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<document id="6012AF8B119F9E120D687D0DCBCC1D62" ID="10.11646/zootaxa.4150.3.3" ID-DOI="10.11646/zootaxa.4150.3.3" ID-GBIF-Dataset="e98edafd-7489-488d-ab8a-276a4a4926a4" ID-ISSN="1175-5326" ID-Zenodo-Dep="265618" ID-ZooBank="79657D7A-324F-4539-A80F-E11BB4189753" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1470984303137" checkinUser="plazi" docAuthor="Patitucci, Luciano Damián, Mulieri, Pablo Ricardo &amp; Mariluis, Juan Carlos" docDate="2016" docId="03F65F6EF053FFD4FF6CFF1FFDA72B53" docLanguage="en" docName="zootaxa.4150.3.3.pdf" docOrigin="Zootaxa 4150 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Helina discolor Stein 1911" docType="treatment" docVersion="7" lastPageNumber="297" masterDocId="FFCF2716F05CFFC4FFFBFF88FFAE2F04" masterDocTitle="Taxonomic review of the species of Helina R. - D. (Diptera: Muscidae) from Andean-Patagonian forests" masterLastPageNumber="313" masterPageNumber="281" pageNumber="296" updateTime="1698381830647" updateUser="plazi">
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<mods:title id="219716E339EEAC74820BE4A456B766CB">Taxonomic review of the species of Helina R. - D. (Diptera: Muscidae) from Andean-Patagonian forests</mods:title>
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<mods:namePart id="D3EE4676F00BB07E0B955161FF18BFE5">Patitucci, Luciano Damián</mods:namePart>
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<mods:namePart id="E085C9AC6CF0F79A0E6B91DA70B7560D">Mulieri, Pablo Ricardo</mods:namePart>
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<mods:namePart id="811BF3BDC48C8F7DEAB3CA3653D28775">Mariluis, Juan Carlos</mods:namePart>
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<treatment id="03F65F6EF053FFD4FF6CFF1FFDA72B53" ID-DOI="http://doi.org/10.5281/zenodo.6088859" ID-GBIF-Taxon="124329285" ID-Zenodo-Dep="6088859" LSID="urn:lsid:plazi:treatment:03F65F6EF053FFD4FF6CFF1FFDA72B53" httpUri="http://treatment.plazi.org/id/03F65F6EF053FFD4FF6CFF1FFDA72B53" lastPageId="16" lastPageNumber="297" pageId="15" pageNumber="296">
<subSubSection id="C345BDF3F053FFCBFF6CFF1FFE8F2FD7" pageId="15" pageNumber="296" type="nomenclature">
<paragraph id="8BE0EE78F053FFCBFF6CFF1FFE5C2FB6" blockId="15.[151,498,151,211]" box="[151,498,151,178]" pageId="15" pageNumber="296">
<heading id="D0A85914F053FFCBFF6CFF1FFE5C2FB6" bold="true" box="[151,498,151,178]" fontSize="11" level="1" pageId="15" pageNumber="296" reason="1">
<taxonomicName id="4C5F95FBF053FFCBFF6CFF1FFE5C2FB6" authority="Stein, 1911" authorityName="Stein" authorityYear="1911" box="[151,498,151,178]" class="Insecta" family="Lonchaeidae" genus="Helina" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="296" phylum="Arthropoda" rank="species" species="discolor">
<emphasis id="B92B326AF053FFCBFF6CFF1FFE5C2FB6" bold="true" box="[151,498,151,178]" pageId="15" pageNumber="296">
<emphasis id="B92B326AF053FFCBFF6CFF1FFEFF2FB5" bold="true" box="[151,337,151,177]" italics="true" pageId="15" pageNumber="296">Helina discolor</emphasis>
(
<bibRefCitation id="EFCE9389F053FFCBFE9AFF10FE472FB6" author="Stein" box="[353,489,152,178]" pageId="15" pageNumber="296" refString="Stein, P. (1911) Die von Schnuse in Sudamerika gefangenen Anthomyiden. Archiv fur Naturgeschichte, 77, 61 - 189." type="journal article" year="1911">Stein, 1911</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BE0EE78F053FFCBFF6CFF32FE8F2FD7" blockId="15.[151,498,151,211]" box="[151,289,186,211]" pageId="15" pageNumber="296">
(
<figureCitation id="1364F2FDF053FFCBFF64FF32FF402FD6" box="[159,238,186,211]" captionStart="FIGURE 7" captionStartId="15.[151,250,1893,1915]" captionTargetBox="[187,1424,934,1854]" captionTargetId="figure@15.[161,1425,934,1871]" captionTargetPageId="15" captionText="FIGURE 7. Helina discolor (Stein), male: A. Lateral view (scale bar: 1 mm). B. Head, frontal view. C. Thorax, dorsal view. D. Abdomen, dorsal view (scale bars: 0.5 mm). E. Sternite 5 (scale bar: 0.2 mm). F. Cercal plate, posterior view. G. Cercal plate and surstylus, lateral view (scale bar: 0.1 mm). H. Phallic complex, lateral view (scale bar: 0.1 mm). (Abbreviations: aed apod, aedeagal apodeme; distiph, distiphallus; epiph, epiphallus; gon, gonopodo; par, paramere.)" httpUri="https://zenodo.org/record/265625/files/figure.png" pageId="15" pageNumber="296">Figs. 7</figureCitation>
,
<figureCitation id="1364F2FDF053FFCBFF01FF33FEA62FD7" box="[250,264,187,211]" captionStart="FIGURE 9" captionStartId="18.[151,250,1937,1959]" captionTargetBox="[168,1423,829,1910]" captionTargetId="figure@18.[163,1423,829,1916]" captionTargetPageId="18" captionText="FIGURE 9. Geographical distribution. A. Helina araucana sp. nov. B. Helina australis Carvalho &amp; Pont. C. Helina bigoti Malloch. D. Helina chilensis Malloch. E. Helina connexa Malloch. F. Helina discolor (Stein). G. Helina dorada sp. nov. (Green area: Andean-Patagonian forests; orange circle: new records; blue triangle: previous distribution.)" httpUri="https://zenodo.org/record/265627/files/figure.png" pageId="15" pageNumber="296">9</figureCitation>
F)
</paragraph>
</subSubSection>
<subSubSection id="C345BDF3F053FFD4FF6CFE8BFB122D80" lastPageId="16" lastPageNumber="297" pageId="15" pageNumber="296" type="description">
<paragraph id="8BE0EE78F053FFCBFF6CFE8BFC222E1C" blockId="15.[151,1436,258,342]" box="[151,908,258,280]" pageId="15" pageNumber="296">
For a complete list of references, see the catalog by
<bibRefCitation id="EFCE9389F053FFCBFD5EFE8AFC292E1C" box="[677,903,258,280]" pageId="15" pageNumber="296" refString="Carvalho, C. J. B. de, Couri, M. S., Pont, A. C., Pamplona, D. &amp; Lopes, S. M. (2005) A Catalogue of the Muscidae (Diptera) of the Neotropical Region. Zootaxa, 860, 1 - 282. http: // dx. doi. org / 10.11646 / zootaxa. 860.1.1" type="journal article">
Carvalho
<emphasis id="B92B326AF053FFCBFCF3FE8BFC922E1C" box="[776,828,258,280]" italics="true" pageId="15" pageNumber="296">et al.</emphasis>
(2005)
</bibRefCitation>
.
</paragraph>
<paragraph id="8BE0EE78F053FFCBFF6CFEA9FDCE2E52" blockId="15.[151,1436,258,342]" pageId="15" pageNumber="296">
Information subsequent to this catalog:
<bibRefCitation id="EFCE9389F053FFCBFDBDFEA9FCDF2E33" author="Lowerberg-Neto" box="[582,881,289,311]" pageId="15" pageNumber="296" refString="Lowerberg-Neto, P., Carvalho, C. J. B. de &amp; Hawkins, B. A. (2011) Tropical niche conservatism as a historical narrative hypothesis for the Neotropics: a case study using the fly family Muscidae. Journal of Biogeography, 38, 1936 - 1947. http: // dx. doi. org / 10.1111 / j. 1365 - 2699.2011.02540. x" type="journal article" year="2011">
Löwerberg-Neto
<emphasis id="B92B326AF053FFCBFD01FEAAFC832E33" box="[762,813,289,311]" italics="true" pageId="15" pageNumber="296">et al</emphasis>
. 2011
</bibRefCitation>
(biogeography);
<bibRefCitation id="EFCE9389F053FFCBFBD7FEA9FA322E33" author="Lowerberg-Neto" box="[1068,1436,289,311]" pageId="15" pageNumber="296" refString="Lowerberg-Neto, P. &amp; Carvalho, C. J. B. de (2013) Muscidae (Insecta: Diptera) of Latin America and the Caribbean: geographic distribution and check-list by country. Zootaxa, 3650 (1), 01 - 147. http: // dx. doi. org / 10.11646 / zootaxa. 3650.1.1" type="journal article" year="2013">Löwerberg-Neto &amp; Carvalho 2013</bibRefCitation>
(checklist),
<bibRefCitation id="EFCE9389F053FFCBFEC5FEC9FE082E52" author="Pont" box="[318,422,320,342]" pageId="15" pageNumber="296" refString="Pont, A. C. (2013). The Fanniidae and Muscidae (Diptera) described by Paul Stein (1852 - 1921). Journal of Zoological Systematics and Evolutionary Research, 89, 31 - 166. http: // dx. doi. org / 10.1002 / zoos. 201300004" type="journal article" year="2013">Pont 2013</bibRefCitation>
(
<typeStatus id="54E450DAF053FFCBFE48FEC8FE4E2E52" box="[435,480,320,342]" pageId="15" pageNumber="296">type</typeStatus>
specimens).
</paragraph>
<paragraph id="8BE0EE78F053FFCBFF6CFE0AFBE02E9E" blockId="15.[151,1436,386,882]" box="[151,1102,386,411]" pageId="15" pageNumber="296">
<emphasis id="B92B326AF053FFCBFF6CFE0AFE932E9F" bold="true" box="[151,317,386,411]" pageId="15" pageNumber="296">Redescription</emphasis>
.
<emphasis id="B92B326AF053FFCBFEB0FE0AFE292E9F" bold="true" box="[331,391,386,411]" italics="true" pageId="15" pageNumber="296">Male</emphasis>
(
<figureCitation id="1364F2FDF053FFCBFE6EFE0AFE772E9F" box="[405,473,386,411]" captionStart="FIGURE 7" captionStartId="15.[151,250,1893,1915]" captionTargetBox="[187,1424,934,1854]" captionTargetId="figure@15.[161,1425,934,1871]" captionTargetPageId="15" captionText="FIGURE 7. Helina discolor (Stein), male: A. Lateral view (scale bar: 1 mm). B. Head, frontal view. C. Thorax, dorsal view. D. Abdomen, dorsal view (scale bars: 0.5 mm). E. Sternite 5 (scale bar: 0.2 mm). F. Cercal plate, posterior view. G. Cercal plate and surstylus, lateral view (scale bar: 0.1 mm). H. Phallic complex, lateral view (scale bar: 0.1 mm). (Abbreviations: aed apod, aedeagal apodeme; distiph, distiphallus; epiph, epiphallus; gon, gonopodo; par, paramere.)" httpUri="https://zenodo.org/record/265625/files/figure.png" pageId="15" pageNumber="296">Fig. 7</figureCitation>
A). Length. Body:
<quantity id="4CA7439DF053FFCBFD57FE0AFCFF2E9E" box="[684,849,386,411]" metricMagnitude="-3" metricUnit="m" metricValue="6.97" metricValueMax="7.68" metricValueMin="6.26" pageId="15" pageNumber="296" unit="mm" value="6.97" valueMax="7.68" valueMin="6.26">6.267.68 mm</quantity>
, wing:
<quantity id="4CA7439DF053FFCBFC58FE0AFBE72E9E" box="[931,1097,386,411]" metricMagnitude="-3" metricUnit="m" metricValue="5.66" metricValueMax="6.42" metricValueMin="4.9" pageId="15" pageNumber="296" unit="mm" value="5.66" valueMax="6.42" valueMin="4.9">4.906.42 mm</quantity>
.
</paragraph>
<paragraph id="8BE0EE78F053FFCBFF3CFE2EFBFE2D2F" blockId="15.[151,1436,386,882]" pageId="15" pageNumber="296">
<emphasis id="B92B326AF053FFCBFF3CFE2EFEAA2EBB" box="[199,260,422,447]" italics="true" pageId="15" pageNumber="296">Head</emphasis>
(
<figureCitation id="1364F2FDF053FFCBFEE9FE2EFEFB2EBB" box="[274,341,422,447]" captionStart="FIGURE 7" captionStartId="15.[151,250,1893,1915]" captionTargetBox="[187,1424,934,1854]" captionTargetId="figure@15.[161,1425,934,1871]" captionTargetPageId="15" captionText="FIGURE 7. Helina discolor (Stein), male: A. Lateral view (scale bar: 1 mm). B. Head, frontal view. C. Thorax, dorsal view. D. Abdomen, dorsal view (scale bars: 0.5 mm). E. Sternite 5 (scale bar: 0.2 mm). F. Cercal plate, posterior view. G. Cercal plate and surstylus, lateral view (scale bar: 0.1 mm). H. Phallic complex, lateral view (scale bar: 0.1 mm). (Abbreviations: aed apod, aedeagal apodeme; distiph, distiphallus; epiph, epiphallus; gon, gonopodo; par, paramere.)" httpUri="https://zenodo.org/record/265625/files/figure.png" pageId="15" pageNumber="296">Fig. 7</figureCitation>
B). Black with silver pollinosity. Holoptic, the shortest distance between eyes is
<quantity id="4CA7439DF053FFCBFB2AFE2FFAD82EBA" box="[1233,1398,423,447]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" metricValueMax="2.7" metricValueMin="2.3" pageId="15" pageNumber="296" unit="mm" value="0.25" valueMax="0.27" valueMin="0.23">0.230.27 mm</quantity>
. 8 10 pairs of frontal setae. Eye hairs long and sparse. Lunule dark brown; fronto-orbital plate, parafacial, gena, postgena, and occiput black with silver pollinosity. Parafrontal plate bare. Postocular setae proclinate. Antenna black; arista with its longest hairs hardly longer than its basal diameter. Palpus black.
</paragraph>
<paragraph id="8BE0EE78F053FFCBFF3CFDBEFE2F2C06" blockId="15.[151,1436,386,882]" pageId="15" pageNumber="296">
<emphasis id="B92B326AF053FFCBFF3CFDBEFEB62D4B" box="[199,280,566,591]" italics="true" pageId="15" pageNumber="296">Thorax</emphasis>
(
<figureCitation id="1364F2FDF053FFCBFED1FDBEFEDF2D4B" box="[298,369,566,591]" captionStart="FIGURE 7" captionStartId="15.[151,250,1893,1915]" captionTargetBox="[187,1424,934,1854]" captionTargetId="figure@15.[161,1425,934,1871]" captionTargetPageId="15" captionText="FIGURE 7. Helina discolor (Stein), male: A. Lateral view (scale bar: 1 mm). B. Head, frontal view. C. Thorax, dorsal view. D. Abdomen, dorsal view (scale bars: 0.5 mm). E. Sternite 5 (scale bar: 0.2 mm). F. Cercal plate, posterior view. G. Cercal plate and surstylus, lateral view (scale bar: 0.1 mm). H. Phallic complex, lateral view (scale bar: 0.1 mm). (Abbreviations: aed apod, aedeagal apodeme; distiph, distiphallus; epiph, epiphallus; gon, gonopodo; par, paramere.)" httpUri="https://zenodo.org/record/265625/files/figure.png" pageId="15" pageNumber="296">Fig. 7</figureCitation>
C). Scutum black with two poorly defined white pollinose vittae; scutellum black with grey pollinosity; anepisternum, anepimeron, katepimeron, katepisternum, proepisternum, proepimeron and meron black; anterior and posterior spiracles dark brown. Chaetotaxy: acrostichal setae 1-2+1; dorsocentral setae 2+4; humeral setae 3; notopleural setae 2, of similar size. Prealar strong, less than one fourth of the length of the anterior supra-alar postsutural seta. Scutellum with short and strong basal and subapical setae. Anepisternum with a series of 810 strong setae.
</paragraph>
<paragraph id="8BE0EE78F053FFCBFF3CFC87FEE22C76" blockId="15.[151,1436,386,882]" pageId="15" pageNumber="296">
<emphasis id="B92B326AF053FFCBFF3CFC87FF512C22" box="[199,255,783,806]" italics="true" pageId="15" pageNumber="296">Wing</emphasis>
. Yellow-brownish; costal spine little longer than the costal bristles; the transverse cross-vein dm-cu strongly curved; vein R 4+5 and vein M straight; cross vein dm-cu at middle of cell r4+5. Both calypters hyaline with yellow margins.
</paragraph>
<caption id="DF20BEF0F053FFCBFF6CF8EDFB8428DB" httpUri="https://zenodo.org/record/265625/files/figure.png" pageId="15" pageNumber="296" targetBox="[187,1424,934,1854]" targetPageId="15">
<paragraph id="8BE0EE78F053FFCBFF6CF8EDFB8428DB" blockId="15.[151,1436,1893,2015]" pageId="15" pageNumber="296">
<emphasis id="B92B326AF053FFCBFF6CF8EDFEBC287F" bold="true" box="[151,274,1893,1915]" pageId="15" pageNumber="296">FIGURE 7.</emphasis>
<taxonomicName id="4C5F95FBF053FFCBFEE3F8EDFDAC287F" authority="Stein" authorityName="Stein" box="[280,514,1893,1915]" class="Insecta" family="Lonchaeidae" genus="Helina" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="296" phylum="Arthropoda" rank="species" species="discolor">
<emphasis id="B92B326AF053FFCBFEE3F8EDFE18287F" box="[280,438,1893,1915]" italics="true" pageId="15" pageNumber="296">Helina discolor</emphasis>
(Stein)
</taxonomicName>
, male:
<emphasis id="B92B326AF053FFCBFDB0F8EDFDCA287F" bold="true" box="[587,612,1893,1915]" pageId="15" pageNumber="296">A.</emphasis>
Lateral view (scale bar: 1 mm).
<emphasis id="B92B326AF053FFCBFC50F8EDFC6C287F" bold="true" box="[939,962,1893,1915]" pageId="15" pageNumber="296">B.</emphasis>
Head, frontal view.
<emphasis id="B92B326AF053FFCBFB6BF8EDFB06287F" bold="true" box="[1168,1192,1893,1915]" pageId="15" pageNumber="296">C.</emphasis>
Thorax, dorsal view.
<emphasis id="B92B326AF053FFCBFA78F8EDFA32287F" bold="true" box="[1411,1436,1893,1915]" pageId="15" pageNumber="296">D.</emphasis>
Abdomen, dorsal view (scale bars: 0.5 mm).
<emphasis id="B92B326AF053FFCBFD9EF80EFDD22898" bold="true" box="[613,636,1926,1948]" pageId="15" pageNumber="296">E.</emphasis>
Sternite 5 (scale bar: 0.2 mm).
<emphasis id="B92B326AF053FFCBFC39F80EFC7D2898" bold="true" box="[962,979,1926,1948]" pageId="15" pageNumber="296">F.</emphasis>
Cercal plate, posterior view.
<emphasis id="B92B326AF053FFCBFAF8F80EFAB62898" bold="true" box="[1283,1304,1926,1948]" pageId="15" pageNumber="296">G.</emphasis>
Cercal plate and surstylus, lateral view (scale bar: 0.1 mm).
<emphasis id="B92B326AF053FFCBFD8CF82FFD3F28B9" bold="true" box="[631,657,1959,1981]" pageId="15" pageNumber="296">H.</emphasis>
Phallic complex, lateral view (scale bar: 0.1 mm). (Abbreviations: aed apod, aedeagal apodeme; distiph, distiphallus; epiph, epiphallus; gon, gonopodo; par, paramere.)
</paragraph>
</caption>
<paragraph id="8BE0EE78F04CFFD4FF3CFF11FDBE2EA9" blockId="16.[151,1437,151,1112]" pageId="16" pageNumber="297">
<emphasis id="B92B326AF04CFFD4FF3CFF11FF522FB4" box="[199,252,153,176]" italics="true" pageId="16" pageNumber="297">Legs</emphasis>
. Coxa and trochanter black. Fore femur and tibia black; mid and hind femora with basal half black and mid and hind tibia brownish-yellow. Fore femur with rows of dorsal, posterodorsal, and posteroventral setae; fore tibia with one posterior seta, four strong preapical setae. Mid femur with 34 setae in the basal third on ventral surface; and 3 preapical setae on anterodorsal to posterior surface; mid tibia with 2 anterodorsal setae, 34 posterior setae, and one strong posteroventral seta at middle; and 4 apical setae (anterodorsal, posteroventral, ventral, and anteroventral). Hind femur with anterodorsal row and 57 setae in the apical middle of anteroventral surface; hind tibia with 4 anteroventral setae, 4 anterodorsal setae and 67 posterodorsal setae, calcar absent. Fore claws and pulvilli longer than mid and hind.
</paragraph>
<paragraph id="8BE0EE78F04CFFD4FF3CFE3FFBC62EF0" blockId="16.[151,1437,151,1112]" pageId="16" pageNumber="297">
<emphasis id="B92B326AF04CFFD4FF3CFE3FFE9F2ED4" box="[199,305,439,464]" italics="true" pageId="16" pageNumber="297">Abdomen</emphasis>
(
<figureCitation id="1364F2FDF04CFFD4FEBFFE3FFE252ED4" box="[324,395,439,464]" captionStart="FIGURE 7" captionStartId="15.[151,250,1893,1915]" captionTargetBox="[187,1424,934,1854]" captionTargetId="figure@15.[161,1425,934,1871]" captionTargetPageId="15" captionText="FIGURE 7. Helina discolor (Stein), male: A. Lateral view (scale bar: 1 mm). B. Head, frontal view. C. Thorax, dorsal view. D. Abdomen, dorsal view (scale bars: 0.5 mm). E. Sternite 5 (scale bar: 0.2 mm). F. Cercal plate, posterior view. G. Cercal plate and surstylus, lateral view (scale bar: 0.1 mm). H. Phallic complex, lateral view (scale bar: 0.1 mm). (Abbreviations: aed apod, aedeagal apodeme; distiph, distiphallus; epiph, epiphallus; gon, gonopodo; par, paramere.)" httpUri="https://zenodo.org/record/265625/files/figure.png" pageId="16" pageNumber="297">Fig. 7</figureCitation>
D). Dark brown with brown-golden pollinosity. Sternite 5 longer than wide, with several strong setae on the apical third; posterior margin with two rounded processes (
<figureCitation id="1364F2FDF04CFFD4FBFFFE54FBE92EF0" box="[1028,1095,476,501]" captionStart="FIGURE 7" captionStartId="15.[151,250,1893,1915]" captionTargetBox="[187,1424,934,1854]" captionTargetId="figure@15.[161,1425,934,1871]" captionTargetPageId="15" captionText="FIGURE 7. Helina discolor (Stein), male: A. Lateral view (scale bar: 1 mm). B. Head, frontal view. C. Thorax, dorsal view. D. Abdomen, dorsal view (scale bars: 0.5 mm). E. Sternite 5 (scale bar: 0.2 mm). F. Cercal plate, posterior view. G. Cercal plate and surstylus, lateral view (scale bar: 0.1 mm). H. Phallic complex, lateral view (scale bar: 0.1 mm). (Abbreviations: aed apod, aedeagal apodeme; distiph, distiphallus; epiph, epiphallus; gon, gonopodo; par, paramere.)" httpUri="https://zenodo.org/record/265625/files/figure.png" pageId="16" pageNumber="297">Fig. 7</figureCitation>
E).
</paragraph>
<paragraph id="8BE0EE78F04CFFD4FF3CFE77FB122D80" blockId="16.[151,1437,151,1112]" pageId="16" pageNumber="297">
<taxonomicName id="4C5F95FBF04CFFD4FF3CFE77FEEF2D1C" box="[199,321,511,536]" class="Magnoliopsida" family="Combretaceae" genus="Terminalia" kingdom="Plantae" order="Myrtales" pageId="16" pageNumber="297" phylum="Tracheophyta" rank="genus">
<emphasis id="B92B326AF04CFFD4FF3CFE77FEEF2D1C" box="[199,321,511,536]" italics="true" pageId="16" pageNumber="297">Terminalia</emphasis>
</taxonomicName>
. Cercal plate with a similar length and width; with a distal incision, and with two protuberances at the apex (
<figureCitation id="1364F2FDF04CFFD4FEF3FDACFEE92D38" box="[264,327,548,573]" captionStart="FIGURE 7" captionStartId="15.[151,250,1893,1915]" captionTargetBox="[187,1424,934,1854]" captionTargetId="figure@15.[161,1425,934,1871]" captionTargetPageId="15" captionText="FIGURE 7. Helina discolor (Stein), male: A. Lateral view (scale bar: 1 mm). B. Head, frontal view. C. Thorax, dorsal view. D. Abdomen, dorsal view (scale bars: 0.5 mm). E. Sternite 5 (scale bar: 0.2 mm). F. Cercal plate, posterior view. G. Cercal plate and surstylus, lateral view (scale bar: 0.1 mm). H. Phallic complex, lateral view (scale bar: 0.1 mm). (Abbreviations: aed apod, aedeagal apodeme; distiph, distiphallus; epiph, epiphallus; gon, gonopodo; par, paramere.)" httpUri="https://zenodo.org/record/265625/files/figure.png" pageId="16" pageNumber="297">Fig 7</figureCitation>
F). Surstylus short than cercal plate, broad, and curving inward distally (
<figureCitation id="1364F2FDF04CFFD4FB71FDACFB772D38" box="[1162,1241,548,573]" captionStart="FIGURE 7" captionStartId="15.[151,250,1893,1915]" captionTargetBox="[187,1424,934,1854]" captionTargetId="figure@15.[161,1425,934,1871]" captionTargetPageId="15" captionText="FIGURE 7. Helina discolor (Stein), male: A. Lateral view (scale bar: 1 mm). B. Head, frontal view. C. Thorax, dorsal view. D. Abdomen, dorsal view (scale bars: 0.5 mm). E. Sternite 5 (scale bar: 0.2 mm). F. Cercal plate, posterior view. G. Cercal plate and surstylus, lateral view (scale bar: 0.1 mm). H. Phallic complex, lateral view (scale bar: 0.1 mm). (Abbreviations: aed apod, aedeagal apodeme; distiph, distiphallus; epiph, epiphallus; gon, gonopodo; par, paramere.)" httpUri="https://zenodo.org/record/265625/files/figure.png" pageId="16" pageNumber="297">Figs. 7</figureCitation>
FG). Aedeagus with aedeagal apodeme curved and strongly sclerotized; epiphalus sclerotized, paramere slightly bent downward with setula; gonopod with short setulae; and distiphallus tubular, slightly sclerotized (
<figureCitation id="1364F2FDF04CFFD4FBAFFDE4FB362D80" box="[1108,1176,620,645]" captionStart="FIGURE 7" captionStartId="15.[151,250,1893,1915]" captionTargetBox="[187,1424,934,1854]" captionTargetId="figure@15.[161,1425,934,1871]" captionTargetPageId="15" captionText="FIGURE 7. Helina discolor (Stein), male: A. Lateral view (scale bar: 1 mm). B. Head, frontal view. C. Thorax, dorsal view. D. Abdomen, dorsal view (scale bars: 0.5 mm). E. Sternite 5 (scale bar: 0.2 mm). F. Cercal plate, posterior view. G. Cercal plate and surstylus, lateral view (scale bar: 0.1 mm). H. Phallic complex, lateral view (scale bar: 0.1 mm). (Abbreviations: aed apod, aedeagal apodeme; distiph, distiphallus; epiph, epiphallus; gon, gonopodo; par, paramere.)" httpUri="https://zenodo.org/record/265625/files/figure.png" pageId="16" pageNumber="297">Fig. 7</figureCitation>
H).
</paragraph>
</subSubSection>
<subSubSection id="C345BDF3F04CFFD4FF3CFD07FBF52C3C" pageId="16" pageNumber="297" type="materials_examined">
<paragraph id="8BE0EE78F04CFFD4FF3CFD07FBF52C3C" blockId="16.[151,1437,151,1112]" pageId="16" pageNumber="297">
<materialsCitation id="3B37E425F04CFFD4FF3CFD07FE352DC8" ID-GBIF-Occurrence="1291551032" collectingDate="2011-02" collectionCode="MACN" collectorName="Mariluis" country="Argentina" location="Cerro Chapelco" municipality="Material" pageId="16" pageNumber="297" specimenCount="3" specimenCount-male="3" stateProvince="Neuquen">
<emphasis id="B92B326AF04CFFD4FF3CFD07FD4A2DAC" bold="true" box="[199,740,655,680]" pageId="16" pageNumber="297">
<collectingMunicipality id="6B847402F04CFFD4FF3CFD07FE812DAC" box="[199,303,655,680]" pageId="16" pageNumber="297">Material</collectingMunicipality>
examined.
<collectingCountry id="F348AEE8F04CFFD4FE47FD07FDC22DAC" box="[444,620,655,680]" name="Argentina" pageId="16" pageNumber="297">ARGENTINA</collectingCountry>
:
<collectingRegion id="499B209AF04CFFD4FD82FD07FD4A2DAC" box="[633,740,655,680]" country="Argentina" name="Neuquen" pageId="16" pageNumber="297">Neuquén</collectingRegion>
</emphasis>
:
<specimenCount id="9D5925F1F04CFFD4FD0EFD18FCFE2DAC" box="[757,848,655,680]" pageId="16" pageNumber="297" type="male">3 males</specimenCount>
,
<location id="8E80B8A3F04CFFD4FCA5FD18FBBA2DAC" LSID="urn:lsid:plazi:treatment:03F65F6EF053FFD4FF6CFF1FFDA72B53:8E80B8A3F04CFFD4FCA5FD18FBBA2DAC" box="[862,1044,655,680]" country="Argentina" municipality="Material" name="Cerro Chapelco" pageId="16" pageNumber="297" stateProvince="Neuquen">Cerro Chapelco</location>
, -40.197050, -71.298453,
<date id="FFE1C8B8F04CFFD4FAB9FD18FA392DAC" box="[1346,1431,656,680]" pageId="16" pageNumber="297" value="2011-02">
<collectingDate id="EFA53150F04CFFD4FAB9FD18FA392DAC" box="[1346,1431,656,680]" pageId="16" pageNumber="297" value="2011-02">II-2011</collectingDate>
</date>
,
<collectorName id="26AA8BAEF04CFFD4FF6CFD3CFF582DC9" box="[151,246,692,717]" pageId="16" pageNumber="297">Mariluis</collectorName>
leg. (
<collectionCode id="ED4E76BDF04CFFD4FEC3FD3CFE3D2DC8" box="[312,403,692,716]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="16" pageNumber="297">MACN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B37E425F04CFFD4FE53FD3CFA362DC8" ID-GBIF-Occurrence="1291551029" box="[424,1432,692,717]" collectingDate="1983-03-21" collectingDateMax="1983-03-28" collectingDateMin="1983-03-21" collectionCode="IFML" collectorName="Gentili" country="Argentina" location="Cerro Chapelco" pageId="16" pageNumber="297" specimenCount="1" specimenCount-female="1" stateProvince="Neuquen">
<specimenCount id="9D5925F1F04CFFD4FE53FD3CFDA02DC9" box="[424,526,692,717]" pageId="16" pageNumber="297" type="female">1 female</specimenCount>
,
<location id="8E80B8A3F04CFFD4FDE0FD3CFD7F2DC9" LSID="urn:lsid:plazi:treatment:03F65F6EF053FFD4FF6CFF1FFDA72B53:8E80B8A3F04CFFD4FDE0FD3CFD7F2DC9" box="[539,721,692,717]" country="Argentina" name="Cerro Chapelco" pageId="16" pageNumber="297" stateProvince="Neuquen">Cerro Chapelco</location>
, -40.197050, -71.298453,
<date id="FFE1C8B8F04CFFD4FC05FD3CFB072DC8" box="[1022,1193,692,716]" pageId="16" pageNumber="297" value="1983-03-21" valueMax="1983-03-28" valueMin="1983-03-21">
<collectingDate id="EFA53150F04CFFD4FC05FD3CFB072DC8" box="[1022,1193,692,716]" pageId="16" pageNumber="297" value="1983-03-21" valueMax="1983-03-28" valueMin="1983-03-21">21-28-III-1983</collectingDate>
</date>
,
<collectorName id="26AA8BAEF04CFFD4FB4DFD3CFAAA2DC9" box="[1206,1284,692,717]" pageId="16" pageNumber="297">Gentili</collectorName>
leg. (
<collectionCode id="ED4E76BDF04CFFD4FABCFD3CFA3E2DC8" box="[1351,1424,692,716]" pageId="16" pageNumber="297">IFML</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B37E425F04CFFD4FF6CFD5FFB682DF4" ID-GBIF-Occurrence="1291551028" box="[151,1222,727,752]" collectingDate="1994-10" collectionCode="MACN" collectorName="El Calafate &amp; Mariluis" country="Argentina" location="El Calafate" pageId="16" pageNumber="297" specimenCount="1" specimenCount-male="1" stateProvince="Santa Cruz">
<emphasis id="B92B326AF04CFFD4FF6CFD5FFE822DF4" bold="true" box="[151,300,727,752]" pageId="16" pageNumber="297">
<collectingRegion id="499B209AF04CFFD4FF6CFD5FFE862DF4" box="[151,296,727,752]" country="Argentina" name="Santa Cruz" pageId="16" pageNumber="297">Santa Cruz</collectingRegion>
:
</emphasis>
<specimenCount id="9D5925F1F04CFFD4FEC3FD50FE242DF4" box="[312,394,727,752]" pageId="16" pageNumber="297" type="male">1 male</specimenCount>
,
<location id="8E80B8A3F04CFFD4FE61FD5FFDB12DF4" LSID="urn:lsid:plazi:treatment:03F65F6EF053FFD4FF6CFF1FFDA72B53:8E80B8A3F04CFFD4FE61FD5FFDB12DF4" box="[410,543,727,752]" country="Argentina" name="El Calafate" pageId="16" pageNumber="297" stateProvince="Santa Cruz">
<collectorName id="26AA8BAEF04CFFD4FE61FD5FFDB12DF4" box="[410,543,727,752]" pageId="16" pageNumber="297">El Calafate</collectorName>
</location>
, -50.321674, -72.263421,
<date id="FFE1C8B8F04CFFD4FCAFFD50FC032DF4" box="[852,941,728,752]" pageId="16" pageNumber="297" value="1994-10">
<collectingDate id="EFA53150F04CFFD4FCAFFD50FC032DF4" box="[852,941,728,752]" pageId="16" pageNumber="297" value="1994-10">X-1994</collectingDate>
</date>
,
<collectorName id="26AA8BAEF04CFFD4FC47FD5FFBB52DF4" box="[956,1051,727,752]" pageId="16" pageNumber="297">Mariluis</collectorName>
leg. (
<collectionCode id="ED4E76BDF04CFFD4FB98FD50FB102DF4" box="[1123,1214,728,752]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="16" pageNumber="297">MACN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B37E425F04CFFD4FB2EFD50FAB92C10" ID-GBIF-Occurrence="1291551043" collectingDate="1995-02" collectionCode="MACN" collectorName="PN Los Glaciares &amp; Mariluis" country="Argentina" location="Rio Mitre" pageId="16" pageNumber="297" specimenCount="3" specimenCount-male="3" stateProvince="Magallanes">
<specimenCount id="9D5925F1F04CFFD4FB2EFD50FA9D2DF4" box="[1237,1331,727,752]" pageId="16" pageNumber="297" type="male">3 males</specimenCount>
,
<collectorName id="26AA8BAEF04CFFD4FAB8FD50FEAA2C11" pageId="16" pageNumber="297">PN Los Glaciares</collectorName>
, Península
<collectingRegion id="499B209AF04CFFD4FE7EFD74FDA72C11" box="[389,521,764,789]" country="Chile" name="Magallanes y de la Antartica Chilena" pageId="16" pageNumber="297">Magallanes</collectingRegion>
,
<location id="8E80B8A3F04CFFD4FDEEFD74FD292C11" LSID="urn:lsid:plazi:treatment:03F65F6EF053FFD4FF6CFF1FFDA72B53:8E80B8A3F04CFFD4FDEEFD74FD292C11" box="[533,647,764,789]" country="Argentina" name="Rio Mitre" pageId="16" pageNumber="297" stateProvince="Magallanes">Río Mitre</location>
, -50.418967, -72.742651,
<date id="FFE1C8B8F04CFFD4FC49FD74FBA62C11" box="[946,1032,764,789]" pageId="16" pageNumber="297" value="1995-02">
<collectingDate id="EFA53150F04CFFD4FC49FD74FBA62C11" box="[946,1032,764,789]" pageId="16" pageNumber="297" value="1995-02">II-1995</collectingDate>
</date>
,
<collectorName id="26AA8BAEF04CFFD4FBEEFD74FBDA2C11" box="[1045,1140,764,789]" pageId="16" pageNumber="297">Mariluis</collectorName>
leg. (
<collectionCode id="ED4E76BDF04CFFD4FB4EFD74FAA12C10" box="[1205,1295,764,788]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="16" pageNumber="297">MACN</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B37E425F04CFFD4FAD8FD74FBF92C3C" ID-GBIF-Occurrence="1291551047" collectingDate="1998-02" collectionCode="MACN" collectorName="Mariluis" country="Argentina" location="Rio Grande" pageId="16" pageNumber="297" specimenCount="1" specimenCount-male="1" stateProvince="Tierra del Fuego">
<emphasis id="B92B326AF04CFFD4FAD8FD74FF4E2C3C" bold="true" pageId="16" pageNumber="297">
<collectingRegion id="499B209AF04CFFD4FAD8FD74FF4E2C3C" country="Argentina" name="Tierra del Fuego - Antartida e Islas del Atlantico Sur" pageId="16" pageNumber="297">Tierra del Fuego</collectingRegion>
</emphasis>
:
<specimenCount id="9D5925F1F04CFFD4FF15FCA8FE922C3C" box="[238,316,799,824]" pageId="16" pageNumber="297" type="male">1 male</specimenCount>
,
<location id="8E80B8A3F04CFFD4FEB3FCA8FE622C3C" LSID="urn:lsid:plazi:treatment:03F65F6EF053FFD4FF6CFF1FFDA72B53:8E80B8A3F04CFFD4FEB3FCA8FE622C3C" box="[328,460,799,824]" country="Argentina" name="Rio Grande" pageId="16" pageNumber="297" stateProvince="Tierra del Fuego">Río Grande</location>
, -53.785843, -67.702577,
<date id="FFE1C8B8F04CFFD4FD0FFCA8FCE52C3C" box="[756,843,800,824]" pageId="16" pageNumber="297" value="1998-02">
<collectingDate id="EFA53150F04CFFD4FD0FFCA8FCE52C3C" box="[756,843,800,824]" pageId="16" pageNumber="297" value="1998-02">II-1998</collectingDate>
</date>
,
<collectorName id="26AA8BAEF04CFFD4FCADFC97FC1B2C3C" box="[854,949,799,824]" pageId="16" pageNumber="297">Mariluis</collectorName>
leg. (
<collectionCode id="ED4E76BDF04CFFD4FC0FFCA8FBE12C3C" box="[1012,1103,800,824]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="16" pageNumber="297">MACN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C345BDF3F04CFFD4FF3CFCCCFDD52C84" pageId="16" pageNumber="297" type="distribution">
<paragraph id="8BE0EE78F04CFFD4FF3CFCCCFDD52C84" blockId="16.[151,1437,151,1112]" pageId="16" pageNumber="297">
<emphasis id="B92B326AF04CFFD4FF3CFCCCFEF72C59" bold="true" box="[199,345,836,861]" pageId="16" pageNumber="297">Distribution</emphasis>
(
<figureCitation id="1364F2FDF04CFFD4FE93FCCCFE052C58" box="[360,427,836,861]" captionStart="FIGURE 9" captionStartId="18.[151,250,1937,1959]" captionTargetBox="[168,1423,829,1910]" captionTargetId="figure@18.[163,1423,829,1916]" captionTargetPageId="18" captionText="FIGURE 9. Geographical distribution. A. Helina araucana sp. nov. B. Helina australis Carvalho &amp; Pont. C. Helina bigoti Malloch. D. Helina chilensis Malloch. E. Helina connexa Malloch. F. Helina discolor (Stein). G. Helina dorada sp. nov. (Green area: Andean-Patagonian forests; orange circle: new records; blue triangle: previous distribution.)" httpUri="https://zenodo.org/record/265627/files/figure.png" pageId="16" pageNumber="297">Fig. 9</figureCitation>
F).
<collectingCountry id="F348AEE8F04CFFD4FE2AFCCCFDD72C58" box="[465,633,836,860]" name="Argentina" pageId="16" pageNumber="297">ARGENTINA</collectingCountry>
:
<collectingRegion id="499B209AF04CFFD4FD78FCCCFD462C58" box="[643,744,836,860]" country="Argentina" name="Neuquen" pageId="16" pageNumber="297">Neuquén</collectingRegion>
(new record),
<collectingRegion id="499B209AF04CFFD4FC77FCCCFBAB2C59" box="[908,1029,836,861]" country="Argentina" name="Rio Negro" pageId="16" pageNumber="297">Río Negro</collectingRegion>
,
<collectingRegion id="499B209AF04CFFD4FBF4FCCCFB252C58" box="[1039,1163,836,860]" country="Argentina" name="Santa Cruz" pageId="16" pageNumber="297">Santa Cruz</collectingRegion>
(new record),
<collectingRegion id="499B209AF04CFFD4FAD4FCCCFF732C84" country="Argentina" name="Tierra del Fuego - Antartida e Islas del Atlantico Sur" pageId="16" pageNumber="297">Tierra del Fuego</collectingRegion>
(new record).
<collectingCountry id="F348AEE8F04CFFD4FE79FCE0FE602C7B" box="[386,462,872,895]" name="Peru" pageId="16" pageNumber="297">PERU</collectingCountry>
:
<collectingRegion id="499B209AF04CFFD4FE23FCE0FD8C2C84" box="[472,546,872,896]" country="Peru" name="Cusco" pageId="16" pageNumber="297">Cuzco</collectingRegion>
, Potosí.
</paragraph>
</subSubSection>
<subSubSection id="C345BDF3F04CFFD4FF3CFC04FE8B2B14" pageId="16" pageNumber="297" type="discussion">
<paragraph id="8BE0EE78F04CFFD4FF3CFC04FE8B2B14" blockId="16.[151,1437,151,1112]" pageId="16" pageNumber="297">
<emphasis id="B92B326AF04CFFD4FF3CFC04FE9A2CA1" bold="true" box="[199,308,908,933]" pageId="16" pageNumber="297">Remarks</emphasis>
.
<bibRefCitation id="EFCE9389F04CFFD4FEBCFC04FE782CA0" author="Stein" box="[327,470,908,933]" pageId="16" pageNumber="297" refString="Stein, P. (1911) Die von Schnuse in Sudamerika gefangenen Anthomyiden. Archiv fur Naturgeschichte, 77, 61 - 189." type="journal article" year="1911">Stein (1911)</bibRefCitation>
described
<taxonomicName id="4C5F95FBF04CFFD4FDA1FC04FCB32CA1" box="[602,797,908,933]" class="Insecta" family="Muscidae" genus="Mydaea" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="297" phylum="Arthropoda" rank="species" species="discolor">
<emphasis id="B92B326AF04CFFD4FDA1FC04FCB32CA1" box="[602,797,908,933]" italics="true" pageId="16" pageNumber="297">Mydaea discolor</emphasis>
</taxonomicName>
with male and females specimens from Cuzco, Perú. Subsecuently,
<bibRefCitation id="EFCE9389F04CFFD4FEC2FC27FE472CCC" author="Malloch" box="[313,489,943,968]" pageId="16" pageNumber="297" refString="Malloch, J. R. (1934) Muscidae. In: Diptera of Patagonia and South Chile. Part 7, Fascicle 2. British Museum (Natural History), London, pp. 171 - 346." type="book chapter" year="1934">Malloch (1934)</bibRefCitation>
recorded this species from Bariloche, Argentina. We provided a redescripton of the male, and for the first time, the description the male
<taxonomicName id="4C5F95FBF04CFFD4FD14FC5CFCCA2CE9" box="[751,868,980,1005]" class="Magnoliopsida" family="Combretaceae" genus="Terminalia" kingdom="Plantae" order="Myrtales" pageId="16" pageNumber="310" phylum="Tracheophyta" rank="genus">terminalia</taxonomicName>
. No females were collected. (see comments
<collectedFrom id="3843AD85F04CFFD4FAA6FC5DFEBB2B14" pageId="16" pageNumber="297">
on
<taxonomicName id="4C5F95FBF04CFFD4FA7AFC5DFEBB2B14" class="Insecta" family="Lonchaeidae" genus="Helina" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="297" phylum="Arthropoda" rank="species" species="neosimplex">
<emphasis id="B92B326AF04CFFD4FA7AFC5DFEBB2B14" italics="true" pageId="16" pageNumber="297">H. neosimplex</emphasis>
</taxonomicName>
</collectedFrom>
).
</paragraph>
</subSubSection>
<subSubSection id="C345BDF3F04CFFD4FF3CFB94FDA72B53" pageId="16" pageNumber="297" type="biology_ecology">
<paragraph id="8BE0EE78F04CFFD4FF3CFB94FDA72B53" blockId="16.[151,1437,151,1112]" pageId="16" pageNumber="297">
<emphasis id="B92B326AF04CFFD4FF3CFB94FEB12B31" bold="true" box="[199,287,1052,1077]" pageId="16" pageNumber="297">Biology</emphasis>
. The specimens captured by JCM in Santa Cruz province and Tierra del Fuego province were collected over rotten meat with a
<collectingMethod id="521E966FF04CFFD4FE59FBB7FDAB2B53" box="[418,517,1087,1112]" pageId="16" pageNumber="297">hand net</collectingMethod>
.
</paragraph>
</subSubSection>
</treatment>
</document>