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<document id="C1F8B237CAC28E24F12CBFFB3850D71A" ID-DOI="10.11646/zootaxa.5383.3.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10361564" ID-ZooBank="20988E42-C14D-4878-A716-8CEAB5E5EF92" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1702374886811" checkinUser="plazi" docAuthor="Tasaku, Yuto, Ono, Hiroki &amp; Maruyama, Munetoshi" docDate="2023" docId="03FE87D6424DFFFB82AA3EC24A29F855" docLanguage="en" docName="zootaxa.5383.3.1.pdf" docOrigin="Zootaxa 5383 (3)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5383.3.1/52454" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Diaulota hokkaidona Ahn &amp; Ohara 2018" docType="treatment" docVersion="2" lastPageNumber="261" masterDocId="FFC7FFAE4247FFF1823D3A504E1BFFA2" masterDocTitle="Review of the intertidal rove beetle tribe Liparocephalini Fenyes (Coleoptera, Staphylinidae, Aleocharinae) from Japan" masterLastPageNumber="296" masterPageNumber="251" pageNumber="261" updateTime="1702375545460" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="6D16919DBE9A653BB87EED5E7ECCF966" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="192CBA46B3C2FB47B791FAABEF344FDB">Review of the intertidal rove beetle tribe Liparocephalini Fenyes (Coleoptera, Staphylinidae, Aleocharinae) from Japan</mods:title>
</mods:titleInfo>
<mods:name id="231AFDBF58EBF96CEA4E986A1F5D2603" type="personal">
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<mods:roleTerm id="43FA3B4451F739358715417473EA4A65">Author</mods:roleTerm>
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<mods:namePart id="4D8BA0227458F6DD29AFC928E35777D7">Tasaku, Yuto</mods:namePart>
<mods:affiliation id="4E9402BB840BFF7BFC8CE9B4DFE7BEBA">Department of Biology, School of Biological Sciences, Tokai University, Hokkaido, 005 - 8601, Japan. &amp; Corresponding author</mods:affiliation>
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<mods:role id="53620EA8B7CB8E98FBB94147A2FDEAB1">
<mods:roleTerm id="21C040D7C8F82797EC9FB0DADC5EC3F5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9A69EEFB8B31AF2D78CF4E064C8BDABF">Ono, Hiroki</mods:namePart>
<mods:affiliation id="B57F5F28E5B6978BCEC95C3C9BAEB9D4">Sounan, Minami-ku, Sagamihara-shi, Kanagawa, 252 - 0312, Japan.</mods:affiliation>
</mods:name>
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<mods:roleTerm id="D17760BFAC053E7980CCCECF4E834C85">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="13C1AB1F3A74EE0392F122D9769674E4">Maruyama, Munetoshi</mods:namePart>
<mods:affiliation id="34DA58778F49BD305AD313ECBFACD89F">The Kyushu University Museum, Hakozaki 6 - 10 - 1, Higashi-ku, Fukuoka, 812 - 8581, Japan.</mods:affiliation>
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<mods:title id="733D7AE36683AD3734B0299606DF5AF6">Zootaxa</mods:title>
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<mods:part id="9E78CD93334B6A525F6A0C6760CAB049">
<mods:date id="E7ADDB58EACE5ABD4028CAB2E5D5594B">2023</mods:date>
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<mods:number id="80CE03025A46EDE9440F97E50FB8C371">2023-12-12</mods:number>
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<mods:number id="A45BE94E5D5545FD848BE6DD123F8982">5383</mods:number>
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<mods:number id="D82C5BA98A42AC08EE8CE9D0D95B94E5">3</mods:number>
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<mods:identifier id="4D9DBC370A2C67DAC03949BA6105B6C4" type="DOI">10.11646/zootaxa.5383.3.1</mods:identifier>
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<treatment id="03FE87D6424DFFFB82AA3EC24A29F855" ID-DOI="http://doi.org/10.5281/zenodo.10361626" ID-Zenodo-Dep="10361626" LSID="urn:lsid:plazi:treatment:03FE87D6424DFFFB82AA3EC24A29F855" httpUri="http://treatment.plazi.org/id/03FE87D6424DFFFB82AA3EC24A29F855" lastPageNumber="261" pageId="10" pageNumber="261">
<subSubSection id="C34D654B424DFFFB82AA3EC24D1EFB72" pageId="10" pageNumber="261" type="nomenclature">
<paragraph id="8BE836C0424DFFFB82AA3EC24C8DFB0E" blockId="10.[151,773,1170,1269]" box="[151,662,1170,1200]" pageId="10" pageNumber="261">
<heading id="D0A081AC424DFFFB82AA3EC24C8DFB0E" bold="true" box="[151,662,1170,1200]" fontSize="11" level="1" pageId="10" pageNumber="261" reason="1">
<taxonomicName id="4C574D43424DFFFB82AA3EC24C8DFB0E" ID-CoL="35GKG" authority="Ahn &amp; Ohara, 2018" authorityName="Ahn &amp; Ohara" authorityYear="2018" box="[151,662,1170,1200]" class="Insecta" family="Staphylinidae" genus="Diaulota" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="261" phylum="Arthropoda" rank="species" species="hokkaidona">
<emphasis id="B923EAD2424DFFFB82AA3EC24C8DFB0E" bold="true" box="[151,662,1170,1200]" pageId="10" pageNumber="261">
<emphasis id="B923EAD2424DFFFB82AA3EC24F8CFB0F" bold="true" box="[151,407,1170,1197]" italics="true" pageId="10" pageNumber="261">Diaulota hokkaidona</emphasis>
Ahn &amp; Ôhara, 2018
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BE836C0424DFFFB82AA3EE74D1EFB72" blockId="10.[151,773,1170,1269]" box="[151,773,1206,1233]" pageId="10" pageNumber="261">[Japanese name: Hokkai-tsutsumune-umi-hanekakushi]</paragraph>
</subSubSection>
<subSubSection id="C34D654B424DFFFB82AA3E8A4FCDFB56" box="[151,470,1242,1269]" pageId="10" pageNumber="261" type="description">
<paragraph id="8BE836C0424DFFFB82AA3E8A4FCDFB56" blockId="10.[151,773,1170,1269]" box="[151,470,1242,1269]" pageId="10" pageNumber="261">
(
<figureCitation id="136C2A45424DFFFB82A23E8A4F19FB57" box="[159,258,1242,1269]" captionStart="FIGURE 5" captionStartId="10.[151,250,1074,1098]" captionTargetBox="[188,1400,284,1040]" captionTargetId="figure-60@10.[177,1411,273,1051]" captionTargetPageId="10" captionText="FIGURE 5 Habitus of Diaulota spp. A, D. aokii; B, D. decipiens sp. n. (holotype); C, D. hokkaidona. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10361578" httpUri="https://zenodo.org/record/10361578/files/figure.png" pageId="10" pageNumber="261">Figs. 5C</figureCitation>
;
<figureCitation id="136C2A45424DFFFB83333E8A4F47FB57" box="[270,348,1242,1269]" captionStart="FIGURE 6" captionStartId="11.[151,250,1943,1967]" captionTargetBox="[191,1395,190,1910]" captionTargetId="figure-19@11.[183,1403,181,1919]" captionTargetPageId="11" captionText="FIGURE 6 Genitalia of Diaulota spp. A–C &amp; F, median lobe of aedeagus (A, D. aokii, lateral view; B, ditto, ventral view; C, D. decipiens sp. n., lateral view; F, D. hokkaidona); D &amp; E, apical lobe of median lobe, lateral view (D, D. aokii; E, D. decipiens sp. n.); G–I, spermatheca (G, D. aokii; H, D. decipiens sp. n.; I, D. hokkaidona). Scale bars: 0.1 mm" figureDoi="http://doi.org/10.5281/zenodo.10361584" httpUri="https://zenodo.org/record/10361584/files/figure.png" pageId="10" pageNumber="261">6F &amp; I</figureCitation>
;
<figureCitation id="136C2A45424DFFFB83553E8A4FD5FB56" box="[360,462,1242,1268]" captionStart="FIGURE 23" captionStartId="35.[151,250,1207,1231]" captionTargetBox="[151,1436,181,1183]" captionTargetId="figure-19@35.[151,1436,181,1183]" captionTargetPageId="35" captionText="FIGURE 23 Distribution and habitat of Diaulota aokii, D. decipiens sp. n., and D. hokkaidona. A, known localities in Japan (red ring, type locarity of D. aokii; red circle, D. aokii; blue square, D. hokkaidona; yellow triangle, D. decipiens sp. n.); B, habitat of D. aokii (Izu-ôshima); C, habitat of D. hokkaidona; white arrow, adults of eating barnacle (Nemuro-shi, Hokkaidô, Photographed by Tomoki Kikuchi)." figureDoi="http://doi.org/10.5281/zenodo.10361663" httpUri="https://zenodo.org/record/10361663/files/figure.png" pageId="10" pageNumber="261">23A &amp; C</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34D654B424DFFFB82AA3F494F24FAF2" pageId="10" pageNumber="261" type="reference_group">
<paragraph id="8BE836C0424DFFFB82AA3F494F24FAF2" blockId="10.[151,1437,1305,1360]" pageId="10" pageNumber="261">
<treatmentCitationGroup id="AB4711EE424DFFFB82AA3F494F24FAF2" pageId="10" pageNumber="261">
<taxonomicName id="4C574D43424DFFFB82AA3F494C0CFA96" ID-CoL="35GKG" authority="Ahn &amp; Ohara" authorityName="Ahn &amp; Ohara" authorityYear="2018" box="[151,535,1305,1332]" class="Insecta" family="Staphylinidae" genus="Diaulota" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="261" phylum="Arthropoda" rank="species" species="hokkaidona">
<emphasis id="B923EAD2424DFFFB82AA3F494F6FFA93" box="[151,372,1305,1329]" italics="true" pageId="10" pageNumber="261">Diaulota hokkaidona</emphasis>
<materialsCitation id="3B3F3C9D424DFFFB83BD3F494C0CFA96" box="[384,535,1305,1332]" location="Tomoshiri" municipality="Nemuro" pageId="10" pageNumber="261" specimenCount="1" stateProvince="Hokkaido">Ahn &amp; Ôhara</materialsCitation>
</taxonomicName>
in
<treatmentCitation id="0AF610D1424DFFFB80793F494D25FA93" author="Song, J. - H. &amp; Yoo, I. - S. &amp; Ahn, K. - J." box="[580,830,1305,1329]" page="932" pageId="10" pageNumber="261" year="2018">
<bibRefCitation id="EFC64B31424DFFFB80793F494D25FA93" author="Song, J. - H. &amp; Yoo, I. - S. &amp; Ahn, K. - J." box="[580,830,1305,1329]" pageId="10" pageNumber="261" pagination="930 - 936" refId="ref23507" refString="Song, J. - H., Yoo, I. - S. &amp; Ahn, K. - J. (2018) Discovery of two new Diaulota Casey species (Coleoptera: Staphylinidae: Aleocharinae) from coastal Hokkaido, Japan based on morphological and molecular characters. Journal of Asia-Pacific Entomology, 21, 930 - 936. https: // doi. org / 10.1016 / j. aspen. 2018.07.002" type="journal article" year="2018">
Song
<emphasis id="B923EAD2424DFFFB80B83F4A4CA3FA93" box="[645,696,1305,1329]" italics="true" pageId="10" pageNumber="261">et al</emphasis>
., 2018: 932
</bibRefCitation>
</treatmentCitation>
(original description;
<typeStatus id="54EC8862424DFFFB860E3F4A4A44FA90" box="[1075,1119,1306,1330]" pageId="10" pageNumber="261">type</typeStatus>
locality:
<collectingRegion id="4993F822424DFFFB86F13F494B28FA93" box="[1228,1331,1305,1329]" country="Japan" name="Hokkaido" pageId="10" pageNumber="261">Hokkaido</collectingRegion>
,
<collectingMunicipality id="6B8CACBA424DFFFB877E3F4A4B82FA93" box="[1347,1433,1306,1329]" pageId="10" pageNumber="261">Nemuro</collectingMunicipality>
,
<location id="8E88601B424DFFFB82FA3F684F2FFAF2" LSID="urn:lsid:plazi:treatment:03FE87D6424DFFFB82AA3EC24A29F855:8E88601B424DFFFB82FA3F684F2FFAF2" box="[199,308,1336,1360]" municipality="Nemuro" name="Tomoshiri" pageId="10" pageNumber="261" stateProvince="Hokkaido">Tomoshiri</location>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C34D654B424DFFFB82AA3F284C61F92D" pageId="10" pageNumber="261" type="materials_examined">
<paragraph id="8BE836C0424DFFFB82AA3F284C61F92D" blockId="10.[151,1437,1400,2039]" pageId="10" pageNumber="261">
<emphasis id="B923EAD2424DFFFB82AA3F284F9FFA30" bold="true" box="[151,388,1400,1426]" pageId="10" pageNumber="261">Material examined.</emphasis>
<materialsCitation id="3B3F3C9D424DFFFB83B33F294F03F9C9" collectingDate="1999-07-25" collectingDateMax="2022-06-20" collectingDateMin="1999-07-25" collectionCode="OMNH, KUM" collectorName="Y. Tasaku &amp; Abashiri-shi &amp; Komaba &amp; Nemuro-shi &amp; Toyosato &amp; Tosamuporo-ko &amp; Hamamatsu &amp; M. Maruyama &amp; Hamanaka-cho &amp; Mochirippu &amp; Akkeshi-cho &amp; Aikappu-misaki &amp; Erimo-cho &amp; Toyo-gyoko &amp; H. Ono" country="Japan" location="Onneto" municipality="Nemuro-shi" pageId="10" pageNumber="261" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Hokkaido">
<emphasis id="B923EAD2424DFFFB83B33F294C71FA30" bold="true" box="[398,618,1400,1427]" pageId="10" pageNumber="261">
<collectingCountry id="F3407650424DFFFB83B33F294FC0FA31" box="[398,475,1401,1427]" name="Japan" pageId="10" pageNumber="261">Japan</collectingCountry>
:
<collectingRegion id="4993F822424DFFFB83D63F284C7DFA30" box="[491,614,1400,1426]" country="Japan" name="Hokkaido" pageId="10" pageNumber="261">Hokkaidô</collectingRegion>
:
</emphasis>
<specimenCount id="9D51FD49424DFFFB80493F294CDDFA31" box="[628,710,1401,1427]" count="1" pageId="10" pageNumber="261" type="male">1 male</specimenCount>
,
<specimenCount id="9D51FD49424DFFFB80E93F294D20FA31" box="[724,827,1401,1427]" count="1" pageId="10" pageNumber="261" type="female">1 female</specimenCount>
, 19 unsexed,
<collectingMunicipality id="6B8CACBA424DFFFB81E03F294A7DFA31" box="[989,1126,1401,1427]" pageId="10" pageNumber="261">Nemuro-shi</collectingMunicipality>
,
<location id="8E88601B424DFFFB86493F294AD2FA31" LSID="urn:lsid:plazi:treatment:03FE87D6424DFFFB82AA3EC24A29F855:8E88601B424DFFFB86493F294AD2FA31" box="[1140,1225,1401,1427]" country="Japan" municipality="Nemuro-shi" name="Onneto" pageId="10" pageNumber="261" stateProvince="Hokkaido">Onnetô</location>
,
<date id="FFE91000424DFFFB86E53F294B6CFA31" box="[1240,1399,1401,1427]" pageId="10" pageNumber="261" value="2020-07-24">
<collectingDate id="EFADE9E8424DFFFB86E53F294B6CFA31" box="[1240,1399,1401,1427]" pageId="10" pageNumber="261" value="2020-07-24">24. VII. 2020</collectingDate>
</date>
,
<collectorName id="26A25316424DFFFB87B83F294EFDFA15" pageId="10" pageNumber="261">Y. Tasaku</collectorName>
(
<collectionCode id="ED46AE05424DFFFB82CA3FCD4F4AFA15" box="[247,337,1437,1463]" country="Japan" httpUri="http://grbio.org/cool/rcjv-ktyc" name="Osaka Museum of Natural History" pageId="10" pageNumber="261" type="Museum">OMNH</collectionCode>
, cYT); 22 unsexed, ditto,
<date id="FFE91000424DFFFB80423FCD4D0FFA15" box="[639,788,1437,1463]" pageId="10" pageNumber="261" value="2020-09-18">
<collectingDate id="EFADE9E8424DFFFB80423FCD4D0FFA15" box="[639,788,1437,1463]" pageId="10" pageNumber="261" value="2020-09-18">18. IX. 2020</collectingDate>
</date>
,
<collectorName id="26A25316424DFFFB811D3FCD4D94FA15" box="[800,911,1437,1463]" pageId="10" pageNumber="261">Y. Tasaku</collectorName>
(cYT); 1 unsexed,
<collectorName id="26A25316424DFFFB86523FCD4AE7FA15" box="[1135,1276,1437,1463]" pageId="10" pageNumber="261">Abashiri-shi</collectorName>
,
<collectorName id="26A25316424DFFFB87373FCD4B77FA15" box="[1290,1388,1437,1463]" pageId="10" pageNumber="261">Komaba</collectorName>
,
<date id="FFE91000424DFFFB87443FCD4EE5FA79" pageId="10" pageNumber="261" value="2022-06-20">
<collectingDate id="EFADE9E8424DFFFB87443FCD4EE5FA79" pageId="10" pageNumber="261" value="2022-06-20">20. VI. 2022</collectingDate>
</date>
,
<collectorName id="26A25316424DFFFB83353F914F6EFA79" box="[264,373,1473,1499]" pageId="10" pageNumber="261">Y. Tasaku</collectorName>
(cYT); 13 unsexed,
<collectorName id="26A25316424DFFFB80613F914CFEFA79" box="[604,741,1473,1499]" pageId="10" pageNumber="261">Nemuro-shi</collectorName>
,
<collectorName id="26A25316424DFFFB80D23F914D43FA79" box="[751,856,1473,1499]" pageId="10" pageNumber="261">Toyosato</collectorName>
,
<collectorName id="26A25316424DFFFB815E3F914A02FA79" box="[867,1049,1473,1499]" pageId="10" pageNumber="261">Tôsamuporo-ko</collectorName>
,
<date id="FFE91000424DFFFB86193F914AA6FA79" box="[1060,1213,1473,1499]" pageId="10" pageNumber="261" value="2020-07-24">
<collectingDate id="EFADE9E8424DFFFB86193F914AA6FA79" box="[1060,1213,1473,1499]" pageId="10" pageNumber="261" value="2020-07-24">24. VII. 2020</collectingDate>
</date>
,
<collectorName id="26A25316424DFFFB86FA3F914B2FFA79" box="[1223,1332,1473,1499]" pageId="10" pageNumber="261">Y. Tasaku</collectorName>
(cYT); 1 unsexed,
<collectorName id="26A25316424DFFFB833D3FB54F92FA5D" box="[256,393,1509,1535]" pageId="10" pageNumber="261">Nemuro-shi</collectorName>
,
<collectorName id="26A25316424DFFFB83AF3FB54C00FA5D" box="[402,539,1509,1535]" pageId="10" pageNumber="261">Hamamatsu</collectorName>
,
<date id="FFE91000424DFFFB80183FB54CA8FA5D" box="[549,691,1509,1535]" pageId="10" pageNumber="261" value="2000-09-28">
<collectingDate id="EFADE9E8424DFFFB80183FB54CA8FA5D" box="[549,691,1509,1535]" pageId="10" pageNumber="261" value="2000-09-28">28. IX. 2000</collectingDate>
</date>
,
<collectorName id="26A25316424DFFFB80813FB54D40FA5D" box="[700,859,1509,1535]" pageId="10" pageNumber="261">M. Maruyama</collectorName>
(
<collectionCode id="ED46AE05424DFFFB81543FB54DABFA5D" box="[873,944,1509,1535]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="10" pageNumber="261" type="Herbarium">KUM</collectionCode>
); 120 unsexed,
<collectorName id="26A25316424DFFFB86663FB54B1CFA5D" box="[1115,1287,1509,1535]" pageId="10" pageNumber="261">Hamanaka-chô</collectorName>
,
<collectorName id="26A25316424DFFFB872C3FB54B83FA5D" box="[1297,1432,1509,1535]" pageId="10" pageNumber="261">Mochirippu</collectorName>
,
<date id="FFE91000424DFFFB82AA3C594F36F981" box="[151,301,1545,1571]" pageId="10" pageNumber="261" value="1999-09-12">
<collectingDate id="EFADE9E8424DFFFB82AA3C594F36F981" box="[151,301,1545,1571]" pageId="10" pageNumber="261" value="1999-09-12">12. IX. 1999</collectingDate>
</date>
,
<collectorName id="26A25316424DFFFB83073C594FC6F981" box="[314,477,1545,1571]" pageId="10" pageNumber="261">M. Maruyama</collectorName>
(
<collectionCode id="ED46AE05424DFFFB83D33C594C2EF981" box="[494,565,1545,1571]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="10" pageNumber="261" type="Herbarium">KUM</collectionCode>
); 1 unsexed, ditto,
<date id="FFE91000424DFFFB812E3C594DB2F981" box="[787,937,1545,1571]" pageId="10" pageNumber="261" value="2020-09-17">
<collectingDate id="EFADE9E8424DFFFB812E3C594DB2F981" box="[787,937,1545,1571]" pageId="10" pageNumber="261" value="2020-09-17">17. IX. 2020</collectingDate>
</date>
,
<collectorName id="26A25316424DFFFB81883C594A3EF981" box="[949,1061,1545,1571]" pageId="10" pageNumber="261">Y. Tasaku</collectorName>
(cYT); 2 unsexed,
<collectorName id="26A25316424DFFFB873B3C594B83F981" box="[1286,1432,1545,1571]" pageId="10" pageNumber="261">Akkeshi-chô</collectorName>
,
<collectorName id="26A25316424DFFFB82AA3C7D4F55F9E5" box="[151,334,1581,1607]" pageId="10" pageNumber="261">Aikappu-misaki</collectorName>
,
<date id="FFE91000424DFFFB83643C7D4FEBF9E5" box="[345,496,1581,1608]" pageId="10" pageNumber="261" value="1999-07-25">
<collectingDate id="EFADE9E8424DFFFB83643C7D4FEBF9E5" box="[345,496,1581,1608]" pageId="10" pageNumber="261" value="1999-07-25">25. VII. 1999</collectingDate>
</date>
,
<collectorName id="26A25316424DFFFB83C63C7D4C81F9E5" box="[507,666,1581,1607]" pageId="10" pageNumber="261">M. Maruyama</collectorName>
(
<collectionCode id="ED46AE05424DFFFB80953C7D4CF4F9E5" box="[680,751,1581,1607]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="10" pageNumber="261" type="Herbarium">KUM</collectionCode>
); 15 unsexed,
<collectorName id="26A25316424DFFFB81B03C7D4A13F9E5" box="[909,1032,1581,1607]" pageId="10" pageNumber="261">Erimo-chô</collectorName>
, near
<collectorName id="26A25316424DFFFB86753C7D4ACEF9E5" box="[1096,1237,1581,1607]" pageId="10" pageNumber="261">Tôyô-gyokô</collectorName>
,
<date id="FFE91000424DFFFB86E23C7D4B6CF9E5" box="[1247,1399,1581,1607]" pageId="10" pageNumber="261" value="2018-07-12">
<collectingDate id="EFADE9E8424DFFFB86E23C7D4B6CF9E5" box="[1247,1399,1581,1607]" pageId="10" pageNumber="261" value="2018-07-12">12. VII. 2018</collectingDate>
</date>
,
<collectorName id="26A25316424DFFFB87BC3C7D4EDCF9C9" pageId="10" pageNumber="261">H. Ono</collectorName>
(cHO)
</materialsCitation>
;
<specimenCount id="9D51FD49424DFFFB83193C014F6AF9C9" box="[292,369,1617,1643]" count="1" pageId="10" pageNumber="261" type="male">1 male</specimenCount>
, Kaminokuni-chô, Harauta,
<date id="FFE91000424DFFFB80903C014D22F9C9" box="[685,825,1617,1643]" pageId="10" pageNumber="261" value="2020-04-11">11. IV. 2020</date>
. K. Mitsuhashi (cYT);
<specimenCount id="9D51FD49424DFFFB86073C014A9CF9C9" box="[1082,1159,1617,1643]" count="1" pageId="10" pageNumber="261" type="male">1 male</specimenCount>
, 3 unsexed, Samani-chô, Fuyushima,
<date id="FFE91000424DFFFB831F3C254FAAF92D" box="[290,433,1653,1679]" pageId="10" pageNumber="261" value="2020-03-28">28. III. 2020</date>
, Y. Tasaku (cYT).
</paragraph>
</subSubSection>
<subSubSection id="C34D654B424DFFFB82FA3CC84C59F871" pageId="10" pageNumber="261" type="diagnosis">
<paragraph id="8BE836C0424DFFFB82FA3CC84C29F8BD" blockId="10.[151,1437,1400,2039]" pageId="10" pageNumber="261">
<emphasis id="B923EAD2424DFFFB82FA3CC84F59F910" bold="true" box="[199,322,1688,1714]" pageId="10" pageNumber="261">Diagnosis.</emphasis>
This species can be distinguished from other species of the genus
<taxonomicName id="4C574D43424DFFFB860A3CC94A80F911" authorityName="Casey" authorityYear="1893" box="[1079,1179,1689,1715]" class="Insecta" family="Staphylinidae" genus="Diaulota" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="261" phylum="Arthropoda" rank="genus">
<emphasis id="B923EAD2424DFFFB860A3CC94A80F911" box="[1079,1179,1689,1715]" italics="true" pageId="10" pageNumber="261">Diaulota</emphasis>
</taxonomicName>
by the combination of the following characteristics: body small and entirely blackish; male head as large as that of female; mentum as long as wide; tarsal formula 4-4-5, apical lobe of the median lobe divided at the apex in lateral view (
<figureCitation id="136C2A45424DFFFB86EA3CB14B30F959" box="[1239,1323,1761,1787]" captionStart="FIGURE 6" captionStartId="11.[151,250,1943,1967]" captionTargetBox="[191,1395,190,1910]" captionTargetId="figure-19@11.[183,1403,181,1919]" captionTargetPageId="11" captionText="FIGURE 6 Genitalia of Diaulota spp. A–C &amp; F, median lobe of aedeagus (A, D. aokii, lateral view; B, ditto, ventral view; C, D. decipiens sp. n., lateral view; F, D. hokkaidona); D &amp; E, apical lobe of median lobe, lateral view (D, D. aokii; E, D. decipiens sp. n.); G–I, spermatheca (G, D. aokii; H, D. decipiens sp. n.; I, D. hokkaidona). Scale bars: 0.1 mm" figureDoi="http://doi.org/10.5281/zenodo.10361584" httpUri="https://zenodo.org/record/10361584/files/figure.png" pageId="10" pageNumber="261">Fig. 6F</figureCitation>
) (see also the diagnosis of
<taxonomicName id="4C574D43424DFFFB83723D564FC7F8BD" authority="Tasaku &amp; Ono &amp; Maruyama, 2023" authorityName="Tasaku &amp; Ono &amp; Maruyama" authorityYear="2023" box="[335,476,1797,1823]" class="Insecta" family="Staphylinidae" genus="Diaulota" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="261" phylum="Arthropoda" rank="species" species="decipiens" status="sp. n.">
<emphasis id="B923EAD2424DFFFB83723D564F78F8BD" box="[335,355,1798,1823]" italics="true" pageId="10" pageNumber="261">D</emphasis>
.
<emphasis id="B923EAD2424DFFFB834F3D554FC7F8BD" box="[370,476,1797,1823]" italics="true" pageId="10" pageNumber="261">decipiens</emphasis>
</taxonomicName>
<emphasis id="B923EAD2424DFFFB83DF3D554C39F8BD" bold="true" box="[482,546,1797,1823]" pageId="10" pageNumber="261">
<taxonomicNameLabel id="A21057A9424DFFFB83DF3D554C39F8BD" box="[482,546,1797,1823]" pageId="10" pageNumber="261" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
).
</paragraph>
<paragraph id="8BE836C0424DFFFB82FA3D784C59F871" blockId="10.[151,1437,1400,2039]" pageId="10" pageNumber="261">
<emphasis id="B923EAD2424DFFFB82FA3D784F31F8E0" bold="true" box="[199,298,1832,1858]" pageId="10" pageNumber="261">Habitat.</emphasis>
<materialsCitation id="3B3F3C9D424DFFFB830C3D794C25F871" location="northern Honshu" pageId="10" pageNumber="261" specimenCount="1" stateProvince="Hokkaido">
This species is distributed in almost all areas of
<collectingRegion id="4993F822424DFFFB81773D794DA7F8E1" box="[842,956,1833,1859]" country="Japan" name="Hokkaido" pageId="10" pageNumber="261">Hokkaidô</collectingRegion>
. In the eastern
<collectingRegion id="4993F822424DFFFB86593D794AC9F8E1" box="[1124,1234,1833,1859]" country="Japan" name="Hokkaido" pageId="10" pageNumber="261">Hokkaidô</collectingRegion>
(around Nemuro-shi), adults were commonly collected from rocky beaches with barnacles (
<figureCitation id="136C2A45424DFFFB81DB3D1D4A57F8C5" box="[998,1100,1869,1895]" captionStart="FIGURE 23" captionStartId="35.[151,250,1207,1231]" captionTargetBox="[151,1436,181,1183]" captionTargetId="figure-19@35.[151,1436,181,1183]" captionTargetPageId="35" captionText="FIGURE 23 Distribution and habitat of Diaulota aokii, D. decipiens sp. n., and D. hokkaidona. A, known localities in Japan (red ring, type locarity of D. aokii; red circle, D. aokii; blue square, D. hokkaidona; yellow triangle, D. decipiens sp. n.); B, habitat of D. aokii (Izu-ôshima); C, habitat of D. hokkaidona; white arrow, adults of eating barnacle (Nemuro-shi, Hokkaidô, Photographed by Tomoki Kikuchi)." figureDoi="http://doi.org/10.5281/zenodo.10361663" httpUri="https://zenodo.org/record/10361663/files/figure.png" pageId="10" pageNumber="261">Fig. 23C</figureCitation>
) and sometimes under stones in mudflats. In central and southern
<collectingRegion id="4993F822424DFFFB80093D214CBEF829" box="[564,677,1905,1931]" country="Japan" name="Hokkaido" pageId="10" pageNumber="261">Hokkaidô</collectingRegion>
, it is less common and adults were collected from rock crevices of rocky shores. Considering its extensive distribution in
<collectingRegion id="4993F822424DFFFB813C3DC54D69F80D" box="[769,882,1941,1967]" country="Japan" name="Hokkaido" pageId="10" pageNumber="261">Hokkaidô</collectingRegion>
, there is a possibility that this species might also be distributed in the
<location id="8E88601B424DFFFB83403DE94C25F871" LSID="urn:lsid:plazi:treatment:03FE87D6424DFFFB82AA3EC24A29F855:8E88601B424DFFFB83403DE94C25F871" box="[381,574,1977,2003]" name="northern Honshu" pageId="10" pageNumber="261" stateProvince="Hokkaido">northern Honshû</location>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34D654B424DFFFB82FA3D8C4A29F855" box="[199,1074,2012,2039]" pageId="10" pageNumber="261" type="distribution">
<paragraph id="8BE836C0424DFFFB82FA3D8C4A29F855" blockId="10.[151,1437,1400,2039]" box="[199,1074,2012,2039]" pageId="10" pageNumber="261">
<emphasis id="B923EAD2424DFFFB82FA3D8C4F7AF854" bold="true" box="[199,353,2012,2038]" pageId="10" pageNumber="261">Distribution.</emphasis>
<collectingCountry id="F3407650424DFFFB835A3D8D4FB0F855" box="[359,427,2013,2039]" name="Japan" pageId="10" pageNumber="261">Japan</collectingCountry>
:
<collectingRegion id="4993F822424DFFFB838B3D8D4C33F855" box="[438,552,2013,2039]" country="Japan" name="Hokkaido" pageId="10" pageNumber="261">Hokkaidô</collectingRegion>
;
<collectingCountry id="F3407650424DFFFB800E3D8D4C99F855" box="[563,642,2013,2039]" name="Russia" pageId="10" pageNumber="261">Russia</collectingCountry>
:
<collectingRegion id="4993F822424DFFFB80B03D8D4D0BF855" box="[653,784,2013,2039]" country="Russia" name="Kamchatka" pageId="10" pageNumber="261">Kamchatka</collectingRegion>
;
<collectingCountry id="F3407650424DFFFB81263D8D4D4CF855" box="[795,855,2013,2039]" name="United States of America" pageId="10" pageNumber="261">USA</collectingCountry>
:
<collectingRegion id="4993F822424DFFFB815D3D8D4DB5F855" box="[864,942,2013,2039]" country="United States of America" name="Alaska" pageId="10" pageNumber="261">Alaska</collectingRegion>
(
<figureCitation id="136C2A45424DFFFB81803D8D4A3DF855" box="[957,1062,2013,2039]" captionStart="FIGURE 23" captionStartId="35.[151,250,1207,1231]" captionTargetBox="[151,1436,181,1183]" captionTargetId="figure-19@35.[151,1436,181,1183]" captionTargetPageId="35" captionText="FIGURE 23 Distribution and habitat of Diaulota aokii, D. decipiens sp. n., and D. hokkaidona. A, known localities in Japan (red ring, type locarity of D. aokii; red circle, D. aokii; blue square, D. hokkaidona; yellow triangle, D. decipiens sp. n.); B, habitat of D. aokii (Izu-ôshima); C, habitat of D. hokkaidona; white arrow, adults of eating barnacle (Nemuro-shi, Hokkaidô, Photographed by Tomoki Kikuchi)." figureDoi="http://doi.org/10.5281/zenodo.10361663" httpUri="https://zenodo.org/record/10361663/files/figure.png" pageId="10" pageNumber="261">Fig. 23A</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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