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<document id="A81D34BD4F32E64B53603C93F63C7B46" ID-CLB-Dataset="303205" ID-DOI="10.25226/bboc.v140i1.2020.a3" ID-GBIF-Dataset="f12ce80e-b597-4bd4-9abb-aa909057cb15" ID-ISSN="2513-9894" ID-Zenodo-Dep="13755707" IM.bibliography_approvedBy="karina" IM.illustrations_approvedBy="karina" IM.materialsCitations_approvedBy="karina" IM.metadata_approvedBy="karina" IM.tables_approvedBy="karina" IM.taxonomicNames_approvedBy="karina" IM.treatments_approvedBy="karina" checkinTime="1718071637822" checkinUser="felipe" docAuthor="Silva, Héctor Gómez de, Pérez Villafaña, Mónica G., Cruz-Nieto, Javier &amp; Cruz-Nieto, Miguel Ángel" docDate="2020" docId="0381A348FFC9CB3536CAAE583980FA53" docLanguage="en" docName="BullBritOrnitholClub.140.1.7-37.pdf" docOrigin="Bulletin of the British Ornithologists Club 140 (1)" docStyle="DocumentStyle:EFA7F293C51FD4806C5290561C16F223.2:BullBritOrnitholClub.2017-.journal_article" docStyleId="EFA7F293C51FD4806C5290561C16F223" docStyleName="BullBritOrnitholClub.2017-.journal_article" docStyleVersion="2" docTitle="Icterus pustulatus subsp. graysonii Cassin 1867" docType="treatment" docVersion="8" lastPageNumber="26" masterDocId="FFB8DB30FFDACB21365AAD593D0DFFC5" masterDocTitle="Are some of the birds endemic to the Tres Marías Islands (Mexico) species?" masterLastPageNumber="37" masterPageNumber="7" pageNumber="25" updateTime="1726425381095" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="B9BBF00EABC900596198EF674641942D">Are some of the birds endemic to the Tres Marías Islands (Mexico) species?</mods:title>
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<mods:namePart id="DFFAA7800A4D04DEEC7E02863EDA296A">Silva, Héctor Gómez de</mods:namePart>
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<heading id="D0DFA532FFC9CB3236CAAE583C78FCFC" fontSize="10" level="2" pageId="19" pageNumber="25" reason="2">
<subSubSection id="C33241D5FFC9CB3236CAAE583E08FCDC" box="[144,773,769,793]" pageId="19" pageNumber="25" type="nomenclature">
<paragraph id="8B97125EFFC9CB3236CAAE583E08FCDC" blockId="19.[144,1165,769,1657]" box="[144,773,769,793]" pageId="19" pageNumber="25">
<emphasis id="B95CCE4CFFC9CB3236CAAE583CD1FCDC" bold="true" box="[144,476,769,793]" pageId="19" pageNumber="25">STREAK-BACKED ORIOLE</emphasis>
<taxonomicName id="4C2869DDFFC9CB3237B8AE5B3E08FCDC" ID-CoL="5J447" authorityName="Cassin" authorityYear="1867" box="[482,773,769,793]" class="Aves" family="Icteridae" genus="Icterus" kingdom="Animalia" order="Passeriformes" pageId="19" pageNumber="25" phylum="Chordata" rank="subSpecies" species="pustulatus" subSpecies="graysonii">
<emphasis id="B95CCE4CFFC9CB3237B8AE5B3E08FCDC" box="[482,773,769,793]" italics="true" pageId="19" pageNumber="25">Icterus pustulatus graysonii</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C33241D5FFC9CB353556AE583980FA53" lastPageId="20" lastPageNumber="26" pageId="19" pageNumber="25" type="description">
<paragraph id="8B97125EFFC9CB323556AE583C78FCFC" blockId="19.[144,1165,769,1657]" pageId="19" pageNumber="25">
(vs.
<taxonomicName id="4C2869DDFFC9CB323561AE5B3EE5FCDC" box="[827,1000,770,793]" pageId="19" pageNumber="25">
<emphasis id="B95CCE4CFFC9CB323561AE5B3EE5FCDC" box="[827,1000,770,793]" italics="true" pageId="19" pageNumber="25">I. p. microstictus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2869DDFFC9CB32327BAE5B3DD4FCFC" pageId="19" pageNumber="25">
<emphasis id="B95CCE4CFFC9CB32327BAE5B3DD4FCFC" italics="true" pageId="19" pageNumber="25">I. p. yaegeri</emphasis>
</taxonomicName>
total score 7).
</paragraph>
<paragraph id="8B97125EFFC9CB3236CAAE183E91FBBC" blockId="19.[144,1165,769,1657]" pageId="19" pageNumber="25">
<emphasis id="B95CCE4CFFC9CB3236CAAE183DF1FC9C" box="[144,252,833,857]" italics="true" pageId="19" pageNumber="25">Coloration</emphasis>
.—Very few short and narrow streaks on back, usually in scapular region vs. more prominent and numerous streaks throughout back (
<figureCitation id="13130EDBFFC9CB323482AE383E28FCBC" box="[728,805,865,889]" captionStart="Figure 15" captionStartId="20.[883,944,130,149]" captionTargetBox="[144,874,128,856]" captionTargetId="figure-330@20.[141,868,125,868]" captionTargetPageId="20" captionText="Figure 15. Back patern, size and intensity of orange in Streak-backed Oriole Icterus pustulatus specimens from Nayarit, from left to right: two typical microstictus / yaegeri, two presumed hybrids (with Phillips specimen at right) and a typical graysoni; note the similar overall size of graysoni and Phillips specimen, which is, however, more orange overall (especially the head), while the other presumed hybrid (which appears almost as long due to specimen preparation) has back streaks intermediate between graysoni and typical mainland Nayarit orioles (Héctor Gómez de Silva)" figureDoi="http://doi.org/10.5281/zenodo.13755733" httpUri="https://zenodo.org/record/13755733/files/figure.png" pageId="19" pageNumber="25">Fig. 15</figureCitation>
;
<bibRefCitation id="EFB96FAFFFC9CB32356AAE383EB7FCBC" author="Grant, P. R." box="[816,954,865,889]" pageId="19" pageNumber="25" pagination="1 - 106" refId="ref14491" refString="Grant, P. R. 1965 a. A systematic study of the terrestrial birds of the Tres Marias Islands, Mexico. Postilla 90: 1 - 106." type="journal article" year="1965">Grant 1965a</bibRefCitation>
, based on
<specimenCount id="9D2ED9D7FFC9CB32326BAE383981FCBC" box="[1073,1164,865,889]" count="39" pageId="19" pageNumber="25" type="adult">39 adult</specimenCount>
male and
<specimenCount id="9D2ED9D7FFC9CB3236A6AED83C5BFC5C" box="[252,342,897,921]" count="13" pageId="19" pageNumber="25" type="adult">13 adult</specimenCount>
female
<taxonomicName id="4C2869DDFFC9CB3237F6AED83F01FC5D" box="[428,524,897,920]" class="Aves" family="Icteridae" genus="Icterus" kingdom="Animalia" order="Passeriformes" pageId="19" pageNumber="25" phylum="Chordata" rank="species" species="graysonii">
<emphasis id="B95CCE4CFFC9CB3237F6AED83F01FC5D" box="[428,524,897,920]" italics="true" pageId="19" pageNumber="25">graysonii</emphasis>
</taxonomicName>
vs.
<specimenCount id="9D2ED9D7FFC9CB32346CAED83F9DFC5C" box="[566,656,897,921]" count="69" pageId="19" pageNumber="25" type="adult">69 adult</specimenCount>
male and
<specimenCount id="9D2ED9D7FFC9CB323558AED83E51FC5C" box="[770,860,897,921]" count="29" pageId="19" pageNumber="25" type="adult">29 adult</specimenCount>
female
<emphasis id="B95CCE4CFFC9CB3235E8AED8393DFC5D" box="[946,1072,897,920]" italics="true" pageId="19" pageNumber="25">microstictus</emphasis>
/
<emphasis id="B95CCE4CFFC9CB32321EAED83980FC5D" box="[1092,1165,897,920]" italics="true" pageId="19" pageNumber="25">yaegeri</emphasis>
[score 2]). Median coverts pale yellow to yellowish-white vs. usually (orangey) yellow in
<emphasis id="B95CCE4CFFC9CB3236CAAE983DD4FC1D" box="[144,217,961,984]" italics="true" pageId="19" pageNumber="25">yaegeri</emphasis>
, the geographically closest mainland taxon, and white in
<emphasis id="B95CCE4CFFC9CB323530AE983EE5FC1D" box="[874,1000,961,984]" italics="true" pageId="19" pageNumber="25">microstictus</emphasis>
, which occurs further inland than
<emphasis id="B95CCE4CFFC9CB32372FAEB83CB3FC3D" box="[373,446,993,1016]" italics="true" pageId="19" pageNumber="25">yaegeri</emphasis>
(
<bibRefCitation id="EFB96FAFFFC9CB323794AEB83F6CFC3C" author="Phillips, A. R." box="[462,609,993,1017]" pageId="19" pageNumber="25" pagination="98 - 105" refId="ref16378" refString="Phillips, A. R. 1995. The northern races of Icterus pustulatus (Icteridae), Scarlet-headed or Streaked-backed Oriole. Bull. Brit. Orn. Cl. 115: 98 - 105." type="journal article" year="1995">Phillips 1995</bibRefCitation>
[score 1]).
<bibRefCitation id="EFB96FAFFFC9CB3234B9AEB83E70FC3C" author="Grant, P. R." box="[739,893,993,1017]" pageId="19" pageNumber="25" pagination="1 - 106" refId="ref14491" refString="Grant, P. R. 1965 a. A systematic study of the terrestrial birds of the Tres Marias Islands, Mexico. Postilla 90: 1 - 106." type="journal article" year="1965">Grant (1965a)</bibRefCitation>
mentioned but did not quantify a tendency to differ in yellow vs. orange plumage. Generally,
<taxonomicName id="4C2869DDFFC9CB323581A9583936FBDD" box="[987,1083,1025,1048]" class="Aves" family="Icteridae" genus="Icterus" kingdom="Animalia" order="Passeriformes" pageId="19" pageNumber="25" phylum="Chordata" rank="species" species="graysonii">
<emphasis id="B95CCE4CFFC9CB323581A9583936FBDD" box="[987,1083,1025,1048]" italics="true" pageId="19" pageNumber="25">graysonii</emphasis>
</taxonomicName>
is less orange overall (consistent with the median coverts character, above, and with
<bibRefCitation id="EFB96FAFFFC9CB32327DA9783DC8FB9C" author="Ridgway, R." pageId="19" pageNumber="25" pagination="1 - 834" refId="ref16712" refString="Ridgway, R. 1902. The birds of North and Middle America. Bull. US Natl. Mus. 50 (2): 1 - 834." type="journal article" year="1902">Ridgway 1902</bibRefCitation>
,
<bibRefCitation id="EFB96FAFFFC9CB32368BA9183CD4FB9C" author="Jaramillo, A. &amp; Burke, P." box="[209,473,1089,1113]" pageId="19" pageNumber="25" refId="ref15460" refString="Jaramillo, A. &amp; Burke, P. 1999. New World blackbirds: the icterids. Princeton Univ. Press." type="book" year="1999">Jaramillo &amp; Burke 1999</bibRefCitation>
) and we support this based on our field work, although
<specimenCount id="9D2ED9D7FFC9CB323239A9183DC6FBBC" count="1" pageId="19" pageNumber="25" type="adult">one adult</specimenCount>
<taxonomicName id="4C2869DDFFC9CB323688A9383C3FFBBD" box="[210,306,1121,1144]" class="Aves" family="Icteridae" genus="Icterus" kingdom="Animalia" order="Passeriformes" pageId="19" pageNumber="25" phylum="Chordata" rank="species" species="graysonii">
<emphasis id="B95CCE4CFFC9CB323688A9383C3FFBBD" box="[210,306,1121,1144]" italics="true" pageId="19" pageNumber="25">graysonii</emphasis>
</taxonomicName>
among the dozens seen was intensely orange (
<figureCitation id="13130EDBFFC9CB323518A9383E82FBBC" box="[834,911,1121,1145]" captionStart="Figure 16" captionStartId="20.[144,205,1232,1251]" captionTargetBox="[144,1164,866,1226]" captionTargetId="figure-219@20.[140,1167,878,1220]" captionTargetPageId="20" captionText="Figure 16. An intensely orange individual of Streak-backed Oriole Icterus pustulatus graysoni, Isla María Cleofas, March 2016 (Javier Cruz Nieto)" figureDoi="http://doi.org/10.5281/zenodo.13765609" httpUri="https://zenodo.org/record/13765609/files/figure.png" pageId="19" pageNumber="25">Fig. 16</figureCitation>
).
</paragraph>
<paragraph id="8B97125EFFC9CB32369AA9D83DFCFADC" blockId="19.[144,1165,769,1657]" pageId="19" pageNumber="25">
<emphasis id="B95CCE4CFFC9CB32369AA9D83C50FB5C" box="[192,349,1153,1177]" italics="true" pageId="19" pageNumber="25">Morphometrics</emphasis>
.—Larger in external measurements (score 2), but with a significantly shorter coracoid both absolutely and relatively (score 2) and a relatively shorter femur (not significantly different whereas all external measurements are significantly different [not scored]). The bill is also differently shaped (longer in relation to width,
<bibRefCitation id="EFB96FAFFFC9CB323595A9B8395BFB3C" author="Grant, P. R." box="[975,1110,1249,1273]" pageId="19" pageNumber="25" pagination="1 - 106" refId="ref14491" refString="Grant, P. R. 1965 a. A systematic study of the terrestrial birds of the Tres Marias Islands, Mexico. Postilla 90: 1 - 106." type="journal article" year="1965">Grant 1965a</bibRefCitation>
[not scored]).
</paragraph>
<paragraph id="8B97125EFFC9CB32369AA8783E32F9BC" blockId="19.[144,1165,769,1657]" pageId="19" pageNumber="25">
<emphasis id="B95CCE4CFFC9CB32369AA8783CDBFAFC" box="[192,470,1313,1337]" italics="true" pageId="19" pageNumber="25">Evidence of hybridisation</emphasis>
.—
<bibRefCitation id="EFB96FAFFFC9CB3237ACA8783F92FAFC" author="Phillips, A. R." box="[502,671,1313,1337]" pageId="19" pageNumber="25" pagination="98 - 105" refId="ref16378" refString="Phillips, A. R. 1995. The northern races of Icterus pustulatus (Icteridae), Scarlet-headed or Streaked-backed Oriole. Bull. Brit. Orn. Cl. 115: 98 - 105." type="journal article" year="1995">Phillips (1995)</bibRefCitation>
collected a mainland specimen that he presumed to be a rare variant of
<emphasis id="B95CCE4CFFC9CB3237ADA8183F4DFA9D" box="[503,576,1345,1368]" italics="true" pageId="19" pageNumber="25">yaegeri</emphasis>
and had plain yellowish interscapulars, with hardly perceptible black streaking (on the back). But the middle wing-coverts are richer, and bill shorter, than
<taxonomicName id="4C2869DDFFC9CB323775A8D83C82FA5D" box="[303,399,1409,1432]" class="Aves" family="Icteridae" genus="Icterus" kingdom="Animalia" order="Passeriformes" pageId="19" pageNumber="25" phylum="Chordata" rank="species" species="graysonii">
<emphasis id="B95CCE4CFFC9CB323775A8D83C82FA5D" box="[303,399,1409,1432]" italics="true" pageId="19" pageNumber="25">graysonii</emphasis>
</taxonomicName>
. This specimen is now in the IBUNAM collection (P022269). Its precise locality, according to the label, is 1¾ km N of Singaita, whereon it is also mentioned Apparently alone in brush, near normal-backed pair. The bird not only resembles
<taxonomicName id="4C2869DDFFC9CB323276A8983981FA1D" box="[1068,1164,1473,1496]" class="Aves" family="Icteridae" genus="Icterus" kingdom="Animalia" order="Passeriformes" pageId="19" pageNumber="25" phylum="Chordata" rank="species" species="graysonii">
<emphasis id="B95CCE4CFFC9CB323276A8983981FA1D" box="[1068,1164,1473,1496]" italics="true" pageId="19" pageNumber="25">graysonii</emphasis>
</taxonomicName>
in its back patern but also in size (
<figureCitation id="13130EDBFFC9CB323454A8B83F56FA3C" box="[526,603,1505,1529]" captionStart="Figure 15" captionStartId="20.[883,944,130,149]" captionTargetBox="[144,874,128,856]" captionTargetId="figure-330@20.[141,868,125,868]" captionTargetPageId="20" captionText="Figure 15. Back patern, size and intensity of orange in Streak-backed Oriole Icterus pustulatus specimens from Nayarit, from left to right: two typical microstictus / yaegeri, two presumed hybrids (with Phillips specimen at right) and a typical graysoni; note the similar overall size of graysoni and Phillips specimen, which is, however, more orange overall (especially the head), while the other presumed hybrid (which appears almost as long due to specimen preparation) has back streaks intermediate between graysoni and typical mainland Nayarit orioles (Héctor Gómez de Silva)" figureDoi="http://doi.org/10.5281/zenodo.13755733" httpUri="https://zenodo.org/record/13755733/files/figure.png" pageId="19" pageNumber="25">Fig. 15</figureCitation>
). Therefore it combines phenotypic characters and is almost surely a hybrid. We found a second mainland specimen (P015891 from Santiago Ixcuintla) that resembles
<taxonomicName id="4C2869DDFFC9CB3237EBAB783F1CF9FD" box="[433,529,1569,1592]" class="Aves" family="Icteridae" genus="Icterus" kingdom="Animalia" order="Passeriformes" pageId="19" pageNumber="25" phylum="Chordata" rank="species" species="graysonii">
<emphasis id="B95CCE4CFFC9CB3237EBAB783F1CF9FD" box="[433,529,1569,1592]" italics="true" pageId="19" pageNumber="25">graysonii</emphasis>
</taxonomicName>
in its fewer and narrower back streaks, but streaks are present even on the central back (
<figureCitation id="13130EDBFFC9CB323456AB183F57F99C" box="[524,602,1601,1625]" captionStart="Figure 15" captionStartId="20.[883,944,130,149]" captionTargetBox="[144,874,128,856]" captionTargetId="figure-330@20.[141,868,125,868]" captionTargetPageId="20" captionText="Figure 15. Back patern, size and intensity of orange in Streak-backed Oriole Icterus pustulatus specimens from Nayarit, from left to right: two typical microstictus / yaegeri, two presumed hybrids (with Phillips specimen at right) and a typical graysoni; note the similar overall size of graysoni and Phillips specimen, which is, however, more orange overall (especially the head), while the other presumed hybrid (which appears almost as long due to specimen preparation) has back streaks intermediate between graysoni and typical mainland Nayarit orioles (Héctor Gómez de Silva)" figureDoi="http://doi.org/10.5281/zenodo.13755733" httpUri="https://zenodo.org/record/13755733/files/figure.png" pageId="19" pageNumber="25">Fig. 15</figureCitation>
). In overall length this specimen is comparable to other mainland specimens, and we believe it is also a hybrid.
</paragraph>
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<paragraph id="8B97125EFFCECB353529ADDB3951FD40" blockId="20.[883,1165,129,645]" pageId="20" pageNumber="26">
Figure 15. Back patern, size and intensity of orange in Streak-backed Oriole
<taxonomicName id="4C2869DDFFCECB35320AADED3EDEFF1A" baseAuthorityName="Wagler" baseAuthorityYear="1829" class="Aves" family="Icteridae" genus="Icterus" kingdom="Animalia" order="Passeriformes" pageId="20" pageNumber="26" phylum="Chordata" rank="species" species="pustulatus">
<emphasis id="B95CCE4CFFCECB35320AADED3EDEFF1A" italics="true" pageId="20" pageNumber="26">Icterus pustulatus</emphasis>
</taxonomicName>
specimens from Nayarit, from left to right: two typical
<emphasis id="B95CCE4CFFCECB35359AADA73927FED5" box="[960,1066,254,272]" italics="true" pageId="20" pageNumber="26">microstictus</emphasis>
/
<emphasis id="B95CCE4CFFCECB353210ADA7398AFED5" box="[1098,1159,254,272]" italics="true" pageId="20" pageNumber="26">yaegeri</emphasis>
, two presumed hybrids (with Phillips specimen at right) and a typical
<taxonomicName id="4C2869DDFFCECB3535B4AC103934FE9E" box="[1006,1081,329,347]" class="Aves" family="Trochilidae" genus="Amazilia" kingdom="Animalia" order="Apodiformes" pageId="20" pageNumber="26" phylum="Chordata" rank="species" species="graysoni">
<emphasis id="B95CCE4CFFCECB3535B4AC103934FE9E" box="[1006,1081,329,347]" italics="true" pageId="20" pageNumber="26">graysoni</emphasis>
</taxonomicName>
; note the similar overall size of
<taxonomicName id="4C2869DDFFCECB35321BAC383981FEB6" box="[1089,1164,353,371]" class="Aves" family="Trochilidae" genus="Amazilia" kingdom="Animalia" order="Apodiformes" pageId="20" pageNumber="26" phylum="Chordata" rank="species" species="graysoni">
<emphasis id="B95CCE4CFFCECB35321BAC383981FEB6" box="[1089,1164,353,371]" italics="true" pageId="20" pageNumber="26">graysoni</emphasis>
</taxonomicName>
and Phillips specimen, which is, however, more orange overall (especially the head), while the other presumed hybrid (which appears almost as long due to specimen preparation) has back streaks intermediate between
<taxonomicName id="4C2869DDFFCECB353591AF18391BFD96" box="[971,1046,577,595]" class="Aves" family="Trochilidae" genus="Amazilia" kingdom="Animalia" order="Apodiformes" pageId="20" pageNumber="26" phylum="Chordata" rank="species" species="graysoni">
<emphasis id="B95CCE4CFFCECB353591AF18391BFD96" box="[971,1046,577,595]" italics="true" pageId="20" pageNumber="26">graysoni</emphasis>
</taxonomicName>
and typical mainland Nayarit orioles (Héctor Gómez de Silva)
</paragraph>
</caption>
<caption id="DF5742D6FFCECB3536CAA9893F07FB39" ID-DOI="http://doi.org/10.5281/zenodo.13765609" ID-Zenodo-Dep="13765609" httpUri="https://zenodo.org/record/13765609/files/figure.png" pageId="20" pageNumber="26" startId="20.[144,205,1232,1251]" targetBox="[144,1164,866,1226]" targetPageId="20" targetType="figure">
<paragraph id="8B97125EFFCECB3536CAA9893F07FB39" blockId="20.[144,1164,1232,1276]" pageId="20" pageNumber="26">
Figure 16. An intensely orange individual of Streak-backed Oriole
<taxonomicName id="4C2869DDFFCECB35357FA9883916FB26" authorityName="Cassin" authorityYear="1867" box="[805,1051,1232,1251]" class="Aves" family="Icteridae" genus="Icterus" kingdom="Animalia" order="Passeriformes" pageId="20" pageNumber="26" phylum="Chordata" rank="subSpecies" species="pustulatus" subSpecies="graysoni">
<emphasis id="B95CCE4CFFCECB35357FA9883916FB26" box="[805,1051,1232,1251]" italics="true" pageId="20" pageNumber="26">Icterus pustulatus graysoni</emphasis>
</taxonomicName>
, Isla María Cleofas, March 2016 (Javier Cruz Nieto)
</paragraph>
</caption>
<paragraph id="8B97125EFFCECB35369AA8473980FA53" blockId="20.[144,1165,1310,1430]" pageId="20" pageNumber="26">
<emphasis id="B95CCE4CFFCECB35369AA8473CBDFAF3" box="[192,432,1310,1334]" italics="true" pageId="20" pageNumber="26">Additional information</emphasis>
.—
<bibRefCitation id="EFB96FAFFFCECB35378AA8473E2FFAF3" author="Cortes-Rodriguez, N. &amp; Hernandez-Banos, B. E. &amp; Navarro-Siguenza, A. G. &amp; Omland, K. E." box="[464,802,1310,1334]" pageId="20" pageNumber="26" pagination="729 - 739" refId="ref13948" refString="Cortes-Rodriguez, N., Hernandez-Banos, B. E., Navarro-Siguenza, A. G. &amp; Omland, K. E. 2008. Geographic variation and genetic structure in the Streak-backed Oriole: low mitochondrial DNA differentiation reveals recent divergence. Condor 110: 729 - 739." type="journal article" year="2008">
Cortés-Rodríguez
<emphasis id="B95CCE4CFFCECB3534FAA8463FC3FAF3" box="[672,718,1310,1334]" italics="true" pageId="20" pageNumber="26">et al</emphasis>
. (2008)
</bibRefCitation>
and
<bibRefCitation id="EFB96FAFFFCECB35353AA8473981FAF3" author="Ortiz-Ramirez, M. F. &amp; Sanchez-Gonzalez, L. A. &amp; Castellanos-Morales, G. &amp; Ornelas, J. F. &amp; Navarro-Siguenza, A. G." box="[864,1164,1310,1334]" pageId="20" pageNumber="26" pagination="716 - 732" refId="ref16237" refString="Ortiz-Ramirez, M. F., Sanchez-Gonzalez, L. A., Castellanos-Morales, G., Ornelas, J. F. &amp; Navarro-Siguenza, A. G. 2018. Concerted Pleistocene dispersal and genetic differentiation in passerine birds from the Tres Marias archipelago, Mexico. Auk 135: 716 - 732." type="journal article" year="2018">
Ortiz-Ramírez
<emphasis id="B95CCE4CFFCECB353250A8463935FAF3" box="[1034,1080,1310,1334]" italics="true" pageId="20" pageNumber="26">et al</emphasis>
. (2018)
</bibRefCitation>
found reciprocal monophyly but shallow genetic divergence between
<taxonomicName id="4C2869DDFFCECB3535AAA866395CFA93" box="[1008,1105,1343,1366]" class="Aves" family="Icteridae" genus="Icterus" kingdom="Animalia" order="Passeriformes" pageId="20" pageNumber="26" phylum="Chordata" rank="species" species="graysonii">
<emphasis id="B95CCE4CFFCECB3535AAA866395CFA93" box="[1008,1105,1343,1366]" italics="true" pageId="20" pageNumber="26">graysonii</emphasis>
</taxonomicName>
and mainland specimens. Shallow genetic divergence is also seen in other sister species of orioles even when they possess distinctly different plumage features (e.g.
<bibRefCitation id="EFB96FAFFFCECB353598A827398FFA53" author="Kondo, B. &amp; Baker, J. M. &amp; Omland, K. E." box="[962,1154,1406,1430]" pageId="20" pageNumber="26" pagination="674 - 680" refId="ref15534" refString="Kondo, B., Baker, J. M. &amp; Omland, K. E. 2004. Recent speciation between the Baltimore Oriole and the Blackbacked Oriole. Condor 106: 674 - 680." type="journal article" year="2004">
Kondo
<emphasis id="B95CCE4CFFCECB35324EA8263933FA53" box="[1044,1086,1406,1430]" italics="true" pageId="20" pageNumber="26">et al</emphasis>
. 2004
</bibRefCitation>
).
</paragraph>
</subSubSection>
</heading>
</treatment>
</document>