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<mods:title id="65F1F03001C1FB549048C8CB2FFDEB7C">Review of the intertidal rove beetle tribe Liparocephalini Fenyes (Coleoptera, Staphylinidae, Aleocharinae) from Japan</mods:title>
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<mods:namePart id="A4DEF32C903718C9B46F27665388145D">Tasaku, Yuto</mods:namePart>
<mods:affiliation id="A4635C91AF6DAAB350A41521BBDCA8DA">Department of Biology, School of Biological Sciences, Tokai University, Hokkaido, 005 - 8601, Japan. &amp; Corresponding author</mods:affiliation>
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<mods:namePart id="A24C3354DBAE37FABB64F67ADBA6CE98">Ono, Hiroki</mods:namePart>
<mods:affiliation id="CF075D2CF3AAA4F8A2E067A163E8D46C">Sounan, Minami-ku, Sagamihara-shi, Kanagawa, 252 - 0312, Japan.</mods:affiliation>
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<mods:namePart id="34D250630E0B2C67E6373076D666ED10">Maruyama, Munetoshi</mods:namePart>
<mods:affiliation id="ACCD37D48C48961141CAC8EA8185F525">The Kyushu University Museum, Hakozaki 6 - 10 - 1, Higashi-ku, Fukuoka, 812 - 8581, Japan.</mods:affiliation>
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<paragraph id="8BE836C0425AFFEC82AA3E494C50FB91" blockId="29.[151,599,1049,1147]" box="[151,587,1049,1076]" pageId="29" pageNumber="280">
<heading id="D0A081AC425AFFEC82AA3E494C50FB91" bold="true" box="[151,587,1049,1076]" fontSize="11" level="1" pageId="29" pageNumber="280" reason="1">
<emphasis id="B923EAD2425AFFEC82AA3E494C50FB91" bold="true" box="[151,587,1049,1076]" pageId="29" pageNumber="280">
Genus
<taxonomicName id="4C574D43425AFFEC82D33E494C50FB91" ID-CoL="8W2X5" authority="Sawada, 1971" authorityName="Sawada" authorityYear="1971" box="[238,587,1049,1076]" class="Insecta" family="Staphylinidae" genus="Halorhadinus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="280" phylum="Arthropoda" rank="genus">
<emphasis id="B923EAD2425AFFEC82D33E494F83FB96" bold="true" box="[238,408,1049,1076]" italics="true" pageId="29" pageNumber="280">Halorhadinus</emphasis>
<bibRefCitation id="EFC64B31425AFFEC83A23E494C50FB91" author="Sawada, K." box="[415,587,1049,1075]" pageId="29" pageNumber="280" pagination="81 - 110" refId="ref23228" refString="Sawada, K. (1971) Aleocharinae (Staphylinidae, Coleoptera) from the Intertidal Zone of Japan, Publications of the Seto Marine Biological Laboratory, 19 (2 - 3), 81 - 110. https: // doi. org / 10.5134 / 175665" type="journal article" year="1971">Sawada, 1971</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE836C0425AFFEC82AA3E6E4C4CFBF5" blockId="29.[151,599,1049,1147]" box="[151,599,1085,1112]" pageId="29" pageNumber="280">[Japanese name: Iso-hanekakushi-zoku]</paragraph>
</subSubSection>
<subSubSection id="C34D654B425AFFEC82AA3E314EE2FBD9" box="[151,249,1121,1147]" pageId="29" pageNumber="280" type="description">
<paragraph id="8BE836C0425AFFEC82AA3E314EE2FBD9" blockId="29.[151,599,1049,1147]" box="[151,249,1121,1147]" pageId="29" pageNumber="280">
(
<figureCitation id="136C2A45425AFFEC82A23E314EEAFBD9" box="[159,241,1121,1147]" captionStart="FIGURE 20" captionStartId="31.[151,250,1874,1898]" captionTargetBox="[160,1426,191,1841]" captionTargetId="figure-19@31.[151,1436,181,1851]" captionTargetPageId="31" captionText="FIGURE 20 Halorhadinus aequalis. A, Habitus; B, labrum (right side, dorsal view; left side, ventral view (epipharynx)); C, mentum; D, labium; E, maxilla; F, right mandible; G, apical lobe of paramere. Scale bars: 1.0 mm for A; 0.1 mm for BG." figureDoi="http://doi.org/10.5281/zenodo.10361647" httpUri="https://zenodo.org/record/10361647/files/figure.png" pageId="29" pageNumber="280">Fig. 20</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34D654B425AFFEC82AA3EF04DF1FB54" pageId="29" pageNumber="280" type="reference_group">
<paragraph id="8BE836C0425AFFEC82AA3EF04DF1FB54" blockId="29.[151,1437,1184,1270]" pageId="29" pageNumber="280">
<treatmentCitationGroup id="AB4711EE425AFFEC82AA3EF04FC5FB1B" box="[151,478,1184,1209]" pageId="29" pageNumber="280">
<treatmentCitation id="0AF610D1425AFFEC82AA3EF04FC5FB1B" author="Sawada, K." box="[151,478,1184,1209]" page="92" pageId="29" pageNumber="280" year="1971">
<taxonomicName id="4C574D43425AFFEC82AA3EF04FC5FB1B" ID-CoL="8W2X5" authority="Sawada, 1971: 92" authorityName="Sawada" authorityPageNumber="92" authorityYear="1971" box="[151,478,1184,1209]" class="Insecta" family="Staphylinidae" genus="Halorhadinus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="280" phylum="Arthropoda" rank="genus">
<emphasis id="B923EAD2425AFFEC82AA3EF04F38FB1A" box="[151,291,1184,1208]" italics="true" pageId="29" pageNumber="280">Halorhadinus</emphasis>
<bibRefCitation id="EFC64B31425AFFEC83173EF04FC5FB1B" author="Sawada, K." box="[298,478,1184,1209]" pageId="29" pageNumber="280" pagination="81 - 110" refId="ref23228" refString="Sawada, K. (1971) Aleocharinae (Staphylinidae, Coleoptera) from the Intertidal Zone of Japan, Publications of the Seto Marine Biological Laboratory, 19 (2 - 3), 81 - 110. https: // doi. org / 10.5134 / 175665" type="journal article" year="1971">Sawada, 1971: 92</bibRefCitation>
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(original description;
<typeStatus id="54EC8862425AFFEC80823EF14CF0FB1B" box="[703,747,1185,1209]" pageId="29" pageNumber="280">type</typeStatus>
species:
<taxonomicName id="4C574D43425AFFEC817A3EF14A48FB1B" ID-CoL="3JFX7" authority="Sawada, 1971" authorityName="Sawada" authorityYear="1971" box="[839,1107,1184,1209]" class="Insecta" family="Staphylinidae" genus="Halorhadinus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="280" phylum="Arthropoda" rank="species" species="aequalis">
<emphasis id="B923EAD2425AFFEC817A3EF14D42FB1A" box="[839,857,1185,1208]" italics="true" pageId="29" pageNumber="280">H</emphasis>
.
<emphasis id="B923EAD2425AFFEC815B3EF04DA7FB1A" box="[870,956,1184,1208]" italics="true" pageId="29" pageNumber="280">aequalis</emphasis>
<bibRefCitation id="EFC64B31425AFFEC81FF3EF04A48FB1B" author="Sawada, K." box="[962,1107,1184,1209]" pageId="29" pageNumber="280" pagination="81 - 110" refId="ref23228" refString="Sawada, K. (1971) Aleocharinae (Staphylinidae, Coleoptera) from the Intertidal Zone of Japan, Publications of the Seto Marine Biological Laboratory, 19 (2 - 3), 81 - 110. https: // doi. org / 10.5134 / 175665" type="journal article" year="1971">Sawada, 1971</bibRefCitation>
</taxonomicName>
);
<treatmentCitation id="0AF610D1425AFFEC86593EF04B19FB1B" author="Ahn, K. - J." box="[1124,1282,1184,1209]" page="123" pageId="29" pageNumber="280" year="2001">
<bibRefCitation id="EFC64B31425AFFEC86593EF04B19FB1B" author="Ahn, K. - J." box="[1124,1282,1184,1209]" pageId="29" pageNumber="280" pagination="123 - 132" refId="ref21270" refString="Ahn, K. - J. (2001) Phylogenetic relationships of the intertidal genus Halorhadinus Sawada and key to the genera of the Liparocephalini (Coleoptera: Staphylinidae: Aleocharinae). Insect Systematics and Evolution, 32, 123 - 132. https: // doi. org / 10.1163 / 187631201 X 00074" type="journal article" year="2001">Ahn, 2001: 123</bibRefCitation>
</treatmentCitation>
(redescription; key to species; phylogenetic analysis);
<treatmentCitation id="0AF610D1425AFFEC80513E904DDDFB75" author="Maruyama, M. &amp; Hayashi, M." box="[620,966,1215,1240]" page="72" pageId="29" pageNumber="280" year="2009">
<bibRefCitation id="EFC64B31425AFFEC80513E904DDDFB75" author="Maruyama, M. &amp; Hayashi, M." box="[620,966,1215,1240]" pageId="29" pageNumber="280" pagination="71 - 82" refId="ref22396" refString="Maruyama, M. &amp; Hayashi, M. (2009) Description of the intertidal aleocharine Halorhadinus sawadai with notes on the genus Halorhadinus Sawada, 1971 (Coleoptera: Staphylinidae). Koleopterologische Rundschau, 79, 71 - 82." type="journal article" year="2009">Maruyama &amp; Hayashi, 2009: 72</bibRefCitation>
</treatmentCitation>
(supplementary description; key to species; systematic position);
<treatmentCitation id="0AF610D1425AFFEC839D3E8E4CA4FB54" author="Ono, H. &amp; Maruyama, M." box="[416,703,1246,1270]" page="42" pageId="29" pageNumber="280" year="2014">
<bibRefCitation id="EFC64B31425AFFEC839D3E8E4CA4FB54" author="Ono, H. &amp; Maruyama, M." box="[416,703,1246,1270]" pageId="29" pageNumber="280" pagination="41 - 50" refId="ref22885" refString="Ono, H. &amp; Maruyama, M. (2014) Five new species of the intertidal genus Halorhadinus Sawada, 1971 (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Esakia, 54, 41 - 50. https: // doi. org / 10.5109 / 1517831" type="journal article" year="2014">Ono &amp; Maruyama, 2014: 42</bibRefCitation>
</treatmentCitation>
(description; key to species).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C34D654B425AFFEC82AA3F4E4FC3F9B3" pageId="29" pageNumber="280" type="diagnosis">
<paragraph id="8BE836C0425AFFEC82AA3F4E4A6EFA6B" blockId="29.[151,1436,1310,1733]" pageId="29" pageNumber="280">
<emphasis id="B923EAD2425AFFEC82AA3F4E4F09FA9A" bold="true" box="[151,274,1310,1336]" pageId="29" pageNumber="280">Diagnosis.</emphasis>
This genus is similar to
<taxonomicName id="4C574D43425AFFEC80233F4F4C87FA9B" authorityName="Casey" authorityYear="1894" box="[542,668,1311,1337]" class="Insecta" family="Staphylinidae" genus="Amblopusa" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="280" phylum="Arthropoda" rank="genus">
<emphasis id="B923EAD2425AFFEC80233F4F4C87FA9B" box="[542,668,1311,1337]" italics="true" pageId="29" pageNumber="280">Amblopusa</emphasis>
</taxonomicName>
in overall bodyform (
<figureCitation id="136C2A45425AFFEC81B03F4F4DFCFA9B" box="[909,999,1311,1337]" captionStart="FIGURE 17" captionStartId="27.[151,250,1647,1671]" captionTargetBox="[163,1424,559,1611]" captionTargetId="figure-163@27.[151,1436,548,1623]" captionTargetPageId="27" captionText="FIGURE 17 Habitus of Amblopusa spp. A, A. magna; B, A. pacifica; C, A. brevipes; D, A.hokkaidona. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10361635" httpUri="https://zenodo.org/record/10361635/files/figure.png" pageId="29" pageNumber="280">Figs. 17</figureCitation>
&amp;
<figureCitation id="136C2A45425AFFEC86343F4F4A26FA9B" box="[1033,1085,1311,1337]" captionStart="FIGURE 20" captionStartId="31.[151,250,1874,1898]" captionTargetBox="[160,1426,191,1841]" captionTargetId="figure-19@31.[151,1436,181,1851]" captionTargetPageId="31" captionText="FIGURE 20 Halorhadinus aequalis. A, Habitus; B, labrum (right side, dorsal view; left side, ventral view (epipharynx)); C, mentum; D, labium; E, maxilla; F, right mandible; G, apical lobe of paramere. Scale bars: 1.0 mm for A; 0.1 mm for BG." figureDoi="http://doi.org/10.5281/zenodo.10361647" httpUri="https://zenodo.org/record/10361647/files/figure.png" pageId="29" pageNumber="280">20A</figureCitation>
), characteristics of mentum and maxilla (
<figureCitation id="136C2A45425AFFEC82C63F134FBFFAFF" box="[251,420,1347,1373]" captionStart="FIGURE 19" captionStartId="30.[151,250,1943,1967]" captionTargetBox="[196,1393,192,1908]" captionTargetId="figure-22@30.[185,1403,181,1919]" captionTargetPageId="30" captionText="FIGURE 19 Mouth parts of Amblopusa brevipes. A, labrum (right side, dorsal view; left side, ventral view (epipharynx)); B, mentum; C, labium; D, maxilla; E, right mandible. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10361643" httpUri="https://zenodo.org/record/10361643/files/figure.png" pageId="29" pageNumber="280">Figs. 19B &amp; D</figureCitation>
;
<figureCitation id="136C2A45425AFFEC838D3F134C0EFAFF" box="[432,533,1347,1373]" captionStart="FIGURE 20" captionStartId="31.[151,250,1874,1898]" captionTargetBox="[160,1426,191,1841]" captionTargetId="figure-19@31.[151,1436,181,1851]" captionTargetPageId="31" captionText="FIGURE 20 Halorhadinus aequalis. A, Habitus; B, labrum (right side, dorsal view; left side, ventral view (epipharynx)); C, mentum; D, labium; E, maxilla; F, right mandible; G, apical lobe of paramere. Scale bars: 1.0 mm for A; 0.1 mm for BG." figureDoi="http://doi.org/10.5281/zenodo.10361647" httpUri="https://zenodo.org/record/10361647/files/figure.png" pageId="29" pageNumber="280">20C &amp; E</figureCitation>
) and apical lobe of paramere (
<figureCitation id="136C2A45425AFFEC81563F134DC0FAFF" box="[875,987,1347,1373]" captionStart="FIGURE 18" captionStartId="28.[151,250,1943,1967]" captionTargetBox="[183,1404,191,1911]" captionTargetId="figure-22@28.[173,1413,181,1920]" captionTargetPageId="28" captionText="FIGURE 18 Genitalia of Amblopusa spp. A &amp; B, A. magna (A, median lobe of aedeagus; B, spermatheca); C &amp; D, A. pacifica (C, median lobe of aedeagus; D, spermatheca); EG, A. brevipes (E, median lobe of aedeagus; F, spermatheca; G, apical lobe of paramere). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10361639" httpUri="https://zenodo.org/record/10361639/files/figure.png" pageId="29" pageNumber="280">Figs. 18G</figureCitation>
&amp;
<figureCitation id="136C2A45425AFFEC81C23F134A2AFAFF" box="[1023,1073,1347,1373]" captionStart="FIGURE 20" captionStartId="31.[151,250,1874,1898]" captionTargetBox="[160,1426,191,1841]" captionTargetId="figure-19@31.[151,1436,181,1851]" captionTargetPageId="31" captionText="FIGURE 20 Halorhadinus aequalis. A, Habitus; B, labrum (right side, dorsal view; left side, ventral view (epipharynx)); C, mentum; D, labium; E, maxilla; F, right mandible; G, apical lobe of paramere. Scale bars: 1.0 mm for A; 0.1 mm for BG." figureDoi="http://doi.org/10.5281/zenodo.10361647" httpUri="https://zenodo.org/record/10361647/files/figure.png" pageId="29" pageNumber="280">20G</figureCitation>
) but can be distinguished by the following characteristics: hind wings sometimes present, female spermatheca coiled many times, mandible elongate (
<figureCitation id="136C2A45425AFFEC82A23FDB4F19FA07" box="[159,258,1419,1445]" captionStart="FIGURE 20" captionStartId="31.[151,250,1874,1898]" captionTargetBox="[160,1426,191,1841]" captionTargetId="figure-19@31.[151,1436,181,1851]" captionTargetPageId="31" captionText="FIGURE 20 Halorhadinus aequalis. A, Habitus; B, labrum (right side, dorsal view; left side, ventral view (epipharynx)); C, mentum; D, labium; E, maxilla; F, right mandible; G, apical lobe of paramere. Scale bars: 1.0 mm for A; 0.1 mm for BG." figureDoi="http://doi.org/10.5281/zenodo.10361647" httpUri="https://zenodo.org/record/10361647/files/figure.png" pageId="29" pageNumber="280">Fig. 20F</figureCitation>
), labrum as long as wide (
<figureCitation id="136C2A45425AFFEC801C3FDB4C9CFA07" box="[545,647,1419,1445]" captionStart="FIGURE 20" captionStartId="31.[151,250,1874,1898]" captionTargetBox="[160,1426,191,1841]" captionTargetId="figure-19@31.[151,1436,181,1851]" captionTargetPageId="31" captionText="FIGURE 20 Halorhadinus aequalis. A, Habitus; B, labrum (right side, dorsal view; left side, ventral view (epipharynx)); C, mentum; D, labium; E, maxilla; F, right mandible; G, apical lobe of paramere. Scale bars: 1.0 mm for A; 0.1 mm for BG." figureDoi="http://doi.org/10.5281/zenodo.10361647" httpUri="https://zenodo.org/record/10361647/files/figure.png" pageId="29" pageNumber="280">Fig. 20B</figureCitation>
), and two medial setae present on prementum (
<figureCitation id="136C2A45425AFFEC86B23FDB4AECFA07" box="[1167,1271,1419,1445]" captionStart="FIGURE 20" captionStartId="31.[151,250,1874,1898]" captionTargetBox="[160,1426,191,1841]" captionTargetId="figure-19@31.[151,1436,181,1851]" captionTargetPageId="31" captionText="FIGURE 20 Halorhadinus aequalis. A, Habitus; B, labrum (right side, dorsal view; left side, ventral view (epipharynx)); C, mentum; D, labium; E, maxilla; F, right mandible; G, apical lobe of paramere. Scale bars: 1.0 mm for A; 0.1 mm for BG." figureDoi="http://doi.org/10.5281/zenodo.10361647" httpUri="https://zenodo.org/record/10361647/files/figure.png" pageId="29" pageNumber="280">Fig. 20D</figureCitation>
). Comparisons with other genera of
<taxonomicName id="4C574D43425AFFEC83BD3FFF4C2FFA6B" authorityName="Casey" authorityYear="1894" box="[384,564,1455,1481]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="280" phylum="Arthropoda" rank="tribe" tribe="Liparocephalini">Liparocephalini</taxonomicName>
, see the key to genera and diagnosis of
<taxonomicName id="4C574D43425AFFEC81CD3FFF4A75FA6B" authorityName="Casey" authorityYear="1894" box="[1008,1134,1455,1481]" class="Insecta" family="Staphylinidae" genus="Amblopusa" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="280" phylum="Arthropoda" rank="genus">
<emphasis id="B923EAD2425AFFEC81CD3FFF4A75FA6B" box="[1008,1134,1455,1481]" italics="true" pageId="29" pageNumber="280">Amblopusa</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BE836C0425AFFEC82FA3F824FC3F9B3" blockId="29.[151,1436,1310,1733]" pageId="29" pageNumber="280">
<emphasis id="B923EAD2425AFFEC82FA3F824F31FA4E" bold="true" box="[199,298,1490,1516]" pageId="29" pageNumber="280">Habitat.</emphasis>
All species of this genus mainly have been collected from gravel beaches and rarely from under driftwood on sandy beaches.
</paragraph>
</subSubSection>
<subSubSection id="C34D654B425AFFEC82FA3C4B4B06F903" pageId="29" pageNumber="280" type="discussion">
<paragraph id="8BE836C0425AFFEC82FA3C4B4B06F903" blockId="29.[151,1436,1310,1733]" pageId="29" pageNumber="280">
<emphasis id="B923EAD2425AFFEC82FA3C4B4F20F997" bold="true" box="[199,315,1563,1589]" pageId="29" pageNumber="280">Remarks.</emphasis>
Eight species of this genus are recorded from
<collectingCountry id="F3407650425AFFEC81693C4B4D8CF997" box="[852,919,1563,1589]" name="Japan" pageId="29" pageNumber="280">Japan</collectingCountry>
. They were relatively recently redescribed or described, and their diagnostic characters are well illustrated (
<bibRefCitation id="EFC64B31425AFFEC81583C6F4AA9F9FB" author="Maruyama, M. &amp; Hayashi, M." box="[869,1202,1599,1625]" pageId="29" pageNumber="280" pagination="71 - 82" refId="ref22396" refString="Maruyama, M. &amp; Hayashi, M. (2009) Description of the intertidal aleocharine Halorhadinus sawadai with notes on the genus Halorhadinus Sawada, 1971 (Coleoptera: Staphylinidae). Koleopterologische Rundschau, 79, 71 - 82." type="journal article" year="2009">Maruyama &amp; Hayashi, 2009</bibRefCitation>
;
<bibRefCitation id="EFC64B31425AFFEC86FC3C6F4ECFF9DF" author="Ono, H. &amp; Maruyama, M." pageId="29" pageNumber="280" pagination="41 - 50" refId="ref22885" refString="Ono, H. &amp; Maruyama, M. (2014) Five new species of the intertidal genus Halorhadinus Sawada, 1971 (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Esakia, 54, 41 - 50. https: // doi. org / 10.5109 / 1517831" type="journal article" year="2014">Ono &amp; Maruyama, 2014</bibRefCitation>
). Therefore, only the diagnosis of the genus is mentioned here. See,
<bibRefCitation id="EFC64B31425AFFEC81F23C334B02F9DF" author="Maruyama, M. &amp; Hayashi, M." box="[975,1305,1635,1661]" pageId="29" pageNumber="280" pagination="71 - 82" refId="ref22396" refString="Maruyama, M. &amp; Hayashi, M. (2009) Description of the intertidal aleocharine Halorhadinus sawadai with notes on the genus Halorhadinus Sawada, 1971 (Coleoptera: Staphylinidae). Koleopterologische Rundschau, 79, 71 - 82." type="journal article" year="2009">Maruyama &amp; Hayashi (2009)</bibRefCitation>
and
<bibRefCitation id="EFC64B31425AFFEC87723C334F79F903" author="Ono, H. &amp; Maruyama, M." pageId="29" pageNumber="280" pagination="41 - 50" refId="ref22885" refString="Ono, H. &amp; Maruyama, M. (2014) Five new species of the intertidal genus Halorhadinus Sawada, 1971 (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Esakia, 54, 41 - 50. https: // doi. org / 10.5109 / 1517831" type="journal article" year="2014">Ono &amp; Maruyama (2014)</bibRefCitation>
for species identification. Our illustrations are based on the
<typeStatus id="54EC8862425AFFEC863B3CD74A2DF903" box="[1030,1078,1671,1697]" pageId="29" pageNumber="280">type</typeStatus>
species
<taxonomicName id="4C574D43425AFFEC86A83CD74B0DF903" authorityName="Sawada" authorityYear="1971" box="[1173,1302,1671,1697]" class="Insecta" family="Staphylinidae" genus="Halorhadinus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="280" phylum="Arthropoda" rank="species" species="aequalis">
<emphasis id="B923EAD2425AFFEC86A83CD74B0DF903" box="[1173,1302,1671,1697]" italics="true" pageId="29" pageNumber="280">H. aequalis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C34D654B425AFFEC82FA3CFA4DAEF967" box="[199,949,1706,1733]" pageId="29" pageNumber="280" type="distribution">
<paragraph id="8BE836C0425AFFEC82FA3CFA4DAEF967" blockId="29.[151,1436,1310,1733]" box="[199,949,1706,1733]" pageId="29" pageNumber="280">
<emphasis id="B923EAD2425AFFEC82FA3CFA4F7AF966" bold="true" box="[199,353,1706,1732]" pageId="29" pageNumber="280">Distribution.</emphasis>
<collectingCountry id="F3407650425AFFEC835A3CFB4FB0F967" box="[359,427,1707,1733]" name="Japan" pageId="29" pageNumber="280">Japan</collectingCountry>
:
<collectingRegion id="4993F822425AFFEC838B3CFB4C3CF967" box="[438,551,1707,1733]" country="Japan" name="Hokkaido" pageId="29" pageNumber="280">Hokkaidô</collectingRegion>
, Honshû, Shikoku, Kyûshû;
<collectingCountry id="F3407650425AFFEC81543CFB4DAAF967" box="[873,945,1707,1733]" name="South Korea" pageId="29" pageNumber="280">Korea</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>