317 lines
45 KiB
XML
317 lines
45 KiB
XML
<document id="86C71F4D756BA62D96F986AA2EE13F99" ID-CLB-Dataset="46361" ID-DOI="10.11646/zootaxa.3768.2.6" ID-GBIF-Dataset="807bb695-5592-4089-9142-010a98add339" ID-ISSN="1175-5326" ID-Zenodo-Dep="225197" ID-ZooBank="12362EE3-1A25-4DC6-8413-4E1264826680" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460388721214" checkinUser="plazi" docAuthor="Sugaya, Kazuki & Yamasako, Junsuke" docDate="2014" docId="BA388799F94BFFE84C9825D9140BF9CE" docLanguage="en" docName="zt03768p195.pdf" docOrigin="Zootaxa 3768 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ishikawatrechus bidilatatus Sugaya, sp. nov." docType="treatment" docUuid="3D1EDA12-629C-4801-B010-5A61A539EB15" docUuidSource="ZooBank" docVersion="10" lastPageNumber="194" masterDocId="4601FFE1F94AFFED4C0F275D1627FFC6" masterDocTitle="A new species of the genus Ishikawatrechus (Coleoptera, Trechinae) from Japan" masterLastPageNumber="195" masterPageNumber="189" pageNumber="190" updateTime="1698297955780" updateUser="ExternalLinkService">
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<mods:mods id="FB553B1E1D68F6CE4BAD6EF42AC4475F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="FB1010407136DDCF1CBF203A81DB8DDB">
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<mods:title id="F489E0C72BACB790AD244F24E81B67C1">A new species of the genus Ishikawatrechus (Coleoptera, Trechinae) from Japan</mods:title>
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</mods:titleInfo>
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<mods:name id="D1456C6DBF39AF7694B8D9BD364993F6" type="personal">
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<mods:roleTerm id="4922A90489F3D8BF5A15E8956C8DF517">Author</mods:roleTerm>
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<mods:namePart id="8C7D76A7C31E52B746493874214D4A70">Sugaya, Kazuki</mods:namePart>
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</mods:name>
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<mods:name id="77FDCDCFEB6F423BC15C8914E1BD4EA9" type="personal">
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<mods:roleTerm id="4B7A68BF43328B5A9BDF908D751C7DE6">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="384781321B5381E276840D5927D7758B">Yamasako, Junsuke</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="EC18C6C2221CA73BC445D4DC2EBBA081">text</mods:typeOfResource>
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<mods:titleInfo id="1C934ADD31418B62212AD27B7C56E181">
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<mods:title id="637711D8AEC256584B5EAE9E30C23C67">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="7318FC87B8AB3148CAD03A1599606334">
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<mods:date id="6E675A6D7A9A0071322184DDB3DC86CB">2014</mods:date>
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<mods:detail id="894EA64BAB1D6BDC40E44EA25749D74C" type="volume">
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<mods:number id="D0FF7CE457F2609250A6939B6668D0F7">3768</mods:number>
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</mods:detail>
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<mods:detail id="80341DAA5BC451E2DD593D02BAF1961F" type="issue">
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<mods:number id="51BA01ABE846FC57273AB476B9694FF0">2</mods:number>
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<mods:extent id="74D18D4742D93E31F154D6212453460E" unit="page">
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<mods:start id="E0552190AD21739ACD28C16D14B99B4F">189</mods:start>
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<mods:end id="C2B6DA51763EA6E0392F549CF5666409">195</mods:end>
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<mods:classification id="B61C162A5C8ACF6D152BB3A0FD5918C2">journal article</mods:classification>
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<mods:identifier id="678BA31378F2C172E4684323C2897A27" type="DOI">10.11646/zootaxa.3768.2.6</mods:identifier>
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<mods:identifier id="E44690D9F18AB4A3890A50C7C67C3F4D" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="4E29E564F92B8BC189F928AC15E93D3B" type="Zenodo-Dep">225197</mods:identifier>
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</mods:mods>
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<treatment id="BA388799F94BFFE84C9825D9140BF9CE" ID-DOI="http://doi.org/10.5281/zenodo.6144104" ID-GBIF-Taxon="119416122" ID-Zenodo-Dep="6144104" LSID="urn:lsid:zoobank.org:act:3D1EDA12-629C-4801-B010-5A61A539EB15" httpUri="http://treatment.plazi.org/id/BA388799F94BFFE84C9825D9140BF9CE" lastPageId="5" lastPageNumber="194" pageId="1" pageNumber="190">
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<subSubSection id="7A8B6504F94BFFEC4C9825D917B2FD04" pageId="1" pageNumber="190" type="nomenclature">
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<paragraph id="322E368FF94BFFEC4C9825D914BBFD58" blockId="1.[151,668,644,706]" box="[151,668,644,670]" pageId="1" pageNumber="190">
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<heading id="696681E3F94BFFEC4C9825D914BBFD58" bold="true" box="[151,668,644,670]" fontSize="10" level="2" pageId="1" pageNumber="190" reason="2">
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<emphasis id="00E5EA9DF94BFFEC4C9825D914BBFD58" bold="true" box="[151,668,644,670]" pageId="1" pageNumber="190">
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<taxonomicName id="F5914D0CF94BFFEC4C9825D91411FD58" ID-CoL="3Q73K" authority="Sugaya" authorityName="Sugaya" box="[151,566,644,670]" class="Insecta" family="Carabidae" genus="Ishikawatrechus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="190" phylum="Arthropoda" rank="species" species="bidilatatus" status="sp. nov.">
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<emphasis id="00E5EA9DF94BFFEC4C9825D917F1FD5B" bold="true" box="[151,470,644,669]" italics="true" pageId="1" pageNumber="190">Ishikawatrechus bidilatatus</emphasis>
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Sugaya
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</taxonomicName>
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,
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<taxonomicNameLabel id="1BD657E6F94BFFEC4E4D25D814BBFD58" box="[578,668,645,670]" pageId="1" pageNumber="190" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="322E368FF94BFFEC4C9825F417B2FD04" blockId="1.[151,668,644,706]" box="[151,405,681,706]" pageId="1" pageNumber="190">
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(
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<figureCitation id="AAAA2A0AF94BFFEC4C9025F4171BFD04" box="[159,316,681,706]" captionStart-0="FIGURES 1 – 2" captionStart-1="FIGURES 3 – 10" captionStartId-0="2.[151,264,1972,1995]" captionStartId-1="3.[151,264,1894,1917]" captionTargetBox-0="[184,1411,206,1928]" captionTargetBox-1="[161,1430,200,1870]" captionTargetId-0="figure@2.[172,1416,193,1938]" captionTargetId-1="figure@3.[151,1436,193,1874]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURES 1 – 2. Habitus of Ishikawatrechus bidilatatus sp. nov., holotype, male. 1, Dorsal view; 2, Left lateral view." captionText-1="FIGURES 3 – 10. Genital organs of Ishikawatrechus spp. 3 – 4, 7 – 8, I. bidilatatus sp. nov.; 5 – 6, 9 – 10, I. ishiharai. 3, 5, Aedeagus in left lateral view; 4, 6, Apex of aedeagus in dorsal view; 7, 9, Left stylus in ventral view; 8, 10, Ditto in dorsal view. Abbreviations; GC 1, basal gonocoxite 1; GC 2, apical gonocoxite 2." httpUri-0="https://zenodo.org/record/225198/files/figure.png" httpUri-1="https://zenodo.org/record/225199/files/figure.png" pageId="1" pageNumber="190">Figs 1–4, 7–8</figureCitation>
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,
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<figureCitation id="AAAA2A0AF94BFFEC4D4825F417AAFD04" box="[327,397,681,706]" captionStart="FIGURES 11 – 19" captionStartId="4.[151,264,1743,1766]" captionTargetBox="[159,1419,200,1714]" captionTargetId="figure@4.[151,1436,193,1722]" captionTargetPageId="4" captionText="FIGURES 11 – 19. Internal sac of Ishikawatrechus spp. 11 – 15, I. bidilatatus sp. nov.; 16 – 19, I. ishiharai. 11, 16, Right lateral view; 12, 17, Left lateral view; 13, 18, Dorsal view; 14, 19, Ventral view; 15, Copulatory piece of I. bidilatatus. Abbreviations; CP, copulatory piece; GP, gonopore." httpUri="https://zenodo.org/record/225200/files/figure.png" pageId="1" pageNumber="190">11–15</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="7A8B6504F94BFFEC4C9825AC1790FC78" pageId="1" pageNumber="190" type="materials_examined">
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<paragraph id="322E368FF94BFFEC4C9825AC172EFC94" blockId="1.[151,1437,753,2038]" pageId="1" pageNumber="190">
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<emphasis id="00E5EA9DF94BFFEC4C9825AC173CFCCC" bold="true" box="[151,283,753,778]" pageId="1" pageNumber="190">
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<typeStatus id="ED2A882DF94BFFEC4C9825AC16F5FCCC" box="[151,210,753,778]" pageId="1" pageNumber="190">Type</typeStatus>
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series
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</emphasis>
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(
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<collectionCode id="5480AE4AF94BFFEC4D2625AF175EFCCC" box="[297,377,754,778]" httpUri="http://grbio.org/cool/0wyw-sfpi" name="Ehime University" pageId="1" pageNumber="190">EUMJ</collectionCode>
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).
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<typeStatus id="ED2A882DF94BFFEC4D8525AC17D0FCCC" box="[394,503,753,778]" pageId="1" pageNumber="190" type="holotype">Holotype</typeStatus>
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:
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<specimenCount id="2497FD06F94BFFEC4E0E25AF1402FCCC" box="[513,549,754,778]" pageId="1" pageNumber="190" type="generic">1♂</specimenCount>
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, “[
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<collectionCode id="5480AE4AF94BFFEC4E4D25AF1484FCCC" box="[578,675,754,778]" pageId="1" pageNumber="190">EHIME</collectionCode>
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:
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<collectingCountry id="4A86761FF94BFFEC4EA225AF1522FCCC" box="[685,773,754,778]" name="Japan" pageId="1" pageNumber="190">JAPAN</collectingCountry>
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] / Ôyashiki, Kawano-uchi / Tôon-shi, alt. ca.
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<quantity id="F5699B6AF94BFFEC490F25AC1360FCCF" box="[1280,1351,753,778]" metricMagnitude="2" metricUnit="m" metricValue="5.3" pageId="1" pageNumber="190" unit="m" value="530.0">530 m</quantity>
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/
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<date id="462F104FF94BFFEC495425AF16E8FCE8" pageId="1" pageNumber="190" value="2012-05-14">14. V. 2012</date>
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/ K. Sugaya leg.” [printed on white label]; “
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<typeStatus id="ED2A882DF94BFFEC4ECA244B157BFCE8" box="[709,860,790,814]" pageId="1" pageNumber="190" type="holotype">HOLOTYPE</typeStatus>
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/
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<taxonomicName id="F5914D0CF94BFFEC4F7C2448128EFCEB" box="[883,1193,789,813]" class="Insecta" family="Carabidae" genus="Ishikawatrechus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="190" phylum="Arthropoda" rank="species" species="bidilatatus" status="sp. nov.">
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<emphasis id="00E5EA9DF94BFFEC4F7C2448128EFCEB" box="[883,1193,789,813]" italics="true" pageId="1" pageNumber="190">Ishikawatrechus bidilatatus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="1BD657E6F94BFFEC48BE2448132CFCE8" box="[1201,1291,789,814]" pageId="1" pageNumber="190" rank="species">
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<emphasis id="00E5EA9DF94BFFEC48BE2448132CFCE8" bold="true" box="[1201,1291,789,814]" pageId="1" pageNumber="190">sp. nov.</emphasis>
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</taxonomicNameLabel>
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” [printed on red label].
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</paragraph>
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<paragraph id="322E368FF94BFFEC4CC824031790FC78" blockId="1.[151,1437,753,2038]" pageId="1" pageNumber="190">
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<typeStatus id="ED2A882DF94BFFEC4CC82403171FFCB0" box="[199,312,862,886]" pageId="1" pageNumber="190" type="paratype">Paratypes</typeStatus>
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:
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<specimenCount id="2497FD06F94BFFEC4D4B24031757FCB3" box="[324,368,862,886]" pageId="1" pageNumber="190" type="generic">1 ♂</specimenCount>
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, same data as
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<typeStatus id="ED2A882DF94BFFEC4E1A24001450FCB0" box="[533,631,861,886]" pageId="1" pageNumber="190" type="holotype">holotype</typeStatus>
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but alt. ca.
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<quantity id="F5699B6AF94BFFEC4EF12400156FFCB3" box="[766,840,861,886]" metricMagnitude="2" metricUnit="m" metricValue="6.3" pageId="1" pageNumber="190" unit="m" value="630.0">630 m</quantity>
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,
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<date id="462F104FF94BFFEC4F5A240315F9FCB0" box="[853,990,862,886]" pageId="1" pageNumber="190" value="2010-10-28">28. X. 2010</date>
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, K. Sugaya leg.;
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<specimenCount id="2497FD06F94BFFEC48AE240312E9FCB3" box="[1185,1230,862,886]" pageId="1" pageNumber="190" type="generic">1 ♂</specimenCount>
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, same locality but
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<date id="462F104FF94BFFEC4C9824DC173BFC5C" box="[151,284,897,922]" pageId="1" pageNumber="190" value="2011-10-16">16. X. 2011</date>
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, K. Sugaya leg.; 1 ♀, same locality but
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<date id="462F104FF94BFFEC4ED624DF1544FC5C" box="[729,867,898,922]" pageId="1" pageNumber="190" value="2011-12-03">3. XII. 2011</date>
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, Y. Hara leg.; 1 ♀, same locality as
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<typeStatus id="ED2A882DF94BFFEC48FC24DC1372FC5C" box="[1267,1365,897,922]" pageId="1" pageNumber="190" type="holotype">holotype</typeStatus>
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but
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<date id="462F104FF94BFFEC498824DC1720FC78" pageId="1" pageNumber="190" value="2012-07-06">6. VII. 2012</date>
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, K. Sugaya leg.
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</paragraph>
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</subSubSection>
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<subSubSection id="7A8B6504F94BFFE84CC8249414A6FCF1" lastPageId="5" lastPageNumber="194" pageId="1" pageNumber="190" type="description">
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<paragraph id="322E368FF94BFFEC4CC824941493FC24" blockId="1.[151,1437,753,2038]" box="[199,692,969,994]" pageId="1" pageNumber="190">
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<emphasis id="00E5EA9DF94BFFEC4CC824941776FC24" bold="true" box="[199,337,969,994]" pageId="1" pageNumber="190">Description</emphasis>
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(n = 5). Habitus as in
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<figureCitation id="AAAA2A0AF94BFFEC4E4324941497FC24" box="[588,688,969,994]" captionStart="FIGURES 1 – 2" captionStartId="2.[151,264,1972,1995]" captionTargetBox="[184,1411,206,1928]" captionTargetId="figure@2.[172,1416,193,1938]" captionTargetPageId="2" captionText="FIGURES 1 – 2. Habitus of Ishikawatrechus bidilatatus sp. nov., holotype, male. 1, Dorsal view; 2, Left lateral view." httpUri="https://zenodo.org/record/225198/files/figure.png" pageId="1" pageNumber="190">Figs 1–2</figureCitation>
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.
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</paragraph>
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<paragraph id="322E368FF94BFFEC4CC824B01723FB88" blockId="1.[151,1437,753,2038]" pageId="1" pageNumber="190">Coloration in dried specimens almost dark reddish brown, but light reddish brown and translucent when alive; palpi, antennomeres 5–11 and legs lighter than the other body parts; pronotal disc and elytral disc markedly iridescent</paragraph>
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<paragraph id="322E368FF94BFFEC4CC823041725FB50" blockId="1.[151,1437,753,2038]" pageId="1" pageNumber="190">Body glabrous; 2 setiferous pores on stria 3 at basal 1/6 and 1/3 respectively; single setiferous pore on stria 5 at basal 3/5.</paragraph>
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<paragraph id="322E368FF94BFFEC4CC823FC15D1FAA8" blockId="1.[151,1437,753,2038]" pageId="1" pageNumber="190">
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Head subquadrate, widest before neck, thence slightly narrower anteriorly; HL/HW 1.05–1.10 (1.08); HW/PW 0.68–0.70 (0.69); HL/PL 0.67–0.73 (0.71); genae weakly convex; frontal furrows deep, but gradually getting shallower and indistinct posteriorly; dorsum with deeply impressed microsculptures composed of fine reticulation, and some deep wrinkles on supraorbital area. Eyes almost effaced, with conspicuous remnants which are elongateoblong and weakly convex laterally. Mentum tooth bifid. Approximate ratio of antennomeres of
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<typeStatus id="ED2A882DF94BFFEC4919226C135FFA8C" box="[1302,1400,1329,1354]" pageId="1" pageNumber="190" type="holotype">holotype</typeStatus>
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as follows; 1.04: 1.00: 1.83: 1.67: 1.67: 1.63: 1.54: 1.38: 1.38: 1.27: 1.44.
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</paragraph>
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<paragraph id="322E368FF94BFFEC4CC82227142AF910" blockId="1.[151,1437,753,2038]" pageId="1" pageNumber="190">Pronotum elongated subcordate, widest at basal 3/4; PW/HW 1.44–1.46 (1.45), PL/PW 1.02–1.08 (1.05), PA/ PW 0.67–0.70 (0.69), PB/PW 0.61–0.63 (0.62); PB/PA 0.88–0.94 (0.91); front margin shallowly emarginate; sides forming edge, arcuately dilated in apical 1/3, widest at apical 2/5, thence straightly narrowed posteriorly toward middle and subparallel sided at basal half; front angles obtuse and rounded; basal margin almost straight; hind angles sharp, weakly projecting backwards; dorsum convex dorsally in apical 2/3, thence declivous posteriorly, depressed near base; median line deep; apical transverse impression distinct; basal transverse impression wide and deep, longitudinally foveolate on each side of median line, laterally merging into basal foveae, which are deep and widely extended anteriorly along side borders; surface even with vague microsculpture composed of transverse finely engraved lines, radially with distinct waving wrinkles extending from median line. Proepipleuron expanding outwards and visible from above.</paragraph>
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<paragraph id="322E368FF94BFFEC4CC821BC17AEF830" blockId="1.[151,1437,753,2038]" pageId="1" pageNumber="190">Elytra oval, widest at basal half; EW/PW 1.78–2.01 (1.84); EL/PL 2.52–2.62 (2.59); EL/EW 1.35–1.54 (1.48); basal peduncle with deep transverse furrow; prehumeral borders slightly short, oblique, slightly emarginate, diminishing anteriorly but reaching basal carina of intervals 5 and 6; humeri very obtuse; lateral sides hardly emarginate behind humeri; disc depressed basal 1/5 which is delimited by the basal part of interval 5 laterally, thence very convex dorsally, with a depression on each lateral side before middle; striae slightly crenulate, deeply impressed but becoming indistinct outwards gradually; apical striole deep, not connected with any stria but directed to the termination of stria 5; each strial interval slightly convex dorsally; microsculpture composed of transverse finely engraved lines.</paragraph>
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<caption id="66EE6607F948FFEF4C9820E91310F80D" box="[151,1335,1972,1995]" httpUri="https://zenodo.org/record/225198/files/figure.png" pageId="2" pageNumber="191" targetBox="[184,1411,206,1928]" targetPageId="2">
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<paragraph id="322E368FF948FFEF4C9820E91310F80D" blockId="2.[151,1335,1972,1995]" box="[151,1335,1972,1995]" pageId="2" pageNumber="191">
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<emphasis id="00E5EA9DF948FFEF4C9820E9171DF80C" bold="true" box="[151,314,1972,1995]" pageId="2" pageNumber="191">FIGURES 1–2.</emphasis>
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Habitus of
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<taxonomicName id="F5914D0CF948FFEF4DBE20E814EFF80D" box="[433,712,1973,1995]" class="Insecta" family="Carabidae" genus="Ishikawatrechus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="191" phylum="Arthropoda" rank="species" species="bidilatatus" status="sp. nov.">
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<emphasis id="00E5EA9DF948FFEF4DBE20E814EFF80D" box="[433,712,1973,1995]" italics="true" pageId="2" pageNumber="191">Ishikawatrechus bidilatatus</emphasis>
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</taxonomicName>
|
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<taxonomicNameLabel id="1BD657E6F948FFEF4EC020EB1538F80D" box="[719,799,1974,1995]" pageId="2" pageNumber="191" rank="species">
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<emphasis id="00E5EA9DF948FFEF4EC020EB1538F80D" bold="true" box="[719,799,1974,1995]" pageId="2" pageNumber="191">sp. nov.</emphasis>
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</taxonomicNameLabel>
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, holotype, male. 1, Dorsal view; 2, Left lateral view.
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</paragraph>
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</caption>
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<caption id="66EE6607F949FFEE4C98203B1563F87C" httpUri="https://zenodo.org/record/225199/files/figure.png" pageId="3" pageNumber="192" targetBox="[161,1430,200,1870]" targetPageId="3">
|
||
<paragraph id="322E368FF949FFEE4C98203B1563F87C" blockId="3.[151,1437,1894,1978]" pageId="3" pageNumber="192">
|
||
<emphasis id="00E5EA9DF949FFEE4C98203B176BF8BA" bold="true" box="[151,332,1894,1917]" pageId="3" pageNumber="192">FIGURES 3–10.</emphasis>
|
||
Genital organs of
|
||
<taxonomicName id="F5914D0CF949FFEE4E14203A14E7F8BB" box="[539,704,1895,1917]" class="Insecta" family="Carabidae" genus="Ishikawatrechus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="192" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="00E5EA9DF949FFEE4E14203A14E7F8BB" box="[539,704,1895,1917]" italics="true" pageId="3" pageNumber="192">Ishikawatrechus</emphasis>
|
||
</taxonomicName>
|
||
spp. 3–4, 7–8,
|
||
<taxonomicName id="F5914D0CF949FFEE4F60203515D2F8BB" box="[879,1013,1895,1917]" class="Insecta" family="Carabidae" genus="Ishikawatrechus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="192" phylum="Arthropoda" rank="species" species="bidilatatus" status="sp. nov.">
|
||
<emphasis id="00E5EA9DF949FFEE4F60203515D2F8BB" box="[879,1013,1895,1917]" italics="true" pageId="3" pageNumber="192">I. bidilatatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="00E5EA9DF949FFEE480F203A1272F8BA" bold="true" box="[1024,1109,1895,1916]" pageId="3" pageNumber="192">
|
||
<taxonomicNameLabel id="1BD657E6F949FFEE480F203A1272F8BA" box="[1024,1109,1895,1916]" pageId="3" pageNumber="192" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 5–6, 9–10,
|
||
<taxonomicName id="F5914D0CF949FFEE48EB2035137EF8BB" box="[1252,1369,1895,1917]" class="Insecta" family="Carabidae" genus="Ishikawatrechus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="192" phylum="Arthropoda" rank="species" species="ishiharai">
|
||
<emphasis id="00E5EA9DF949FFEE48EB2035137EF8BB" box="[1252,1369,1895,1917]" italics="true" pageId="3" pageNumber="192">I. ishiharai</emphasis>
|
||
</taxonomicName>
|
||
. 3, 5, Aedeagus in left lateral view; 4, 6, Apex of aedeagus in dorsal view; 7, 9, Left stylus in ventral view; 8, 10, Ditto in dorsal view. Abbreviations; GC1, basal gonocoxite 1; GC2, apical gonocoxite 2.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="66EE6607F94EFFE94C9821921421F8E4" httpUri="https://zenodo.org/record/225200/files/figure.png" pageId="4" pageNumber="193" targetBox="[159,1419,200,1714]" targetPageId="4">
|
||
<paragraph id="322E368FF94EFFE94C9821921421F8E4" blockId="4.[151,1436,1743,1826]" pageId="4" pageNumber="193">
|
||
<emphasis id="00E5EA9DF94EFFE94C9821921774F922" bold="true" box="[151,339,1743,1766]" pageId="4" pageNumber="193">FIGURES 11–19.</emphasis>
|
||
Internal sac of
|
||
<taxonomicName id="F5914D0CF94EFFE94DFC219214BFF923" box="[499,664,1743,1765]" class="Insecta" family="Carabidae" genus="Ishikawatrechus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="193" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="00E5EA9DF94EFFE94DFC219214BFF923" box="[499,664,1743,1765]" italics="true" pageId="4" pageNumber="193">Ishikawatrechus</emphasis>
|
||
</taxonomicName>
|
||
spp. 11–15,
|
||
<taxonomicName id="F5914D0CF94EFFE94F13218C15B9F923" box="[796,926,1743,1766]" class="Insecta" family="Carabidae" genus="Ishikawatrechus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="193" phylum="Arthropoda" rank="species" species="bidilatatus" status="sp. nov.">
|
||
<emphasis id="00E5EA9DF94EFFE94F13218C15B9F923" box="[796,926,1743,1766]" italics="true" pageId="4" pageNumber="193">I. bidilatatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="1BD657E6F94EFFE94FAA218D15D1F923" box="[933,1014,1744,1765]" pageId="4" pageNumber="193" rank="species">
|
||
<emphasis id="00E5EA9DF94EFFE94FAA218D15D1F923" bold="true" box="[933,1014,1744,1765]" pageId="4" pageNumber="193">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
; 16–19,
|
||
<taxonomicName id="F5914D0CF94EFFE9485D218C12E4F923" box="[1106,1219,1743,1766]" class="Insecta" family="Carabidae" genus="Ishikawatrechus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="193" phylum="Arthropoda" rank="species" species="ishiharai">
|
||
<emphasis id="00E5EA9DF94EFFE9485D218C12E4F923" box="[1106,1219,1743,1766]" italics="true" pageId="4" pageNumber="193">I. ishiharai</emphasis>
|
||
</taxonomicName>
|
||
. 11, 16, Right lateral view; 12, 17, Left lateral view; 13, 18, Dorsal view; 14, 19, Ventral view; 15, Copulatory piece of
|
||
<taxonomicName id="F5914D0CF94EFFE9487721B212DFF8C2" box="[1144,1272,1774,1796]" class="Insecta" family="Carabidae" genus="Ishikawatrechus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="193" phylum="Arthropoda" rank="species" species="bidilatatus">
|
||
<emphasis id="00E5EA9DF94EFFE9487721B212DFF8C2" box="[1144,1272,1774,1796]" italics="true" pageId="4" pageNumber="193">I. bidilatatus</emphasis>
|
||
</taxonomicName>
|
||
. Abbreviations; CP, copulatory piece; GP, gonopore.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="322E368FF94EFFE84CC820121235FF12" blockId="4.[151,1436,1871,2004]" lastBlockId="5.[151,1437,151,1545]" lastPageId="5" lastPageNumber="194" pageId="4" pageNumber="193">
|
||
Male genitalia (n = 3,
|
||
<figureCitation id="AAAA2A0AF94EFFE94DC220121413F8AE" box="[461,564,1871,1896]" captionStart="FIGURES 3 – 10" captionStartId="3.[151,264,1894,1917]" captionTargetBox="[161,1430,200,1870]" captionTargetId="figure@3.[151,1436,193,1874]" captionTargetPageId="3" captionText="FIGURES 3 – 10. Genital organs of Ishikawatrechus spp. 3 – 4, 7 – 8, I. bidilatatus sp. nov.; 5 – 6, 9 – 10, I. ishiharai. 3, 5, Aedeagus in left lateral view; 4, 6, Apex of aedeagus in dorsal view; 7, 9, Left stylus in ventral view; 8, 10, Ditto in dorsal view. Abbreviations; GC 1, basal gonocoxite 1; GC 2, apical gonocoxite 2." httpUri="https://zenodo.org/record/225199/files/figure.png" pageId="4" pageNumber="193">Figs 3–4</figureCitation>
|
||
,
|
||
<figureCitation id="AAAA2A0AF94EFFE94E4D201214ADF8AE" box="[578,650,1871,1896]" captionStart="FIGURES 11 – 19" captionStartId="4.[151,264,1743,1766]" captionTargetBox="[159,1419,200,1714]" captionTargetId="figure@4.[151,1436,193,1722]" captionTargetPageId="4" captionText="FIGURES 11 – 19. Internal sac of Ishikawatrechus spp. 11 – 15, I. bidilatatus sp. nov.; 16 – 19, I. ishiharai. 11, 16, Right lateral view; 12, 17, Left lateral view; 13, 18, Dorsal view; 14, 19, Ventral view; 15, Copulatory piece of I. bidilatatus. Abbreviations; CP, copulatory piece; GP, gonopore." httpUri="https://zenodo.org/record/225200/files/figure.png" pageId="4" pageNumber="193">11–15</figureCitation>
|
||
). Aedeagus in lateral view (
|
||
<figureCitation id="AAAA2A0AF94EFFE94FDE20121231F8AE" box="[977,1046,1871,1896]" captionStart="FIGURES 3 – 10" captionStartId="3.[151,264,1894,1917]" captionTargetBox="[161,1430,200,1870]" captionTargetId="figure@3.[151,1436,193,1874]" captionTargetPageId="3" captionText="FIGURES 3 – 10. Genital organs of Ishikawatrechus spp. 3 – 4, 7 – 8, I. bidilatatus sp. nov.; 5 – 6, 9 – 10, I. ishiharai. 3, 5, Aedeagus in left lateral view; 4, 6, Apex of aedeagus in dorsal view; 7, 9, Left stylus in ventral view; 8, 10, Ditto in dorsal view. Abbreviations; GC 1, basal gonocoxite 1; GC 2, apical gonocoxite 2." httpUri="https://zenodo.org/record/225199/files/figure.png" pageId="4" pageNumber="193">Fig. 3</figureCitation>
|
||
) robust, about 1/3 as long as EL, curved ventrally at basal 1/3, thence hardly arcuate apically; basal part markedly curved ventrally behind basal orifice, roundly pointed at apex, with small sagittal aileron; apical part in dorsal view (
|
||
<figureCitation id="AAAA2A0AF94EFFE9488620CA12F7F876" box="[1161,1232,1943,1968]" captionStart="FIGURES 3 – 10" captionStartId="3.[151,264,1894,1917]" captionTargetBox="[161,1430,200,1870]" captionTargetId="figure@3.[151,1436,193,1874]" captionTargetPageId="3" captionText="FIGURES 3 – 10. Genital organs of Ishikawatrechus spp. 3 – 4, 7 – 8, I. bidilatatus sp. nov.; 5 – 6, 9 – 10, I. ishiharai. 3, 5, Aedeagus in left lateral view; 4, 6, Apex of aedeagus in dorsal view; 7, 9, Left stylus in ventral view; 8, 10, Ditto in dorsal view. Abbreviations; GC 1, basal gonocoxite 1; GC 2, apical gonocoxite 2." httpUri="https://zenodo.org/record/225199/files/figure.png" pageId="4" pageNumber="193">Fig. 4</figureCitation>
|
||
) tapered apically, faintly twisted leftwards; apical orifice wide, deeply emarginated at left side; apical lobe in dorsal view (
|
||
<figureCitation id="AAAA2A0AF94EFFE9492920E6134EF812" box="[1318,1385,1979,2004]" captionStart="FIGURES 3 – 10" captionStartId="3.[151,264,1894,1917]" captionTargetBox="[161,1430,200,1870]" captionTargetId="figure@3.[151,1436,193,1874]" captionTargetPageId="3" captionText="FIGURES 3 – 10. Genital organs of Ishikawatrechus spp. 3 – 4, 7 – 8, I. bidilatatus sp. nov.; 5 – 6, 9 – 10, I. ishiharai. 3, 5, Aedeagus in left lateral view; 4, 6, Apex of aedeagus in dorsal view; 7, 9, Left stylus in ventral view; 8, 10, Ditto in dorsal view. Abbreviations; GC 1, basal gonocoxite 1; GC 2, apical gonocoxite 2." httpUri="https://zenodo.org/record/225199/files/figure.png" pageId="4" pageNumber="193">Fig. 4</figureCitation>
|
||
) not so extended forward, with rounded apex; ventral hook elongate. Styli (
|
||
<figureCitation id="AAAA2A0AF94FFFE84FE927CA1209FF76" box="[998,1070,151,176]" captionStart="FIGURES 3 – 10" captionStartId="3.[151,264,1894,1917]" captionTargetBox="[161,1430,200,1870]" captionTargetId="figure@3.[151,1436,193,1874]" captionTargetPageId="3" captionText="FIGURES 3 – 10. Genital organs of Ishikawatrechus spp. 3 – 4, 7 – 8, I. bidilatatus sp. nov.; 5 – 6, 9 – 10, I. ishiharai. 3, 5, Aedeagus in left lateral view; 4, 6, Apex of aedeagus in dorsal view; 7, 9, Left stylus in ventral view; 8, 10, Ditto in dorsal view. Abbreviations; GC 1, basal gonocoxite 1; GC 2, apical gonocoxite 2." httpUri="https://zenodo.org/record/225199/files/figure.png" pageId="5" pageNumber="194">Fig. 3</figureCitation>
|
||
) in lateral view feebly curved ventrally, bearing 4 setae on each apex; left one obviously larger than right one.
|
||
</paragraph>
|
||
<paragraph id="322E368FF94FFFE84CC827821374FDFB" blockId="5.[151,1437,151,1545]" pageId="5" pageNumber="194">
|
||
Internal sac in fully inflated condition through eversion (n = 2,
|
||
<figureCitation id="AAAA2A0AF94FFFE84FB027821266FF3E" box="[959,1089,223,248]" captionStart="FIGURES 11 – 19" captionStartId="4.[151,264,1743,1766]" captionTargetBox="[159,1419,200,1714]" captionTargetId="figure@4.[151,1436,193,1722]" captionTargetPageId="4" captionText="FIGURES 11 – 19. Internal sac of Ishikawatrechus spp. 11 – 15, I. bidilatatus sp. nov.; 16 – 19, I. ishiharai. 11, 16, Right lateral view; 12, 17, Left lateral view; 13, 18, Dorsal view; 14, 19, Ventral view; 15, Copulatory piece of I. bidilatatus. Abbreviations; CP, copulatory piece; GP, gonopore." httpUri="https://zenodo.org/record/225200/files/figure.png" pageId="5" pageNumber="194">Figs 11–14</figureCitation>
|
||
) about 1.50 times as long as aedeagus, pyriform, nearly symmetrical though twisted rightwards behind middle; basal half swollen in campanular shape, with two projections on dorsal side and a copulatory piece (CP,
|
||
<figureCitation id="AAAA2A0AF94FFFE84859267A1325FE86" box="[1110,1282,295,320]" captionStart="FIGURES 11 – 19" captionStartId="4.[151,264,1743,1766]" captionTargetBox="[159,1419,200,1714]" captionTargetId="figure@4.[151,1436,193,1722]" captionTargetPageId="4" captionText="FIGURES 11 – 19. Internal sac of Ishikawatrechus spp. 11 – 15, I. bidilatatus sp. nov.; 16 – 19, I. ishiharai. 11, 16, Right lateral view; 12, 17, Left lateral view; 13, 18, Dorsal view; 14, 19, Ventral view; 15, Copulatory piece of I. bidilatatus. Abbreviations; CP, copulatory piece; GP, gonopore." httpUri="https://zenodo.org/record/225200/files/figure.png" pageId="5" pageNumber="194">Figs 11, 13–15</figureCitation>
|
||
) on left side, covered with spicules except for elongated narrow area on basal 1/5 of left latero-ventral side; apical half well swollen in bulb-shape, with spicules on longitudinal area extending from basal half successively to periphery of gonopore through right latero-ventral side; gonopore (GP,
|
||
<figureCitation id="AAAA2A0AF94FFFE84F3526C91599FE6A" box="[826,958,404,429]" captionStart="FIGURES 11 – 19" captionStartId="4.[151,264,1743,1766]" captionTargetBox="[159,1419,200,1714]" captionTargetId="figure@4.[151,1436,193,1722]" captionTargetPageId="4" captionText="FIGURES 11 – 19. Internal sac of Ishikawatrechus spp. 11 – 15, I. bidilatatus sp. nov.; 16 – 19, I. ishiharai. 11, 16, Right lateral view; 12, 17, Left lateral view; 13, 18, Dorsal view; 14, 19, Ventral view; 15, Copulatory piece of I. bidilatatus. Abbreviations; CP, copulatory piece; GP, gonopore." httpUri="https://zenodo.org/record/225200/files/figure.png" pageId="5" pageNumber="194">Figs 11, 14</figureCitation>
|
||
) opening ventral side. Dorsal projections (
|
||
<figureCitation id="AAAA2A0AF94FFFE84C9026EA1701FE16" box="[159,294,439,464]" captionStart="FIGURES 11 – 19" captionStartId="4.[151,264,1743,1766]" captionTargetBox="[159,1419,200,1714]" captionTargetId="figure@4.[151,1436,193,1722]" captionTargetPageId="4" captionText="FIGURES 11 – 19. Internal sac of Ishikawatrechus spp. 11 – 15, I. bidilatatus sp. nov.; 16 – 19, I. ishiharai. 11, 16, Right lateral view; 12, 17, Left lateral view; 13, 18, Dorsal view; 14, 19, Ventral view; 15, Copulatory piece of I. bidilatatus. Abbreviations; CP, copulatory piece; GP, gonopore." httpUri="https://zenodo.org/record/225200/files/figure.png" pageId="5" pageNumber="194">Figs. 11–14</figureCitation>
|
||
) composed of large and small triangular swellings; large one just before the middle of internal sac, small one before the large one. CP hyaline, elongated paddle-shaped, attached on the middle of right side of internal sac, about 1/5 as long as internal sac, rounded at apex. Spicules (
|
||
<figureCitation id="AAAA2A0AF94FFFE8480D26A212ACFDDE" box="[1026,1163,511,536]" captionStart="FIGURES 11 – 19" captionStartId="4.[151,264,1743,1766]" captionTargetBox="[159,1419,200,1714]" captionTargetId="figure@4.[151,1436,193,1722]" captionTargetPageId="4" captionText="FIGURES 11 – 19. Internal sac of Ishikawatrechus spp. 11 – 15, I. bidilatatus sp. nov.; 16 – 19, I. ishiharai. 11, 16, Right lateral view; 12, 17, Left lateral view; 13, 18, Dorsal view; 14, 19, Ventral view; 15, Copulatory piece of I. bidilatatus. Abbreviations; CP, copulatory piece; GP, gonopore." httpUri="https://zenodo.org/record/225200/files/figure.png" pageId="5" pageNumber="194">Figs. 11–15</figureCitation>
|
||
) fine, scaly and feebly sclerotized, gradually getting larger and more spinous dorsally; those on apical half rudimentary sclerotized.
|
||
</paragraph>
|
||
<paragraph id="322E368FF94FFFE84CC8251A1458FD0A" blockId="5.[151,1437,151,1545]" pageId="5" pageNumber="194">
|
||
Female genitalia (n = 2,
|
||
<figureCitation id="AAAA2A0AF94FFFE84DD4251A1467FDA6" box="[475,576,583,608]" captionStart="FIGURES 3 – 10" captionStartId="3.[151,264,1894,1917]" captionTargetBox="[161,1430,200,1870]" captionTargetId="figure@3.[151,1436,193,1874]" captionTargetPageId="3" captionText="FIGURES 3 – 10. Genital organs of Ishikawatrechus spp. 3 – 4, 7 – 8, I. bidilatatus sp. nov.; 5 – 6, 9 – 10, I. ishiharai. 3, 5, Aedeagus in left lateral view; 4, 6, Apex of aedeagus in dorsal view; 7, 9, Left stylus in ventral view; 8, 10, Ditto in dorsal view. Abbreviations; GC 1, basal gonocoxite 1; GC 2, apical gonocoxite 2." httpUri="https://zenodo.org/record/225199/files/figure.png" pageId="5" pageNumber="194">Figs 7–8</figureCitation>
|
||
). GC1 rectangular, with 4–5 short setae on ventral side of apical margin and 2 short setae on dorsal side of inner margin near apex; GC2 thick, slightly curved outwardly, roundly pointed at apex, weakly emarginated near base on outer margin, with 2 ensiform setae on dorsal side near base and 2 nematiform setae on ventral side near apex.
|
||
</paragraph>
|
||
<paragraph id="322E368FF94FFFE84CC8258A14A6FCF1" blockId="5.[151,1437,151,1545]" pageId="5" pageNumber="194">
|
||
<emphasis id="00E5EA9DF94FFFE84CC8258A174CFD36" bold="true" box="[199,363,727,752]" pageId="5" pageNumber="194">Measurement</emphasis>
|
||
(n = 5). TL: 5.34–6.23 (5.75) mm, HL: 0.86–0.99 (0.95) mm, HW: 0.82–0.94(0.88) mm, PL: 1.27–1.45 (1.34) mm, PW: 1.18–1.36 (1.27) mm, PA: 0.82–0.92 (0.88) mm, PB: 0.74–0.84 (0.79) mm, EL: 3.21– 3.79 (3.47) mm, EW: 2.10–2.51 (2.34) mm.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7A8B6504F94FFFE84CC824191586FBD6" pageId="5" pageNumber="194" type="diagnosis">
|
||
<paragraph id="322E368FF94FFFE84CC824191586FBD6" blockId="5.[151,1437,151,1545]" pageId="5" pageNumber="194">
|
||
<emphasis id="00E5EA9DF94FFFE84CC824191766FC9B" bold="true" box="[199,321,836,861]" pageId="5" pageNumber="194">Diagnosis.</emphasis>
|
||
This new species can be easily distinguished from the other congeners of
|
||
<taxonomicName id="F5914D0CF94FFFE8489324191375FC9A" box="[1180,1362,836,860]" class="Insecta" family="Carabidae" genus="Ishikawatrechus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="194" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="00E5EA9DF94FFFE8489324191375FC9A" box="[1180,1362,836,860]" italics="true" pageId="5" pageNumber="194">Ishikawatrechus</emphasis>
|
||
</taxonomicName>
|
||
by its dark coloration and pronotum with subparallel sides at basal 3/7 and short hind angles projected posteriorly. It is similar to
|
||
<taxonomicName id="F5914D0CF94FFFE84D0524D017A0FC62" box="[266,391,908,932]" class="Insecta" family="Carabidae" genus="Ishikawatrechus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="ishiharai">
|
||
<emphasis id="00E5EA9DF94FFFE84D0524D017A0FC62" box="[266,391,908,932]" italics="true" pageId="5" pageNumber="194">I. ishiharai</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="AAAA2A0AF94FFFE84D9824D11464FC62" box="[407,579,908,933]" captionStart="FIGURES 3 – 10" captionStartId="3.[151,264,1894,1917]" captionTargetBox="[161,1430,200,1870]" captionTargetId="figure@3.[151,1436,193,1874]" captionTargetPageId="3" captionText="FIGURES 3 – 10. Genital organs of Ishikawatrechus spp. 3 – 4, 7 – 8, I. bidilatatus sp. nov.; 5 – 6, 9 – 10, I. ishiharai. 3, 5, Aedeagus in left lateral view; 4, 6, Apex of aedeagus in dorsal view; 7, 9, Left stylus in ventral view; 8, 10, Ditto in dorsal view. Abbreviations; GC 1, basal gonocoxite 1; GC 2, apical gonocoxite 2." httpUri="https://zenodo.org/record/225199/files/figure.png" pageId="5" pageNumber="194">Figs 5–6, 9–10</figureCitation>
|
||
,
|
||
<figureCitation id="AAAA2A0AF94FFFE84E5F24D114B1FC63" box="[592,662,908,933]" captionStart="FIGURES 11 – 19" captionStartId="4.[151,264,1743,1766]" captionTargetBox="[159,1419,200,1714]" captionTargetId="figure@4.[151,1436,193,1722]" captionTargetPageId="4" captionText="FIGURES 11 – 19. Internal sac of Ishikawatrechus spp. 11 – 15, I. bidilatatus sp. nov.; 16 – 19, I. ishiharai. 11, 16, Right lateral view; 12, 17, Left lateral view; 13, 18, Dorsal view; 14, 19, Ventral view; 15, Copulatory piece of I. bidilatatus. Abbreviations; CP, copulatory piece; GP, gonopore." httpUri="https://zenodo.org/record/225200/files/figure.png" pageId="5" pageNumber="194">16–19</figureCitation>
|
||
) in general appearance, but well differentiated from the latter by the following characteristics: large body size, pronotum peculiarly elongated, nearly parallel-sided in basal 3/7; hind angles of pronotum protrudent posteriorly; aedeagus hardly arcuate in apical half; internal sac with two remarkable projections on dorsal side and a copulatory piece on right lateral side.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7A8B6504F94FFFE84CC8234113ABFB66" pageId="5" pageNumber="194" type="etymology">
|
||
<paragraph id="322E368FF94FFFE84CC82341136EFBF2" blockId="5.[151,1437,151,1545]" box="[199,1353,1052,1077]" pageId="5" pageNumber="194">
|
||
<emphasis id="00E5EA9DF94FFFE84CC823411769FBF3" bold="true" box="[199,334,1052,1077]" pageId="5" pageNumber="194">Etymology.</emphasis>
|
||
The specific name is referring to the two remarkable dorsal projections on the internal sac.
|
||
</paragraph>
|
||
<paragraph id="322E368FF94FFFE84CC8236213ABFB66" blockId="5.[151,1437,151,1545]" pageId="5" pageNumber="194">
|
||
<emphasis id="00E5EA9DF94FFFE84CC8236217AEFB9E" bold="true" box="[199,393,1087,1112]" pageId="5" pageNumber="194">Biological notes.</emphasis>
|
||
The
|
||
<typeStatus id="ED2A882DF94FFFE84DCD231D17D5FB9E" box="[450,498,1088,1112]" pageId="5" pageNumber="194">type</typeStatus>
|
||
locality is located at the headwaters of a small branch of the Shigenobu River in the northwestern part of Shikoku. All materials in this study were collected from an upper hypogean zone, approximately
|
||
<quantity id="F5699B6AF94FFFE84D3023D517B4FB59" box="[319,403,1160,1184]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="194" unit="cm" value="100.0">100 cm</quantity>
|
||
below ground, under a thick layer of volcanic and sedimentary rocks covered with humus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7A8B6504F94FFFE84CC823F1140BF9CE" pageId="5" pageNumber="194" type="discussion">
|
||
<paragraph id="322E368FF94FFFE84CC823F1140BF9CE" blockId="5.[151,1437,151,1545]" pageId="5" pageNumber="194">
|
||
<emphasis id="00E5EA9DF94FFFE84CC823F1171CFB03" bold="true" box="[199,315,1196,1221]" pageId="5" pageNumber="194">Remarks.</emphasis>
|
||
The genus
|
||
<taxonomicName id="F5914D0CF94FFFE84DDC23F114AEFB02" box="[467,649,1196,1220]" class="Insecta" family="Carabidae" genus="Ishikawatrechus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="194" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="00E5EA9DF94FFFE84DDC23F114AEFB02" box="[467,649,1196,1220]" italics="true" pageId="5" pageNumber="194">Ishikawatrechus</emphasis>
|
||
</taxonomicName>
|
||
has been divided into 7 species groups based on the combined characteristics of the external features including the male genital structure (
|
||
<bibRefCitation id="56004B7EF94FFFE84FE8238D124FFB2E" author="Ueno" box="[999,1128,1231,1256]" pageId="5" pageNumber="194" refString="Ueno, S. - I. (1957) Studies on the Japanese Trechinae (VI) (Coleoptera, Harpalidae). Memoirs of the College of Science, University of Kyoto, series B, (24), 179 - 218." type="journal article" year="1957">Uéno 1957</bibRefCitation>
|
||
, 1996, 1999a, 2002, 2008). The new species,
|
||
<taxonomicName id="F5914D0CF94FFFE84D5023A817C9FACA" box="[351,494,1268,1292]" class="Insecta" family="Carabidae" genus="Ishikawatrechus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="bidilatatus">
|
||
<emphasis id="00E5EA9DF94FFFE84D5023A817C9FACA" box="[351,494,1268,1292]" italics="true" pageId="5" pageNumber="194">I. bidilatatus</emphasis>
|
||
</taxonomicName>
|
||
, should belong to the
|
||
<taxonomicName id="F5914D0CF94FFFE84EE623A91569FACA" box="[745,846,1268,1292]" class="Insecta" family="Carabidae" genus="Ishikawatrechus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="ishiharai">
|
||
<emphasis id="00E5EA9DF94FFFE84EE623A91569FACA" box="[745,846,1268,1292]" italics="true" pageId="5" pageNumber="194">ishiharai</emphasis>
|
||
</taxonomicName>
|
||
species group (sensu
|
||
<bibRefCitation id="56004B7EF94FFFE8484A23A912E2FACA" author="Ueno" box="[1093,1221,1268,1292]" pageId="5" pageNumber="194" refString="Ueno, S. - I. (1997) New Anophthalmic Trechines of the Genus Ishikawatrechus (Coleoptera, Trechinae) Discovered by the Late Yoshiteru MurakamiI. Journal of the Speleological Society of Japan, 22, 1 - 13." type="journal article" year="1997">Uéno 1997</bibRefCitation>
|
||
), which consists of 8 species with 1 subspecies and distributed widely in the Ishizuchi Range (
|
||
<bibRefCitation id="56004B7EF94FFFE84815224512B9FAF6" author="Ueno" box="[1050,1182,1304,1328]" pageId="5" pageNumber="194" refString="Ueno, S. - I. (1994) Ishikawatrechus ishiharai, a new anophthalmic trechine beetle from the Ishizuchi Mountains, Southwest Japan (Coleoptera, Trechinae). Transactions of the Shikoku Entomological Society, 20, 339 - 344." type="journal article" year="1994">Uéno 1994</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="56004B7EF94FFFE848A0224512CEFAF6" author="Ueno" box="[1199,1257,1304,1328]" pageId="5" pageNumber="194" refString="Ueno, S. - I. (1997) New Anophthalmic Trechines of the Genus Ishikawatrechus (Coleoptera, Trechinae) Discovered by the Late Yoshiteru MurakamiI. Journal of the Speleological Society of Japan, 22, 1 - 13." type="journal article" year="1997">1997</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="56004B7EF94FFFE848F5224A1364FAF6" author="Ueno" box="[1274,1347,1303,1328]" pageId="5" pageNumber="194" refString="Ueno, S. - I. (1999 b) Ishikawatrechus kusamai (Coleoptera, Trechinae), a new anophthalmic cave trechine from western Shikoku, Japan. Elytra, 27, 81 - 85." type="journal article" year="1999" yearSuffix="b">1999b</bibRefCitation>
|
||
, 2008), because of its close relationship with
|
||
<taxonomicName id="F5914D0CF94FFFE84E58226014FFFA92" box="[599,728,1340,1364]" class="Insecta" family="Carabidae" genus="Ishikawatrechus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="ishiharai">
|
||
<emphasis id="00E5EA9DF94FFFE84E58226014FFFA92" box="[599,728,1340,1364]" italics="true" pageId="5" pageNumber="194">I. ishiharai</emphasis>
|
||
</taxonomicName>
|
||
. The new species shares with
|
||
<taxonomicName id="F5914D0CF94FFFE84847226012EEFA92" box="[1096,1225,1340,1364]" class="Insecta" family="Carabidae" genus="Ishikawatrechus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="ishiharai">
|
||
<emphasis id="00E5EA9DF94FFFE84847226012EEFA92" box="[1096,1225,1340,1364]" italics="true" pageId="5" pageNumber="194">I. ishiharai</emphasis>
|
||
</taxonomicName>
|
||
not only external features but also the basic structure of the internal sac, and their
|
||
<typeStatus id="ED2A882DF94FFFE84F77223C158FFABF" box="[888,936,1377,1401]" pageId="5" pageNumber="194">type</typeStatus>
|
||
localities are extremely close to each other. On the other hand, in spite of those affinities,
|
||
<taxonomicName id="F5914D0CF94FFFE84E9A22D81503FA5A" box="[661,804,1412,1436]" class="Insecta" family="Carabidae" genus="Ishikawatrechus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="bidilatatus">
|
||
<emphasis id="00E5EA9DF94FFFE84E9A22D81503FA5A" box="[661,804,1412,1436]" italics="true" pageId="5" pageNumber="194">I. bidilatatus</emphasis>
|
||
</taxonomicName>
|
||
is well differentiated from
|
||
<taxonomicName id="F5914D0CF94FFFE8485B22D812F7FA5A" box="[1108,1232,1412,1436]" class="Insecta" family="Carabidae" genus="Ishikawatrechus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="ishiharai">
|
||
<emphasis id="00E5EA9DF94FFFE8485B22D812F7FA5A" box="[1108,1232,1412,1436]" italics="true" pageId="5" pageNumber="194">I. ishiharai</emphasis>
|
||
</taxonomicName>
|
||
in the structure of the everted internal sac in a fully inflated condition, having appendixes as mentioned in the diagnosis above. The structures of the everted internal sac in a fully inflated condition are therefore valuable characters for the taxonomic study on the genus
|
||
<taxonomicName id="F5914D0CF94FFFE84D6022AD1402F9CE" box="[367,549,1520,1544]" class="Insecta" family="Carabidae" genus="Ishikawatrechus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="194" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="00E5EA9DF94FFFE84D6022AD1402F9CE" box="[367,549,1520,1544]" italics="true" pageId="5" pageNumber="194">Ishikawatrechus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |