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<mods:title id="999989555A1825D864278CB8911B0881">On the identity of the species described in the genus Lithobius Leach, 1814 by L. J. Dobroruka from the former Czechoslovakia (Czech and Slovak Republics) (Chilopoda: Lithobiomorpha)</mods:title>
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<mods:namePart id="DFE4BCCD0F0C29CF9E3CD5B958BDB51B">Tuf, Ivan Hadrián</mods:namePart>
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<mods:namePart id="613B3B047751FA28C52687D3DEC8C1DE">Wytwer, Jolanta</mods:namePart>
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<treatment id="0392AA42FFC8FFEF34C4FA8D509082AA" ID-DOI="http://doi.org/10.5281/zenodo.6229881" ID-GBIF-Taxon="119369026" ID-Zenodo-Dep="6229881" LSID="urn:lsid:plazi:treatment:0392AA42FFC8FFEF34C4FA8D509082AA" httpUri="http://treatment.plazi.org/id/0392AA42FFC8FFEF34C4FA8D509082AA" lastPageId="5" lastPageNumber="42" pageId="4" pageNumber="41">
<subSubSection id="C32148DFFFC8FFEE34C4FA8D50798614" box="[151,634,1373,1399]" pageId="4" pageNumber="41" type="nomenclature">
<paragraph id="8B841B54FFC8FFEE34C4FA8D50798614" blockId="4.[151,634,1373,1399]" box="[151,634,1373,1399]" pageId="4" pageNumber="41">
<heading id="D0CCAC38FFC8FFEE34C4FA8D50798614" bold="true" box="[151,634,1373,1399]" fontSize="11" level="1" pageId="4" pageNumber="41" reason="1">
<taxonomicName id="4C3B60D7FFC8FFEE34C4FA8D50798614" ID-CoL="3VJ2S" authority="Dobroruka, 1971" authorityName="Dobroruka" authorityYear="1971" box="[151,634,1373,1399]" class="Chilopoda" family="Lithobiidae" genus="Lithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="magurensis">
<emphasis id="B94FC746FFC8FFEE34C4FA8D50798614" bold="true" box="[151,634,1373,1399]" pageId="4" pageNumber="41">
<emphasis id="B94FC746FFC8FFEE34C4FA8D539D8614" bold="true" box="[151,414,1373,1399]" italics="true" pageId="4" pageNumber="41">Lithobius magurensis</emphasis>
<bibRefCitation id="EFAA66A5FFC8FFEE35F6FA8D50798614" author="Dobroruka" box="[421,634,1373,1399]" pageId="4" pageNumber="45" refString="Dobroruka, L. J. (1971) Chilopoda von Belanske Tatry und Spisska Magura. V e stnik C eskoslovenske spole c nosti zoologicke, 25, 261 - 264." type="journal article" year="1971">Dobroruka, 1971</bibRefCitation>
</emphasis>
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<subSubSection id="C32148DFFFC8FFEE34C4FA7A50E786A3" box="[151,740,1448,1474]" pageId="4" pageNumber="41" type="reference_group">
<paragraph id="8B841B54FFC8FFEE34C4FA7A50E786A3" blockId="4.[151,740,1448,1474]" box="[151,740,1448,1474]" pageId="4" pageNumber="41">
<treatmentCitationGroup id="AB2B3C7AFFC8FFEE34C4FA7A50E786A3" box="[151,740,1448,1474]" pageId="4" pageNumber="41">
<taxonomicName id="4C3B60D7FFC8FFEE34C4FA7A506B86A3" ID-CoL="3VJ2S" authority="Dobroruka 1971: 261" authorityName="Dobroruka" authorityPageNumber="261" authorityYear="1971" box="[151,616,1449,1473]" class="Chilopoda" family="Lithobiidae" genus="Lithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="magurensis">
<emphasis id="B94FC746FFC8FFEE34C4FA7A537F86A2" box="[151,380,1450,1473]" italics="true" pageId="4" pageNumber="41">Lithobius magurensis</emphasis>
<treatmentCitation id="0A9A3D45FFC8FFEE35D0FA79506B86A3" author="Dobroruka" box="[387,616,1449,1472]" page="261" pageId="4" pageNumber="41" year="1971">
<bibRefCitation id="EFAA66A5FFC8FFEE35D0FA79503586A3" author="Dobroruka" box="[387,566,1449,1472]" pageId="4" pageNumber="45" refString="Dobroruka, L. J. (1971) Chilopoda von Belanske Tatry und Spisska Magura. V e stnik C eskoslovenske spole c nosti zoologicke, 25, 261 - 264." type="journal article" year="1971">Dobroruka 1971</bibRefCitation>
: 261
</treatmentCitation>
</taxonomicName>
262, fig. 1
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</paragraph>
</subSubSection>
<subSubSection id="C32148DFFFC8FFEE34C4FA22567785CF" pageId="4" pageNumber="41" type="materials_examined">
<paragraph id="8B841B54FFC8FFEE34C4FA225105856F" blockId="4.[151,1437,1522,2028]" box="[151,774,1522,1548]" pageId="4" pageNumber="41">
<emphasis id="B94FC746FFC8FFEE34C4FA22533E856F" bold="true" box="[151,317,1522,1548]" pageId="4" pageNumber="41">
<typeStatus id="5480A5F6FFC8FFEE34C4FA2252D7856F" box="[151,212,1522,1548]" pageId="4" pageNumber="41">Type</typeStatus>
locality.
</emphasis>
Spišská Magura Mts,
<collectingCountry id="F32C5BC4FFC8FFEE3610FA225105856F" box="[579,774,1522,1548]" name="Slovakia" pageId="4" pageNumber="41">Slovak Republic</collectingCountry>
</paragraph>
<paragraph id="8B841B54FFC8FFEE3495F9CA5687853F" blockId="4.[151,1437,1522,2028]" pageId="4" pageNumber="41">
<emphasis id="B94FC746FFC8FFEE3495F9CA53908557" bold="true" box="[198,403,1562,1588]" pageId="4" pageNumber="41">
<typeStatus id="5480A5F6FFC8FFEE3495F9CA53008557" box="[198,259,1562,1588]" pageId="4" pageNumber="41">Type</typeStatus>
specimens.
</emphasis>
The original description of
<taxonomicName id="4C3B60D7FFC8FFEE36B6F9CB518F8557" box="[741,908,1563,1588]" class="Chilopoda" family="Lithobiidae" genus="Lithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="magurensis">
<emphasis id="B94FC746FFC8FFEE36B6F9CB518F8557" box="[741,908,1563,1588]" italics="true" pageId="4" pageNumber="41">L. magurensis</emphasis>
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was based on the
<typeStatus id="5480A5F6FFC8FFEE3038F9CA56D18557" box="[1131,1234,1562,1588]" pageId="4" pageNumber="41" type="holotype">holotype</typeStatus>
(ɗ), in the
<collectionCode id="ED2A8391FFC8FFEE310FF9CA57988557" box="[1372,1435,1562,1588]" httpUri="http://grbio.org/cool/fx6k-enrh" name="Natal Museum" pageId="4" pageNumber="41">NMP</collectionCode>
collection, and 1
<typeStatus id="5480A5F6FFC8FFEE3532F99253C5853F" box="[353,454,1602,1628]" pageId="4" pageNumber="41" type="paratype">paratype</typeStatus>
(Ψ), mentioned as deposited in the collection of Dobroruka.
</paragraph>
<paragraph id="8B841B54FFC8FFEE3495F9BA567785CF" blockId="4.[151,1437,1522,2028]" pageId="4" pageNumber="41">
<emphasis id="B94FC746FFC8FFEE3495F9BA53BE85E7" bold="true" box="[198,445,1642,1668]" pageId="4" pageNumber="41">Material examined.</emphasis>
The
<typeStatus id="5480A5F6FFC8FFEE35ACF9BA506585E7" box="[511,614,1642,1668]" pageId="4" pageNumber="41" type="holotype">holotype</typeStatus>
was restudied. Neither the
<typeStatus id="5480A5F6FFC8FFEE37E6F9BA561985E7" box="[949,1050,1642,1668]" pageId="4" pageNumber="41" type="paratype">paratype</typeStatus>
nor the other specimens of this species referred to in the original descriptions were found in Dobrorukas collection.
</paragraph>
</subSubSection>
<subSubSection id="C32148DFFFC8FFEF3495F96A509082AA" lastPageId="5" lastPageNumber="42" pageId="4" pageNumber="41" type="discussion">
<paragraph id="8B841B54FFC8FFEF3495F96A564A8249" blockId="4.[151,1437,1522,2028]" lastBlockId="5.[151,1437,152,458]" lastPageId="5" lastPageNumber="42" pageId="4" pageNumber="41">
<emphasis id="B94FC746FFC8FFEE3495F96A534385B7" bold="true" box="[198,320,1722,1748]" pageId="4" pageNumber="41">Remarks.</emphasis>
Apart from the
<typeStatus id="5480A5F6FFC8FFEE3656F96A503B85B7" box="[517,568,1722,1748]" pageId="4" pageNumber="41">type</typeStatus>
material, only 3 Ψ of
<taxonomicName id="4C3B60D7FFC8FFEE3717F96B51EF85B7" box="[836,1004,1723,1748]" class="Chilopoda" family="Lithobiidae" genus="Lithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="magurensis">
<emphasis id="B94FC746FFC8FFEE3717F96B51EF85B7" box="[836,1004,1723,1748]" italics="true" pageId="4" pageNumber="41">L. magurensis</emphasis>
</taxonomicName>
were recorded by Dobroruka in the Podyjí NP,
<collectingCountry id="F32C5BC4FFC8FFEE354EF93253DE859F" box="[285,477,1762,1788]" name="Czech Republic" pageId="4" pageNumber="41">Czech Republic</collectingCountry>
. The diagnosis of this species was based on a combination of the following characters: tergites without projections, a single claw on the last legs, and several specific sexual features of the 14th and 15th male legs. These were a dorsal sulcus on the 14th femur and a swelling with a tuft of setae on the 14th tibia. The tibia of the 15th leg was stated as being flattened, with a sulcus on the femur as well. However, the re-examined
<typeStatus id="5480A5F6FFC8FFEE35F3F852500584FF" box="[416,518,1922,1948]" pageId="4" pageNumber="41" type="holotype">holotype</typeStatus>
differed fundamentally from Dobrorukas description. The spinulation of the 14th legs is D: 1,0,3,1,1 instead of 1,0,1-2,0-1,0 while the spinulation of the 15th legs is D: 1,0,3,0,0 instead of 1,0,2,1,0. The 14th tibia shows a thin sulcus, as well as a swelling. In addition, the head is broad, resembling that of
<taxonomicName id="4C3B60D7FFC9FFEF34BEFF48503F83D1" authority="C.L. Koch, 1847" authorityName="C.L. Koch" authorityYear="1847" box="[237,572,152,178]" class="Chilopoda" family="Lithobiidae" genus="Lithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="5" pageNumber="42" phylum="Arthropoda" rank="species" species="muticus">
<emphasis id="B94FC746FFC9FFEF34BEFF48536B83D2" box="[237,360,152,177]" italics="true" pageId="5" pageNumber="42">L. muticus</emphasis>
<bibRefCitation id="EFAA66A5FFC9FFEF3522FF48503F83D1" author="Koch" box="[369,572,152,178]" pageId="5" pageNumber="46" refString="Koch, C. L. (1847) System der Myriapoden. In: Panzer, G. &amp; Herrich-Schaffer, L. (Eds.), Kritische Revision der Insectenfauna Deutschlands 3. Friedrich Pustet, Regensburg, pp. 1 - 270." type="journal article" year="1847">C.L. Koch, 1847</bibRefCitation>
</taxonomicName>
. The spinulation pattern found falls well within the variation range of
<taxonomicName id="4C3B60D7FFC9FFEF31D7FF4852F183BA" class="Chilopoda" family="Lithobiidae" genus="Lithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="5" pageNumber="42" phylum="Arthropoda" rank="species" species="muticus">
<emphasis id="B94FC746FFC9FFEF31D7FF4852F183BA" italics="true" pageId="5" pageNumber="42">L. muticus</emphasis>
</taxonomicName>
as described by
<bibRefCitation id="EFAA66A5FFC9FFEF35E0FF10505783B9" author="Latzel" box="[435,596,192,218]" pageId="5" pageNumber="46" refString="Latzel, R. (1880) Die Myriapoden der Osterreichisch-Ungarischen Monarchie. I.: Die Chilopoden. A. Holder, Wien, 228 pp." type="book" year="1880">Latzel (1880)</bibRefCitation>
,
<bibRefCitation id="EFAA66A5FFC9FFEF360CFF1050F883B9" author="Eason" box="[607,763,192,218]" pageId="5" pageNumber="45" refString="Eason, E. H. (1964) Centipedes of the British Isles. Frederick Warne &amp; Co Ltd, London &amp; New York, 294 pp." type="book" year="1964">Eason (1964)</bibRefCitation>
and
<bibRefCitation id="EFAA66A5FFC9FFEF3760FF1051D683B9" author="Koren" box="[819,981,192,218]" pageId="5" pageNumber="46" refString="Koren, A. (1992) Die Chilopoden-Fauna von Karnten und Osttirol. Teil 2. Lithobiomorpha. Carintia II, 51, 1 - 138." type="journal article" year="1992">Koren (1992)</bibRefCitation>
. Although thin sulci are not commonly met with in the available descriptions of this species, specimens of
<taxonomicName id="4C3B60D7FFC9FFEF37EEFF38563A8262" box="[957,1081,232,257]" class="Chilopoda" family="Lithobiidae" genus="Lithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="5" pageNumber="42" phylum="Arthropoda" rank="species" species="muticus">
<emphasis id="B94FC746FFC9FFEF37EEFF38563A8262" box="[957,1081,232,257]" italics="true" pageId="5" pageNumber="42">L. muticus</emphasis>
</taxonomicName>
from Central Europe, at least those from the
<collectingCountry id="F32C5BC4FFC9FFEF3514FEC050068249" box="[327,517,272,298]" name="Czech Republic" pageId="5" pageNumber="42">Czech Republic</collectingCountry>
, do show these characters (cf
<bibRefCitation id="EFAA66A5FFC9FFEF3730FEC0563F8249" author="Folkmanova" box="[867,1084,272,298]" pageId="5" pageNumber="45" refString="Folkmanova, B. (1959) StonoZky (Chilopoda). [Key of centipedes (Chilopoda).] In: Kratochvil, J. (Ed), Kli c zvi r eny C SR III. NCSAV, Praha, p. 49 - 66. (in Czech)." type="book chapter" year="1959">Folkmanová 1959</bibRefCitation>
).
</paragraph>
<paragraph id="8B841B54FFC9FFEF3495FEE856D98219" blockId="5.[151,1437,152,458]" pageId="5" pageNumber="42">
Two Ψ in the material from the SMMZ collection identified and labelled by Dobroruka as
<taxonomicName id="4C3B60D7FFC9FFEF30A5FEE8579F8232" box="[1270,1436,312,337]" class="Chilopoda" family="Lithobiidae" genus="Lithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="5" pageNumber="42" phylum="Arthropoda" rank="species" species="magurensis">
<emphasis id="B94FC746FFC9FFEF30A5FEE8579F8232" box="[1270,1436,312,337]" italics="true" pageId="5" pageNumber="42">L. magurensis</emphasis>
</taxonomicName>
are in fact 1 Ψ of
<taxonomicName id="4C3B60D7FFC9FFEF3533FEB053D9821A" box="[352,474,352,377]" class="Chilopoda" family="Lithobiidae" genus="Lithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="5" pageNumber="42" phylum="Arthropoda" rank="species" species="muticus">
<emphasis id="B94FC746FFC9FFEF3533FEB053D9821A" box="[352,474,352,377]" italics="true" pageId="5" pageNumber="42">L. muticus</emphasis>
</taxonomicName>
and 1 immature ɗ of
<taxonomicName id="4C3B60D7FFC9FFEF368BFEB056D58219" authority="Fanzago, 1874" authorityName="Fanzago" authorityYear="1874" box="[728,1238,352,378]" class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" kingdom="Animalia" order="Lithobiomorpha" pageId="5" pageNumber="42" phylum="Arthropoda" rank="species" species="tridentinus">
<emphasis id="B94FC746FFC9FFEF368BFEB05608821A" box="[728,1035,352,377]" italics="true" pageId="5" pageNumber="42">Eupolybothrus tridentinus</emphasis>
(
<bibRefCitation id="EFAA66A5FFC9FFEF3049FEB056CE8219" author="Fanzago" box="[1050,1229,352,378]" pageId="5" pageNumber="45" refString="Fanzago, F. (1874) I chilopodi Italiani. Atti della Societa Veneto-Trentina di Scienze Naturali in Padova, 3, 17 - 64." type="journal article" year="1874">Fanzago, 1874</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="8B841B54FFC9FFEF3495FE57509082AA" blockId="5.[151,1437,152,458]" pageId="5" pageNumber="42">
<emphasis id="B94FC746FFC9FFEF3495FE57539682C2" bold="true" box="[198,405,391,417]" pageId="5" pageNumber="42">Taxonomic note.</emphasis>
We consider
<taxonomicName id="4C3B60D7FFC9FFEF3664FE58560682C1" authority="Dobroruka, 1971" authorityName="Dobroruka" authorityYear="1971" box="[567,1029,392,418]" class="Chilopoda" family="Lithobiidae" genus="Lithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="5" pageNumber="42" phylum="Arthropoda" rank="species" species="magurensis">
<emphasis id="B94FC746FFC9FFEF3664FE58513782C2" box="[567,820,392,417]" italics="true" pageId="5" pageNumber="42">Lithobius magurensis</emphasis>
<bibRefCitation id="EFAA66A5FFC9FFEF376FFE58560682C1" author="Dobroruka" box="[828,1029,392,418]" pageId="5" pageNumber="45" refString="Dobroruka, L. J. (1971) Chilopoda von Belanske Tatry und Spisska Magura. V e stnik C eskoslovenske spole c nosti zoologicke, 25, 261 - 264." type="journal article" year="1971">Dobroruka, 1971</bibRefCitation>
</taxonomicName>
as a junior subjective synonym of
<taxonomicName id="4C3B60D7FFC9FFEF34C4FE60503582A9" authority="C.L. Koch, 1847" authorityName="C.L. Koch" authorityYear="1847" box="[151,566,432,458]" class="Chilopoda" family="Lithobiidae" genus="Lithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="5" pageNumber="42" phylum="Arthropoda" rank="species" species="muticus">
<emphasis id="B94FC746FFC9FFEF34C4FE60536B82AA" box="[151,360,432,457]" italics="true" pageId="5" pageNumber="42">Lithobius muticus</emphasis>
<bibRefCitation id="EFAA66A5FFC9FFEF353CFE60503582A9" author="Koch" box="[367,566,432,458]" pageId="5" pageNumber="46" refString="Koch, C. L. (1847) System der Myriapoden. In: Panzer, G. &amp; Herrich-Schaffer, L. (Eds.), Kritische Revision der Insectenfauna Deutschlands 3. Friedrich Pustet, Regensburg, pp. 1 - 270." type="journal article" year="1847">C.L. Koch, 1847</bibRefCitation>
</taxonomicName>
,
<taxonomicNameLabel id="A27C7A3DFFC9FFEF3611FE7F509082AA" box="[578,659,431,457]" pageId="5" pageNumber="42">
<emphasis id="B94FC746FFC9FFEF3611FE7F509082AA" bold="true" box="[578,659,431,457]" pageId="5" pageNumber="42">syn. n.</emphasis>
</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>