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<document id="5C51EFA41CCEB1B0A92B9C8BD56771AF" ID-DOI="10.11646/zootaxa.5383.3.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10361564" ID-ZooBank="20988E42-C14D-4878-A716-8CEAB5E5EF92" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1702374886811" checkinUser="plazi" docAuthor="Tasaku, Yuto, Ono, Hiroki &amp; Maruyama, Munetoshi" docDate="2023" docId="03FE87D64249FFE082AA3AC74FA9FD27" docLanguage="en" docName="zootaxa.5383.3.1.pdf" docOrigin="Zootaxa 5383 (3)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5383.3.1/52454" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Diaulota uenoi" docType="treatment" docVersion="2" lastPageNumber="268" masterDocId="FFC7FFAE4247FFF1823D3A504E1BFFA2" masterDocTitle="Review of the intertidal rove beetle tribe Liparocephalini Fenyes (Coleoptera, Staphylinidae, Aleocharinae) from Japan" masterLastPageNumber="296" masterPageNumber="251" pageNumber="265" updateTime="1702375545460" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="F35313D07E08FEE5026EAEB85B74274E">Review of the intertidal rove beetle tribe Liparocephalini Fenyes (Coleoptera, Staphylinidae, Aleocharinae) from Japan</mods:title>
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<mods:namePart id="BE040E69800C38604AF4E285AFEA1077">Tasaku, Yuto</mods:namePart>
<mods:affiliation id="3CDC8EFBE854F0485846EA375A048589">Department of Biology, School of Biological Sciences, Tokai University, Hokkaido, 005 - 8601, Japan. &amp; Corresponding author</mods:affiliation>
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<mods:namePart id="378690A265253A0AA1B5AEFAE43C8370">Ono, Hiroki</mods:namePart>
<mods:affiliation id="65357C0AEBEFB35A77106F204D6C0CBF">Sounan, Minami-ku, Sagamihara-shi, Kanagawa, 252 - 0312, Japan.</mods:affiliation>
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<mods:namePart id="CECD839BA45AF91E11B476736ECB828E">Maruyama, Munetoshi</mods:namePart>
<mods:affiliation id="0039EC5C4C064AB0BAE16B698F3403F3">The Kyushu University Museum, Hakozaki 6 - 10 - 1, Higashi-ku, Fukuoka, 812 - 8581, Japan.</mods:affiliation>
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<mods:date id="E74E015CDD16B8363536EFAE95067BFE">2023</mods:date>
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<treatment id="03FE87D64249FFE082AA3AC74FA9FD27" ID-DOI="http://doi.org/10.5281/zenodo.10361631" ID-Zenodo-Dep="10361631" LSID="urn:lsid:plazi:treatment:03FE87D64249FFE082AA3AC74FA9FD27" httpUri="http://treatment.plazi.org/id/03FE87D64249FFE082AA3AC74FA9FD27" lastPageId="17" lastPageNumber="268" pageId="14" pageNumber="265">
<subSubSection id="C34D654B4249FFFF82AA3AC74C89FF77" pageId="14" pageNumber="265" type="nomenclature">
<paragraph id="8BE836C04249FFFF82AA3AC74C08FF13" blockId="14.[151,658,151,249]" box="[151,531,151,178]" pageId="14" pageNumber="265">
<heading id="D0A081AC4249FFFF82AA3AC74C08FF13" bold="true" box="[151,531,151,178]" fontSize="11" level="1" pageId="14" pageNumber="265" reason="1">
<taxonomicName id="4C574D434249FFFF82AA3AC74C08FF13" ID-CoL="35GKM" authority="(Sawada, 1955)" baseAuthorityName="Sawada" baseAuthorityYear="1955" box="[151,531,151,178]" class="Insecta" family="Staphylinidae" genus="Diaulota" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="uenoi">
<emphasis id="B923EAD24249FFFF82AA3AC74C08FF13" bold="true" box="[151,531,151,178]" pageId="14" pageNumber="265">
<emphasis id="B923EAD24249FFFF82AA3AC74F57FF13" bold="true" box="[151,332,151,178]" italics="true" pageId="14" pageNumber="265">Diaulota uenoi</emphasis>
(
<bibRefCitation id="EFC64B314249FFFF83613AC74C11FF13" author="Sawada, K." box="[348,522,151,177]" pageId="14" pageNumber="265" pagination="81 - 87" refId="ref23139" refString="Sawada, K. (1955) Marine Insects of the Tokara Islands VII. Fam. Staphylinidae. Publications of the Seto Marine Biological Laboratory, 5 (1), 81 - 87. https: // doi. org / 10.5134 / 174533" type="journal article" year="1955">Sawada, 1955</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BE836C04249FFFF82AA3AEC4C89FF77" blockId="14.[151,658,151,249]" box="[151,658,187,214]" pageId="14" pageNumber="265">[Japanese name: Shiributo-umi-hanekakushi]</paragraph>
</subSubSection>
<subSubSection id="C34D654B4249FFFF82AA3A8F4F49FF5B" box="[151,338,223,249]" pageId="14" pageNumber="265" type="description">
<paragraph id="8BE836C04249FFFF82AA3A8F4F49FF5B" blockId="14.[151,658,151,249]" box="[151,338,223,249]" pageId="14" pageNumber="265">
(
<figureCitation id="136C2A454249FFFF82A23A8F4EF5FF5B" box="[159,238,223,249]" captionStart="FIGURE 8" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[184,1403,191,1909]" captionTargetId="figure-19@13.[174,1413,181,1919]" captionTargetPageId="13" captionText="FIGURE 8 Diaulota uenoi. A, habitus, male; B, ditto, female; C, labium; D, mentum; E, median lode of aedeagus, lateral view; F, internal structure of median lobe, dorsal view; G, ditto, ventral view; H, spermatheca. Scale bars: 1.0 mm for A, B; 0.1 mm for CH. Abbreviation: as, apical sclerite; su, suspensonia." figureDoi="http://doi.org/10.5281/zenodo.10361594" httpUri="https://zenodo.org/record/10361594/files/figure.png" pageId="14" pageNumber="265">Figs. 8</figureCitation>
,
<figureCitation id="136C2A454249FFFF82C63A8F4F12FF5B" box="[251,265,223,249]" captionStart="FIGURE 9" captionStartId="14.[151,250,1956,1980]" captionTargetBox="[164,1383,828,1922]" captionTargetId="figure-227@14.[164,1423,828,1933]" captionTargetPageId="14" captionText="FIGURE 9 Holotype of Diaulota uenoi. A, dissected body parts and label on slide glass; B, labrum; C, median lobe, lateral view. Scale bars: 0.1 mm for A &amp; B." figureDoi="http://doi.org/10.5281/zenodo.10361598" httpUri="https://zenodo.org/record/10361598/files/figure.png" pageId="14" pageNumber="265">9</figureCitation>
&amp;
<figureCitation id="136C2A454249FFFF83103A8F4F51FF5B" box="[301,330,223,249]" captionStart="FIGURE 24" captionStartId="36.[151,250,1780,1804]" captionTargetBox="[151,1436,345,1757]" captionTargetId="figure-99@36.[151,1436,345,1757]" captionTargetPageId="36" captionText="FIGURE 24 Distribution of Diaulota uenoi (red circle (red ring, type locality)), D. submarina (blue square), and D. orientalis sp. n. (yellow triangle)." figureDoi="http://doi.org/10.5281/zenodo.10361669" httpUri="https://zenodo.org/record/10361669/files/figure.png" pageId="14" pageNumber="265">24</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34D654B4249FFFF82AA3B4E4CE8FED6" pageId="14" pageNumber="265" type="reference_group">
<paragraph id="8BE836C04249FFFF82AA3B4E4F7FFEF7" blockId="14.[151,1436,286,372]" pageId="14" pageNumber="265">
<treatmentCitationGroup id="AB4711EE4249FFFF82AA3B4E4F7FFEF7" pageId="14" pageNumber="265">
<treatmentCitation id="0AF610D14249FFFF82AA3B4E4C35FE94" author="Sawada, K." box="[151,558,286,310]" page="82" pageId="14" pageNumber="265" year="1955">
<taxonomicName id="4C574D434249FFFF82AA3B4E4C35FE94" ID-CoL="3FLL6" authority="Sawada, 1955: 82" authorityName="Sawada" authorityPageNumber="82" authorityYear="1955" box="[151,558,286,310]" class="Insecta" family="Staphylinidae" genus="Genoplectes" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="uenoi">
<emphasis id="B923EAD24249FFFF82AA3B4E4F40FE94" box="[151,347,286,310]" italics="true" pageId="14" pageNumber="265">Genoplectes uenoi</emphasis>
<bibRefCitation id="EFC64B314249FFFF83543B4E4C35FE94" author="Sawada, K." box="[361,558,286,310]" pageId="14" pageNumber="265" pagination="81 - 87" refId="ref23139" refString="Sawada, K. (1955) Marine Insects of the Tokara Islands VII. Fam. Staphylinidae. Publications of the Seto Marine Biological Laboratory, 5 (1), 81 - 87. https: // doi. org / 10.5134 / 174533" type="journal article" year="1955">Sawada, 1955: 82</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description;
<typeStatus id="54EC88624249FFFF81153B4F4D4FFE95" box="[808,852,287,311]" pageId="14" pageNumber="265">type</typeStatus>
locality: Nakanoshima (Yoriki));
<treatmentCitation id="0AF610D14249FFFF86F63B4E4B87FE94" author="Sawada, K." box="[1227,1436,286,310]" page="106" pageId="14" pageNumber="265" year="1971">
<bibRefCitation id="EFC64B314249FFFF86F63B4E4B87FE94" author="Sawada, K." box="[1227,1436,286,310]" pageId="14" pageNumber="265" pagination="81 - 110" refId="ref23228" refString="Sawada, K. (1971) Aleocharinae (Staphylinidae, Coleoptera) from the Intertidal Zone of Japan, Publications of the Seto Marine Biological Laboratory, 19 (2 - 3), 81 - 110. https: // doi. org / 10.5134 / 175665" type="journal article" year="1971">Sawada, 1971: 106</bibRefCitation>
</treatmentCitation>
(redescription).
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</paragraph>
<paragraph id="8BE836C04249FFFF82AA3B0C4CE8FED6" blockId="14.[151,1436,286,372]" box="[151,755,347,372]" pageId="14" pageNumber="265">
<taxonomicName id="4C574D434249FFFF82AA3B0C4F2BFED1" ID-CoL="35GKM" baseAuthorityName="Sawada" baseAuthorityYear="1955" box="[151,304,348,372]" class="Insecta" family="Staphylinidae" genus="Diaulota" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="uenoi">
<emphasis id="B923EAD24249FFFF82AA3B0C4F2BFED1" box="[151,304,348,372]" italics="true" pageId="14" pageNumber="265">Diaulota uenoi</emphasis>
</taxonomicName>
: Ahn, 1996: 284 (partim, misidentification).
</paragraph>
</subSubSection>
<subSubSection id="C34D654B4249FFFF82AA3BF24F86FC86" pageId="14" pageNumber="265" type="materials_examined">
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<emphasis id="B923EAD24249FFFF82AA3BF24F5CFE19" bold="true" box="[151,327,417,444]" pageId="14" pageNumber="265">Type material.</emphasis>
<materialsCitation id="3B3F3C9D4249FFFF836D3BF24C72FDA6" collectionCode="VI, OMNH" collectorName="S. Ueno &amp; K. Sawada &amp; Y. Tasaku" pageId="14" pageNumber="265" specimenCount="2" typeStatus="holotype">
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<typeStatus id="54EC88624249FFFF836D3BF24FA4FE1E" box="[336,447,418,444]" pageId="14" pageNumber="265" type="holotype">Holotype</typeStatus>
.
</emphasis>
Male, “
<taxonomicName id="4C574D434249FFFF801C3BF24CB5FE1E" authorityName="Sawada" authorityYear="1955" box="[545,686,418,444]" class="Insecta" family="Staphylinidae" genus="Genoplectes" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="265" phylum="Arthropoda" rank="genus">Genoplectes</taxonomicName>
/
<taxonomicName id="4C574D434249FFFF80F43BF24D97FE1E" authority="K. Sawada" authorityName="K. Sawada" authorityYear="1955" box="[713,908,418,444]" class="Insecta" family="Staphylinidae" genus="Diaulota" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="uenoi">uenoi K. Sawada</taxonomicName>
/
<emphasis id="B923EAD24249FFFF819B3BF34A13FE18" bold="true" box="[934,1032,419,442]" pageId="14" pageNumber="265">中ノ島,</emphasis>
TOKARA Is. / 4
<collectionCode id="ED46AE054249FFFF86DE3BF24B1AFE1E" box="[1251,1281,418,444]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="14" pageNumber="265">VI</collectionCode>
52.
<collectorName id="26A253164249FFFF877D3BF24B86FE1E" box="[1344,1437,418,444]" pageId="14" pageNumber="265">S. Uéno</collectorName>
/ (
<typeStatus id="54EC88624249FFFF828D3B964F05FE42" box="[176,286,454,480]" pageId="14" pageNumber="265" type="holotype">Holotype</typeStatus>
)” (internal structure of median lobe and sternite VIII lost) (whole body dissected by
<collectorName id="26A253164249FFFF86D23B964B71FE42" box="[1263,1386,454,480]" pageId="14" pageNumber="265">K. Sawada</collectorName>
and mounted by
<collectorName id="26A253164249FFFF831F3BBA4F8AFDA6" box="[290,401,490,516]" pageId="14" pageNumber="265">Y. Tasaku</collectorName>
) (
<collectionCode id="ED46AE054249FFFF839A3BBA4C1BFDA6" box="[423,512,490,516]" country="Japan" httpUri="http://grbio.org/cool/rcjv-ktyc" name="Osaka Museum of Natural History" pageId="14" pageNumber="265" type="Museum">OMNH</collectionCode>
) (
<figureCitation id="136C2A454249FFFF802A3BBA4C40FDA6" box="[535,603,490,516]" captionStart="FIGURE 9" captionStartId="14.[151,250,1956,1980]" captionTargetBox="[164,1383,828,1922]" captionTargetId="figure-227@14.[164,1423,828,1933]" captionTargetPageId="14" captionText="FIGURE 9 Holotype of Diaulota uenoi. A, dissected body parts and label on slide glass; B, labrum; C, median lobe, lateral view. Scale bars: 0.1 mm for A &amp; B." figureDoi="http://doi.org/10.5281/zenodo.10361598" httpUri="https://zenodo.org/record/10361598/files/figure.png" pageId="14" pageNumber="265">Fig. 9</figureCitation>
).
</materialsCitation>
</paragraph>
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<collectingRegion id="4993F8224249FFFF82FA385D4F15FD85" box="[199,270,525,551]" pageId="14" pageNumber="265">Other</collectingRegion>
material examined.
</emphasis>
<materialsCitation id="3B3F3C9D4249FFFF8035385E4EB3FDEE" collectionCode="V" country="Japan" location="Tenjin-zaki" municipality="Tanabe-shi" pageId="14" pageNumber="265" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Honshu">
<emphasis id="B923EAD24249FFFF8035385E4D8CFD8A" bold="true" box="[520,919,525,552]" pageId="14" pageNumber="265">
<collectingCountry id="F34076504249FFFF8035385E4C4EFD8A" box="[520,597,526,552]" name="Japan" pageId="14" pageNumber="265">Japan</collectingCountry>
:
<collectingRegion id="4993F8224249FFFF805E385D4CDEFD8A" box="[611,709,525,552]" pageId="14" pageNumber="265">Honshû</collectingRegion>
:
<collectingRegion id="4993F8224249FFFF80EF385E4D88FD8A" box="[722,915,526,552]" pageId="14" pageNumber="265">Wakayama-ken</collectingRegion>
:
</emphasis>
<specimenCount id="9D51FD494249FFFF819D385E4DEBFD8A" box="[928,1008,526,552]" count="1" pageId="14" pageNumber="265" type="male">1 male</specimenCount>
,
<specimenCount id="9D51FD494249FFFF81C0385E4A79FD8A" box="[1021,1122,526,552]" count="1" pageId="14" pageNumber="265" type="female">1 female</specimenCount>
,
<collectingMunicipality id="6B8CACBA4249FFFF8652385E4AF7FD8A" box="[1135,1260,526,552]" pageId="14" pageNumber="265">Tanabe-shi</collectingMunicipality>
,
<location id="8E88601B4249FFFF86C5385E4B61FD8A" LSID="urn:lsid:plazi:treatment:03FE87D64249FFE082AA3AC74FA9FD27:8E88601B4249FFFF86C5385E4B61FD8A" box="[1272,1402,526,552]" country="Japan" municipality="Tanabe-shi" name="Tenjin-zaki" pageId="14" pageNumber="265" stateProvince="Honshu">Tenjin-zaki</location>
,
<date id="FFE910004249FFFF87BA385E4EB3FDEE" pageId="14" pageNumber="265" value="2019-05-03">
3.
<collectionCode id="ED46AE054249FFFF82AA38624EB3FDEE" box="[151,168,562,588]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="14" pageNumber="265" type="Museum">V</collectionCode>
</date>
</materialsCitation>
. 2019, M. Shimono;
<materialsCitation id="3B3F3C9D4249FFFF83A638624DC2FDEE" box="[411,985,562,588]" collectionCode="V" collectorName="Wakayama-shi" country="Japan" location="Kada-wan" pageId="14" pageNumber="265" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Wakayama-ken">
<specimenCount id="9D51FD494249FFFF83A638624FF1FDEE" box="[411,490,562,588]" count="1" pageId="14" pageNumber="265" type="male">1 male</specimenCount>
,
<specimenCount id="9D51FD494249FFFF83CB38624C40FDEE" box="[502,603,562,588]" count="1" pageId="14" pageNumber="265" type="female">1 female</specimenCount>
,
<collectorName id="26A253164249FFFF805A38624D0BFDEE" box="[615,784,562,588]" pageId="14" pageNumber="265">Wakayama-shi</collectorName>
,
<location id="8E88601B4249FFFF812138624D89FDEE" LSID="urn:lsid:plazi:treatment:03FE87D64249FFE082AA3AC74FA9FD27:8E88601B4249FFFF812138624D89FDEE" box="[796,914,562,588]" country="Japan" name="Kada-wan" pageId="14" pageNumber="265" stateProvince="Wakayama-ken">Kada-wan</location>
,
<date id="FFE910004249FFFF81A338624DC2FDEE" box="[926,985,562,588]" pageId="14" pageNumber="265" value="2007-05-18">
18.
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</date>
</materialsCitation>
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. 2007, Y.
<location id="8E88601B4249FFFF867338624ADCFDEE" LSID="urn:lsid:plazi:treatment:03FE87D64249FFE082AA3AC74FA9FD27:8E88601B4249FFFF867338624ADCFDEE" box="[1102,1223,562,588]" country="Japan" name="Kawakami" pageId="14" pageNumber="265" stateProvince="Wakayama-ken">Kawakami</location>
(
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)
</materialsCitation>
;
<materialsCitation id="3B3F3C9D4249FFFF870E38624FC5FDD2" collectionCode="III" collectorName="Shirahama-cho &amp; Seto" country="Japan" pageId="14" pageNumber="265" specimenCount="1" specimenCount-female="1" stateProvince="Wakayama-ken">
<specimenCount id="9D51FD494249FFFF870E38624B83FDEE" box="[1331,1432,562,588]" count="1" pageId="14" pageNumber="265" type="female">1 female</specimenCount>
,
<collectorName id="26A253164249FFFF82AA38064F58FDD2" box="[151,323,598,624]" pageId="14" pageNumber="265">Shirahama-chô</collectorName>
,
<collectorName id="26A253164249FFFF836C38064F9EFDD2" box="[337,389,598,624]" pageId="14" pageNumber="265">Seto</collectorName>
,
<date id="FFE910004249FFFF83AE38064FC5FDD2" box="[403,478,598,624]" pageId="14" pageNumber="265" value="2006-03-29">
29.
<collectionCode id="ED46AE054249FFFF838238064FC5FDD2" box="[447,478,598,624]" pageId="14" pageNumber="265">III</collectionCode>
</date>
</materialsCitation>
<materialsCitation id="3B3F3C9D4249FFFF83E338064D1EFDD2" box="[478,773,598,624]" collectionCode="KUM" country="Japan" location="Nakase" pageId="14" pageNumber="265" specimenCount="1" stateProvince="Wakayama-ken">
. 2006, Y.
<location id="8E88601B4249FFFF806F38064CBFFDD2" LSID="urn:lsid:plazi:treatment:03FE87D64249FFE082AA3AC74FA9FD27:8E88601B4249FFFF806F38064CBFFDD2" box="[594,676,598,624]" country="Japan" name="Nakase" pageId="14" pageNumber="265" stateProvince="Wakayama-ken">Nakase</location>
(
<collectionCode id="ED46AE054249FFFF808B38064CE6FDD2" box="[694,765,598,624]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="14" pageNumber="265" type="Herbarium">KUM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B3F3C9D4249FFFF812E38064B55FDD2" box="[787,1358,598,624]" collectionCode="V" collectorName="Wakayama-shi" country="Japan" location="Kabuto" pageId="14" pageNumber="265" specimenCount="11" specimenCount-female="7" specimenCount-male="4" stateProvince="Wakayama-ken">
<specimenCount id="9D51FD494249FFFF812E38064D75FDD2" box="[787,878,598,624]" count="4" pageId="14" pageNumber="265" type="male">4 males</specimenCount>
,
<specimenCount id="9D51FD494249FFFF814638064DF7FDD2" box="[891,1004,598,624]" count="7" pageId="14" pageNumber="265" type="female">7 females</specimenCount>
,
<collectorName id="26A253164249FFFF81C438064AB9FDD2" box="[1017,1186,598,624]" pageId="14" pageNumber="265">Wakayama-shi</collectorName>
,
<location id="8E88601B4249FFFF869338064B18FDD2" LSID="urn:lsid:plazi:treatment:03FE87D64249FFE082AA3AC74FA9FD27:8E88601B4249FFFF869338064B18FDD2" box="[1198,1283,598,624]" country="Japan" name="Kabuto" pageId="14" pageNumber="265" stateProvince="Wakayama-ken">Kabuto</location>
,
<date id="FFE910004249FFFF872C38064B55FDD2" box="[1297,1358,598,624]" pageId="14" pageNumber="265" value="2007-05-20">
20.
<collectionCode id="ED46AE054249FFFF870038064B55FDD2" box="[1341,1358,598,624]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="14" pageNumber="265" type="Museum">V</collectionCode>
</date>
</materialsCitation>
<materialsCitation id="3B3F3C9D4249FFFF877038064F7EFD36" collectionCode="OMNH" country="Japan" location="Fujie" pageId="14" pageNumber="265" specimenCount="1" stateProvince="Wakayama-ken">
. 2007, S.
<location id="8E88601B4249FFFF828A382A4EEAFD36" LSID="urn:lsid:plazi:treatment:03FE87D64249FFE082AA3AC74FA9FD27:8E88601B4249FFFF828A382A4EEAFD36" box="[183,241,634,660]" country="Japan" name="Fujie" pageId="14" pageNumber="265" stateProvince="Wakayama-ken">Fujie</location>
(
<collectionCode id="ED46AE054249FFFF833C382A4F46FD36" box="[257,349,634,660]" country="Japan" httpUri="http://grbio.org/cool/rcjv-ktyc" name="Osaka Museum of Natural History" pageId="14" pageNumber="265" type="Museum">OMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B3F3C9D4249FFFF834C38294D65FD36" box="[369,894,633,660]" collectionCode="IV" collectorName="Shiroyama" country="Japan" location="Kyushu" pageId="14" pageNumber="265" specimenCount="1" stateProvince="Kyushu">
<emphasis id="B923EAD24249FFFF834C38294CBDFD31" bold="true" box="[369,678,633,660]" pageId="14" pageNumber="265">
<collectingRegion id="4993F8224249FFFF834C38294FC8FD36" box="[369,467,633,660]" pageId="14" pageNumber="265">Kyûshû</collectingRegion>
:
<collectingRegion id="4993F8224249FFFF83DD38294CBAFD31" box="[480,673,633,659]" pageId="14" pageNumber="265">Kagoshima-ken</collectingRegion>
:
</emphasis>
<collectorName id="26A253164249FFFF8092382A4D30FD36" box="[687,811,634,660]" pageId="14" pageNumber="265">Shiroyama</collectorName>
,
<date id="FFE910004249FFFF8105382A4D65FD36" box="[824,894,634,660]" pageId="14" pageNumber="265" value="1983-04-13">
13.
<collectionCode id="ED46AE054249FFFF8159382A4D65FD36" box="[868,894,634,660]" pageId="14" pageNumber="265">IV</collectionCode>
</date>
</materialsCitation>
<materialsCitation id="3B3F3C9D4249FFFF8140382A4AB9FD36" box="[893,1186,634,662]" collectionCode="KUM" country="Japan" location="Ohara" pageId="14" pageNumber="265" specimenCount="1" stateProvince="Kagoshima-ken">
. 1983, M.
<location id="8E88601B4249FFFF81C3382B4A59FD34" LSID="urn:lsid:plazi:treatment:03FE87D64249FFE082AA3AC74FA9FD27:8E88601B4249FFFF81C3382B4A59FD34" box="[1022,1090,635,662]" country="Japan" name="Ohara" pageId="14" pageNumber="265" stateProvince="Kagoshima-ken">Ôhara</location>
(
<collectionCode id="ED46AE054249FFFF866E382A4A81FD36" box="[1107,1178,634,660]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="14" pageNumber="265" type="Herbarium">KUM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B3F3C9D4249FFFF8692382A4F9CFD1A" collectionCode="IV" collectorName="Ibusuki-shi &amp; Miyaga-hama" country="Japan" pageId="14" pageNumber="265" specimenCount="3" specimenCount-male="3" stateProvince="Kagoshima-ken">
<specimenCount id="9D51FD494249FFFF8692382A4B11FD36" box="[1199,1290,634,660]" count="3" pageId="14" pageNumber="265" type="male">3 males</specimenCount>
,
<collectorName id="26A253164249FFFF872A382A4B83FD36" box="[1303,1432,634,660]" pageId="14" pageNumber="265">Ibusuki-shi</collectorName>
,
<collectorName id="26A253164249FFFF82AA38CE4F2EFD1A" box="[151,309,670,696]" pageId="14" pageNumber="265">Miyaga-hama</collectorName>
,
<date id="FFE910004249FFFF837F38CE4F9CFD1A" box="[322,391,670,696]" pageId="14" pageNumber="265" value="2008-04-19">
19.
<collectionCode id="ED46AE054249FFFF835038CE4F9CFD1A" box="[365,391,670,696]" pageId="14" pageNumber="265">IV</collectionCode>
</date>
</materialsCitation>
<materialsCitation id="3B3F3C9D4249FFFF83BA38CE4D05FD1A" box="[391,798,670,696]" collectionCode="KUM" collectorName="Maruyama" country="Japan" pageId="14" pageNumber="265" specimenCount="1" stateProvince="Kagoshima-ken">
. 2008, M.
<collectorName id="26A253164249FFFF803938CE4C66FD1A" box="[516,637,670,696]" pageId="14" pageNumber="265">Maruyama</collectorName>
<emphasis id="B923EAD24249FFFF80BB38CF4CA3FD1A" box="[646,696,670,696]" italics="true" pageId="14" pageNumber="265">et al</emphasis>
. (
<collectionCode id="ED46AE054249FFFF80F338CE4D0EFD1A" box="[718,789,670,696]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="14" pageNumber="265" type="Herbarium">KUM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B3F3C9D4249FFFF811738CE4F8CFD7E" collectionCode="IV" collectorName="Minamisatsuma-shi &amp; Kasasa-machi" country="Japan" location="Koura" municipality="Ko-ura" pageId="14" pageNumber="265" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Kagoshima-ken">
<specimenCount id="9D51FD494249FFFF811738CE4D9FFD1A" box="[810,900,670,696]" count="3" pageId="14" pageNumber="265" type="male">3 males</specimenCount>
,
<specimenCount id="9D51FD494249FFFF81AD38CE4A1BFD1A" box="[912,1024,670,696]" count="2" pageId="14" pageNumber="265" type="female">2 females</specimenCount>
,
<collectorName id="26A253164249FFFF863138CE4AF7FD1A" box="[1036,1260,670,696]" pageId="14" pageNumber="265">Minamisatsuma-shi</collectorName>
,
<collectorName id="26A253164249FFFF86C538CE4B83FD1A" box="[1272,1432,670,696]" pageId="14" pageNumber="265">Kasasa-machi</collectorName>
,
<collectingMunicipality id="6B8CACBA4249FFFF82AA38924EFDFD7E" box="[151,230,706,732]" pageId="14" pageNumber="265">Ko-ura</collectingMunicipality>
(
<location id="8E88601B4249FFFF82C838924F5AFD7E" LSID="urn:lsid:plazi:treatment:03FE87D64249FFE082AA3AC74FA9FD27:8E88601B4249FFFF82C838924F5AFD7E" box="[245,321,706,732]" country="Japan" municipality="Ko-ura" name="Koura" pageId="14" pageNumber="265" stateProvince="Kagoshima-ken">Koura</location>
),
<date id="FFE910004249FFFF836E38924F8CFD7E" box="[339,407,706,732]" pageId="14" pageNumber="265" value="2008-04-20">
20.
<collectionCode id="ED46AE054249FFFF834038924F8CFD7E" box="[381,407,706,732]" pageId="14" pageNumber="265">IV</collectionCode>
</date>
</materialsCitation>
<materialsCitation id="3B3F3C9D4249FFFF83AA38924D3DFD7E" box="[407,806,706,732]" collectionCode="KUM" country="Japan" location="Maruyama" pageId="14" pageNumber="265" specimenCount="1" stateProvince="Kagoshima-ken">
. 2008, M.
<location id="8E88601B4249FFFF802D38924C92FD7E" LSID="urn:lsid:plazi:treatment:03FE87D64249FFE082AA3AC74FA9FD27:8E88601B4249FFFF802D38924C92FD7E" box="[528,649,706,732]" country="Japan" name="Maruyama" pageId="14" pageNumber="265" stateProvince="Kagoshima-ken">Maruyama</location>
(
<collectionCode id="ED46AE054249FFFF80A538924CC5FD7E" box="[664,734,706,732]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="14" pageNumber="265" type="Herbarium">KUM</collectionCode>
, cYT)
</materialsCitation>
;
<materialsCitation id="3B3F3C9D4249FFFF810C38924EC0FCA2" collectionCode="IV" collectorName="Minamisatsuma-shi &amp; Aiboshi" country="Japan" location="Aiboshi-gawa" pageId="14" pageNumber="265" specimenCount="2" specimenCount-female="2" stateProvince="Kagoshima-ken">
<specimenCount id="9D51FD494249FFFF810C38924DBBFD7E" box="[817,928,706,732]" count="2" pageId="14" pageNumber="265" type="female">2 females</specimenCount>
,
<collectorName id="26A253164249FFFF819638924A90FD7E" box="[939,1163,706,732]" pageId="14" pageNumber="265">Minamisatsuma-shi</collectorName>
,
<collectorName id="26A253164249FFFF86AE38924AF4FD7E" box="[1171,1263,706,732]" pageId="14" pageNumber="265">Aiboshi</collectorName>
,
<location id="8E88601B4249FFFF86C538924B83FD7E" LSID="urn:lsid:plazi:treatment:03FE87D64249FFE082AA3AC74FA9FD27:8E88601B4249FFFF86C538924B83FD7E" box="[1272,1432,706,732]" country="Japan" name="Aiboshi-gawa" pageId="14" pageNumber="265" stateProvince="Kagoshima-ken">Aiboshi-gawa</location>
,
<date id="FFE910004249FFFF82AA38B64EC0FCA2" box="[151,219,742,768]" pageId="14" pageNumber="265" value="2008-04-20">
20.
<collectionCode id="ED46AE054249FFFF82FC38B64EC0FCA2" box="[193,219,742,768]" pageId="14" pageNumber="265">IV</collectionCode>
</date>
</materialsCitation>
<materialsCitation id="3B3F3C9D4249FFFF82E638B64C7DFCA2" box="[219,614,742,768]" collectionCode="KUM" collectorName="Maruyama" country="Japan" pageId="14" pageNumber="265" specimenCount="1" stateProvince="Kagoshima-ken">
. 2008 M,
<collectorName id="26A253164249FFFF837238B64FD3FCA2" box="[335,456,742,768]" pageId="14" pageNumber="265">Maruyama</collectorName>
<emphasis id="B923EAD24249FFFF83ED38B74C13FCA2" box="[464,520,742,768]" italics="true" pageId="14" pageNumber="265">et al.</emphasis>
(
<collectionCode id="ED46AE054249FFFF802A38B64C45FCA2" box="[535,606,742,768]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="14" pageNumber="265" type="Herbarium">KUM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B3F3C9D4249FFFF804C38B54B83FCA2" box="[625,1432,741,768]" collectionCode="III" collectorName="Nishinoomote-shi" country="Japan" location="Asakawa" pageId="14" pageNumber="265" specimenCount="1" specimenCount-female="1" stateProvince="Ryukyu">
<emphasis id="B923EAD24249FFFF804C38B54D94FD5D" bold="true" box="[625,911,741,768]" pageId="14" pageNumber="265">
<collectingRegion id="4993F8224249FFFF804C38B54CC8FCA2" box="[625,723,741,768]" pageId="14" pageNumber="265">Ryûkyû</collectingRegion>
:
<collectingRegion id="4993F8224249FFFF80E238B54D90FD5D" box="[735,907,741,767]" pageId="14" pageNumber="265">Tanega-shima</collectingRegion>
:
</emphasis>
<specimenCount id="9D51FD494249FFFF81AB38B64DE1FCA2" box="[918,1018,742,768]" count="1" pageId="14" pageNumber="265" type="female">1 female</specimenCount>
,
<collectorName id="26A253164249FFFF863838B64ACAFCA2" box="[1029,1233,742,768]" pageId="14" pageNumber="265">Nishinoomote-shi</collectorName>
,
<location id="8E88601B4249FFFF86E638B64B5EFCA2" LSID="urn:lsid:plazi:treatment:03FE87D64249FFE082AA3AC74FA9FD27:8E88601B4249FFFF86E638B64B5EFCA2" box="[1243,1349,742,768]" country="Japan" name="Asakawa" pageId="14" pageNumber="265" stateProvince="Ryukyu">Asakawa</location>
,
<date id="FFE910004249FFFF877238B64B83FCA2" box="[1359,1432,742,768]" pageId="14" pageNumber="265" value="2010-03-27">
27.
<collectionCode id="ED46AE054249FFFF874438B64B83FCA2" box="[1401,1432,742,768]" pageId="14" pageNumber="265">III</collectionCode>
</date>
</materialsCitation>
. 2010, M. Asano (cYT).
</paragraph>
</subSubSection>
<caption id="DF2866484249FFFF82AA3DF44C1CF842" ID-DOI="http://doi.org/10.5281/zenodo.10361598" ID-Zenodo-Dep="10361598" httpUri="https://zenodo.org/record/10361598/files/figure.png" pageId="14" pageNumber="265" startId="14.[151,250,1956,1980]" targetBox="[164,1383,828,1922]" targetPageId="14" targetType="figure">
<paragraph id="8BE836C04249FFFF82AA3DF44C1CF842" blockId="14.[151,1436,1956,2017]" pageId="14" pageNumber="265">
<emphasis id="B923EAD24249FFFF82AA3DF44F14F81E" bold="true" box="[151,271,1956,1980]" pageId="14" pageNumber="265">FIGURE 9</emphasis>
Holotype of
<taxonomicName id="4C574D434249FFFF83A43DF44C28F81E" authorityName="K. Sawada" authorityYear="1955" box="[409,563,1956,1980]" class="Insecta" family="Staphylinidae" genus="Diaulota" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="uenoi">
<emphasis id="B923EAD24249FFFF83A43DF44C28F81E" box="[409,563,1956,1980]" italics="true" pageId="14" pageNumber="265">Diaulota uenoi</emphasis>
</taxonomicName>
. A, dissected body parts and label on slide glass; B, labrum; C, median lobe, lateral view. Scale bars: 0.1 mm for A &amp; B.
</paragraph>
</caption>
<subSubSection id="C34D654B4248FFFE82FA3AC64A73FE73" pageId="15" pageNumber="266" type="diagnosis">
<paragraph id="8BE836C04248FFFE82FA3AC64A73FE73" blockId="15.[151,1437,150,465]" pageId="15" pageNumber="266">
<emphasis id="B923EAD24248FFFE82FA3AC64F59FF12" bold="true" box="[199,322,150,176]" pageId="15" pageNumber="266">Diagnosis.</emphasis>
It is distinguished from the other species of the genus except
<taxonomicName id="4C574D434248FFFE81CA3AC74A89FF13" authorityName="Ahn" authorityYear="2023" box="[1015,1170,151,177]" class="Insecta" family="Staphylinidae" genus="Diaulota" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="266" phylum="Arthropoda" rank="species" species="submarina">
<emphasis id="B923EAD24248FFFE81CA3AC74A10FF12" box="[1015,1035,151,176]" italics="true" pageId="15" pageNumber="266">D</emphasis>
.
<emphasis id="B923EAD24248FFFE86273AC74A89FF13" box="[1050,1170,151,177]" italics="true" pageId="15" pageNumber="266">submarina</emphasis>
</taxonomicName>
and
<taxonomicName id="4C574D434248FFFE86F53AC74B4DFF13" authority="Tasaku &amp; Ono &amp; Maruyama, 2023" authorityName="Tasaku &amp; Ono &amp; Maruyama" authorityYear="2023" box="[1224,1366,151,177]" class="Insecta" family="Staphylinidae" genus="Diaulota" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="266" phylum="Arthropoda" rank="species" species="orientalis" status="sp. n.">
<emphasis id="B923EAD24248FFFE86F53AC74AC7FF12" box="[1224,1244,151,176]" italics="true" pageId="15" pageNumber="266">D</emphasis>
.
<emphasis id="B923EAD24248FFFE86D63AC74B4DFF13" box="[1259,1366,151,177]" italics="true" pageId="15" pageNumber="266">orientalis</emphasis>
</taxonomicName>
<emphasis id="B923EAD24248FFFE87603AC74B86FF13" bold="true" box="[1373,1437,151,177]" pageId="15" pageNumber="266">
<taxonomicNameLabel id="A21057A94248FFFE87603AC74B86FF13" box="[1373,1437,151,177]" pageId="15" pageNumber="266" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by the following combination of characteristics: ground color reddish brown, abdominal segments VI and VII black (
<figureCitation id="136C2A454248FFFE82A23A8F4F2DFF5B" box="[159,310,223,249]" captionStart="FIGURE 8" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[184,1403,191,1909]" captionTargetId="figure-19@13.[174,1413,181,1919]" captionTargetPageId="13" captionText="FIGURE 8 Diaulota uenoi. A, habitus, male; B, ditto, female; C, labium; D, mentum; E, median lode of aedeagus, lateral view; F, internal structure of median lobe, dorsal view; G, ditto, ventral view; H, spermatheca. Scale bars: 1.0 mm for A, B; 0.1 mm for CH. Abbreviation: as, apical sclerite; su, suspensonia." figureDoi="http://doi.org/10.5281/zenodo.10361594" httpUri="https://zenodo.org/record/10361594/files/figure.png" pageId="15" pageNumber="266">Figs. 8A &amp; B</figureCitation>
); male head larger than that of female; mentum transverse (
<figureCitation id="136C2A454248FFFE81F13A8F4A3DFF5B" box="[972,1062,223,249]" captionStart="FIGURE 8" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[184,1403,191,1909]" captionTargetId="figure-19@13.[174,1413,181,1919]" captionTargetPageId="13" captionText="FIGURE 8 Diaulota uenoi. A, habitus, male; B, ditto, female; C, labium; D, mentum; E, median lode of aedeagus, lateral view; F, internal structure of median lobe, dorsal view; G, ditto, ventral view; H, spermatheca. Scale bars: 1.0 mm for A, B; 0.1 mm for CH. Abbreviation: as, apical sclerite; su, suspensonia." figureDoi="http://doi.org/10.5281/zenodo.10361594" httpUri="https://zenodo.org/record/10361594/files/figure.png" pageId="15" pageNumber="266">Fig. 8D</figureCitation>
); infraorbital carina short, shorter than half of head; tarsal formula 4-4-4. This species very similar to
<taxonomicName id="4C574D434248FFFE819E3B534A24FEBF" authorityName="Ahn" authorityYear="2023" box="[931,1087,259,285]" class="Insecta" family="Staphylinidae" genus="Diaulota" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="266" phylum="Arthropoda" rank="species" species="submarina">
<emphasis id="B923EAD24248FFFE819E3B534DACFEBE" box="[931,951,259,284]" italics="true" pageId="15" pageNumber="266">D</emphasis>
.
<emphasis id="B923EAD24248FFFE81FA3B534A24FEBF" box="[967,1087,259,285]" italics="true" pageId="15" pageNumber="266">submarina</emphasis>
</taxonomicName>
and
<taxonomicName id="4C574D434248FFFE86443B534B13FEBF" authority="Tasaku &amp; Ono &amp; Maruyama, 2023" authorityName="Tasaku &amp; Ono &amp; Maruyama" authorityYear="2023" box="[1145,1288,259,285]" class="Insecta" family="Staphylinidae" genus="Diaulota" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="266" phylum="Arthropoda" rank="species" species="orientalis" status="sp. n.">
<emphasis id="B923EAD24248FFFE86443B534A96FEBE" box="[1145,1165,259,284]" italics="true" pageId="15" pageNumber="266">D</emphasis>
.
<emphasis id="B923EAD24248FFFE86A03B534B13FEBF" box="[1181,1288,259,285]" italics="true" pageId="15" pageNumber="266">orientalis</emphasis>
</taxonomicName>
<emphasis id="B923EAD24248FFFE872C3B534B48FEBF" bold="true" box="[1297,1363,259,285]" pageId="15" pageNumber="266">
<taxonomicNameLabel id="A21057A94248FFFE872C3B534B48FEBF" box="[1297,1363,259,285]" pageId="15" pageNumber="266" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
and it is very difficult to distinguish by external morphological characters (we call them as the
<taxonomicName id="4C574D434248FFFE86B23B774B22FEE2" authorityName="K. Sawada" authorityYear="1955" box="[1167,1337,295,321]" class="Insecta" family="Staphylinidae" genus="Diaulota" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="266" phylum="Arthropoda" rank="species" species="uenoi">
<emphasis id="B923EAD24248FFFE86B23B774B22FEE2" box="[1167,1337,295,321]" italics="true" pageId="15" pageNumber="266">Diaulota uenoi</emphasis>
</taxonomicName>
species-complex). Compared to
<taxonomicName id="4C574D434248FFFE839B3B1B4C5BFEC7" authorityName="Ahn" authorityYear="2023" box="[422,576,331,357]" class="Insecta" family="Staphylinidae" genus="Diaulota" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="266" phylum="Arthropoda" rank="species" species="submarina">
<emphasis id="B923EAD24248FFFE839B3B1B4FA1FEC6" box="[422,442,331,356]" italics="true" pageId="15" pageNumber="266">D</emphasis>
.
<emphasis id="B923EAD24248FFFE83F53B1B4C5BFEC7" box="[456,576,331,357]" italics="true" pageId="15" pageNumber="266">submarina</emphasis>
</taxonomicName>
, it can be distinguished by the characteristics: apex of apical sclerite rounded in lateral view (
<figureCitation id="136C2A454248FFFE83793B3F4F81FE2B" box="[324,410,367,393]" captionStart="FIGURE 8" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[184,1403,191,1909]" captionTargetId="figure-19@13.[174,1413,181,1919]" captionTargetPageId="13" captionText="FIGURE 8 Diaulota uenoi. A, habitus, male; B, ditto, female; C, labium; D, mentum; E, median lode of aedeagus, lateral view; F, internal structure of median lobe, dorsal view; G, ditto, ventral view; H, spermatheca. Scale bars: 1.0 mm for A, B; 0.1 mm for CH. Abbreviation: as, apical sclerite; su, suspensonia." figureDoi="http://doi.org/10.5281/zenodo.10361594" httpUri="https://zenodo.org/record/10361594/files/figure.png" pageId="15" pageNumber="266">Fig. 8E</figureCitation>
); suspensonia broad (
<figureCitation id="136C2A454248FFFE80B53B3F4D05FE2B" box="[648,798,367,393]" captionStart="FIGURE 8" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[184,1403,191,1909]" captionTargetId="figure-19@13.[174,1413,181,1919]" captionTargetPageId="13" captionText="FIGURE 8 Diaulota uenoi. A, habitus, male; B, ditto, female; C, labium; D, mentum; E, median lode of aedeagus, lateral view; F, internal structure of median lobe, dorsal view; G, ditto, ventral view; H, spermatheca. Scale bars: 1.0 mm for A, B; 0.1 mm for CH. Abbreviation: as, apical sclerite; su, suspensonia." figureDoi="http://doi.org/10.5281/zenodo.10361594" httpUri="https://zenodo.org/record/10361594/files/figure.png" pageId="15" pageNumber="266">Figs. 8F &amp; G</figureCitation>
). Compared to
<taxonomicName id="4C574D434248FFFE81FA3B3F4A48FE2B" authority="Tasaku &amp; Ono &amp; Maruyama, 2023" authorityName="Tasaku &amp; Ono &amp; Maruyama" authorityYear="2023" box="[967,1107,367,393]" class="Insecta" family="Staphylinidae" genus="Diaulota" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="266" phylum="Arthropoda" rank="species" species="orientalis" status="sp. n.">
<emphasis id="B923EAD24248FFFE81FA3B3F4DC0FE2A" box="[967,987,367,392]" italics="true" pageId="15" pageNumber="266">D</emphasis>
.
<emphasis id="B923EAD24248FFFE81D53B3F4A48FE2B" box="[1000,1107,367,393]" italics="true" pageId="15" pageNumber="266">orientalis</emphasis>
</taxonomicName>
<emphasis id="B923EAD24248FFFE86643B3F4A83FE2B" bold="true" box="[1113,1176,367,393]" pageId="15" pageNumber="266">
<taxonomicNameLabel id="A21057A94248FFFE86643B3F4A83FE2B" box="[1113,1176,367,393]" pageId="15" pageNumber="266" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, it can be distinguished by the following characteristics: apex of median lobe pointed in ventral view; apical sclerite extend toward apical margin in ventral view (
<figureCitation id="136C2A454248FFFE83993BE74C26FE73" box="[420,573,439,465]" captionStart="FIGURE 8" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[184,1403,191,1909]" captionTargetId="figure-19@13.[174,1413,181,1919]" captionTargetPageId="13" captionText="FIGURE 8 Diaulota uenoi. A, habitus, male; B, ditto, female; C, labium; D, mentum; E, median lode of aedeagus, lateral view; F, internal structure of median lobe, dorsal view; G, ditto, ventral view; H, spermatheca. Scale bars: 1.0 mm for A, B; 0.1 mm for CH. Abbreviation: as, apical sclerite; su, suspensonia." figureDoi="http://doi.org/10.5281/zenodo.10361594" httpUri="https://zenodo.org/record/10361594/files/figure.png" pageId="15" pageNumber="266">Figs. 8F &amp; G</figureCitation>
); spermatheca once coiled around base (
<figureCitation id="136C2A454248FFFE863C3BE74A40FE73" box="[1025,1115,439,465]" captionStart="FIGURE 8" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[184,1403,191,1909]" captionTargetId="figure-19@13.[174,1413,181,1919]" captionTargetPageId="13" captionText="FIGURE 8 Diaulota uenoi. A, habitus, male; B, ditto, female; C, labium; D, mentum; E, median lode of aedeagus, lateral view; F, internal structure of median lobe, dorsal view; G, ditto, ventral view; H, spermatheca. Scale bars: 1.0 mm for A, B; 0.1 mm for CH. Abbreviation: as, apical sclerite; su, suspensonia." figureDoi="http://doi.org/10.5281/zenodo.10361594" httpUri="https://zenodo.org/record/10361594/files/figure.png" pageId="15" pageNumber="266">Fig. 8H</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF2866484248FFFE82AA3CA24C4CF88C" ID-DOI="http://doi.org/10.5281/zenodo.10361603" ID-Zenodo-Dep="10361603" httpUri="https://zenodo.org/record/10361603/files/figure.png" pageId="15" pageNumber="266" startId="15.[151,250,1778,1802]" targetBox="[162,1425,500,1744]" targetPageId="15" targetType="figure">
<paragraph id="8BE836C04248FFFE82AA3CA24C4CF88C" blockId="15.[151,1436,1778,1839]" pageId="15" pageNumber="266">
<emphasis id="B923EAD24248FFFE82AA3CA24F01F8A8" bold="true" box="[151,282,1778,1802]" pageId="15" pageNumber="266">FIGURE 10</emphasis>
Habitus and mouthparts of
<taxonomicName id="4C574D434248FFFE80053CA24D1EF8A8" authorityName="Ahn" authorityYear="2023" box="[568,773,1778,1802]" class="Insecta" family="Staphylinidae" genus="Diaulota" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="266" phylum="Arthropoda" rank="species" species="submarina">
<emphasis id="B923EAD24248FFFE80053CA24D1EF8A8" box="[568,773,1778,1802]" italics="true" pageId="15" pageNumber="266">Diaulota submarina</emphasis>
</taxonomicName>
. A, habitus, male; B, ditto, female; C, labium, D, mentum. Scale bars: 1.0 mm for A &amp; B; 0.1 mm for C &amp; D.
</paragraph>
</caption>
<subSubSection id="C34D654B4248FFE082FA3D0D4C84FD9F" lastPageId="17" lastPageNumber="268" pageId="15" pageNumber="266" type="discussion">
<paragraph id="8BE836C04248FFE082FA3D0D4C61FF77" blockId="15.[151,1437,1884,2019]" lastBlockId="17.[151,1443,151,645]" lastPageId="17" lastPageNumber="268" pageId="15" pageNumber="266">
<emphasis id="B923EAD24248FFFE82FA3D0D4F20F8D5" bold="true" box="[199,315,1885,1911]" pageId="15" pageNumber="266">Remarks.</emphasis>
This species, initially described by
<bibRefCitation id="EFC64B314248FFFE80F33D0C4D6CF8D5" author="Sawada, K." box="[718,887,1884,1911]" pageId="15" pageNumber="266" pagination="81 - 87" refId="ref23139" refString="Sawada, K. (1955) Marine Insects of the Tokara Islands VII. Fam. Staphylinidae. Publications of the Seto Marine Biological Laboratory, 5 (1), 81 - 87. https: // doi. org / 10.5134 / 174533" type="journal article" year="1955">Sawada (1955)</bibRefCitation>
as
<taxonomicName id="4C574D434248FFFE81A13D0C4A32F8D4" authorityName="Sawada" authorityYear="1955" box="[924,1065,1884,1910]" class="Insecta" family="Staphylinidae" genus="Genoplectes" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="266" phylum="Arthropoda" rank="genus">Genoplectes</taxonomicName>
, was originally based on a single male specimen collected from Nakano-shima, Tokara-rettô, in the northern Ryûkyû. Subsequently,
<bibRefCitation id="EFC64B314248FFFE86CE3DD04B87F839" author="Sawada, K." box="[1267,1436,1920,1947]" pageId="15" pageNumber="266" pagination="81 - 110" refId="ref23228" refString="Sawada, K. (1971) Aleocharinae (Staphylinidae, Coleoptera) from the Intertidal Zone of Japan, Publications of the Seto Marine Biological Laboratory, 19 (2 - 3), 81 - 110. https: // doi. org / 10.5134 / 175665" type="journal article" year="1971">Sawada (1971)</bibRefCitation>
undertook a redescription and recorded this species from Wakayama-ken, the southern Honshû. Following these contributions, Ahn (1996) synonymized
<taxonomicName id="4C574D434248FFFE805B3D984CEBF840" authorityName="Sawada" authorityYear="1955" box="[614,752,1992,2018]" class="Insecta" family="Staphylinidae" genus="Genoplectes" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="266" phylum="Arthropoda" rank="genus">Genoplectes</taxonomicName>
under
<taxonomicName id="4C574D434248FFFE817F3D984DBDF840" authorityName="Casey" authorityYear="1893" box="[834,934,1992,2018]" class="Insecta" family="Staphylinidae" genus="Diaulota" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="266" phylum="Arthropoda" rank="genus">
<emphasis id="B923EAD24248FFFE817F3D984DBDF840" box="[834,934,1992,2018]" italics="true" pageId="15" pageNumber="266">Diaulota</emphasis>
</taxonomicName>
, extending its known distribution to
<collectingCountry id="F34076504248FFFE876C3D994B82F841" box="[1361,1433,1993,2019]" name="South Korea" pageId="15" pageNumber="266">Korea</collectingCountry>
. Notably, the Korean population was recently described as
<taxonomicName id="4C574D434256FFE081113AC74DDCFF13" authorityName="Ahn" authorityYear="2023" box="[812,967,151,177]" class="Insecta" family="Staphylinidae" genus="Diaulota" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="268" phylum="Arthropoda" rank="species" species="submarina">
<emphasis id="B923EAD24256FFE081113AC74DDCFF13" box="[812,967,151,177]" italics="true" pageId="17" pageNumber="268">D. submarina</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFC64B314256FFE081CC3AC74A6CFF13" author="Ahn, K. - J." box="[1009,1143,151,177]" pageId="17" pageNumber="268" pagination="141 - 147" refId="ref21503" refString="Ahn, K. - J. (2023) Description of Diaulota submarina sp. nov. (Coleoptera: Staphylinidae: Aleocharinae) on Korean coasts. Zootaxa, 5336 (1), 141 - 147. https: // doi. org / 10.11646 / zootaxa. 5336.1.8" type="journal article" year="2023">Ahn (2023)</bibRefCitation>
. Consequently, its current distribution is exclusively limited to
<collectingCountry id="F34076504256FFE0800E3AEB4C6DFF77" box="[563,630,187,213]" name="Japan" pageId="17" pageNumber="268">Japan</collectingCountry>
.
</paragraph>
<caption id="DF2866484257FFE182AA3D2D4F7AF87E" ID-DOI="http://doi.org/10.5281/zenodo.10361606" ID-Zenodo-Dep="10361606" httpUri="https://zenodo.org/record/10361606/files/figure.png" pageId="16" pageNumber="267" startId="16.[151,250,1917,1941]" targetBox="[160,1427,192,1882]" targetPageId="16" targetType="figure">
<paragraph id="8BE836C04257FFE182AA3D2D4F7AF87E" blockId="16.[151,1437,1917,2013]" pageId="16" pageNumber="267">
<emphasis id="B923EAD24257FFE182AA3D2D4F07F837" bold="true" box="[151,284,1917,1941]" pageId="16" pageNumber="267">FIGURE 11</emphasis>
Genitalia of
<taxonomicName id="4C574D434257FFE183963D2D4C60F837" authorityName="Ahn" authorityYear="2023" box="[427,635,1917,1941]" class="Insecta" family="Staphylinidae" genus="Diaulota" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="267" phylum="Arthropoda" rank="species" species="submarina">
<emphasis id="B923EAD24257FFE183963D2D4C60F837" box="[427,635,1917,1941]" italics="true" pageId="16" pageNumber="267">Diaulota submarina</emphasis>
</taxonomicName>
(AD, from Tsushima; EH, from Hiroshima-ken). A &amp; E, median lobe of aedeagus, lateral view; B &amp; F, ditto, ventral view; C &amp; G, internal structure of median lobe, ventral view; D &amp; H, spermatheca. Scale bars: 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BE836C04256FFE082FA3A8F4A51FEE3" blockId="17.[151,1443,151,645]" pageId="17" pageNumber="268">
<materialsCitation id="3B3F3C9D4256FFE082FA3A8F4A5DFEE3" country="Japan" location="Japan" pageId="17" pageNumber="268" specimenCount="1" typeStatus="holotype">
In this study, we conducted an extensive examination of numerous specimens collected from
<collectingCountry id="F34076504256FFE086D93A8F4B3CFF5B" box="[1252,1319,223,249]" name="Japan" pageId="17" pageNumber="268">Japan</collectingCountry>
, including the
<typeStatus id="54EC88624256FFE082FF3B534F33FEBF" box="[194,296,259,285]" pageId="17" pageNumber="268" type="holotype">holotype</typeStatus>
. Our findings indicate that
<taxonomicName id="4C574D434256FFE0805F3B534CDFFEBE" authorityName="K. Sawada" authorityYear="1955" box="[610,708,259,284]" class="Insecta" family="Staphylinidae" genus="Diaulota" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="268" phylum="Arthropoda" rank="species" species="uenoi">
<emphasis id="B923EAD24256FFE0805F3B534CDFFEBE" box="[610,708,259,284]" italics="true" pageId="17" pageNumber="268">D. uenoi</emphasis>
</taxonomicName>
is presently confined to the Pacific side of southern
<collectingCountry id="F34076504256FFE087193B534B7CFEBF" box="[1316,1383,259,285]" name="Japan" pageId="17" pageNumber="268">Japan</collectingCountry>
, and populations from other regions do not correspond to this species (as detailed below)
</materialsCitation>
.
</paragraph>
<paragraph id="8BE836C04256FFE082FA3B1B4FB1FE73" blockId="17.[151,1443,151,645]" pageId="17" pageNumber="268">
When comparing the median lobe of the specimen collected from Nakano-shima (
<figureCitation id="136C2A454256FFE086463B1B4ACFFEC7" box="[1147,1236,331,357]" captionStart="FIGURE 9" captionStartId="14.[151,250,1956,1980]" captionTargetBox="[164,1383,828,1922]" captionTargetId="figure-227@14.[164,1423,828,1933]" captionTargetPageId="14" captionText="FIGURE 9 Holotype of Diaulota uenoi. A, dissected body parts and label on slide glass; B, labrum; C, median lobe, lateral view. Scale bars: 0.1 mm for A &amp; B." figureDoi="http://doi.org/10.5281/zenodo.10361598" httpUri="https://zenodo.org/record/10361598/files/figure.png" pageId="17" pageNumber="268">Fig. 9C</figureCitation>
) with those from other regions (
<figureCitation id="136C2A454256FFE0837F3B3F4F80FE2B" box="[322,411,367,393]" captionStart="FIGURE 8" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[184,1403,191,1909]" captionTargetId="figure-19@13.[174,1413,181,1919]" captionTargetPageId="13" captionText="FIGURE 8 Diaulota uenoi. A, habitus, male; B, ditto, female; C, labium; D, mentum; E, median lode of aedeagus, lateral view; F, internal structure of median lobe, dorsal view; G, ditto, ventral view; H, spermatheca. Scale bars: 1.0 mm for A, B; 0.1 mm for CH. Abbreviation: as, apical sclerite; su, suspensonia." figureDoi="http://doi.org/10.5281/zenodo.10361594" httpUri="https://zenodo.org/record/10361594/files/figure.png" pageId="17" pageNumber="268">Fig. 8E</figureCitation>
), it becomes apparent that the apical lobe of the Nakano-shima specimen is marginally shorter. Nevertheless, the remaining morphological characteristics closely align. Consequently, we have classified them as a single species.
</paragraph>
<paragraph id="8BE836C04256FFE082FA3B8A4C84FD9F" blockId="17.[151,1443,151,645]" pageId="17" pageNumber="268">
<emphasis id="B923EAD24256FFE082FA3B8A4F31FE56" bold="true" box="[199,298,474,500]" pageId="17" pageNumber="268">Habitat.</emphasis>
This species has been mainly collected from rocky shores. Members of the
<taxonomicName id="4C574D434256FFE086B23B8B4B22FE56" authorityName="K. Sawada" authorityYear="1955" box="[1167,1337,475,501]" class="Insecta" family="Staphylinidae" genus="Diaulota" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="268" phylum="Arthropoda" rank="species" species="uenoi">
<emphasis id="B923EAD24256FFE086B23B8B4B22FE56" box="[1167,1337,475,501]" italics="true" pageId="17" pageNumber="268">Diaulota uenoi</emphasis>
</taxonomicName>
species-complex exhibit a preference for environments abundant in seaweed; nonetheless, they are occasionally encountered in rocky shores with sparse seaweed presence.
</paragraph>
</subSubSection>
<subSubSection id="C34D654B4256FFE082FA38164FA9FD27" pageId="17" pageNumber="268" type="distribution">
<paragraph id="8BE836C04256FFE082FA38164FA9FD27" blockId="17.[151,1443,151,645]" pageId="17" pageNumber="268">
<emphasis id="B923EAD24256FFE082FA38164F7AFDC2" bold="true" box="[199,353,582,608]" pageId="17" pageNumber="268">Distribution.</emphasis>
<collectingCountry id="F34076504256FFE0835538174FB7FDC3" box="[360,428,583,609]" name="Japan" pageId="17" pageNumber="268">Japan</collectingCountry>
: Honshû, Shikoku, Kyûshû (Pacific Ocean side), Ryûkyû (Tanega-shima, Nakano-shima, Tokuno-shima) (
<figureCitation id="136C2A454256FFE0836C383B4FBDFD27" box="[337,422,619,645]" captionStart="FIGURE 24" captionStartId="36.[151,250,1780,1804]" captionTargetBox="[151,1436,345,1757]" captionTargetId="figure-99@36.[151,1436,345,1757]" captionTargetPageId="36" captionText="FIGURE 24 Distribution of Diaulota uenoi (red circle (red ring, type locality)), D. submarina (blue square), and D. orientalis sp. n. (yellow triangle)." figureDoi="http://doi.org/10.5281/zenodo.10361669" httpUri="https://zenodo.org/record/10361669/files/figure.png" pageId="17" pageNumber="268">Fig. 24</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>