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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1092.75831" ID-GBIF-Dataset="c2f1268d-f466-4148-990f-2b948efcf6a7" ID-PMC="PMC9005484" ID-Pensoft-Pub="1313-2970-1092-31" ID-Pensoft-UUID="B3F12D9131AD5C878EAC9A12C068EA6D" ID-PubMed="35586293" ID-ZooBank="28BB93BFA3D54EF5A91867AECD12B808" ModsDocID="1313-2970-1092-31" checkinTime="1649136097056" checkinUser="pensoft" docAuthor="Zhao, Ming-Zhi &amp; Liu, Wei-Xin" docDate="2022" docId="17B31633B7945791930A48C3A0263F07" docLanguage="en" docName="ZooKeys 1092: 31-45" docOrigin="ZooKeys 1092" docPubDate="2022-04-04" docSource="http://dx.doi.org/10.3897/zookeys.1092.75831" docTitle="Didrepanephorus birmanicus Zhao &amp; Liu 2022, comb. nov." docType="treatment" docVersion="4" id="B3F12D9131AD5C878EAC9A12C068EA6D" lastPageNumber="31" masterDocId="B3F12D9131AD5C878EAC9A12C068EA6D" masterDocTitle="A contribution to the genus Didrepanephorus Wood-Mason, 1878 (Coleoptera, Scarabaeidae, Rutelinae)" masterLastPageNumber="45" masterPageNumber="31" pageNumber="31" updateTime="1668151609506" updateUser="ExternalLinkService">
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<mods:title>A contribution to the genus Didrepanephorus Wood-Mason, 1878 (Coleoptera, Scarabaeidae, Rutelinae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zhao, Ming-Zhi</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-3158-6177</mods:nameIdentifier>
<mods:affiliation>College of Plant Protection, South China Agricultural University, Guangzhou, 510642, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Liu, Wei-Xin</mods:namePart>
<mods:affiliation>College of Plant Protection, South China Agricultural University, Guangzhou, 510642, China</mods:affiliation>
<mods:nameIdentifier type="email">da2000wei@163.com</mods:nameIdentifier>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-04-04</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1092</mods:number>
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<mods:start>31</mods:start>
<mods:end>45</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1092.75831</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1092.75831</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1092-31</mods:identifier>
<mods:identifier type="ZooBank">28BB93BFA3D54EF5A91867AECD12B808</mods:identifier>
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<treatment ID-GBIF-Taxon="194444416" LSID="urn:lsid:plazi:treatment:17B31633B7945791930A48C3A0263F07" httpUri="http://treatment.plazi.org/id/17B31633B7945791930A48C3A0263F07" lastPageNumber="31" pageId="0" pageNumber="31">
<subSubSection pageId="0" pageNumber="31" type="nomenclature">
<paragraph pageId="0" pageNumber="31">
<taxonomicName LSID="17B31633-B794-5791-930A-48C3A0263F07" authority="(Arrow, 1907)" authorityName="Zhao &amp; Liu" authorityYear="2022" baseAuthorityName="Arrow" baseAuthorityYear="1907" class="Insecta" family="Scarabaeidae" genus="Didrepanephorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Didrepanephorus birmanicus" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="birmanicus" status="comb. nov.">Didrepanephorus birmanicus (Arrow, 1907)</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="31">comb. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="31" type="description">
<paragraph pageId="0" pageNumber="31">
<figureCitation captionStart="Figures 710" captionStartId="F2" captionText="Figures 7 - 10. Morphological details of Didrepanephorus Didrepanephorus species 7 head of holotype of D. tangzhaoyangi Zhao &amp; Liu, sp. nov., oblique lateral view 8 head of D. birmanicus (Arrow, 1907) (Lectotype of Fruhstorferia birmanica Arrow, 1907), oblique lateral view, photo © Keita Matsumoto (NHMUK) 9 - 10 male genitalia of holotype of D. tangzhaoyangi 9 dorsal view 10 ventral view. Scale bar for genitalia only." figureDoi="10.3897/zookeys.1092.75831.figures7-10" httpUri="https://binary.pensoft.net/fig/668068" pageId="0" pageNumber="31">Figs 8</figureCitation>
<figureCitation captionStart="Figures 1118" captionStartId="F3" captionText="Figures 11 - 18. Type material of Didrepanephorus species. 11 - 15 male of D. birmanicus (Arrow, 1907) (Lectotype of Fruhstorferia birmanica Arrow, 1907) 16 - 18 female of D. mizunumai Nagai &amp; Hirasawa, 1991 (paralectotype of F. birmanica), habitus in dorsal view 11, 16 habitus in dorsal view 12, 17 habitus in lateral view 13 aedeagus showing parameres in dorsal view 14 aedeagus in dorsal view 15, 18 attached labels. All photos © Keita Matsumoto (NHMUK)." figureDoi="10.3897/zookeys.1092.75831.figures11-18" httpUri="https://binary.pensoft.net/fig/668069" pageId="0" pageNumber="31">, 11-15</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="31" type="reference_group">
<paragraph pageId="0" pageNumber="31">
<taxonomicName authorityName="Zhao &amp; Liu" authorityYear="2022" baseAuthorityName="Zhao &amp; Liu" baseAuthorityYear="2022" class="Insecta" family="Scarabaeidae" genus="Fruhstorferia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fruhstorferia birmanica" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="birmanica" status="comb. nov.">Fruhstorferia birmanica</taxonomicName>
Arrow, 1907: 354 [original description];
<bibRefCitation author="Arrow, GJ" journalOrPublisher="Taylor &amp; Francis, London" pageId="0" pageNumber="31" refId="B2" refString="Arrow, GJ, 1917. The Fauna of British India, Including Ceylon and Burma. ColeopteraLamellicornia part II (Rutelinae, Desmonycinae, and Euchirinae). Taylor &amp; Francis, London" title="The Fauna of British India, Including Ceylon and Burma. ColeopteraLamellicornia part II (Rutelinae, Desmonycinae, and Euchirinae)." year="1917">Arrow 1917</bibRefCitation>
: 49 [partim], fig. 15;
<bibRefCitation author="Ohaus, F" journalOrPublisher="Junk, s' - Gravenhage, Berlin" pageId="0" pageNumber="31" refId="B30" refString="Ohaus, F, 1918. Scarabaeidae: Euchirinae, Phaenomerinae, Rutelinae. In: Schenkling S (Ed.) Coleopterorum catalogus. Pars 66. Dr. W. Junk, s'-Gravenhage, Berlin" title="Scarabaeidae: Euchirinae, Phaenomerinae, Rutelinae. In: Schenkling S (Ed.) Coleopterorum catalogus. Pars 66. Dr. W." year="1918">Ohaus 1918</bibRefCitation>
: 43 [catalogued];
<bibRefCitation author="Ohaus, F" journalOrPublisher="Desmet-Verteneuil, Bruxelles" pageId="0" pageNumber="31" refId="B32" refString="Ohaus, F, 1934b. ColeopteraLamellicornia Fam. Scarabaeidae Subfam. Rutelinae. Erster Teil. In: Wytsman PAG (Ed.) Genera Insectorum, Fasc. 199 (A). Desmet-Verteneuil, Bruxelles" title="ColeopteraLamellicornia Fam. Scarabaeidae Subfam. Rutelinae. Erster Teil. In: Wytsman PAG (Ed.) Genera Insectorum, Fasc. 199 (A)." year="1934 b">Ohaus 1934b</bibRefCitation>
: 121 [catalogued];
<bibRefCitation author="Machatschke, JW" journalOrPublisher="Junk, s' - Gravenhage, Berlin" pageId="0" pageNumber="31" refId="B13" refString="Machatschke, JW, 1972. Coleopterorum catalogus supplementa. Pars 66, fascicle 1. Superfamilie Scarabaeoidea, Familie Melolonthidae,Subfamilie Rutelinae. Editio Secunda. Dr. W. Junk, s'-Gravenhage, Berlin" title="Coleopterorum catalogus supplementa. Pars 66, fascicle 1. Superfamilie Scarabaeoidea, Familie Melolonthidae, Subfamilie Rutelinae. Editio Secunda. Dr. W." year="1972">Machatschke 1972</bibRefCitation>
: 53 [catalogued];
<bibRefCitation author="Muramoto, R" journalOrPublisher="Kogane" pageId="0" pageNumber="31" pagination="12 - 16" refId="B21" refString="Muramoto, R, Araya, K, 2000. A new species of the genus Fruhstorferia Kolbe, 1894 from northern Thailand. Kogane 1: 12 - 16" title="A new species of the genus Fruhstorferia Kolbe, 1894 from northern Thailand." volume="1" year="2000">Muramoto and Araya 2000</bibRefCitation>
: 12, figs 3-4 [syntype male figured];
<bibRefCitation author="Krajcik, M" journalOrPublisher="Stettiner Entomologische Zeitung" pageId="0" pageNumber="31" refId="B11" refString="Krajcik, M, 2007. Checklist of the Scarabaeoidea of the World. 2. Rutelinae (Coleoptera: Scarabaeidae: Rutelinae). Animma. x(Supplementum 4): 1-139." title="Checklist of the Scarabaeoidea of the World. 2. Rutelinae (Coleoptera: Scarabaeidae: Rutelinae). Animma. x (Supplementum 4): 1 - 139." year="2007">
<normalizedToken originalValue="Krajčík">Krajcik</normalizedToken>
2007
</bibRefCitation>
: 70 [catalogued];
<bibRefCitation author="Krajcik, M" journalOrPublisher="Stettiner Entomologische Zeitung" pageId="0" pageNumber="31" refId="B12" refString="Krajcik, M, 2012. Checklist of the world Scarabaeoidea. Animma. x(Supplementum 5): 1-139." title="Checklist of the world Scarabaeoidea. Animma. x (Supplementum 5): 1 - 139." year="2012">
<normalizedToken originalValue="Krajčík">Krajcik</normalizedToken>
2012
</bibRefCitation>
: 110 [catalogued, subgeneric placement unnoted].
</paragraph>
<paragraph pageId="0" pageNumber="31">
<taxonomicName authorityName="Zhao &amp; Liu" authorityYear="2022" baseAuthorityName="Zhao &amp; Liu" baseAuthorityYear="2022" class="Insecta" family="Scarabaeidae" genus="Fruhstorferia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fruhstorferia (Kibakoganea) birmanica" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="birmanica" status="comb. nov." subGenus="Kibakoganea">Fruhstorferia (Kibakoganea) birmanica</taxonomicName>
Arrow, 1907:
<bibRefCitation author="Nagai, S" journalOrPublisher="Transactions of the Shikoku Entomological Society" pageId="0" pageNumber="31" pagination="25 - 31" refId="B24" refString="Nagai, S, 1984. Two new may-beetles of the genus Fruhstorferia from Malaysia, with the descriptions of a new subgenus. Transactions of the Shikoku Entomological Society 16 (4): 25 - 31" title="Two new may-beetles of the genus Fruhstorferia from Malaysia, with the descriptions of a new subgenus." volume="16" year="1984">Nagai 1984</bibRefCitation>
: 29 [catalogued].
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="31" type="type locality">
<paragraph pageId="0" pageNumber="31">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="31">&quot;Burma, Ruby Mines&quot;, in currently Mogok City of Mandalay, Myanmar.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="31" type="type material">
<paragraph pageId="0" pageNumber="31">Type material</paragraph>
<paragraph pageId="0" pageNumber="31">
(1 specimen).
<emphasis bold="true" italics="true" pageId="0" pageNumber="31">Lectotype</emphasis>
of
<taxonomicName authorityName="Zhao &amp; Liu" authorityYear="2022" baseAuthorityName="Zhao &amp; Liu" baseAuthorityYear="2022" class="Insecta" family="Scarabaeidae" genus="Fruhstorferia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fruhstorferia birmanica" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="birmanica">
<emphasis italics="true" pageId="0" pageNumber="31">Fruhstorferia birmanica</emphasis>
</taxonomicName>
Arrow, 1907 (hereby designated). ♂ (BMNH), &quot;Birmah [R]uby Mes // Doherty // 64607 // Fry Coll. 1905-100. //
<taxonomicName authorityName="Zhao &amp; Liu" authorityYear="2022" baseAuthorityName="Zhao &amp; Liu" baseAuthorityYear="2022" class="Insecta" family="Scarabaeidae" genus="Fruhstorferia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fruhstorferia birmanica" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="birmanica">
<emphasis italics="true" pageId="0" pageNumber="31">Fruhstorferia birmanica</emphasis>
</taxonomicName>
, Arrow type ♂ A. a M., 1907. // Type // HOLOTYPE
<taxonomicName authorityName="Zhao &amp; Liu" authorityYear="2022" baseAuthorityName="Zhao &amp; Liu" baseAuthorityYear="2022" class="Insecta" family="Scarabaeidae" genus="Fruhstorferia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fruhstorferia birmanica" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="birmanica">
<emphasis italics="true" pageId="0" pageNumber="31">Fruhstorferia birmanica</emphasis>
</taxonomicName>
A. M. SOULA det 1994 // NHMUK014379787&quot;. It will be provided with an additional red label: LECTOTYPE
<taxonomicName authorityName="Zhao &amp; Liu" authorityYear="2022" baseAuthorityName="Zhao &amp; Liu" baseAuthorityYear="2022" class="Insecta" family="Scarabaeidae" genus="Fruhstorferia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fruhstorferia birmanica" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="birmanica">
<emphasis italics="true" pageId="0" pageNumber="31">Fruhstorferia birmanica</emphasis>
</taxonomicName>
Arrow, 1907 des. Zhao Ming-Zhi 2021.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="31" type="remarks">
<paragraph pageId="0" pageNumber="31">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="31">
The taxon
<taxonomicName authorityName="Zhao &amp; Liu" authorityYear="2022" baseAuthorityName="Zhao &amp; Liu" baseAuthorityYear="2022" class="Insecta" family="Scarabaeidae" genus="Fruhstorferia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fruhstorferia birmanica" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="birmanica">
<emphasis italics="true" pageId="0" pageNumber="31">Fruhstorferia birmanica</emphasis>
</taxonomicName>
was originally described based on a pair of specimens from Ruby Mines, Burma (
<bibRefCitation DOI="https://doi.org/10.1080/00222930709487274" author="Arrow, GJ" pageId="0" pageNumber="31" refId="B1" refString="Arrow, GJ, 1907. Some new species and genera of lamellicorn Coleoptera from the Indian Empire (part 1). The Annals and Magazine of natural History, including Zoology, Botany and Geology Series 7 19(112): 347-359. https://doi.org/10.1080/00222930709487274" title="Some new species and genera of lamellicorn Coleoptera from the Indian Empire (part 1). The Annals and Magazine of natural History, including Zoology, Botany and Geology Series 7 19 (112): 347 - 359." url="https://doi.org/10.1080/00222930709487274" year="1907">Arrow 1907</bibRefCitation>
). The lectotype has not yet been designated and the male illustrated by
<bibRefCitation author="Muramoto, R" journalOrPublisher="Kogane" pageId="0" pageNumber="31" pagination="12 - 16" refId="B21" refString="Muramoto, R, Araya, K, 2000. A new species of the genus Fruhstorferia Kolbe, 1894 from northern Thailand. Kogane 1: 12 - 16" title="A new species of the genus Fruhstorferia Kolbe, 1894 from northern Thailand." volume="1" year="2000">Muramoto and Araya (2000)</bibRefCitation>
has been erroneously fixed as holotype. According to Art. 74.7.1 (ICZN 1999), both specimens should be regarded as syntypes. The examination of both syntypes reveals that the female is conspecific with
<taxonomicName lsidName="D. mizunumai" pageId="0" pageNumber="31" rank="species" species="mizunumai">
<emphasis italics="true" pageId="0" pageNumber="31">D. mizunumai</emphasis>
</taxonomicName>
Nagai &amp; Hirasawa, 1991. Therefore, a lectotype designation is necessary.
</paragraph>
<paragraph pageId="0" pageNumber="31">
The male genitalia of the lectotype (Figs
<figureCitation captionStart="Figures 710" captionStartId="F2" captionText="Figures 7 - 10. Morphological details of Didrepanephorus Didrepanephorus species 7 head of holotype of D. tangzhaoyangi Zhao &amp; Liu, sp. nov., oblique lateral view 8 head of D. birmanicus (Arrow, 1907) (Lectotype of Fruhstorferia birmanica Arrow, 1907), oblique lateral view, photo © Keita Matsumoto (NHMUK) 9 - 10 male genitalia of holotype of D. tangzhaoyangi 9 dorsal view 10 ventral view. Scale bar for genitalia only." figureDoi="10.3897/zookeys.1092.75831.figures7-10" httpUri="https://binary.pensoft.net/fig/668068" pageId="0" pageNumber="31">8</figureCitation>
,
<figureCitation captionStart="Figures 1118" captionStartId="F3" captionText="Figures 11 - 18. Type material of Didrepanephorus species. 11 - 15 male of D. birmanicus (Arrow, 1907) (Lectotype of Fruhstorferia birmanica Arrow, 1907) 16 - 18 female of D. mizunumai Nagai &amp; Hirasawa, 1991 (paralectotype of F. birmanica), habitus in dorsal view 11, 16 habitus in dorsal view 12, 17 habitus in lateral view 13 aedeagus showing parameres in dorsal view 14 aedeagus in dorsal view 15, 18 attached labels. All photos © Keita Matsumoto (NHMUK)." figureDoi="10.3897/zookeys.1092.75831.figures11-18" httpUri="https://binary.pensoft.net/fig/668069" pageId="0" pageNumber="31">11-15</figureCitation>
) is partly damaged, which greatly complicates the comparison to its related
<taxonomicName class="Insecta" family="Scarabaeidae" genus="Didrepanephorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Didrepanephorus species" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species">Didrepanephorus species</taxonomicName>
. However, the similarity of this
<taxonomicName class="Insecta" family="Scarabaeidae" genus="Didrepanephorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Didrepanephorus species" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species">Didrepanephorus species</taxonomicName>
to small-sized male of
<taxonomicName lsidName="D. yunnanus" pageId="0" pageNumber="31" rank="species" species="yunnanus">
<emphasis italics="true" pageId="0" pageNumber="31">D. yunnanus</emphasis>
</taxonomicName>
(Ohaus, 1911) is still apparent in many external features, especially in the allied shape of terminal segment of maxillary palp, which is more expanded and compressed than other
<taxonomicName authorityName="Wood-mason" authorityYear="1878" class="Insecta" family="Scarabaeidae" genus="Didrepanephorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Didrepanephorus" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="31">Didrepanephorus</emphasis>
</taxonomicName>
<taxonomicName class="Insecta" family="Scarabaeidae" genus="Didrepanephorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Didrepanephorus species" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species">Didrepanephorus species</taxonomicName>
. There is a strong concavity at the base of lower margin of mandible. The protrusion at lower margin is partly hidden under the labrum. These characters match the above definitions for
<taxonomicName authorityName="Wood-mason" authorityYear="1878" class="Insecta" family="Scarabaeidae" genus="Didrepanephorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Didrepanephorus" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="31">Didrepanephorus</emphasis>
</taxonomicName>
. Thus, this
<taxonomicName class="Insecta" family="Scarabaeidae" genus="Didrepanephorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Didrepanephorus species" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species">Didrepanephorus species</taxonomicName>
is transferred to the genus
<taxonomicName authorityName="Wood-mason" authorityYear="1878" class="Insecta" family="Scarabaeidae" genus="Didrepanephorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Didrepanephorus" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="31">Didrepanephorus</emphasis>
</taxonomicName>
herein. Among the five subspecies of
<taxonomicName lsidName="D. yunnanus" pageId="0" pageNumber="31" rank="species" species="yunnanus">
<emphasis italics="true" pageId="0" pageNumber="31">D. yunnanus</emphasis>
</taxonomicName>
, i.e.,
<taxonomicName lsidName="D. y. subsp. yunnanus" pageId="0" pageNumber="31" rank="subSpecies" species="y." subSpecies="yunnanus">
<emphasis italics="true" pageId="0" pageNumber="31">D. y. yunnanus</emphasis>
</taxonomicName>
(Ohaus, 1911),
<taxonomicName lsidName="D. y. subsp. clermonti" pageId="0" pageNumber="31" rank="subSpecies" species="y." subSpecies="clermonti">
<emphasis italics="true" pageId="0" pageNumber="31">D. y. clermonti</emphasis>
</taxonomicName>
Benderitter, 1929,
<taxonomicName lsidName="D. y. subsp. piaoacensis" pageId="0" pageNumber="31" rank="subSpecies" species="y." subSpecies="piaoacensis">
<emphasis italics="true" pageId="0" pageNumber="31">D. y. piaoacensis</emphasis>
</taxonomicName>
Nagai, 2004,
<taxonomicName lsidName="D. y. subsp. wakaharai" pageId="0" pageNumber="31" rank="subSpecies" species="y." subSpecies="wakaharai">
<emphasis italics="true" pageId="0" pageNumber="31">D. y. wakaharai</emphasis>
</taxonomicName>
Nagai, 2004, and
<taxonomicName lsidName="D. y. subsp. kachinensis" pageId="0" pageNumber="31" rank="subSpecies" species="y." subSpecies="kachinensis">
<emphasis italics="true" pageId="0" pageNumber="31">D. y. kachinensis</emphasis>
</taxonomicName>
Muramoto, 2005, recognized by
<bibRefCitation author="Muramoto, R" journalOrPublisher="Kogane" pageId="0" pageNumber="31" pagination="23 - 32" refId="B17" refString="Muramoto, R, 2005. Notes on Didrepanephorus yunnanus (Ohaus, 1911) and the genus Kibakoganea Nagai, 1984. Kogane 6: 23 - 32" title="Notes on Didrepanephorus yunnanus (Ohaus, 1911) and the genus Kibakoganea Nagai, 1984." volume="6" year="2005">Muramoto (2005)</bibRefCitation>
,
<taxonomicName lsidName="D. y. subsp. kachinensis" pageId="0" pageNumber="31" rank="subSpecies" species="y." subSpecies="kachinensis">
<emphasis italics="true" pageId="0" pageNumber="31">D. y. kachinensis</emphasis>
</taxonomicName>
appears to be most similar to
<taxonomicName lsidName="D. birmanicus" pageId="0" pageNumber="31" rank="species" species="birmanicus">
<emphasis italics="true" pageId="0" pageNumber="31">D. birmanicus</emphasis>
</taxonomicName>
due to the generally reddish brown body in combination to the closest geographical distance. To fully understand the relation between the two
<taxonomicName class="Insecta" family="Scarabaeidae" genus="Didrepanephorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Didrepanephorus species" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species">Didrepanephorus species</taxonomicName>
, examination of topotypical specimens of
<taxonomicName lsidName="D. birmanicus" pageId="0" pageNumber="31" rank="species" species="birmanicus">
<emphasis italics="true" pageId="0" pageNumber="31">D. birmanicus</emphasis>
</taxonomicName>
with undamaged male genitalia is needed.
</paragraph>
<paragraph pageId="0" pageNumber="31">
This
<taxonomicName class="Insecta" family="Scarabaeidae" genus="Didrepanephorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Didrepanephorus species" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species">Didrepanephorus species</taxonomicName>
was partly misinterpreted and the name was previously applied to two different
<taxonomicName class="Insecta" family="Scarabaeidae" genus="Didrepanephorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Didrepanephorus species" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species">Didrepanephorus species</taxonomicName>
(see below). This
<taxonomicName class="Insecta" family="Scarabaeidae" genus="Didrepanephorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Didrepanephorus species" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species">Didrepanephorus species</taxonomicName>
is so far only known from the lectotype.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="31" type="distribution">
<paragraph pageId="0" pageNumber="31">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="31">Myanmar (Mandalay Region: Mogok).</paragraph>
</subSubSection>
</treatment>
</document>