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<document id="598400CC50BFA9554154CF0E0B3B2E9F" ID-DOI="10.11646/zootaxa.5383.3.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10361564" ID-ZooBank="20988E42-C14D-4878-A716-8CEAB5E5EF92" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="karina" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="karina" IM.treatments_approvedBy="karina" checkinTime="1702374886811" checkinUser="plazi" docAuthor="Tasaku, Yuto, Ono, Hiroki &amp; Maruyama, Munetoshi" docDate="2023" docId="03FE87D6425DFFEA82AA3D144CE4FDAE" docLanguage="en" docName="zootaxa.5383.3.1.pdf" docOrigin="Zootaxa 5383 (3)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5383.3.1/52454" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Amblopusa pacifica" docType="treatment" docVersion="3" lastPageNumber="278" masterDocId="FFC7FFAE4247FFF1823D3A504E1BFFA2" masterDocTitle="Review of the intertidal rove beetle tribe Liparocephalini Fenyes (Coleoptera, Staphylinidae, Aleocharinae) from Japan" masterLastPageNumber="296" masterPageNumber="251" pageNumber="277" updateTime="1702492195905" updateUser="karina" zenodo-license-document="CLOSED">
<mods:mods id="97A40B16E763270A33C6A32229A35196" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="BA4DC8BF506079D0606DB16AEDB1F56B">
<mods:title id="3B1FE57A744E6075B854986C8271BC15">Review of the intertidal rove beetle tribe Liparocephalini Fenyes (Coleoptera, Staphylinidae, Aleocharinae) from Japan</mods:title>
</mods:titleInfo>
<mods:name id="DE48E49695B7FCD97590B5B96BC6FDEA" type="personal">
<mods:role id="F9AC208E8234D772BA21FCEBAFECB553">
<mods:roleTerm id="E78AEB8B18A4890CCD29CAFD4AA9ABED">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6431C0801E31AE76FADBD42E7E0FEC4C">Tasaku, Yuto</mods:namePart>
<mods:affiliation id="52531B941230E2E35C2025DF6EE5E8E7">Department of Biology, School of Biological Sciences, Tokai University, Hokkaido, 005 - 8601, Japan. &amp; Corresponding author</mods:affiliation>
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<mods:roleTerm id="2BEBFE54A8ECEF9D52E2171D190A34A0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="46B8BDD1686159F609A309B6B09CCED0">Ono, Hiroki</mods:namePart>
<mods:affiliation id="8B7ED4EC93078FC172F2A272A8B95A86">Sounan, Minami-ku, Sagamihara-shi, Kanagawa, 252 - 0312, Japan.</mods:affiliation>
</mods:name>
<mods:name id="76122D6E9D60A3F08D2C4EB055B5207B" type="personal">
<mods:role id="355FC79A11CF9B1827A3F1CD187BE7A6">
<mods:roleTerm id="184DF7B7D972104D82C4EF817347DC50">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="39D6464D65257CB67690612DCE735CC8">Maruyama, Munetoshi</mods:namePart>
<mods:affiliation id="BA832A60BD33B314DE9498029A17B8CB">The Kyushu University Museum, Hakozaki 6 - 10 - 1, Higashi-ku, Fukuoka, 812 - 8581, Japan.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="A9BAB4280C6D31E1E31EA123AB40D5BC">text</mods:typeOfResource>
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<mods:title id="A5F4A1B984E1557B2A5F3489CE26A2AE">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="22FCA4D039A576A195ED7AC6CF23E87F">
<mods:date id="35C97F56BDF38FD22E62FB4073E78F57">2023</mods:date>
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<mods:number id="5AD148F7DB2FD7886E05D0C0E8B567AA">2023-12-12</mods:number>
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<mods:detail id="D7A84A28079C9A84EC16D684EEFB0344" type="volume">
<mods:number id="78A3A92B29A2B1E7C333CAF8F4B953FA">5383</mods:number>
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<mods:detail id="971530D78BBF35968203BF69ECEEAE9A" type="issue">
<mods:number id="D87574F92819578A2C2F1565E6A409A3">3</mods:number>
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<mods:start id="C7759BF526303642D373667F55FD4EDE">251</mods:start>
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<mods:url id="A90067DA1FEB2D112F8D61D332F0E4AD">https://www.mapress.com/zt/article/download/zootaxa.5383.3.1/52454</mods:url>
</mods:location>
<mods:classification id="7F0972A0FB411B2B7E344C502582E3D1">journal article</mods:classification>
<mods:identifier id="9ABB26DE8439FFC59A0FBD8E599D8F8F" type="DOI">10.11646/zootaxa.5383.3.1</mods:identifier>
<mods:identifier id="45DB32083B382AFAF5133625FAB7DA49" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
<treatment id="03FE87D6425DFFEA82AA3D144CE4FDAE" LSID="urn:lsid:plazi:treatment:03FE87D6425DFFEA82AA3D144CE4FDAE" httpUri="http://treatment.plazi.org/id/03FE87D6425DFFEA82AA3D144CE4FDAE" lastPageId="27" lastPageNumber="278" pageId="26" pageNumber="277">
<subSubSection id="C34D654B425DFFEB82AA3D144C98F820" pageId="26" pageNumber="277" type="nomenclature">
<paragraph id="8BE836C0425DFFEB82AA3D144C56F8FC" blockId="26.[151,643,1860,1959]" box="[151,589,1860,1887]" pageId="26" pageNumber="277">
<heading id="D0A081AC425DFFEB82AA3D144C56F8FC" bold="true" box="[151,589,1860,1887]" fontSize="11" level="1" pageId="26" pageNumber="277" reason="1">
<taxonomicName id="4C574D43425DFFEB82AA3D144C56F8FC" ID-CoL="CLB3" authority="(Sawada, 1991)" baseAuthorityName="Sawada" baseAuthorityYear="1991" box="[151,589,1860,1887]" class="Insecta" family="Staphylinidae" genus="Amblopusa" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="277" phylum="Arthropoda" rank="species" species="pacifica">
<emphasis id="B923EAD2425DFFEB82AA3D144C56F8FC" bold="true" box="[151,589,1860,1887]" pageId="26" pageNumber="277">
<emphasis id="B923EAD2425DFFEB82AA3D144F9CF8FC" bold="true" box="[151,391,1860,1887]" italics="true" pageId="26" pageNumber="277">Amblopusa pacifica</emphasis>
(
<bibRefCitation id="EFC64B31425DFFEB83AA3D144C5EF8FC" author="Sawada, K." box="[407,581,1860,1886]" pageId="26" pageNumber="277" pagination="141 - 152" refId="ref23310" refString="Sawada, K. (1991) On a new genus and species of intertidal Aleocharinae (Coleoptera: Staphylinidae) and Goniacerinae (Pselaphidae) from Singapore and Japan. Raffles Bulletin of Zoology, 39, 141 - 152." type="journal article" year="1991">Sawada, 1991</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BE836C0425DFFEB82AA3D394C98F820" blockId="26.[151,643,1860,1959]" box="[151,643,1896,1923]" pageId="26" pageNumber="277">[Japanese name: Nokoba-umi-hanekakushi]</paragraph>
</subSubSection>
<subSubSection id="C34D654B425DFFEB82AA3DDC4C1AF805" box="[151,513,1932,1959]" pageId="26" pageNumber="277" type="description">
<paragraph id="8BE836C0425DFFEB82AA3DDC4C1AF805" blockId="26.[151,643,1860,1959]" box="[151,513,1932,1959]" pageId="26" pageNumber="277">
(
<figureCitation id="136C2A45425DFFEB82A23DDC4F0BF805" box="[159,272,1932,1959]" captionStart="FIGURE 17" captionStartId="27.[151,250,1647,1671]" captionTargetBox="[163,1424,559,1611]" captionTargetId="figure-163@27.[151,1436,548,1623]" captionTargetPageId="27" captionText="FIGURE 17 Habitus of Amblopusa spp. A, A. magna; B, A. pacifica; C, A. brevipes; D, A.hokkaidona. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10361635" httpUri="https://zenodo.org/record/10361635/files/figure.png" pageId="26" pageNumber="277">Figs. 17B</figureCitation>
;
<figureCitation id="136C2A45425DFFEB83213DDD4F9DF805" box="[284,390,1933,1959]" captionStart="FIGURE 18" captionStartId="28.[151,250,1943,1967]" captionTargetBox="[183,1404,191,1911]" captionTargetId="figure-22@28.[173,1413,181,1920]" captionTargetPageId="28" captionText="FIGURE 18 Genitalia of Amblopusa spp. A &amp; B, A. magna (A, median lobe of aedeagus; B, spermatheca); C &amp; D, A. pacifica (C, median lobe of aedeagus; D, spermatheca); E–G, A. brevipes (E, median lobe of aedeagus; F, spermatheca; G, apical lobe of paramere). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10361639" httpUri="https://zenodo.org/record/10361639/files/figure.png" pageId="26" pageNumber="277">18C &amp; D</figureCitation>
;
<figureCitation id="136C2A45425DFFEB83AF3DDC4FECF805" box="[402,503,1932,1959]" captionStart="FIGURE 26" captionStartId="38.[151,250,1588,1612]" captionTargetBox="[151,1436,181,1564]" captionTargetId="figure-22@38.[151,1436,181,1564]" captionTargetPageId="38" captionText="FIGURE 26 Distribution and habitat of Amblopusa spp. of Japan. A, known localities in Japan (red circle, A. magna; blue square, A. pacifica; yellow triangle, A. brevipes; green inverted triangle, A. hokkaidona); B, habitat of A. magna and A. pacifica (Wakkainai-shi, Hokkaidô, photographed by Daisuke Sumikawa); C, habitat of A. magna (Abashiri-shi, Hokkaidô)." figureDoi="http://doi.org/10.5281/zenodo.10361675" httpUri="https://zenodo.org/record/10361675/files/figure.png" pageId="26" pageNumber="277">26A &amp; B</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34D654B425DFFEA82AA3D814B11FF6F" lastPageId="27" lastPageNumber="278" pageId="26" pageNumber="277" type="reference_group">
<paragraph id="8BE836C0425DFFEA82AA3D814A1AFF0D" blockId="26.[151,1292,2001,2025]" lastBlockId="27.[151,1290,150,205]" lastPageId="27" lastPageNumber="278" pageId="26" pageNumber="277">
<treatmentCitationGroup id="AB4711EE425DFFEB82AA3D814C4CF84B" box="[151,599,2001,2025]" pageId="26" pageNumber="277">
<treatmentCitation id="0AF610D1425DFFEB82AA3D814C4CF84B" author="Sawada, K." box="[151,599,2001,2025]" page="147" pageId="26" pageNumber="277" year="1991">
<taxonomicName id="4C574D43425DFFEB82AA3D814C4CF84B" ID-CoL="MJVT" authority="Sawada, 1991: 147" authorityName="Sawada" authorityPageNumber="147" authorityYear="1991" box="[151,599,2001,2025]" class="Insecta" family="Staphylinidae" genus="Boreorhadinus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="277" phylum="Arthropoda" rank="species" species="pacificus">
<emphasis id="B923EAD2425DFFEB82AA3D814F94F84B" box="[151,399,2001,2025]" italics="true" pageId="26" pageNumber="277">Boreorhadinus pacificus</emphasis>
<bibRefCitation id="EFC64B31425DFFEB83AB3D814C4CF84B" author="Sawada, K." box="[406,599,2001,2025]" pageId="26" pageNumber="277" pagination="141 - 152" refId="ref23310" refString="Sawada, K. (1991) On a new genus and species of intertidal Aleocharinae (Coleoptera: Staphylinidae) and Goniacerinae (Pselaphidae) from Singapore and Japan. Raffles Bulletin of Zoology, 39, 141 - 152." type="journal article" year="1991">Sawada, 1991: 147</bibRefCitation>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
<treatmentCitationGroup id="AB4711EE425DFFEA80633D814C9EFF0D" lastPageId="27" lastPageNumber="278" pageId="26" pageNumber="277">
(original description;
<typeStatus id="54EC8862425DFFEB81043D814D7EF84B" box="[825,869,2001,2025]" pageId="26" pageNumber="277">type</typeStatus>
locality: Akkeshi, east
<collectingRegion id="4993F822425DFFEB866F3D814AA2F84B" box="[1106,1209,2001,2025]" country="Japan" name="Hokkaido" pageId="26" pageNumber="277">Hokkaido</collectingRegion>
,
<collectingCountry id="F3407650425DFFEB86FE3D814B1AF84B" box="[1219,1281,2001,2025]" name="Japan" pageId="26" pageNumber="277">Japan</collectingCountry>
).
<emphasis id="B923EAD2425CFFEA82AA3AC74F9CFF0D" box="[151,391,151,175]" italics="true" pageId="27" pageNumber="278">
<taxonomicName id="4C574D43425CFFEA82AA3AC74F64FF0D" ID-CoL="MJVT" authorityName="Ueno" authorityYear="1978" box="[151,383,151,175]" class="Insecta" family="Staphylinidae" genus="Amblopusa" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="278" phylum="Arthropoda" rank="species" species="pacificus">Amblopusa “pacificus</taxonomicName>
”
</emphasis>
: Ahn &amp; Ashe, 1995: 143
</treatmentCitationGroup>
<treatmentCitationGroup id="AB4711EE425CFFEA80B63AC74A1AFF0D" box="[651,1025,151,175]" pageId="27" pageNumber="278">
(as a junior synonym of
<taxonomicName id="4C574D43425CFFEA81BF3AC74DE8FF0D" ID-CoL="65ZWV" authorityName="Casey" authorityYear="1894" box="[898,1011,151,175]" class="Insecta" family="Staphylinidae" genus="Amblopusa" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="278" phylum="Arthropoda" rank="species" species="brevipes">
<emphasis id="B923EAD2425CFFEA81BF3AC74D8AFF0C" box="[898,913,151,174]" italics="true" pageId="27" pageNumber="278">A</emphasis>
.
<emphasis id="B923EAD2425CFFEA81A33AC74DE8FF0D" box="[926,1011,151,175]" italics="true" pageId="27" pageNumber="278">brevipes</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BE836C0425CFFEA82AA3AE54B11FF6F" blockId="27.[151,1290,150,205]" box="[151,1290,181,205]" pageId="27" pageNumber="278">
<treatmentCitationGroup id="AB4711EE425CFFEA82AA3AE54B11FF6F" box="[151,1290,181,205]" pageId="27" pageNumber="278">
<taxonomicName id="4C574D43425CFFEA82AA3AE54C11FF6F" ID-CoL="CLB3" authority="(Sawada, 1991)" baseAuthorityName="Sawada" baseAuthorityYear="1991" box="[151,522,181,205]" class="Insecta" family="Staphylinidae" genus="Amblopusa" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="278" phylum="Arthropoda" rank="species" species="pacifica">
<emphasis id="B923EAD2425CFFEA82AA3AE54F79FF6F" box="[151,354,181,205]" italics="true" pageId="27" pageNumber="278">Amblopusa pacifica</emphasis>
(
<bibRefCitation id="EFC64B31425CFFEA83523AE54C19FF6F" author="Sawada, K." box="[367,514,181,205]" pageId="27" pageNumber="278" pagination="141 - 152" refId="ref23310" refString="Sawada, K. (1991) On a new genus and species of intertidal Aleocharinae (Coleoptera: Staphylinidae) and Goniacerinae (Pselaphidae) from Singapore and Japan. Raffles Bulletin of Zoology, 39, 141 - 152." type="journal article" year="1991">Sawada, 1991</bibRefCitation>
)
</taxonomicName>
:
<treatmentCitation id="0AF610D1425CFFEA80293AE54CD6FF6F" author="Zerche, L." box="[532,717,181,205]" page="106" pageId="27" pageNumber="278" year="1998">
<bibRefCitation id="EFC64B31425CFFEA80293AE54CD6FF6F" author="Zerche, L." box="[532,717,181,205]" pageId="27" pageNumber="278" pagination="103 - 113" refId="ref23977" refString="Zerche, L. (1998) Amblopusa magna sp. n. - eine neue Art der amphipazifischen Gattung Amblopusa Casey, 1893 aus dem Fernen Osten RuSSlands und ihre Stellung im phylogenetischen System (Coleoptera: Staphylinidae, Aleocharinae, Liparocephalini). Beitrage zur Entomologie, 48, 103 - 113." type="journal article" year="1998">Zerche, 1998: 106</bibRefCitation>
</treatmentCitation>
(characters; habitus photograph; phylogenetic analysis).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C34D654B425CFFEA82AA3AA64D46FEDE" pageId="27" pageNumber="278" type="materials_examined">
<paragraph id="8BE836C0425CFFEA82AA3AA64D46FEDE" blockId="27.[151,1437,246,525]" pageId="27" pageNumber="278">
<emphasis id="B923EAD2425CFFEA82AA3AA64F98FEB2" bold="true" box="[151,387,246,272]" pageId="27" pageNumber="278">Material examined.</emphasis>
<materialsCitation id="3B3F3C9D425CFFEA83B13AA64B2BFEB3" box="[396,1328,246,273]" collectionCode="VII" country="Japan" location="Odaito" municipality="Betsukai-cho" pageId="27" pageNumber="278" specimenCount="5" specimenCount-female="3" specimenCount-male="2" stateProvince="Hokkaido">
<emphasis id="B923EAD2425CFFEA83B13AA64C7CFEB2" bold="true" box="[396,615,246,272]" pageId="27" pageNumber="278">
<collectingCountry id="F3407650425CFFEA83B13AA64FC2FEB2" box="[396,473,246,272]" name="Japan" pageId="27" pageNumber="278">Japan</collectingCountry>
:
<collectingRegion id="4993F822425CFFEA83DA3AA64C79FEB2" box="[487,610,246,272]" country="Japan" name="Hokkaido" pageId="27" pageNumber="278">Hokkaidô</collectingRegion>
:
</emphasis>
<specimenCount id="9D51FD49425CFFEA804D3AA64CD0FEB2" box="[624,715,246,272]" count="2" pageId="27" pageNumber="278" type="male">2 males</specimenCount>
,
<specimenCount id="9D51FD49425CFFEA80E43AA64D51FEB2" box="[729,842,246,272]" count="3" pageId="27" pageNumber="278" type="female">3 females</specimenCount>
, 5 unsexed,
<collectingMunicipality id="6B8CACBA425CFFEA81E73AA64A6AFEB2" box="[986,1137,246,272]" pageId="27" pageNumber="278">Betsukai-chô</collectingMunicipality>
,
<location id="8E88601B425CFFEA86433AA64AD6FEB2" LSID="urn:lsid:plazi:treatment:03FE87D6425DFFEA82AA3D144CE4FDAE:8E88601B425CFFEA86433AA64AD6FEB2" box="[1150,1229,246,272]" country="Japan" municipality="Betsukai-cho" name="Odaito" pageId="27" pageNumber="278" stateProvince="Hokkaido">Odaitô</location>
,
<date id="FFE91000425CFFEA86E63AA64B2BFEB3" box="[1243,1328,246,273]" pageId="27" pageNumber="278" value="2020-07-23">
23.
<collectionCode id="ED46AE05425CFFEA873B3AA74B2BFEB3" box="[1286,1328,247,273]" pageId="27" pageNumber="278">VII</collectionCode>
</date>
</materialsCitation>
. 2020, Y. Tasaku (cYT);
<materialsCitation id="3B3F3C9D425CFFEA837E3B4A4DC7FEFB" collectingDate="2022-08-03" collectionCode="KUM, IV" collectorName="D. Sumikawa &amp; Y. Tasaku &amp; Wakkainai-shi &amp; Souya-mura &amp; Higashiura" country="Japan" location="Akkeshi-ko" municipality="Akkeshi-cho" pageId="27" pageNumber="278" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Hokkaido">
<specimenCount id="9D51FD49425CFFEA837E3B4A4F89FE96" box="[323,402,282,308]" count="1" pageId="27" pageNumber="278" type="male">1 male</specimenCount>
,
<specimenCount id="9D51FD49425CFFEA83A33B4A4C1FFE96" box="[414,516,282,308]" count="1" pageId="27" pageNumber="278" type="female">1 female</specimenCount>
, 25 unsexed,
<collectingMunicipality id="6B8CACBA425CFFEA80A03B4A4D34FE96" box="[669,815,282,308]" pageId="27" pageNumber="278">Akkeshi-chô</collectingMunicipality>
,
<location id="8E88601B425CFFEA81073B4A4DDBFE96" LSID="urn:lsid:plazi:treatment:03FE87D6425DFFEA82AA3D144CE4FDAE:8E88601B425CFFEA81073B4A4DDBFE96" box="[826,960,282,308]" country="Japan" municipality="Akkeshi-cho" name="Akkeshi-ko" pageId="27" pageNumber="278" stateProvince="Hokkaido">Akkeshi-ko</location>
,
<date id="FFE91000425CFFEA81F13B4A4A78FE96" box="[972,1123,282,309]" pageId="27" pageNumber="278" value="2022-08-03">
<collectingDate id="EFADE9E8425CFFEA81F13B4A4A78FE96" box="[972,1123,282,309]" pageId="27" pageNumber="278" value="2022-08-03">3. VIII. 2022</collectingDate>
</date>
,
<collectorName id="26A25316425CFFEA86523B4B4B12FE96" box="[1135,1289,282,309]" pageId="27" pageNumber="278">D. Sumikawa</collectorName>
&amp;
<collectorName id="26A25316425CFFEA87133B4B4B87FE96" box="[1326,1436,282,309]" pageId="27" pageNumber="278">Y. Tasaku</collectorName>
(
<collectionCode id="ED46AE05425CFFEA82A23B6E4EFEFEFA" box="[159,229,318,344]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="27" pageNumber="278" type="Herbarium">KUM</collectionCode>
, cYT); 15 unsexed,
<collectorName id="26A25316425CFFEA83FA3B6E4C70FEFA" box="[455,619,318,344]" pageId="27" pageNumber="278">Wakkainai-shi</collectorName>
,
<collectorName id="26A25316425CFFEA804A3B6E4D18FEFA" box="[631,771,318,344]" pageId="27" pageNumber="278">Souya-mura</collectorName>
,
<collectorName id="26A25316425CFFEA81333B6E4D97FEFA" box="[782,908,318,344]" pageId="27" pageNumber="278">Higashiura</collectorName>
,
<date id="FFE91000425CFFEA81AA3B6E4DC7FEFB" box="[919,988,318,345]" pageId="27" pageNumber="278" value="2022-04-20">
20.
<collectionCode id="ED46AE05425CFFEA81FF3B6F4DC7FEFB" box="[962,988,319,345]" pageId="27" pageNumber="278">IV</collectionCode>
</date>
</materialsCitation>
<materialsCitation id="3B3F3C9D425CFFEA81E13B6F4D42FEDE" collectingDate="1999-09-11" collectionCode="KUM" collectorName="D. Sumikawa &amp; M. Maruyama" country="Japan" location="Shunkunitai" municipality="Nemuro-shi" pageId="27" pageNumber="278" specimenCount="1" stateProvince="Hokkaido">
. 2022,
<collectorName id="26A25316425CFFEA860D3B6F4AD2FEFA" box="[1072,1225,318,345]" pageId="27" pageNumber="278">D. Sumikawa</collectorName>
(cYT); 2 unsexed,
<collectingMunicipality id="6B8CACBA425CFFEA82AA3B324F3BFEDE" box="[151,288,354,380]" pageId="27" pageNumber="278">Nemuro-shi</collectingMunicipality>
,
<location id="8E88601B425CFFEA83163B324FAEFEDE" LSID="urn:lsid:plazi:treatment:03FE87D6425DFFEA82AA3D144CE4FDAE:8E88601B425CFFEA83163B324FAEFEDE" box="[299,437,354,380]" country="Japan" municipality="Nemuro-shi" name="Shunkunitai" pageId="27" pageNumber="278" stateProvince="Hokkaido">Shunkunitai</location>
,
<date id="FFE91000425CFFEA83FD3B324C4BFEDE" box="[448,592,354,381]" pageId="27" pageNumber="278" value="1999-09-11">
<collectingDate id="EFADE9E8425CFFEA83FD3B324C4BFEDE" box="[448,592,354,381]" pageId="27" pageNumber="278" value="1999-09-11">11. IX. 1999</collectingDate>
</date>
,
<collectorName id="26A25316425CFFEA80663B334CE0FEDF" box="[603,763,355,381]" pageId="27" pageNumber="278">M. Maruyama</collectorName>
(
<collectionCode id="ED46AE05425CFFEA81373B324D4AFEDE" box="[778,849,354,380]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="27" pageNumber="278" type="Herbarium">KUM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34D654B425CFFEA82FA3BD64B1FFE4A" pageId="27" pageNumber="278" type="diagnosis">
<paragraph id="8BE836C0425CFFEA82FA3BD64B1FFE4A" blockId="27.[151,1437,246,525]" pageId="27" pageNumber="278">
<emphasis id="B923EAD2425CFFEA82FA3BD64F59FE02" bold="true" box="[199,322,390,416]" pageId="27" pageNumber="278">Diagnosis.</emphasis>
This species can be distinguished from other
<taxonomicName id="4C574D43425CFFEA810B3BD64A10FE02" box="[822,1035,390,416]" pageId="27" pageNumber="278">
<emphasis id="B923EAD2425CFFEA810B3BD64DAFFE02" box="[822,948,390,416]" italics="true" pageId="27" pageNumber="278">Amblopusa</emphasis>
species
</taxonomicName>
by the combination of the following characteristics: body small (about
<quantity id="4CAF9B25425CFFEA80243BFA4C6BFE67" box="[537,624,426,453]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="27" pageNumber="278" unit="mm" value="2.5">2.5 mm</quantity>
long); pronotum rather flattened with anterolateral margin angular (
<figureCitation id="136C2A45425CFFEA87523BFA4ED2FE4A" captionStart="FIGURE 17" captionStartId="27.[151,250,1647,1671]" captionTargetBox="[163,1424,559,1611]" captionTargetId="figure-163@27.[151,1436,548,1623]" captionTargetPageId="27" captionText="FIGURE 17 Habitus of Amblopusa spp. A, A. magna; B, A. pacifica; C, A. brevipes; D, A.hokkaidona. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10361635" httpUri="https://zenodo.org/record/10361635/files/figure.png" pageId="27" pageNumber="278">Fig. 17B</figureCitation>
); median lobe of aedeagus with apical lobe slightly swollen near apex in lateral view (
<figureCitation id="136C2A45425CFFEA86AD3B9E4AECFE4A" box="[1168,1271,462,488]" captionStart="FIGURE 18" captionStartId="28.[151,250,1943,1967]" captionTargetBox="[183,1404,191,1911]" captionTargetId="figure-22@28.[173,1413,181,1920]" captionTargetPageId="28" captionText="FIGURE 18 Genitalia of Amblopusa spp. A &amp; B, A. magna (A, median lobe of aedeagus; B, spermatheca); C &amp; D, A. pacifica (C, median lobe of aedeagus; D, spermatheca); E–G, A. brevipes (E, median lobe of aedeagus; F, spermatheca; G, apical lobe of paramere). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10361639" httpUri="https://zenodo.org/record/10361639/files/figure.png" pageId="27" pageNumber="278">Fig. 18C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34D654B425CFFEA82FA3BA24CE4FDAE" box="[199,767,498,525]" pageId="27" pageNumber="278" type="distribution">
<paragraph id="8BE836C0425CFFEA82FA3BA24CE4FDAE" blockId="27.[151,1437,246,525]" box="[199,767,498,525]" pageId="27" pageNumber="278">
<emphasis id="B923EAD2425CFFEA82FA3BA24F7AFDAE" bold="true" box="[199,353,498,524]" pageId="27" pageNumber="278">Distribution.</emphasis>
<collectingCountry id="F3407650425CFFEA835A3BA34FB0FDAF" box="[359,427,499,525]" name="Japan" pageId="27" pageNumber="278">Japan</collectingCountry>
: eastern
<collectingRegion id="4993F822425CFFEA80313BA24C61FDAE" box="[524,634,498,524]" country="Japan" name="Hokkaido" pageId="27" pageNumber="278">Hokkaidô</collectingRegion>
(
<figureCitation id="136C2A45425CFFEA80B73BA24CE8FDAE" box="[650,755,498,524]" captionStart="FIGURE 26" captionStartId="38.[151,250,1588,1612]" captionTargetBox="[151,1436,181,1564]" captionTargetId="figure-22@38.[151,1436,181,1564]" captionTargetPageId="38" captionText="FIGURE 26 Distribution and habitat of Amblopusa spp. of Japan. A, known localities in Japan (red circle, A. magna; blue square, A. pacifica; yellow triangle, A. brevipes; green inverted triangle, A. hokkaidona); B, habitat of A. magna and A. pacifica (Wakkainai-shi, Hokkaidô, photographed by Daisuke Sumikawa); C, habitat of A. magna (Abashiri-shi, Hokkaidô)." figureDoi="http://doi.org/10.5281/zenodo.10361675" httpUri="https://zenodo.org/record/10361675/files/figure.png" pageId="27" pageNumber="278">Fig. 26A</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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