treatments-xml/data/65/88/A2/6588A24D1B2659B9ADD46C0850DDA833.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1124.91156" ID-Pensoft-Pub="1313-2970-1124-161" ID-Pensoft-UUID="9EED96A198EF5BE1B29FAA9DA80C5F36" ID-ZooBank="517DD5E896D64D5A99D9086809DB6110" ModsDocID="1313-2970-1124-161" checkinTime="1666056858948" checkinUser="pensoft" docAuthor="Bento, Matheus, Jameson, Mary Liz &amp; Seidel, Matthias" docDate="2022" docId="6588A24D1B2659B9ADD46C0850DDA833" docLanguage="en" docName="ZooKeys 1124: 161-189" docOrigin="ZooKeys 1124" docPubDate="2022-10-17" docSource="http://dx.doi.org/10.3897/zookeys.1124.91156" docTitle="Macraspis maculata Burmeister 1844" docType="treatment" docVersion="1" id="9EED96A198EF5BE1B29FAA9DA80C5F36" lastPageNumber="161" masterDocId="9EED96A198EF5BE1B29FAA9DA80C5F36" masterDocTitle="New species and illustrated key of Macraspis (Scarabaeidae, Rutelinae, Rutelini) from the Amazon biome of Brazil" masterLastPageNumber="189" masterPageNumber="161" pageNumber="161" updateTime="1666056858948" updateUser="pensoft">
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<mods:titleInfo>
<mods:title>New species and illustrated key of Macraspis (Scarabaeidae, Rutelinae, Rutelini) from the Amazon biome of Brazil</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Bento, Matheus</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-9846-9728</mods:nameIdentifier>
<mods:affiliation>Instituto Nacional de Pesquisas da Amazonia (INPA), Coordenacao de Biodiversidade, Laboratorio de Sistematica e Ecologia de Coleoptera (LASEC), Manaus, Brazil</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jameson, Mary Liz</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-3076-2028</mods:nameIdentifier>
<mods:affiliation>Wichita State University, Wichita, Kansas, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Seidel, Matthias</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-4913-8778</mods:nameIdentifier>
<mods:affiliation>Naturhistorisches Museum Wien, Zweite Zoologische Abteilung, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
<mods:nameIdentifier type="email">matth.seidel@gmail.com</mods:nameIdentifier>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-10-17</mods:number>
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<mods:detail type="volume">
<mods:number>1124</mods:number>
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<mods:extent unit="page">
<mods:start>161</mods:start>
<mods:end>189</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1124.91156</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1124.91156</mods:identifier>
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<paragraph pageId="0" pageNumber="161">
<taxonomicName LSID="6588A24D-1B26-59B9-ADD4-6C0850DDA833" authority="Burmeister, 1844" authorityName="Burmeister" authorityYear="1844" class="Insecta" family="Scarabaeidae" genus="Macraspis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macraspis maculata" order="Coleoptera" pageId="0" pageNumber="161" phylum="Arthropoda" rank="species" species="maculata">Macraspis maculata Burmeister, 1844</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="161" type="description">
<paragraph pageId="0" pageNumber="161">
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Macraspis spp. (A-O). Dorsal view of head of A M. oblonga B M. olivieri and C M. xanthosticta. Lateral view of mesothorax of D M. festiva E M. maculata and F M. c. chloraspis. Female pygidium of G M. oblonga H M. olivieri and I M. morio. Dorsal view of female abdomen of J M. p. pseudochrysis and K M. lepiouffi paratype. Metatarsus of L M. morio and O M. olivieri. Female last abdominal ventrites of M M. morio and N M. p. pseudochrysis. Scale bars: 0.5 mm (A-C), 1 mm (D-O)." figureDoi="10.3897/zookeys.1124.91156.figure2" httpUri="https://binary.pensoft.net/fig/758064" pageId="0" pageNumber="161">Figs 2E</figureCitation>
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Macraspis maculata Burmeister, 1844, lectotype male (A-D) A dorsal view B ventral view C aedeagus D lectotype label. M. maculata crosarai Soula, 1998, holotype male (E-H) E dorsal view F ventral view G aedeagus H labels." figureDoi="10.3897/zookeys.1124.91156.figure8" httpUri="https://binary.pensoft.net/fig/758070" pageId="0" pageNumber="161">, 8A-D</figureCitation>
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Macraspis opala sp. nov. (A-H) and Macraspis maculata Burmeister (I-M): Macraspis opala sp. nov., holotype male A dorsal view B ventral view C pygidium in caudal view D spiculum gastrale E aedeagus, dorsal view F aedeagus, caudal view G lateral view, paramera (of paratype) and H caudal view. Macraspis maculata Burmeister, 1844 from Linhares, Espirito Santo state, Brazil, I spiculum gastrale J aedeagus in dorsal view K caudal view and L lateral view M pygidium in caudal view. Scale bars: 2 mm (A, B); 1 mm (C, M); 0.5 mm (E, G, J, L); 0.2 mm (F, H, K)." figureDoi="10.3897/zookeys.1124.91156.figure9" httpUri="https://binary.pensoft.net/fig/758071" pageId="0" pageNumber="161">, 9I-M</figureCitation>
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Distribution of Brazilian species of Macraspis treated in this publication. The Brazilian Amazon biome is colored lime green. Brazilian states are abbreviated as follows: AC = Acre, AM = Amazonas, AP = Amapa, BA = Bahia, ES = Espirito Santo, GO = Goias, MA = Maranhao, MG = Minas Gerais, MT = Mato Grosso = PA = Piaui, RO = Rondonia, RJ = Rio de Janiero, RR = Roraima, SP = Sao Paulo, TO = Tocantins." figureDoi="10.3897/zookeys.1124.91156.figure12" httpUri="https://binary.pensoft.net/fig/758074" pageId="0" pageNumber="161">, 12</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="161" type="reference_group">
<paragraph pageId="0" pageNumber="161">
<taxonomicName authorityName="Burmeister" authorityYear="1844" class="Insecta" family="Scarabaeidae" genus="Macraspis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macraspis maculata" order="Coleoptera" pageId="0" pageNumber="161" phylum="Arthropoda" rank="species" species="maculata">Macraspis maculata</taxonomicName>
Burmeister, 1844: 351 (original description); 
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.101716" author="Lucas, PH" journalOrPublisher="Bertrand, Paris" pageId="0" pageNumber="161" refId="B14" refString="Lucas, PH, 1857. Entomologie. In: Lucas MH (Ed.) Animaux Nouveaux ou Rares Recueillis pendant l'Expedition dans les parties centrales de l'Amerique du Sud, de Rio de Janeiro a Lima, et de Lima au Para; executee par ordre du Government francais pendant les annees 1843 a 1847, sous la direction du Comte Francis de Castelnau. P. Bertrand, Paris, DOI: https://doi.org/10.5962/bhl.title.101716" title="Entomologie. In: Lucas MH (Ed.) Animaux Nouveaux ou Rares Recueillis pendant l'Expedition dans les parties centrales de l'Amerique du Sud, de Rio de Janeiro a Lima, et de Lima au Para; executee par ordre du Government francais pendant les annees 1843 a 1847, sous la direction du Comte Francis de Castelnau. P." url="https://doi.org/10.5962/bhl.title.101716" year="1857">Lucas 1857</bibRefCitation>
: 124, pl. 7 fig. 6; 
<bibRefCitation author="Ohaus, F" journalOrPublisher="Stettiner Entomologische Zeitung" pageId="0" pageNumber="161" pagination="42 - 63" refId="B18" refString="Ohaus, F, 1898. Ruteliden der neuen Welt. Stettiner Entomologische Zeitung 59: 42 - 63" title="Ruteliden der neuen Welt." volume="59" year="1898">Ohaus 1898</bibRefCitation>
: 52; 
<bibRefCitation author="Ohaus, F" journalOrPublisher="Coleopterorum Catalogus" pageId="0" pageNumber="161" pagination="1 - 241" refId="B20" refString="Ohaus, F, 1918. Scarabaeidae: Euchirinae, Phaenomerinae, Rutelinae. Coleopterorum Catalogus 20: 1 - 241" title="Scarabaeidae: Euchirinae, Phaenomerinae, Rutelinae." volume="20" year="1918">1918</bibRefCitation>
: 55; 
<bibRefCitation author="Ohaus, F" journalOrPublisher="Coleopterorum Catalogus" pageId="0" pageNumber="161" refId="B21" refString="Ohaus, F, 1934. Coleoptera lamellicornia, Fam. Scarabaeidae, Subfam. Rutelinae. T.1: Tribus Rutelini. Genera Insectorum Fasc. 199A: 1-172." title="Coleoptera lamellicornia, Fam. Scarabaeidae, Subfam. Rutelinae. T. 1: Tribus Rutelini. Genera Insectorum Fasc. 199 A: 1 - 172." year="1934">1934</bibRefCitation>
: 154; 
<bibRefCitation DOI="https://doi.org/10.5479/si.03629236.185.2" author="Blackwelder, RE" journalOrPublisher="Bulletin - United States National Museum" pageId="0" pageNumber="161" pagination="189 - 265" refId="B3" refString="Blackwelder, RE, 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Bulletin - United States National Museum 185: 189 - 265, DOI: https://doi.org/10.5479/si.03629236.185.2" title="Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America." url="https://doi.org/10.5479/si.03629236.185.2" volume="185" year="1944">Blackwelder 1944</bibRefCitation>
: 242; 
<bibRefCitation author="Machatschke, JW" journalOrPublisher="Coleopterorum Catalogus Supplementa" pageId="0" pageNumber="161" pagination="1 - 361" refId="B15" refString="Machatschke, JW, 1972. Scarabaeoidea: Melolonthidae, Rutelinae. Coleopterorum Catalogus Supplementa 66: 1 - 361" title="Scarabaeoidea: Melolonthidae, Rutelinae." volume="66" year="1972">Machatschke 1972</bibRefCitation>
: 71; 
<bibRefCitation author="Soula, M" journalOrPublisher="Hillside Books, Canterbury" pageId="0" pageNumber="161" refId="B25" refString="Soula, M, 1998. Les Coleopteres du Monde, 26.1: Rutelini 2. Revision des Anthicheirina 1. Hillside Books, Canterbury" title="Les Coleopteres du Monde, 26.1: Rutelini 2. Revision des Anthicheirina 1." year="1998">Soula 1998</bibRefCitation>
: 37; 
<bibRefCitation author="Soula, M" journalOrPublisher="Hillside Books, Canterbury" pageId="0" pageNumber="161" refId="B27" refString="Soula, M, 2003. Les Coleopteres du Monde, 29: Rutelini 3. Pseudoptenomela - Paraptenomela - Exoptenomela - Ptenomela - Calomacraspis - Paramacraspis - Paradorysthetus - Macraspis - Pseudorysthetus - Dorysthetus - Anticheiroides - Anticheira - Pseudoanthicheiroides. Hillside Books, Canterbury" title="Les Coleopteres du Monde, 29: Rutelini 3. Pseudoptenomela - Paraptenomela - Exoptenomela - Ptenomela - Calomacraspis - Paramacraspis - Paradorysthetus - Macraspis - Pseudorysthetus - Dorysthetus - Anticheiroides - Anticheira - Pseudoanthicheiroides." year="2003">2003</bibRefCitation>
: 39; 
<bibRefCitation author="Krajcik, M" journalOrPublisher="Insecta Mundi" pageId="0" pageNumber="161" refId="B13" refString="Krajcik, M, 2007. Checklist of the Scarabaeoidea of the World. 2. Rutelinae (Coleoptera: Scarabaeidae: Rutelinae). Animma.X. Supplement 4. 2nd Edn. Plzen, 142 pp." title="Checklist of the Scarabaeoidea of the World. 2. Rutelinae (Coleoptera: Scarabaeidae: Rutelinae). Animma. X. Supplement 4. 2 nd Edn. Plzen, 142 pp." year="2007">Krajcik 2007</bibRefCitation>
: 80; 
<bibRefCitation author="Brule, S" journalOrPublisher="SEAG, PAG" pageId="0" pageNumber="161" refId="B4" refString="Brule, S, Touroult, J, Poirier, E, Dalens, PH, 2014. Resultats de l'etude-inventaire entomologique du site de la Montagne Pelee, Sauel (Guyane), 2010-2012. Rapport de la Societe entomologique Antilles-Guyane. SEAG, PAG" title="Resultats de l'etude-inventaire entomologique du site de la Montagne Pelee, Sauel (Guyane), 2010 - 2012. Rapport de la Societe entomologique Antilles-Guyane." year="2014">
<normalizedToken originalValue="Brûlé">Brule</normalizedToken>
et al. 2014
</bibRefCitation>
: 182; 
<bibRefCitation DOI="https://doi.org/10.2317/kent-88-02-186-207.1" author="Ratcliffe, BC" journalOrPublisher="Journal of the Kansas Entomological Society" pageId="0" pageNumber="161" pagination="186 - 207" refId="B23" refString="Ratcliffe, BC, Jameson, ML, Figueroa, L, Cave, RD, Paulsen, MJ, Cano, EB, Beza-Beza, C, Jimenez-Ferbans, L, Reyes-Castillo, R, 2015. Beetles (Coleoptera) of Peru: A survey of families. Scarabaeoidea. Journal of the Kansas Entomological Society 88 (2): 186 - 207, DOI: https://doi.org/10.2317/kent-88-02-186-207.1" title="Beetles (Coleoptera) of Peru: A survey of families. Scarabaeoidea." url="https://doi.org/10.2317/kent-88-02-186-207.1" volume="88" year="2015">Ratcliffe et al. 2015</bibRefCitation>
: 199; 
<bibRefCitation author="Hielkema, AJ" journalOrPublisher="Insecta Mundi" pageId="0" pageNumber="161" pagination="1 - 306" refId="B9" refString="Hielkema, AJ, Hielkema, MA, 2019. An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas. Insecta Mundi 0732: 1 - 306" title="An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas." volume="0732" year="2019">Hielkema and Hielkema 2019</bibRefCitation>
: 143.
</paragraph>
<paragraph pageId="0" pageNumber="161">
<taxonomicName authorityName="Soula" authorityYear="1998" class="Insecta" family="Scarabaeidae" genus="Macraspis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macraspis maculata subsp. crosarai" order="Coleoptera" pageId="0" pageNumber="161" phylum="Arthropoda" rank="subSpecies" species="maculata" subSpecies="crosarai">Macraspis maculata crosarai</taxonomicName>
Soula, 1998: 38 (original description); 
<bibRefCitation author="Soula, M" journalOrPublisher="Hillside Books, Canterbury" pageId="0" pageNumber="161" refId="B27" refString="Soula, M, 2003. Les Coleopteres du Monde, 29: Rutelini 3. Pseudoptenomela - Paraptenomela - Exoptenomela - Ptenomela - Calomacraspis - Paramacraspis - Paradorysthetus - Macraspis - Pseudorysthetus - Dorysthetus - Anticheiroides - Anticheira - Pseudoanthicheiroides. Hillside Books, Canterbury" title="Les Coleopteres du Monde, 29: Rutelini 3. Pseudoptenomela - Paraptenomela - Exoptenomela - Ptenomela - Calomacraspis - Paramacraspis - Paradorysthetus - Macraspis - Pseudorysthetus - Dorysthetus - Anticheiroides - Anticheira - Pseudoanthicheiroides." year="2003">Soula 2003</bibRefCitation>
: 39; 
<bibRefCitation author="Krajcik, M" journalOrPublisher="Insecta Mundi" pageId="0" pageNumber="161" refId="B13" refString="Krajcik, M, 2007. Checklist of the Scarabaeoidea of the World. 2. Rutelinae (Coleoptera: Scarabaeidae: Rutelinae). Animma.X. Supplement 4. 2nd Edn. Plzen, 142 pp." title="Checklist of the Scarabaeoidea of the World. 2. Rutelinae (Coleoptera: Scarabaeidae: Rutelinae). Animma. X. Supplement 4. 2 nd Edn. Plzen, 142 pp." year="2007">Krajcik 2007</bibRefCitation>
: 80. Syn. nov.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="161" type="type material">
<paragraph pageId="0" pageNumber="161">Type material examined</paragraph>
<paragraph pageId="0" pageNumber="161">
<emphasis bold="true" pageId="0" pageNumber="161">
(2 males, 1 female). 
<emphasis italics="true" pageId="0" pageNumber="161">Lectotype</emphasis>
</emphasis>
male of 
<taxonomicName authorityName="Burmeister" authorityYear="1844" class="Insecta" family="Scarabaeidae" genus="Macraspis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macraspis maculata" order="Coleoptera" pageId="0" pageNumber="161" phylum="Arthropoda" rank="species" species="maculata">
<emphasis italics="true" pageId="0" pageNumber="161">Macraspis maculata</emphasis>
</taxonomicName>
(designated by Soula, 1998) deposited at MLUH, labeled: &quot;maculata / Dej. / Bras. Coll. (green, handwritten) // LECTOTYPE ♂ / Macraspis / maculata B. / M. Soula 1994&quot;. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="161">Holotype</emphasis>
male of 
<taxonomicName authorityName="Soula" authorityYear="1998" class="Insecta" family="Scarabaeidae" genus="Macraspis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macraspis maculata subsp. crosarai" order="Coleoptera" pageId="0" pageNumber="161" phylum="Arthropoda" rank="subSpecies" species="maculata" subSpecies="crosarai">
<emphasis italics="true" pageId="0" pageNumber="161">Macraspis maculata crosarai</emphasis>
</taxonomicName>
Soula deposited at MNHN, labeled: 
<normalizedToken originalValue="“BRÉSIL">&quot;BRESIL</normalizedToken>
/ 
<normalizedToken originalValue="ÉT">ET</normalizedToken>
. DE GOYAZ / JATAHY / PUJOL, 12-97; 1-98&quot; (white, printed) // &quot;HOLOTYPE / Macraspis / maculata / crosarai / So. / M. Soula det.&quot; (red, printed and handwritten). 
<emphasis bold="true" italics="true" pageId="0" pageNumber="161">Paratype</emphasis>
female of 
<taxonomicName lsidName="M. maculata subsp. crosarai" pageId="0" pageNumber="161" rank="subSpecies" species="maculata" subSpecies="crosarai">
<emphasis italics="true" pageId="0" pageNumber="161">M. maculata crosarai</emphasis>
</taxonomicName>
at MNHN, labeled: &quot;Jathay / 
<normalizedToken originalValue="État">Etat</normalizedToken>
de Goyaz / Ch. Pujol 1895-96&quot; (white, printed) // 
<normalizedToken originalValue="“Muséum">&quot;Museum</normalizedToken>
Paris / ex Coll. / R. 
<normalizedToken originalValue="Oberthür">Oberthuer</normalizedToken>
/ 1952&quot; (bluish white, printed) // &quot;MACRASPIS / maculata Bu / Soula det.&quot; (white, printed) // &quot;ALLOTYPE / Macraspis / maculata / crosarai / So. / M. Soula det.&quot; (red, printed) // &quot; MNHN / EC10776&quot; (white, printed).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="161" type="materials_examined">
<paragraph pageId="0" pageNumber="161">Additional material examined</paragraph>
<paragraph pageId="0" pageNumber="161">
<emphasis bold="true" pageId="0" pageNumber="161">
(
<specimenCount type="male">18 males</specimenCount>
, 
<specimenCount type="female">13 females</specimenCount>
).
</emphasis>
<materialsCitation collectingDate="1971-03-05" collectorName="Elias" country="Brazil" location="Prado" specimenCount="♀" stateProvince="Bahia">
<collectingCountry name="Brazil">Brazil</collectingCountry>
, 
<collectingRegion country="Brazil" name="Bahia">Bahia</collectingRegion>
, 
<location LSID="urn:lsid:plazi:treatment:6588A24D1B2659B9ADD46C0850DDA833:45D946E375AEB910911A40E55F1AB78F" country="Brazil" name="Prado" stateProvince="Bahia">Prado</location>
, 
<collectingDate value="1971-03-05">05.III.1971</collectingDate>
, C. 
<collectorName>Elias</collectorName>
(leg.) (
<specimenCount>♀</specimenCount>
, DZUP)
</materialsCitation>
; 
<materialsCitation collectingDate="1968-09-26" collectingDateMax="1969-10-04" collectingDateMin="1968-09-26" collectorName="Elias" country="Brazil" location="Conceicao da Barra" specimenCount="2" specimenCount-male="2" stateProvince="Espirito Santo">
<collectingCountry name="Brazil">Brazil</collectingCountry>
, 
<collectingRegion country="Brazil" name="Espirito Santo">
<normalizedToken originalValue="Espírito">Espirito</normalizedToken>
Santo
</collectingRegion>
, 
<location LSID="urn:lsid:plazi:treatment:6588A24D1B2659B9ADD46C0850DDA833:31559B01890B3DED16B628B63646D367" country="Brazil" name="Conceicao da Barra" stateProvince="Espirito Santo">
<normalizedToken originalValue="Conceição">Conceicao</normalizedToken>
da Barra
</location>
, 
<collectingDate value="1969-10-04">04.X.1969</collectingDate>
, C. T. &amp; C. 
<collectorName>Elias</collectorName>
(legs.) (
<specimenCount>♂</specimenCount>
, 
<specimenCount>♀</specimenCount>
, DZUP); idem, but 
<collectingDate value="1968-09-26">26.IX.1968</collectingDate>
(
<specimenCount type="male">2♂</specimenCount>
, 
<specimenCount>♀</specimenCount>
, DZUP); idem, but 
<collectingDate value="1968-11-19" valueMax="1968-11-25" valueMin="1968-11-19">19-25.XI.1968</collectingDate>
(
<specimenCount>♂</specimenCount>
, 
<specimenCount>♀</specimenCount>
, DZUP)
</materialsCitation>
; 
<materialsCitation collectingDate="1906-10-26" collectingDateMax="1966-11" collectingDateMin="1906-10-26" collectorName="Maller, J. F. Zikan Coll." country="Brazil" location="Rio Itabapoana" municipality="Linhares" specimenCount="♂" stateProvince="Espirito Santo">
<collectingCountry name="Brazil">Brazil</collectingCountry>
, 
<collectingRegion country="Brazil" name="Espirito Santo">
<normalizedToken originalValue="Espírito">Espirito</normalizedToken>
Santo
</collectingRegion>
, 
<collectingMunicipality>Linhares</collectingMunicipality>
, 
<collectingDate value="1965-10">X.1965</collectingDate>
(
<specimenCount>♂</specimenCount>
, DZUP); idem, but 
<collectingDate value="1965-11">XI.1965</collectingDate>
(
<specimenCount>♀</specimenCount>
, DZUP); idem, but no date (
<specimenCount>♂</specimenCount>
, DZUP); idem, but 
<collectingDate value="1966-11">XI.1966</collectingDate>
, A. 
<collectorName>Maller</collectorName>
(leg.) (
<specimenCount>♂</specimenCount>
, 
<specimenCount>♀</specimenCount>
, DZUP); idem, but 
<location LSID="urn:lsid:plazi:treatment:6588A24D1B2659B9ADD46C0850DDA833:7BB261827C56728EA7380ACEC93286C1" country="Brazil" municipality="Linhares" name="Rio Itabapoana" stateProvince="Espirito Santo">Rio Itabapoana</location>
, 
<collectingDate value="1906-10-26">26.X.1906</collectingDate>
, 
<collectorName>
J. F. 
<normalizedToken originalValue="Zikán">Zikan</normalizedToken>
Coll.
</collectorName>
(
<specimenCount>♀</specimenCount>
, CEIOC)
</materialsCitation>
; 
<materialsCitation collectingDate="1961-12" collectorName="Seabra, Alvarenga" country="Brazil" location="Corcovado" specimenCount="♀" stateProvince="Rio de Janeiro">
<collectingCountry name="Brazil">Brazil</collectingCountry>
, 
<collectingRegion country="Brazil" name="Rio de Janeiro">Rio de Janeiro</collectingRegion>
, 
<collectingRegion country="Brazil" name="Rio de Janeiro">Rio de Janeiro</collectingRegion>
, 
<location LSID="urn:lsid:plazi:treatment:6588A24D1B2659B9ADD46C0850DDA833:ACB82DA89AAAF890360FBAD4BC2B2035" country="Brazil" name="Corcovado" stateProvince="Rio de Janeiro">Corcovado</location>
, 
<collectingDate value="1961-12">XII.1961</collectingDate>
, 
<collectorName>Seabra</collectorName>
&amp; 
<collectorName>Alvarenga</collectorName>
(legs.) (
<specimenCount>♀</specimenCount>
, DZUP)
</materialsCitation>
; 
<materialsCitation collectingDate="1926-11-28" collectorName="Zikan Coll." country="Brazil" location="Rio Jose Pedro" specimenCount="♀" stateProvince="Minas Gerais">
<collectingCountry name="Brazil">Brazil</collectingCountry>
, 
<collectingRegion country="Brazil" name="Minas Gerais">Minas Gerais</collectingRegion>
, 
<location LSID="urn:lsid:plazi:treatment:6588A24D1B2659B9ADD46C0850DDA833:6BFCFF2DE546C31D95AFA1BF87A16439" country="Brazil" name="Rio Jose Pedro" stateProvince="Minas Gerais">
Rio 
<normalizedToken originalValue="José">Jose</normalizedToken>
Pedro
</location>
, 
<collectingDate value="1926-11-28">28.XI.1926</collectingDate>
, J. F. 
<collectorName>
<normalizedToken originalValue="Zikán">Zikan</normalizedToken>
Coll.
</collectorName>
(
<specimenCount>♀</specimenCount>
, CEIOC)
</materialsCitation>
; 
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="Tris., Trisanga, Irisanga" country="Brazil" location="Brazil" specimenCount="3" specimenCount-male="3">
<collectingCountry name="Brazil">Brazil</collectingCountry>
, 
<collectorName>Tris.</collectorName>
[= 
<collectorName>Trisanga</collectorName>
(= 
<collectorName>Irisanga</collectorName>
)], N.8 (
<specimenCount type="male">3♂</specimenCount>
, NHMW)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="161" type="distribution">
<paragraph pageId="0" pageNumber="161">Distribution</paragraph>
<paragraph pageId="0" pageNumber="161">
<emphasis bold="true" pageId="0" pageNumber="161">
(Fig. 
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Distribution of Brazilian species of Macraspis treated in this publication. The Brazilian Amazon biome is colored lime green. Brazilian states are abbreviated as follows: AC = Acre, AM = Amazonas, AP = Amapa, BA = Bahia, ES = Espirito Santo, GO = Goias, MA = Maranhao, MG = Minas Gerais, MT = Mato Grosso = PA = Piaui, RO = Rondonia, RJ = Rio de Janiero, RR = Roraima, SP = Sao Paulo, TO = Tocantins." figureDoi="10.3897/zookeys.1124.91156.figure12" httpUri="https://binary.pensoft.net/fig/758074" pageId="0" pageNumber="161">12</figureCitation>
).
</emphasis>
Brazil. 
<normalizedToken originalValue="São">Sao</normalizedToken>
Paulo (Irisanga = 
<normalizedToken originalValue="Oriçanga">Oricanga</normalizedToken>
; 
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.8135" author="Burmeister, HCC" journalOrPublisher="Enslin, Berlin" pageId="0" pageNumber="161" refId="B5" refString="Burmeister, HCC, 1844. Handbuch der Entomologie (ColeopteraLamellicornia Anthobia et Phyllophaga Systellochela). Volume 4, Part 1. T.C.F. Enslin, Berlin, DOI: https://doi.org/10.5962/bhl.title.8135" title="Handbuch der Entomologie (ColeopteraLamellicornia Anthobia et Phyllophaga Systellochela). Volume 4, Part 1. T. C. F." url="https://doi.org/10.5962/bhl.title.8135" year="1844">Burmeister 1844</bibRefCitation>
), Rio de Janeiro, 
<normalizedToken originalValue="Goiás">Goias</normalizedToken>
(
<bibRefCitation author="Soula, M" journalOrPublisher="Hillside Books, Canterbury" pageId="0" pageNumber="161" refId="B25" refString="Soula, M, 1998. Les Coleopteres du Monde, 26.1: Rutelini 2. Revision des Anthicheirina 1. Hillside Books, Canterbury" title="Les Coleopteres du Monde, 26.1: Rutelini 2. Revision des Anthicheirina 1." year="1998">Soula 1998</bibRefCitation>
, 
<bibRefCitation author="Soula, M" journalOrPublisher="Hillside Books, Canterbury" pageId="0" pageNumber="161" refId="B27" refString="Soula, M, 2003. Les Coleopteres du Monde, 29: Rutelini 3. Pseudoptenomela - Paraptenomela - Exoptenomela - Ptenomela - Calomacraspis - Paramacraspis - Paradorysthetus - Macraspis - Pseudorysthetus - Dorysthetus - Anticheiroides - Anticheira - Pseudoanthicheiroides. Hillside Books, Canterbury" title="Les Coleopteres du Monde, 29: Rutelini 3. Pseudoptenomela - Paraptenomela - Exoptenomela - Ptenomela - Calomacraspis - Paramacraspis - Paradorysthetus - Macraspis - Pseudorysthetus - Dorysthetus - Anticheiroides - Anticheira - Pseudoanthicheiroides." year="2003">2003</bibRefCitation>
), Bahia, 
<normalizedToken originalValue="Espírito">Espirito</normalizedToken>
Santo, Minas Gerais (new state records).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="161" type="remarks">
<paragraph pageId="0" pageNumber="161">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="161">
<taxonomicName authorityName="Burmeister" authorityYear="1844" class="Insecta" family="Scarabaeidae" genus="Macraspis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macraspis maculata" order="Coleoptera" pageId="0" pageNumber="161" phylum="Arthropoda" rank="species" species="maculata">
<emphasis italics="true" pageId="0" pageNumber="161">Macraspis maculata</emphasis>
</taxonomicName>
has been recorded from the Brazilian Amazon (
<bibRefCitation author="Ohaus, F" journalOrPublisher="Coleopterorum Catalogus" pageId="0" pageNumber="161" pagination="1 - 241" refId="B20" refString="Ohaus, F, 1918. Scarabaeidae: Euchirinae, Phaenomerinae, Rutelinae. Coleopterorum Catalogus 20: 1 - 241" title="Scarabaeidae: Euchirinae, Phaenomerinae, Rutelinae." volume="20" year="1918">Ohaus 1918</bibRefCitation>
, 
<bibRefCitation author="Ohaus, F" journalOrPublisher="Coleopterorum Catalogus" pageId="0" pageNumber="161" refId="B21" refString="Ohaus, F, 1934. Coleoptera lamellicornia, Fam. Scarabaeidae, Subfam. Rutelinae. T.1: Tribus Rutelini. Genera Insectorum Fasc. 199A: 1-172." title="Coleoptera lamellicornia, Fam. Scarabaeidae, Subfam. Rutelinae. T. 1: Tribus Rutelini. Genera Insectorum Fasc. 199 A: 1 - 172." year="1934">1934</bibRefCitation>
; 
<bibRefCitation author="Machatschke, JW" journalOrPublisher="Coleopterorum Catalogus Supplementa" pageId="0" pageNumber="161" pagination="1 - 361" refId="B15" refString="Machatschke, JW, 1972. Scarabaeoidea: Melolonthidae, Rutelinae. Coleopterorum Catalogus Supplementa 66: 1 - 361" title="Scarabaeoidea: Melolonthidae, Rutelinae." volume="66" year="1972">Machatschke 1972</bibRefCitation>
; 
<bibRefCitation author="Brule, S" journalOrPublisher="SEAG, PAG" pageId="0" pageNumber="161" refId="B4" refString="Brule, S, Touroult, J, Poirier, E, Dalens, PH, 2014. Resultats de l'etude-inventaire entomologique du site de la Montagne Pelee, Sauel (Guyane), 2010-2012. Rapport de la Societe entomologique Antilles-Guyane. SEAG, PAG" title="Resultats de l'etude-inventaire entomologique du site de la Montagne Pelee, Sauel (Guyane), 2010 - 2012. Rapport de la Societe entomologique Antilles-Guyane." year="2014">
<normalizedToken originalValue="Brûlé">Brule</normalizedToken>
et al. 2014
</bibRefCitation>
) but similarities with other species have posed problems with identification. In his treatment for 
<taxonomicName lsidName="M. maculata" pageId="0" pageNumber="161" rank="species" species="maculata">
<emphasis italics="true" pageId="0" pageNumber="161">M. maculata</emphasis>
</taxonomicName>
, 
<bibRefCitation author="Soula, M" journalOrPublisher="Hillside Books, Canterbury" pageId="0" pageNumber="161" refId="B25" refString="Soula, M, 1998. Les Coleopteres du Monde, 26.1: Rutelini 2. Revision des Anthicheirina 1. Hillside Books, Canterbury" title="Les Coleopteres du Monde, 26.1: Rutelini 2. Revision des Anthicheirina 1." year="1998">Soula (1998)</bibRefCitation>
considered the species to be widely distributed, and he justified the creation of related taxa because: 
<normalizedToken originalValue="“…">&quot;...</normalizedToken>
some populations that are more geographically isolated are already genetically isolated enough to describe subspecies (or even new species ...)&quot; (translated from French). 
<bibRefCitation author="Soula, M" journalOrPublisher="Hillside Books, Canterbury" pageId="0" pageNumber="161" refId="B25" refString="Soula, M, 1998. Les Coleopteres du Monde, 26.1: Rutelini 2. Revision des Anthicheirina 1. Hillside Books, Canterbury" title="Les Coleopteres du Monde, 26.1: Rutelini 2. Revision des Anthicheirina 1." year="1998">Soula (1998</bibRefCitation>
, 
<bibRefCitation author="Soula, M" journalOrPublisher="Hillside Books, Canterbury" pageId="0" pageNumber="161" refId="B27" refString="Soula, M, 2003. Les Coleopteres du Monde, 29: Rutelini 3. Pseudoptenomela - Paraptenomela - Exoptenomela - Ptenomela - Calomacraspis - Paramacraspis - Paradorysthetus - Macraspis - Pseudorysthetus - Dorysthetus - Anticheiroides - Anticheira - Pseudoanthicheiroides. Hillside Books, Canterbury" title="Les Coleopteres du Monde, 29: Rutelini 3. Pseudoptenomela - Paraptenomela - Exoptenomela - Ptenomela - Calomacraspis - Paramacraspis - Paradorysthetus - Macraspis - Pseudorysthetus - Dorysthetus - Anticheiroides - Anticheira - Pseudoanthicheiroides." year="2003">2003</bibRefCitation>
) restricted 
<taxonomicName authority="maculata" lsidName="M. maculata subsp. maculata" pageId="0" pageNumber="161" rank="subSpecies" species="maculata" subSpecies="maculata">
<emphasis italics="true" pageId="0" pageNumber="161">M. maculata maculata</emphasis>
</taxonomicName>
to Brazilian Atlantic Forest and described 
<taxonomicName lsidName="M. maculata subsp. crosarai" pageId="0" pageNumber="161" rank="subSpecies" species="maculata" subSpecies="crosarai">
<emphasis italics="true" pageId="0" pageNumber="161">M. maculata crosarai</emphasis>
</taxonomicName>
from Brazilian Cerrado (distinguished from the nominotypical subspecies by means of coloration). Influenced by Antonio 
<normalizedToken originalValue="Martínez’s">Martinez's</normalizedToken>
identification labels, 
<bibRefCitation author="Soula, M" journalOrPublisher="Hillside Books, Canterbury" pageId="0" pageNumber="161" refId="B25" refString="Soula, M, 1998. Les Coleopteres du Monde, 26.1: Rutelini 2. Revision des Anthicheirina 1. Hillside Books, Canterbury" title="Les Coleopteres du Monde, 26.1: Rutelini 2. Revision des Anthicheirina 1." year="1998">Soula (1998)</bibRefCitation>
described 
<taxonomicName lsidName="M. martinezi" pageId="0" pageNumber="161" rank="species" species="martinezi">
<emphasis italics="true" pageId="0" pageNumber="161">M. martinezi</emphasis>
</taxonomicName>
and compared it with 
<taxonomicName lsidName="M. maculata" pageId="0" pageNumber="161" rank="species" species="maculata">
<emphasis italics="true" pageId="0" pageNumber="161">M. maculata</emphasis>
</taxonomicName>
(perhaps constituting one of 
<normalizedToken originalValue="Soula’s">Soula's</normalizedToken>
&quot;genetically isolated&quot; populations of 
<taxonomicName lsidName="M. maculata" pageId="0" pageNumber="161" rank="species" species="maculata">
<emphasis italics="true" pageId="0" pageNumber="161">M. maculata</emphasis>
</taxonomicName>
). He included three subspecies, 
<taxonomicName lsidName="M. martinezi subsp. auzerali" pageId="0" pageNumber="161" rank="subSpecies" species="martinezi" subSpecies="auzerali">
<emphasis italics="true" pageId="0" pageNumber="161">M. martinezi auzerali</emphasis>
</taxonomicName>
, 
<taxonomicName lsidName="M. martinezi subsp. curoei" pageId="0" pageNumber="161" rank="subSpecies" species="martinezi" subSpecies="curoei">
<emphasis italics="true" pageId="0" pageNumber="161">M. martinezi curoei</emphasis>
</taxonomicName>
Soula, 1998, and 
<taxonomicName lsidName="M. martinezi subsp. colombica" pageId="0" pageNumber="161" rank="subSpecies" species="martinezi" subSpecies="colombica">
<emphasis italics="true" pageId="0" pageNumber="161">M. martinezi colombica</emphasis>
</taxonomicName>
Soula, 1998, all of which are distributed in the Amazon biome (only the first subspecies is known from the Brazilian Amazon) and similar to 
<taxonomicName lsidName="M. maculata" pageId="0" pageNumber="161" rank="species" species="maculata">
<emphasis italics="true" pageId="0" pageNumber="161">M. maculata</emphasis>
</taxonomicName>
in coloration, form, and genitalic form. The validity of 
<taxonomicName lsidName="M. martinezi" pageId="0" pageNumber="161" rank="species" species="martinezi">
<emphasis italics="true" pageId="0" pageNumber="161">M. martinezi</emphasis>
</taxonomicName>
and its subspecies-complex requires evaluation in future studies. Examination of the primary types of 
<taxonomicName authority="maculata" lsidName="M. maculata subsp. maculata" pageId="0" pageNumber="161" rank="subSpecies" species="maculata" subSpecies="maculata">
<emphasis italics="true" pageId="0" pageNumber="161">M. maculata maculata</emphasis>
</taxonomicName>
(Fig. 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Macraspis maculata Burmeister, 1844, lectotype male (A-D) A dorsal view B ventral view C aedeagus D lectotype label. M. maculata crosarai Soula, 1998, holotype male (E-H) E dorsal view F ventral view G aedeagus H labels." figureDoi="10.3897/zookeys.1124.91156.figure8" httpUri="https://binary.pensoft.net/fig/758070" pageId="0" pageNumber="161">8A-D</figureCitation>
) and 
<taxonomicName lsidName="M. maculata subsp. crosarai" pageId="0" pageNumber="161" rank="subSpecies" species="maculata" subSpecies="crosarai">
<emphasis italics="true" pageId="0" pageNumber="161">M. maculata crosarai</emphasis>
</taxonomicName>
(Fig. 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Macraspis maculata Burmeister, 1844, lectotype male (A-D) A dorsal view B ventral view C aedeagus D lectotype label. M. maculata crosarai Soula, 1998, holotype male (E-H) E dorsal view F ventral view G aedeagus H labels." figureDoi="10.3897/zookeys.1124.91156.figure8" httpUri="https://binary.pensoft.net/fig/758070" pageId="0" pageNumber="161">8E-H</figureCitation>
) revealed no conspicuous differences between these taxa. Based on this examination, we synonymize 
<taxonomicName lsidName="M. maculata subsp. crosarai" pageId="0" pageNumber="161" rank="subSpecies" species="maculata" subSpecies="crosarai">
<emphasis italics="true" pageId="0" pageNumber="161">M. maculata crosarai</emphasis>
</taxonomicName>
with the nominotypical 
<taxonomicName lsidName="M. maculata" pageId="0" pageNumber="161" rank="species" species="maculata">
<emphasis italics="true" pageId="0" pageNumber="161">M. maculata</emphasis>
</taxonomicName>
. Now, 
<taxonomicName lsidName="M. maculata" pageId="0" pageNumber="161" rank="species" species="maculata">
<emphasis italics="true" pageId="0" pageNumber="161">M. maculata</emphasis>
</taxonomicName>
no longer includes subspecies. Therefore, although the taxon has been recorded to the Brazilian Amazon and French Guiana by past authors (
<bibRefCitation author="Ohaus, F" journalOrPublisher="Coleopterorum Catalogus" pageId="0" pageNumber="161" pagination="1 - 241" refId="B20" refString="Ohaus, F, 1918. Scarabaeidae: Euchirinae, Phaenomerinae, Rutelinae. Coleopterorum Catalogus 20: 1 - 241" title="Scarabaeidae: Euchirinae, Phaenomerinae, Rutelinae." volume="20" year="1918">Ohaus 1918</bibRefCitation>
, 
<bibRefCitation author="Ohaus, F" journalOrPublisher="Coleopterorum Catalogus" pageId="0" pageNumber="161" refId="B21" refString="Ohaus, F, 1934. Coleoptera lamellicornia, Fam. Scarabaeidae, Subfam. Rutelinae. T.1: Tribus Rutelini. Genera Insectorum Fasc. 199A: 1-172." title="Coleoptera lamellicornia, Fam. Scarabaeidae, Subfam. Rutelinae. T. 1: Tribus Rutelini. Genera Insectorum Fasc. 199 A: 1 - 172." year="1934">1934</bibRefCitation>
; 
<bibRefCitation author="Machatschke, JW" journalOrPublisher="Coleopterorum Catalogus Supplementa" pageId="0" pageNumber="161" pagination="1 - 361" refId="B15" refString="Machatschke, JW, 1972. Scarabaeoidea: Melolonthidae, Rutelinae. Coleopterorum Catalogus Supplementa 66: 1 - 361" title="Scarabaeoidea: Melolonthidae, Rutelinae." volume="66" year="1972">Machatschke 1972</bibRefCitation>
; 
<bibRefCitation author="Brule, S" journalOrPublisher="SEAG, PAG" pageId="0" pageNumber="161" refId="B4" refString="Brule, S, Touroult, J, Poirier, E, Dalens, PH, 2014. Resultats de l'etude-inventaire entomologique du site de la Montagne Pelee, Sauel (Guyane), 2010-2012. Rapport de la Societe entomologique Antilles-Guyane. SEAG, PAG" title="Resultats de l'etude-inventaire entomologique du site de la Montagne Pelee, Sauel (Guyane), 2010 - 2012. Rapport de la Societe entomologique Antilles-Guyane." year="2014">
<normalizedToken originalValue="Brûlé">Brule</normalizedToken>
et al. 2014
</bibRefCitation>
), 
<taxonomicName lsidName="M. maculata" pageId="0" pageNumber="161" rank="species" species="maculata">
<emphasis italics="true" pageId="0" pageNumber="161">M. maculata</emphasis>
</taxonomicName>
is treated 
<emphasis italics="true" pageId="0" pageNumber="161">sensu</emphasis>
<bibRefCitation author="Soula, M" journalOrPublisher="Hillside Books, Canterbury" pageId="0" pageNumber="161" refId="B25" refString="Soula, M, 1998. Les Coleopteres du Monde, 26.1: Rutelini 2. Revision des Anthicheirina 1. Hillside Books, Canterbury" title="Les Coleopteres du Monde, 26.1: Rutelini 2. Revision des Anthicheirina 1." year="1998">Soula (1998</bibRefCitation>
, 
<bibRefCitation author="Soula, M" journalOrPublisher="Hillside Books, Canterbury" pageId="0" pageNumber="161" refId="B27" refString="Soula, M, 2003. Les Coleopteres du Monde, 29: Rutelini 3. Pseudoptenomela - Paraptenomela - Exoptenomela - Ptenomela - Calomacraspis - Paramacraspis - Paradorysthetus - Macraspis - Pseudorysthetus - Dorysthetus - Anticheiroides - Anticheira - Pseudoanthicheiroides. Hillside Books, Canterbury" title="Les Coleopteres du Monde, 29: Rutelini 3. Pseudoptenomela - Paraptenomela - Exoptenomela - Ptenomela - Calomacraspis - Paramacraspis - Paradorysthetus - Macraspis - Pseudorysthetus - Dorysthetus - Anticheiroides - Anticheira - Pseudoanthicheiroides." year="2003">2003</bibRefCitation>
) in this paper and considered to be distributed in the Brazilian Cerrado and Atlantic Forest rather than the Brazilian Amazon region. For this reason, this species was excluded from the key provided here.
</paragraph>
<caption doi="10.3897/zookeys.1124.91156.figure8" httpUri="https://binary.pensoft.net/fig/758070" pageId="0" pageNumber="161" start="Figure 8" startId="F8">
<paragraph pageId="0" pageNumber="161">
<emphasis bold="true" pageId="0" pageNumber="161">Figure 8.</emphasis>
<taxonomicName authorityName="Burmeister" authorityYear="1844" class="Insecta" family="Scarabaeidae" genus="Macraspis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macraspis maculata" order="Coleoptera" pageId="0" pageNumber="161" phylum="Arthropoda" rank="species" species="maculata">
<emphasis italics="true" pageId="0" pageNumber="161">Macraspis maculata</emphasis>
</taxonomicName>
Burmeister, 1844, lectotype male (
<emphasis bold="true" pageId="0" pageNumber="161">A-D</emphasis>
) 
<emphasis bold="true" pageId="0" pageNumber="161">A</emphasis>
dorsal view 
<emphasis bold="true" pageId="0" pageNumber="161">B</emphasis>
ventral view 
<emphasis bold="true" pageId="0" pageNumber="161">C</emphasis>
aedeagus 
<emphasis bold="true" pageId="0" pageNumber="161">D</emphasis>
lectotype label. 
<taxonomicName lsidName="M. maculata subsp. crosarai" pageId="0" pageNumber="161" rank="subSpecies" species="maculata" subSpecies="crosarai">
<emphasis italics="true" pageId="0" pageNumber="161">M. maculata crosarai</emphasis>
</taxonomicName>
Soula, 1998, holotype male (
<emphasis bold="true" pageId="0" pageNumber="161">E-H</emphasis>
) 
<emphasis bold="true" pageId="0" pageNumber="161">E</emphasis>
dorsal view 
<emphasis bold="true" pageId="0" pageNumber="161">F</emphasis>
ventral view 
<emphasis bold="true" pageId="0" pageNumber="161">G</emphasis>
aedeagus 
<emphasis bold="true" pageId="0" pageNumber="161">H</emphasis>
labels.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>