box="[767,923,1119,1143]" pageId="9" pageNumber="56" type="generic">13 specimens</specimenCount>
, 93.3115.0 mm SL) were excluded from the
<typeStatus id="54B31E98FFBFFFA116AFD68D8DA8D7DF" box="[151,199,1156,1180]" pageId="9" pageNumber="56">type</typeStatus>
series of the new species. Although we regard the Okinawa specimens as conspecific with the
<typeStatus id="54B31E98FFBFFFA112C8D68D884FD7DF" box="[1264,1312,1156,1180]" pageId="9" pageNumber="56">type</typeStatus>
specimens of
<taxonomicName id="4C08DBB9FFBFFFA1168DD6A18C65D7FC" ID-CoL="7DM3Y" box="[181,266,1192,1215]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="56" phylum="Chordata" rank="species" species="itoui">
<emphasis id="B97C7C28FFBFFFA1168DD6A18C65D7FC" box="[181,266,1192,1215]" italics="true" pageId="9" pageNumber="56">U. itoui</emphasis>
</taxonomicName>
from Kagoshima and Miyazaki Prefectures, no color photographs of the Okinawa specimens when fresh or alive were available and we could not confirm the specimens indisputable identifications.
</paragraph>
</subSubSection>
</treatment>
</document>