183 lines
17 KiB
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183 lines
17 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zse.97.59143" ID-GBIF-Dataset="83d5b180-4e0b-44d1-ad83-67ad55ee0355" ID-GBIF-Taxon="182236441" ID-Pensoft-Pub="1860-0743-1-1" ID-Pensoft-UUID="BD006A3AD2C6516AB596CDA05E0F375D" ID-ZooBank="1D5553BA24894F2C9527B7E46B9B432B" ModsDocID="1860-0743-97-1-1" checkinTime="1621423297205" checkinUser="pensoft" docAuthor="Jirapatrasilp, Parin, Pall-Gergely, Barna, Sutcharit, Chirasak & Tongkerd, Piyoros" docDate="2021" docId="E6B6BF891D4A5897B6A04011D0E21EBA" docLanguage="en" docName="ZoosystEvol 97(1): 1-20" docOrigin="Zoosystematics and Evolution 97 (1)" docPubDate="2021-01-05" docSource="http://dx.doi.org/10.3897/zse.97.59143" docTitle="Dicharax pratatensis" docType="treatment" docVersion="3" id="BD006A3AD2C6516AB596CDA05E0F375D" lastPageNumber="1" masterDocId="BD006A3AD2C6516AB596CDA05E0F375D" masterDocTitle="The operculate micro land snail genus Dicharax Kobelt & Moellendorff, 1900 (Caenogastropoda, Alycaeidae) in Thailand, with description of new species" masterLastPageNumber="20" masterPageNumber="1" pageId="0" pageNumber="1" updateTime="1632012559486" updateUser="ExternalLinkService">
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<mods:titleInfo>
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<mods:title>The operculate micro land snail genus Dicharax Kobelt & Moellendorff, 1900 (Caenogastropoda, Alycaeidae) in Thailand, with description of new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Jirapatrasilp, Parin</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-5591-6724</mods:nameIdentifier>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Pall-Gergely, Barna</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6167-7221</mods:nameIdentifier>
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<mods:affiliation>Plant Protection Institute, Centre for Agricultural Research, Herman Otto ut 15, H- 1022 Budapest, Hungary</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Tongkerd, Piyoros</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-9221-9293</mods:nameIdentifier>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:nameIdentifier type="email">piyorose@hotmail.com</mods:nameIdentifier>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>Zoosystematics and Evolution</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-01-05</mods:number>
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<mods:detail type="volume">
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<mods:number>97</mods:number>
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<mods:number>1</mods:number>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>20</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zse.97.59143</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zse.97.59143</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1860-0743-1-1</mods:identifier>
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<mods:identifier type="ZooBank">1D5553BA24894F2C9527B7E46B9B432B</mods:identifier>
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<mods:identifier type="Pensoft-UUID">BD006A3AD2C6516AB596CDA05E0F375D</mods:identifier>
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<treatment ID-GBIF-Taxon="182236441" LSID="urn:lsid:plazi:treatment:E6B6BF891D4A5897B6A04011D0E21EBA" httpUri="http://treatment.plazi.org/id/E6B6BF891D4A5897B6A04011D0E21EBA" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="E6B6BF89-1D4A-5897-B6A0-4011D0E21EBA" authority="(Panha & Burch, 1997)" baseAuthorityName="Panha & Burch" baseAuthorityYear="1997" class="Gastropoda" family="Cyclophoridae" genus="Dicharax" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Dicharax pratatensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="pratatensis">Dicharax pratatensis (Panha & Burch, 1997)</taxonomicName>
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Synoptic view of the eight Dicharax taxa in Thailand. A. Dicharax omissus, specimen CUMZ 7426 (gold plated); B. Dicharax pratatensis, specimen CUMZ 7427; C. Dicharax panhai sp. nov., paratype CUMZ 7429 / 2; D. Dicharax pongrati sp. nov., paratype CUMZ 7430 / 2; E. Dicharax notus, specimen CUMZ 7425; F. Dicharax burchi sp. nov., paratype CUMZ 7428 / 2; G. Dicharax borealis sp. nov., paratype CUMZ 5072 / 2; H. Dicharax cucullatus, specimen CUMZ 7421." figureDoi="10.3897/zse.97.59143.figure6" httpUri="https://binary.pensoft.net/fig/494370" pageId="0" pageNumber="1">Figs 6B</figureCitation>
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. A, B. Dicharax omissus A. Syntype NHMUK 1903.7.1.1228; B. Specimen CUMZ 7426 from Tham Mae La Na, Mae Hong Son Province C, D. Dicharax pratatensis C. Holotype CUMZ Cy 001; D. Paratype CUMZ Cy 002 from Doi Chiang Dao, Chiang Mai Province. Close-up images of the aperture not to scale." figureDoi="10.3897/zse.97.59143.figure8" httpUri="https://binary.pensoft.net/fig/494372" pageId="0" pageNumber="1">, 8C, D</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="reference_group">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus pratatensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="pratatensis">Alycaeus pratatensis</taxonomicName>
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Panha & Burch, 1997: 119-122, fig. 2a-c. Type locality: "Tham Pratat, Erawan National Park, Kanchanaburi Province, Thailand", hereby emended to: "Doi Chiang Dao, Chiang Mai Province, Northern Thailand".
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type material">
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<paragraph pageId="0" pageNumber="1">Type material examined.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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CUMZ Cy 001 (Fig.
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. A, B. Dicharax omissus A. Syntype NHMUK 1903.7.1.1228; B. Specimen CUMZ 7426 from Tham Mae La Na, Mae Hong Son Province C, D. Dicharax pratatensis C. Holotype CUMZ Cy 001; D. Paratype CUMZ Cy 002 from Doi Chiang Dao, Chiang Mai Province. Close-up images of the aperture not to scale." figureDoi="10.3897/zse.97.59143.figure8" httpUri="https://binary.pensoft.net/fig/494372" pageId="0" pageNumber="1">8C</figureCitation>
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),
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">paratypes</emphasis>
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CUMZ Cy 002 (2 shells; Fig.
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. A, B. Dicharax omissus A. Syntype NHMUK 1903.7.1.1228; B. Specimen CUMZ 7426 from Tham Mae La Na, Mae Hong Son Province C, D. Dicharax pratatensis C. Holotype CUMZ Cy 001; D. Paratype CUMZ Cy 002 from Doi Chiang Dao, Chiang Mai Province. Close-up images of the aperture not to scale." figureDoi="10.3897/zse.97.59143.figure8" httpUri="https://binary.pensoft.net/fig/494372" pageId="0" pageNumber="1">8D</figureCitation>
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), FMNH (1 shell), NHMUK 20200324 (2 shells) and UMMZ (1 shell) from Doi Chiang Dao, Chiang Mai Province, Thailand,
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<geoCoordinate degrees="19" direction="north" minutes="14" orientation="latitude" precision="1" seconds="26.7" value="19.24075">19°14'26.7"N</geoCoordinate>
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,
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<geoCoordinate degrees="98" direction="east" minutes="52" orientation="longitude" precision="1" seconds="49.8" value="98.8805">98°52'49.8"E</geoCoordinate>
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, 24-28 May 1997, coll. S. Panha, J.B. Burch, P. Dumrongrojwattana.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Other material examined.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<materialsCitation collectingDate="1997-05-24" collectingDateMax="1997-05-28" collectingDateMin="1997-05-24" collectorName="S. Panha, Panha, J. B, Burch, P" country="Thailand" latitude="19.24075" location="Doi Chiang Dao" longLatPrecision="1" longitude="98.8805" specimenCount="1" stateProvince="Chiang Mai Province">
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CUMZ 7427 from
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<location LSID="urn:lsid:plazi:treatment:E6B6BF891D4A5897B6A04011D0E21EBA:42334BE520C6FEB26077729357997FC9" country="Thailand" latitude="19.24075" longLatPrecision="1" longitude="98.8805" name="Doi Chiang Dao" stateProvince="Chiang Mai Province">Doi Chiang Dao</location>
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,
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<collectingRegion country="Thailand" name="Chiang Mai">Chiang Mai Province</collectingRegion>
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,
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<collectingCountry name="Thailand">Thailand</collectingCountry>
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,
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<geoCoordinate degrees="19" direction="north" minutes="14" orientation="latitude" precision="1" seconds="26.7" value="19.24075">19°14'26.7"N</geoCoordinate>
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,
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<geoCoordinate degrees="98" direction="east" minutes="52" orientation="longitude" precision="1" seconds="49.8" value="98.8805">98°52'49.8"E</geoCoordinate>
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,
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<collectingDate value="1997-05-24" valueMax="1997-05-28" valueMin="1997-05-24">24-28 May 1997</collectingDate>
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, coll.
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<collectorName>S.
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<collectorName>Panha, J.B</collectorName>
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</collectorName>
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.
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<collectorName>Burch, P</collectorName>
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. Dumrongrojwattana (16 shells; Fig.
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Synoptic view of the eight Dicharax taxa in Thailand. A. Dicharax omissus, specimen CUMZ 7426 (gold plated); B. Dicharax pratatensis, specimen CUMZ 7427; C. Dicharax panhai sp. nov., paratype CUMZ 7429 / 2; D. Dicharax pongrati sp. nov., paratype CUMZ 7430 / 2; E. Dicharax notus, specimen CUMZ 7425; F. Dicharax burchi sp. nov., paratype CUMZ 7428 / 2; G. Dicharax borealis sp. nov., paratype CUMZ 5072 / 2; H. Dicharax cucullatus, specimen CUMZ 7421." figureDoi="10.3897/zse.97.59143.figure6" httpUri="https://binary.pensoft.net/fig/494370" pageId="0" pageNumber="1">6B</figureCitation>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">Shell small (SH up to 2 mm, SW up to 3.2 mm), depressed-conical. Spire ca. 1/6 of shell height. R1 with regular, very strong ribs; R2 shorter than R3; R2 with 44-51 ribs. R3 with prominent, blunt swelling near aperture, anterior slope of swelling steeper than posterior slope. Aperture round. Inner peristome thickened, protruding, with slight parieto-columellar angle. Outer peristome expanded, not reflected. Umbilicus round. Operculum multispiral.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="distribution">
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<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="1">Known only from Doi Chiang Dao, Chiang Mai Province, Thailand.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="remarks">
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<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Re-examination of the type materials (holotype Cy 001: Fig.
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. A, B. Dicharax omissus A. Syntype NHMUK 1903.7.1.1228; B. Specimen CUMZ 7426 from Tham Mae La Na, Mae Hong Son Province C, D. Dicharax pratatensis C. Holotype CUMZ Cy 001; D. Paratype CUMZ Cy 002 from Doi Chiang Dao, Chiang Mai Province. Close-up images of the aperture not to scale." figureDoi="10.3897/zse.97.59143.figure8" httpUri="https://binary.pensoft.net/fig/494372" pageId="0" pageNumber="1">8C</figureCitation>
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and paratypes Cy 002: Fig.
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. A, B. Dicharax omissus A. Syntype NHMUK 1903.7.1.1228; B. Specimen CUMZ 7426 from Tham Mae La Na, Mae Hong Son Province C, D. Dicharax pratatensis C. Holotype CUMZ Cy 001; D. Paratype CUMZ Cy 002 from Doi Chiang Dao, Chiang Mai Province. Close-up images of the aperture not to scale." figureDoi="10.3897/zse.97.59143.figure8" httpUri="https://binary.pensoft.net/fig/494372" pageId="0" pageNumber="1">8D</figureCitation>
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) deposited in CUMZ and figured in the original description (
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<bibRefCitation author="Panha, S" journalOrPublisher="Malacological Review" pageId="0" pageNumber="1" pagination="119 - 122" refId="B48" refString="Panha, S, Burch, JB, 1997. A new cave dweller of the genus Alycaeus in Thailand (Prosobranchia: Cyclophoracea: Family Cyclophoridae). Malacological Review 30: 119 - 122" title="A new cave dweller of the genus Alycaeus in Thailand (Prosobranchia: Cyclophoracea: Family Cyclophoridae)." volume="30" year="1997">Panha and Burch 1997</bibRefCitation>
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: fig. 2) revealed that the locality code associated with the types points to "Doi Chiang Dao". However, the locality of the paratypes of
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<taxonomicName class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus pratatensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="pratatensis">
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<emphasis italics="true" pageId="0" pageNumber="1">Alycaeus pratatensis</emphasis>
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</taxonomicName>
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deposited in SMF (inv. number: 331452) appears as "Tham Pratat, Erawan National Park, Kanchanaburi Province". After we compared all the type materials to the same specimen lots of both localities housed in the general CUMZ collections and communicated with S. Panha, it became evident that the type material might have been mis-allocated. We conclude that the published type locality of
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<taxonomicName genus="A." lsidName="A. pratatensis" pageId="0" pageNumber="1" rank="species" species="pratatensis">
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<emphasis italics="true" pageId="0" pageNumber="1">A. pratatensis</emphasis>
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</taxonomicName>
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(Tham Pratat, Erawan National Park, Kanchanaburi Province, Thailand) is incorrect, and another species (
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<taxonomicName authorityName="Jirapatrasilp & Pall-Gergely" authorityYear="2021" class="Gastropoda" family="Cyclophoridae" genus="Dicharax" kingdom="Animalia" lsidName="Dicharax panhai" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="panhai">
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<emphasis italics="true" pageId="0" pageNumber="1">Dicharax panhai</emphasis>
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</taxonomicName>
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sp. nov.) lives there. Thus, the type locality of
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<taxonomicName genus="A." lsidName="A. pratatensis" pageId="0" pageNumber="1" rank="species" species="pratatensis">
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<emphasis italics="true" pageId="0" pageNumber="1">A. pratatensis</emphasis>
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</taxonomicName>
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is hereby amended to Doi Chiang Dao, Chiang Mai Province, Thailand.
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<bibRefCitation DOI="https://doi.org/10.3897/zookeys.981.53583" author="Pall-Gergely, B" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="1 - 220" refId="B47" refString="Pall-Gergely, B, Sajan, S, Tripathy, B, Meng, K, Takahiro, A, Ablett, J, 2020b. Genus-level revision of the Alycaeidae (Gastropoda: Cyclophoroidea), with an annotated species catalogue. ZooKeys 981: 1 - 220, DOI: https://doi.org/10.3897/zookeys.981.53583" title="Genus-level revision of the Alycaeidae (Gastropoda: Cyclophoroidea), with an annotated species catalogue." url="https://doi.org/10.3897/zookeys.981.53583" volume="981" year="2020 b">
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<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
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et al. (2020b
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</bibRefCitation>
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: 71) transferred
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<taxonomicName genus="A." lsidName="A. pratatensis" pageId="0" pageNumber="1" rank="species" species="pratatensis">
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<emphasis italics="true" pageId="0" pageNumber="1">A. pratatensis</emphasis>
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</taxonomicName>
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to
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<taxonomicName authorityName="Kobelt & Moellendorff" authorityYear="1900" class="Gastropoda" family="Cyclophoridae" genus="Dicharax" kingdom="Animalia" lsidName="Dicharax" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Dicharax</emphasis>
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</taxonomicName>
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based on the examination of paratype specimens deposited in the Senckenberg Museum (SMF 331452). However, the present study revealed that those shells and all other CUMZ specimens from Tham Pratat, Erawan National Park, Kanchanaburi Province, Thailand belong to
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<taxonomicName genus="D." lsidName="D. panhai" pageId="0" pageNumber="1" rank="species" species="panhai">
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<emphasis italics="true" pageId="0" pageNumber="1">D. panhai</emphasis>
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</taxonomicName>
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sp. nov. (see below).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |