303 lines
43 KiB
XML
303 lines
43 KiB
XML
<document id="A8D294BB63DC4EBD27878CF02D970F37" ID-DOI="10.5281/zenodo.273746" ID-GBIF-Dataset="f2a7ea5a-521a-4012-9e5e-f46e028bfb40" ID-ISSN="1175-5326" ID-Zenodo-Dep="273746" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459836960861" checkinUser="plazi" docAuthor="Lin, Qi-Bin & Huang, Di-Ying" docDate="2007" docId="03FE87BDF16AFF9865D8DB2B50F2AE38" docLanguage="en" docName="zt01469p064.pdf" docOrigin="Zootaxa 1469" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Nodalula dalinghensis Lin & Huang, 2007, sp. nov." docType="treatment" docVersion="7" lastPageNumber="63" masterDocId="FFC7FFC5F16BFF9D654FDF015017AF3A" masterDocTitle="A new family of Cavilabiata from the Lower Cretaceous Yixian Formation, China (Odonata: Anisoptera)" masterLastPageNumber="64" masterPageNumber="59" pageNumber="60" updateTime="1698223851924" updateUser="plazi">
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<mods:titleInfo id="98CD37AAB6F5CA6188732A93C5388737">
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<mods:title id="154487CB09028FA8F9F53A00A21094DC">A new family of Cavilabiata from the Lower Cretaceous Yixian Formation, China (Odonata: Anisoptera)</mods:title>
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<mods:name id="339A9F7353C09D6C249413B6CC44D0AF" type="personal">
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<mods:roleTerm id="6812098B678344FAA984651917A7BCCB">Author</mods:roleTerm>
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<mods:namePart id="144799A75882D1C4A267C61661F7E8EA">Lin, Qi-Bin</mods:namePart>
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<mods:roleTerm id="F5AFC35AFF4D5B966D86BC30583DED26">Author</mods:roleTerm>
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<mods:namePart id="6E5FEBA4D14BD337176CE860DF142305">Huang, Di-Ying</mods:namePart>
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<mods:typeOfResource id="65A48D1FBED778E1129D261A401D6F34">text</mods:typeOfResource>
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<mods:title id="30E952EA9BCB6473A813E059B7FE8EE3">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="CE3CB51A5E36803BB2FD7B344486F85C">
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<mods:date id="AB2A9C282764589C660D1F04FCE8560C">2007</mods:date>
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<mods:detail id="5ACED7EBD42CCD63C8E79C2D524E8B5B" type="volume">
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<mods:number id="9E7652A250662477AD729E5E47EF60E5">1469</mods:number>
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<mods:start id="BB185816DC7116E6956DCA6B64F4634F">59</mods:start>
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<mods:classification id="8769318352A00DB092DA60614A91EF16">journal article</mods:classification>
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<mods:identifier id="3399D1277BAA7C97863C113E532D0DE6" type="DOI">10.5281/zenodo.273746</mods:identifier>
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<mods:identifier id="5463B532B841EC0B89E8371E2DD67FE5" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="F78C406609471BDB83138F9B2CC89332" type="Zenodo-Dep">273746</mods:identifier>
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<treatment id="03FE87BDF16AFF9865D8DB2B50F2AE38" ID-DOI="http://doi.org/10.5281/zenodo.6250451" ID-GBIF-Taxon="119359232" ID-Zenodo-Dep="6250451" LSID="urn:lsid:plazi:treatment:03FE87BDF16AFF9865D8DB2B50F2AE38" httpUri="http://treatment.plazi.org/id/03FE87BDF16AFF9865D8DB2B50F2AE38" lastPageId="5" lastPageNumber="63" pageId="1" pageNumber="60">
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<subSubSection id="C34D6520F16AFF9C65D8DB2B5113AB5D" pageId="1" pageNumber="60" type="nomenclature">
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<paragraph id="8BE836ABF16AFF9C65D8DB2B521BAB7E" blockId="1.[151,524,1066,1127]" box="[151,524,1066,1092]" pageId="1" pageNumber="60">
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<heading id="D0A081C7F16AFF9C65D8DB2B521BAB7E" bold="true" box="[151,524,1066,1092]" fontSize="11" level="1" pageId="1" pageNumber="60" reason="1">
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<emphasis id="B923EAB9F16AFF9C65D8DB2B521BAB7E" bold="true" box="[151,524,1066,1092]" pageId="1" pageNumber="60">
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<taxonomicName id="4C574D28F16AFF9C65D8DB2B51B0AB7E" box="[151,423,1066,1092]" class="Insecta" family="Nodalulaidae" genus="Nodalula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="60" phylum="Arthropoda" rank="species" species="dalinghensis" status="sp. nov.">
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<emphasis id="B923EAB9F16AFF9C65D8DB2B51B0AB7E" bold="true" box="[151,423,1066,1092]" italics="true" pageId="1" pageNumber="60">Nodalula dalinghensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21057C2F16AFF9C64E0DB2B521BAB7E" box="[431,524,1066,1092]" pageId="1" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BE836ABF16AFF9C65D8DB4C5113AB5D" blockId="1.[151,524,1066,1127]" box="[151,260,1101,1127]" pageId="1" pageNumber="60">
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<figureCitation id="136C2A2EF16AFF9C65D8DB4C5113AB5D" box="[151,260,1101,1127]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,255,1321,1345]" captionStartId-1="3.[151,255,1610,1634]" captionStartId-2="5.[151,255,594,618]" captionTargetBox-0="[220,1368,197,1301]" captionTargetBox-1="[252,1331,723,1588]" captionTargetBox-2="[184,1402,317,567]" captionTargetId-0="figure@2.[220,1369,197,1303]" captionTargetId-1="figure@3.[252,1332,723,1590]" captionTargetId-2="figure@5.[184,1403,317,568]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="5" captionText-0="FIGURE 1. Nodalula dalinghensis gen. et sp. nov., holotype Cat. No. 123517 (female), photograph of dorsal view (scale bar represents 10 mm)." captionText-1="FIGURE 2. Nodalula dalinghensis gen. et sp. nov., holotype Cat. No. 123517, Enlarged view showing the left fore and hind wings venation (scale bar represents 10 mm)." captionText-2="FIGURE 3. Nodalula dalinghensis gen. et sp. nov., holotype Cat. No. 123517, Enlarged view showing the apex of abdomen (scale bar represents 10 mm)." httpUri-0="https://zenodo.org/record/273747/files/figure.png" httpUri-1="https://zenodo.org/record/273748/files/figure.png" httpUri-2="https://zenodo.org/record/273749/files/figure.png" pageId="1" pageNumber="60">Figs. 1–3</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34D6520F16AFF9C65D8DB9C5177AA3D" pageId="1" pageNumber="60" type="materials_examined">
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<paragraph id="8BE836ABF16AFF9C65D8DB9C5177AA3D" blockId="1.[151,1437,1181,2007]" pageId="1" pageNumber="60">
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<emphasis id="B923EAB9F16AFF9C65D8DB9C511AAB8D" bold="true" box="[151,269,1181,1207]" pageId="1" pageNumber="60">Material:</emphasis>
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A nearly complete adult dragonfly with the four wings and the trunk (except the head) preserved with brown colour, without counterpart. The wings are strongly distorted, especially in their anterior parts. Cat. No. 123517.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34D6520F16AFF9C6588DA1452BDAA45" pageId="1" pageNumber="60" type="etymology">
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<paragraph id="8BE836ABF16AFF9C6588DA1453C2AA15" blockId="1.[151,1437,1181,2007]" box="[199,981,1301,1327]" pageId="1" pageNumber="60">
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<emphasis id="B923EAB9F16AFF9C6588DA145140AA15" bold="true" box="[199,343,1301,1327]" pageId="1" pageNumber="60">Etymology:</emphasis>
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Derived from Dalinghe, a name of a river in this area.
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</paragraph>
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<paragraph id="8BE836ABF16AFF9C6588DA3C52BDAA45" blockId="1.[151,1437,1181,2007]" pageId="1" pageNumber="60">
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<emphasis id="B923EAB9F16AFF9C6588DA3C51C3AA6D" bold="true" box="[199,468,1341,1367]" pageId="1" pageNumber="60">Locality and horizon:</emphasis>
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Yixian Formation, Barremian, Early Cretaceous, Dalinghou, in the Jianshangou of Beipiao City, Liaoning Province (
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<bibRefCitation id="EFC64B5AF16AFF9C676EDA64528AAA45" author="Sha" box="[545,669,1381,1407]" pageId="1" pageNumber="64" refString="Sha, J. - G. (2007) Cretaceous stratigraphy of northeast China: non-marine and marine correlation. Cretaceous Research, in press." type="book" year="2007">Sha, 2007</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C34D6520F16AFF9E6588DA8C5231ADAD" lastPageId="3" lastPageNumber="62" pageId="1" pageNumber="60" type="description">
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<paragraph id="8BE836ABF16AFF9C6588DA8C5463AAF5" blockId="1.[151,1437,1181,2007]" pageId="1" pageNumber="60">
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<emphasis id="B923EAB9F16AFF9C6588DA8C5176AA9D" bold="true" box="[199,353,1421,1447]" pageId="1" pageNumber="60">Description:</emphasis>
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A nearly complete body of an adult female with the thorax, abdomen, the six legs and four wings in connection, but with most part of head missing; whole body
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<quantity id="4CAF9B4EF16AFF9C6687DAB45426AAF5" box="[968,1073,1461,1487]" metricMagnitude="-2" metricUnit="m" metricValue="6.22" pageId="1" pageNumber="60" unit="mm" value="62.2">62.2 mm</quantity>
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long.
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</paragraph>
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<paragraph id="8BE836ABF16AFF9C6588DADC54D7A925" blockId="1.[151,1437,1181,2007]" pageId="1" pageNumber="60">
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Thorax
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<quantity id="4CAF9B4EF16AFF9C646CDADC519AAACD" box="[291,397,1501,1527]" metricMagnitude="-2" metricUnit="m" metricValue="1.3599999999999999" pageId="1" pageNumber="60" unit="mm" value="13.6">13.6 mm</quantity>
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long, 8.0 mm wide; prothorax small,
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<quantity id="4CAF9B4EF16AFF9C6603DADC53B0AACD" box="[844,935,1501,1527]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="1" pageNumber="60" unit="mm" value="1.6">1.6 mm</quantity>
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long,
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<quantity id="4CAF9B4EF16AFF9C66BDDADC5421AACD" box="[1010,1078,1501,1527]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="60" unit="mm" value="8.0">8 mm</quantity>
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wide, with its anterior margin bend forward, narrower than posterior margin; synthorax large,
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<quantity id="4CAF9B4EF16AFF9C66CDD90453C3A925" box="[898,980,1541,1567]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="60" unit="mm" value="12.0">12 mm</quantity>
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long, 8.0 mm wide.
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</paragraph>
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<paragraph id="8BE836ABF16AFF9C6588D92C52FEA985" blockId="1.[151,1437,1181,2007]" pageId="1" pageNumber="60">Coxa of fore leg large and stout; femur strong, with one row of regular spines along inner margin; tibia as long as femur, furnished with strong spines; mid leg larger and stouter than fore leg, with stout femur, tibia incompletely preserved, femur and tibia with spines along their inner margins; hind leg the largest, femur large, tibia incomplete, both furnished with spines.</paragraph>
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<paragraph id="8BE836ABF16AFF9E6588D9CC517EAE10" blockId="1.[151,1437,1181,2007]" lastBlockId="3.[151,1437,152,663]" lastPageId="3" lastPageNumber="62" pageId="1" pageNumber="60">
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Fore wing
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<quantity id="4CAF9B4EF16AFF9C6403D9CC51B5A9DD" box="[332,418,1741,1767]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="60" unit="mm" value="40.0">40 mm</quantity>
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long,
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<quantity id="4CAF9B4EF16AFF9C64BCD9CC522DA9DD" box="[499,570,1741,1767]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="60" unit="mm" value="8.0">8 mm</quantity>
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wide, approximately four times as long as wide; distance from base to nodus
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<quantity id="4CAF9B4EF16AFF9C65ACD9F4512EA835" box="[227,313,1781,1807]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="1" pageNumber="60" unit="mm" value="17.0">17 mm</quantity>
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, nodus to pterostigma
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<quantity id="4CAF9B4EF16AFF9C6770D9F452BCA835" box="[575,683,1781,1807]" metricMagnitude="-2" metricUnit="m" metricValue="1.23" pageId="1" pageNumber="60" unit="mm" value="12.3">12.3 mm</quantity>
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, pterostigma to wing apex
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<quantity id="4CAF9B4EF16AFF9C66ABD9F45456A835" box="[996,1089,1781,1807]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="1" pageNumber="60" unit="mm" value="3.5">3.5 mm</quantity>
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, base to arculus
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<quantity id="4CAF9B4EF16AFF9C604FD9F4554BA835" box="[1280,1372,1781,1807]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="1" pageNumber="60" unit="mm" value="3.4">3.4 mm</quantity>
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; hind wing broader than fore wing,
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<quantity id="4CAF9B4EF16AFF9C64B3D81C5270A80D" box="[508,615,1821,1847]" metricMagnitude="-2" metricUnit="m" metricValue="3.86" pageId="1" pageNumber="60" unit="mm" value="38.6">38.6 mm</quantity>
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long,
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<quantity id="4CAF9B4EF16AFF9C67FBD81C5309A80D" box="[692,798,1821,1847]" metricMagnitude="-2" metricUnit="m" metricValue="1.16" pageId="1" pageNumber="60" unit="mm" value="11.6">11.6 mm</quantity>
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wide, approximately four times as long as wide; distance from base to nodus
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<quantity id="4CAF9B4EF16AFF9C648BD8445238A865" box="[452,559,1861,1887]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" pageId="1" pageNumber="60" unit="mm" value="14.5">14.5 mm</quantity>
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, nodus to pterostigma
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<quantity id="4CAF9B4EF16AFF9C6677D84453B3A865" box="[824,932,1861,1887]" metricMagnitude="-2" metricUnit="m" metricValue="1.52" pageId="1" pageNumber="60" unit="mm" value="15.2">15.2 mm</quantity>
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, pterostigma to wing apex
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<quantity id="4CAF9B4EF16AFF9C61AED8445529A865" box="[1249,1342,1861,1887]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="1" pageNumber="60" unit="mm" value="3.5">3.5 mm</quantity>
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, base to arculus
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<quantity id="4CAF9B4EF16AFF9C65BCD86C5147A8BD" box="[243,336,1901,1927]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="1" pageNumber="60" unit="mm" value="5.5">5.5 mm</quantity>
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; pterostigma
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<quantity id="4CAF9B4EF16AFF9C64A2D86C5250A8BD" box="[493,583,1901,1927]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="1" pageNumber="60" unit="mm" value="3.1">3.1 mm</quantity>
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||
long,
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<quantity id="4CAF9B4EF16AFF9C67DED86C52C2A8BD" box="[657,725,1901,1927]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="60" unit="mm" value="1.0">1 mm</quantity>
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wide, approximately 3 times as long as wide, covering three cross-veins, pterostigmal brace strong and distinctly oblique; no cross-vein in distal antesubnodal area (‘cordulegastrid’ gap); ScP with a strong curve at nodus; nodal cross-vein aligned with ScP at nodus, perpendicular to C and RA; subnodus not well-preserved; in fore wing about 14 secondary antenodal and antesubnodal cross-veins, not aligned; in hind wing about 12 secondary antenodal and antesubnodal cross-veins, not aligned; Ax1 well basal and Ax2 distal of discoidal triangle, with two secondary cross-veins between them; base of
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<collectingCountry id="F340763BF169FF9F65BBD90E510EA913" box="[244,281,1551,1577]" name="Poland" pageId="2" pageNumber="61">RP</collectingCountry>
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2 aligned with subnodus;
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<collectingCountry id="F340763BF169FF9F6708D90E527BA913" box="[583,620,1551,1577]" name="Poland" pageId="2" pageNumber="61">RP</collectingCountry>
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2 and IR2 with a double curve;
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<collectingCountry id="F340763BF169FF9F66ACD90E541FA913" box="[995,1032,1551,1577]" name="Poland" pageId="2" pageNumber="61">RP</collectingCountry>
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3/4 and MA also undulate but less distinctly than
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<collectingCountry id="F340763BF169FF9F6403D9365164A96B" box="[332,371,1591,1617]" name="Poland" pageId="2" pageNumber="61">RP</collectingCountry>
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2; Rspl well-defined and parallel to IR2, with one row of cells between it and IR2; IR1 short, with its base below middle of pterostigma; arculus straight, perpendicular to RA, with its costal part distinctly longer than its ventral part;
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<collectingCountry id="F340763BF169FF9F6772D9865276A99B" box="[573,609,1671,1697]" name="Poland" pageId="2" pageNumber="61">RP</collectingCountry>
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and MA well-separated in the arculus; discoidal triangle in fore and hind wings transverse, of similar shape, although the fore wing one is slightly narrower than that of hind wing, both divided in two cells by a cross-vein, distal side MAb slightly curved; anterior margin (MA) of hypertriangle more or less straight; hypertriangle elongate and narrow, free of cross-veins; subtriangles of fore and hind wings unicellular, but well-defined by PsA vein, that of fore wing being broader than that of hind wing, with its posterior margin more rounded; postdiscoidal area of fore wing with two rows of cells just distal of triangle, three in hind wing; Mspl well defined and parallel to MA, with one row of cells between them; hind wing MP somewhat shortened and distinctly curved towards hind margin, reaching this margin nearly opposite nodus; hind wing area between MP and CuAa not very broad, with only one row of transverse cells near discoidal triangle; CuAa curved, with only four posterior branches; basal part of CuA straight but rather long,
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<quantity id="4CAF9B4EF168FF9E65D8DFC150E3AFE0" box="[151,244,192,218]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="3" pageNumber="62" unit="mm" value="1.6">1.6 mm</quantity>
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long; anal loop complete rather transverse, rhombus-like, divided into seven cells; anal area rather broad, with five rows of cells between anal loop and posterior wing margin; hind wing base of anal margin rounded (female).
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</paragraph>
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<caption id="DF286623F169FF9F65D8DA2851C5AA59" httpUri="https://zenodo.org/record/273747/files/figure.png" pageId="2" pageNumber="61" targetBox="[220,1368,197,1301]" targetPageId="2">
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||
<paragraph id="8BE836ABF169FF9F65D8DA2851C5AA59" blockId="2.[151,1436,1321,1379]" pageId="2" pageNumber="61">
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<emphasis id="B923EAB9F169FF9F65D8DA28510BAA7B" bold="true" box="[151,284,1321,1345]" pageId="2" pageNumber="61">FIGURE 1.</emphasis>
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<taxonomicName id="4C574D28F169FF9F6469DA28520CAA7A" box="[294,539,1321,1344]" class="Insecta" family="Nodalulaidae" genus="Nodalula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="dalinghensis">
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<emphasis id="B923EAB9F169FF9F6469DA28520CAA7A" box="[294,539,1321,1344]" italics="true" pageId="2" pageNumber="61">Nodalula dalinghensis</emphasis>
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</taxonomicName>
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<emphasis id="B923EAB9F169FF9F676ADA2852C2AA7B" bold="true" box="[549,725,1321,1345]" pageId="2" pageNumber="61">gen. et sp. nov.</emphasis>
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, holotype Cat. No. 123517 (female), photograph of dorsal view (scale bar represents 10 mm).
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</paragraph>
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</caption>
|
||
<paragraph id="8BE836ABF168FF9E6588DE395231ADAD" blockId="3.[151,1437,152,663]" pageId="3" pageNumber="62">
|
||
Abdomen
|
||
<quantity id="4CAF9B4EF168FF9E640ADE3951A7AE68" box="[325,432,312,338]" metricMagnitude="-2" metricUnit="m" metricValue="4.86" pageId="3" pageNumber="62" unit="mm" value="48.6">48.6 mm</quantity>
|
||
long,
|
||
<quantity id="4CAF9B4EF168FF9E674FDE39524AAE68" box="[512,605,312,338]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="3" pageNumber="62" unit="mm" value="4.6">4.6 mm</quantity>
|
||
wide; first abdominal segment well developed and large, obconic in form, its anterior margin about 1.5 time as wide as its posterior margin; second abdominal segment longer, but narrower than first, about three times as long as wide; third segment similar to second, about 2.3 times as long as wide; fourth and fifth similar, slightly broader than third; but slight shorter than the 3rd; sixth similar to fourth in width, but shorter, about two times as long as wide; seventh shorter than sixth, longer than wide; eighth shorter than seventh, as long as wide; a median longitudinal carina on second to eighth terga; a pattern of light coloration on second to sixth segments, consisting in two longitudinal stripes and two outer apical spots; two superior anal appendages normally developed; median appendage shorter than superior anal appendages; no visible ovipositor.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF286623F168FF9E65D8D94B52A6A9BE" httpUri="https://zenodo.org/record/273748/files/figure.png" pageId="3" pageNumber="62" targetBox="[252,1331,723,1588]" targetPageId="3">
|
||
<paragraph id="8BE836ABF168FF9E65D8D94B52A6A9BE" blockId="3.[151,1435,1610,1668]" pageId="3" pageNumber="62">
|
||
<emphasis id="B923EAB9F168FF9E65D8D94B510EA958" bold="true" box="[151,281,1610,1634]" pageId="3" pageNumber="62">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C574D28F168FF9E646FD94B5205A95B" box="[288,530,1610,1633]" class="Insecta" family="Nodalulaidae" genus="Nodalula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="dalinghensis">
|
||
<emphasis id="B923EAB9F168FF9E646FD94B5205A95B" box="[288,530,1610,1633]" italics="true" pageId="3" pageNumber="62">Nodalula dalinghensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B923EAB9F168FF9E6757D94B52A8A958" bold="true" box="[536,703,1610,1634]" pageId="3" pageNumber="62">gen. et sp. nov.</emphasis>
|
||
, holotype Cat. No. 123517, Enlarged view showing the left fore and hind wings venation (scale bar represents 10 mm).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C34D6520F168FF986588D9B350F2AE38" lastPageId="5" lastPageNumber="64" pageId="3" pageNumber="62" type="discussion">
|
||
<paragraph id="8BE836ABF168FF9E6588D9B351FDA856" blockId="3.[151,1436,1714,2020]" pageId="3" pageNumber="62">
|
||
<emphasis id="B923EAB9F168FF9E6588D9B35143A9F6" bold="true" box="[199,340,1714,1740]" pageId="3" pageNumber="62">Discussion:</emphasis>
|
||
The attribution of
|
||
<taxonomicName id="4C574D28F168FF9E677FD9B2528AA9F6" box="[560,669,1715,1740]" class="Insecta" family="Nodalulaidae" genus="Nodalula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="62" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B923EAB9F168FF9E677FD9B2528AA9F6" box="[560,669,1715,1740]" italics="true" pageId="3" pageNumber="62">Nodalula</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B923EAB9F168FF9E67EBD9B35306A9F6" bold="true" box="[676,785,1714,1740]" pageId="3" pageNumber="62">
|
||
<taxonomicNameLabel id="A21057C2F168FF9E67EBD9B35306A9F6" box="[676,785,1714,1740]" pageId="3" pageNumber="62" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
to the Cavilabiata
|
||
<bibRefCitation id="EFC64B5AF168FF9E66A2D9B3549FA9F6" author="Bechly" box="[1005,1160,1714,1740]" pageId="3" pageNumber="64" refString="Bechly, G., Nel, A. & Martinez-Delclos, X. (1996) Redescription of Nannogomphus bavaricus Handlirsch from the Upper Jurassic of Germany, with an analysis of its phylogenetic position (Odonata, Anisoptera: Gomphidae or Libellulidae). Archaeopteryx, 14, 51 - 66." type="journal article" year="1996">Bechly, 1996</bibRefCitation>
|
||
is supported by the following characters: distal part of antesubnodal area free of cross-veins (‘cordulegastrid gap’); gaff (= basal CuA before its branching) of the hind wing distinctly prolonged. The third venational character, viz. the
|
||
<collectingCountry id="F340763BF168FF9E602ED8035591A826" box="[1377,1414,1794,1820]" name="Poland" pageId="3" pageNumber="62">RP</collectingCountry>
|
||
3/ 4 and MA slightly undulating in both pairs of wings, is impossible to determine because of the important deformations of these wings.
|
||
</paragraph>
|
||
<paragraph id="8BE836ABF168FF996588D87B51E0A998" blockId="3.[151,1436,1714,2020]" lastBlockId="4.[151,1437,152,2018]" lastPageId="4" lastPageNumber="63" pageId="3" pageNumber="62">
|
||
Affinities with the Cordulegastrida
|
||
<bibRefCitation id="EFC64B5AF168FF9E6722D87B531DA8AE" author="Bechly" box="[621,778,1914,1940]" pageId="3" pageNumber="64" refString="Bechly, G., Nel, A. & Martinez-Delclos, X. (1996) Redescription of Nannogomphus bavaricus Handlirsch from the Upper Jurassic of Germany, with an analysis of its phylogenetic position (Odonata, Anisoptera: Gomphidae or Libellulidae). Archaeopteryx, 14, 51 - 66." type="journal article" year="1996">Bechly, 1996</bibRefCitation>
|
||
can be excluded because
|
||
<taxonomicName id="4C574D28F168FF9E610ED87A54B9A8AE" box="[1089,1198,1915,1940]" class="Insecta" family="Nodalulaidae" genus="Nodalula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="62" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B923EAB9F168FF9E610ED87A54B9A8AE" box="[1089,1198,1915,1940]" italics="true" pageId="3" pageNumber="62">Nodalula</emphasis>
|
||
</taxonomicName>
|
||
has not the ‘female ovipositor strongly elongated into a gently tapered spade’, and the venational synapomorphies of this clade, viz. wings with a basal furcation of IR2 basal of the lestine oblique vein; pterostigmal brace vein reduced; discoidal triangles of similar elongate shape in both pairs of wings, correlated with a secondarily less distinct pseudo-anal vein PsA and subdiscoidal triangle. On the contrary, it has the venational synapomorphies of the Brachystigmata
|
||
<bibRefCitation id="EFC64B5AF16FFF996416DFE951EDAE38" author="Bechly" box="[345,506,232,258]" pageId="4" pageNumber="64" refString="Bechly, G., Nel, A. & Martinez-Delclos, X. (1996) Redescription of Nannogomphus bavaricus Handlirsch from the Upper Jurassic of Germany, with an analysis of its phylogenetic position (Odonata, Anisoptera: Gomphidae or Libellulidae). Archaeopteryx, 14, 51 - 66." type="journal article" year="1996">Bechly, 1996</bibRefCitation>
|
||
, i.e. relatively short pterostigmata that cover only 1–3 complete cells; in the hind wing the gaff (= basal CuA before its branching) is strongly prolonged and very straight (in the ground plan); area between MP and CuA basally widened; terminal branch of CuAa seems to be secondarily branched on CuA;
|
||
<collectingCountry id="F340763BF16FFF996437DE61518AAE40" box="[376,413,352,378]" name="Poland" pageId="4" pageNumber="63">RP</collectingCountry>
|
||
3/4 and MA strictly parallel up to the hind margin; area between
|
||
<collectingCountry id="F340763BF16FFF9961EBDE6154DEAE40" box="[1188,1225,352,378]" name="Poland" pageId="4" pageNumber="63">RP</collectingCountry>
|
||
2 and IR2 distally distinctly widened, with more than one cell row in the distal half. Within this clade, the comparison with the Mesozoic Nannogomphidae
|
||
<bibRefCitation id="EFC64B5AF16FFF9964A8DEB15296AEF0" author="Bechly" box="[487,641,432,458]" pageId="4" pageNumber="64" refString="Bechly, G., Nel, A. & Martinez-Delclos, X. (1996) Redescription of Nannogomphus bavaricus Handlirsch from the Upper Jurassic of Germany, with an analysis of its phylogenetic position (Odonata, Anisoptera: Gomphidae or Libellulidae). Archaeopteryx, 14, 51 - 66." type="journal article" year="1996">Bechly, 1996</bibRefCitation>
|
||
is not easy because there is no strong synapomorphy for this family. Affinities with
|
||
<taxonomicName id="4C574D28F16FFF996405DED952C3AEC8" authority="Handlirsch, 1906" authorityName="Handlirsch" authorityYear="1906" box="[330,724,472,498]" class="Insecta" family="Gomphidae" genus="Nannogomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="63" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B923EAB9F16FFF996405DED95215AECB" box="[330,514,472,497]" italics="true" pageId="4" pageNumber="63">Nannogomphus</emphasis>
|
||
Handlirsch, 1906
|
||
</taxonomicName>
|
||
are excluded because the latter is characterized by the synapomorphy ‘CuAa without posterior branches’ (
|
||
<bibRefCitation id="EFC64B5AF16FFF99678EDD0153B0AD20" author="Bechly" box="[705,935,512,538]" pageId="4" pageNumber="64" refString="Bechly, G., Nel, A. & Martinez-Delclos, X. (1996) Redescription of Nannogomphus bavaricus Handlirsch from the Upper Jurassic of Germany, with an analysis of its phylogenetic position (Odonata, Anisoptera: Gomphidae or Libellulidae). Archaeopteryx, 14, 51 - 66." type="journal article" year="1996">
|
||
Bechly
|
||
<emphasis id="B923EAB9F16FFF996652DD01534EAD23" box="[797,857,512,537]" italics="true" pageId="4" pageNumber="63">et al.</emphasis>
|
||
, 1996
|
||
</bibRefCitation>
|
||
). Affinities with
|
||
<emphasis id="B923EAB9F16FFF996120DD015556AD23" box="[1135,1345,512,537]" italics="true" pageId="4" pageNumber="63">Prohemeroscopus</emphasis>
|
||
Bechly
|
||
<emphasis id="B923EAB9F16FFF9965D8DD2950C5AD7B" box="[151,210,552,577]" italics="true" pageId="4" pageNumber="63">et al.</emphasis>
|
||
, 1998 (originally in
|
||
<taxonomicName id="4C574D28F16FFF99648BDD29529BAD78" box="[452,652,552,578]" class="Insecta" family="Hemeroscopidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="63" phylum="Arthropoda" rank="family">Hemeroscopidae</taxonomicName>
|
||
, but latter transferred into the Nannogomphidae) are more delicate to discuss. Nevertheless,
|
||
<taxonomicName id="4C574D28F16FFF9964F2DD51523DAD53" box="[445,554,592,617]" class="Insecta" family="Nodalulaidae" genus="Nodalula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="62" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B923EAB9F16FFF9964F2DD51523DAD53" box="[445,554,592,617]" italics="true" pageId="4" pageNumber="63">Nodalula</emphasis>
|
||
</taxonomicName>
|
||
can be attributed to the Eubrachystigmata
|
||
<bibRefCitation id="EFC64B5AF16FFF996150DD5154ABAD50" author="Bechly" box="[1055,1212,592,618]" pageId="4" pageNumber="64" refString="Bechly, G., Nel, A. & Martinez-Delclos, X. (1996) Redescription of Nannogomphus bavaricus Handlirsch from the Upper Jurassic of Germany, with an analysis of its phylogenetic position (Odonata, Anisoptera: Gomphidae or Libellulidae). Archaeopteryx, 14, 51 - 66." type="journal article" year="1996">Bechly, 1996</bibRefCitation>
|
||
, sister group of the Nannogomphidae, on the basis of the apomorphy ‘hind wing CuAa more distinctly curved towards the hind margin, and thus further shortened with less than five posterior branches’. Within this group,
|
||
<taxonomicName id="4C574D28F16FFF996193DDA1555EAD83" box="[1244,1353,672,697]" class="Insecta" family="Nodalulaidae" genus="Nodalula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="62" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B923EAB9F16FFF996193DDA1555EAD83" box="[1244,1353,672,697]" italics="true" pageId="4" pageNumber="63">Nodalula</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<emphasis id="B923EAB9F16FFF996598DDC95193ADDB" box="[215,388,712,737]" italics="true" pageId="4" pageNumber="63">Hemeroscopus</emphasis>
|
||
Pritykina, 1977 (
|
||
<taxonomicName id="4C574D28F16FFF996701DDC953C5ADD8" authority="Pritykina, 1977" authorityName="Pritykina" authorityYear="1977" box="[590,978,712,738]" class="Insecta" family="Hemeroscopidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="63" phylum="Arthropoda" rank="family">Hemeroscopidae Pritykina, 1977</taxonomicName>
|
||
) in its crossed discoidal cells, presence of well-defined Rspl and Mspl, presence of a distinct pterostigmal brace; shorter pterostigma, lestine oblique cross-vein shifted basally one cell distal of the subnodus, and narrower and smaller hind wing anal cell. The monophyly of the Neobrachystigmata
|
||
<bibRefCitation id="EFC64B5AF16FFF996712DC4152ECAC60" author="Bechly" box="[605,763,832,858]" pageId="4" pageNumber="64" refString="Bechly, G., Nel, A. & Martinez-Delclos, X. (1996) Redescription of Nannogomphus bavaricus Handlirsch from the Upper Jurassic of Germany, with an analysis of its phylogenetic position (Odonata, Anisoptera: Gomphidae or Libellulidae). Archaeopteryx, 14, 51 - 66." type="journal article" year="1996">Bechly, 1996</bibRefCitation>
|
||
, sister group of the
|
||
<taxonomicName id="4C574D28F16FFF9966A8DC4154B8AC60" box="[999,1199,832,858]" class="Insecta" family="Hemeroscopidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="63" phylum="Arthropoda" rank="family">Hemeroscopidae</taxonomicName>
|
||
, is supported by the following synapomorphies: ‘arculus not distinctly angled but more or less straight, with posterior part (crossvein) of arculus distinctly shorter than anterior part’. This character seems to be present in
|
||
<taxonomicName id="4C574D28F16FFF99618EDC915539AC93" box="[1217,1326,912,937]" class="Insecta" family="Nodalulaidae" genus="Nodalula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="62" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B923EAB9F16FFF99618EDC915539AC93" box="[1217,1326,912,937]" italics="true" pageId="4" pageNumber="63">Nodalula</emphasis>
|
||
</taxonomicName>
|
||
although it is not obvious because the arculus are partly broken; ‘hind wing MP distinctly curved towards the hind margin and thus somewhat shortened, ending basal of the level of the nodus’. Despite the deformation of the costal region of its wings, it seems that
|
||
<taxonomicName id="4C574D28F16FFF99671DDB0952A8AB1B" box="[594,703,1032,1057]" class="Insecta" family="Nodalulaidae" genus="Nodalula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="62" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B923EAB9F16FFF99671DDB0952A8AB1B" box="[594,703,1032,1057]" italics="true" pageId="4" pageNumber="63">Nodalula</emphasis>
|
||
</taxonomicName>
|
||
has a significant curvature of the hind wing MP. Within this group, our fossil has not the synapomorphy of the Chlorogomphida
|
||
<bibRefCitation id="EFC64B5AF16FFF99668EDB315448AB70" author="Bechly" box="[961,1119,1072,1098]" pageId="4" pageNumber="64" refString="Bechly, G., Nel, A. & Martinez-Delclos, X. (1996) Redescription of Nannogomphus bavaricus Handlirsch from the Upper Jurassic of Germany, with an analysis of its phylogenetic position (Odonata, Anisoptera: Gomphidae or Libellulidae). Archaeopteryx, 14, 51 - 66." type="journal article" year="1996">Bechly, 1996</bibRefCitation>
|
||
, viz. ‘hypertriangles much longer than discoidal triangles in both pairs of wings’.
|
||
<taxonomicName id="4C574D28F16FFF996606DB5953A1AB4B" box="[841,950,1112,1137]" class="Insecta" family="Nodalulaidae" genus="Nodalula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="62" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B923EAB9F16FFF996606DB5953A1AB4B" box="[841,950,1112,1137]" italics="true" pageId="4" pageNumber="63">Nodalula</emphasis>
|
||
</taxonomicName>
|
||
shares with the Paneurypalpidomorpha
|
||
<bibRefCitation id="EFC64B5AF16FFF9965D8DB815121ABA0" author="Bechly" box="[151,310,1152,1178]" pageId="4" pageNumber="64" refString="Bechly, G., Nel, A. & Martinez-Delclos, X. (1996) Redescription of Nannogomphus bavaricus Handlirsch from the Upper Jurassic of Germany, with an analysis of its phylogenetic position (Odonata, Anisoptera: Gomphidae or Libellulidae). Archaeopteryx, 14, 51 - 66." type="journal article" year="1996">Bechly, 1996</bibRefCitation>
|
||
, sister group of the Chlorogomphida, two synapomorphies, i.e. ‘lestine oblique vein only 1–2 cells distad of the subnodus in both pairs of wings’, ‘fore wing ax1 shifted basal of the level of the distal angle of the discoidal triangle’, but not several others, viz. ‘anterior margin of hypertriangle distinctly convex, especially in the hind wings, because of a basally arched MA’, ‘fore wing discoidal triangle more or less transverse instead of elongate’, and ‘in both pairs of wings Ax1 and Ax2 relatively close together with not more than one secondary antenodal cross-vein between them’. The Paneurypalpidomorpha comprises the Mesozoic Juracorduliidae Bechly, 1998 and the Eurypalpidomorpha.
|
||
<taxonomicName id="4C574D28F16FFF9967B3DA71537EAAB3" box="[764,873,1392,1417]" class="Insecta" family="Nodalulaidae" genus="Nodalula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="62" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B923EAB9F16FFF9967B3DA71537EAAB3" box="[764,873,1392,1417]" italics="true" pageId="4" pageNumber="63">Nodalula</emphasis>
|
||
</taxonomicName>
|
||
also differs from
|
||
<emphasis id="B923EAB9F16FFF99610FDA7154CFAAB3" box="[1088,1240,1392,1417]" italics="true" pageId="4" pageNumber="63">Juracordulia</emphasis>
|
||
Bechly,
|
||
<quantity id="4CAF9B4EF16FFF99600DDA71558BAAB0" box="[1346,1436,1392,1418]" metricMagnitude="1" metricUnit="m" metricValue="5.07492" pageId="4" pageNumber="63" unit="in" value="1998.0">1998 in</quantity>
|
||
its less elongate anal loop and CuAa with more branches (plesiomorphies) (Bechly, 1998). It has not the main synapomorphy of the Eurypalpidomorpha
|
||
<bibRefCitation id="EFC64B5AF16FFF9967DADAC15325AAE0" author="Bechly" box="[661,818,1472,1498]" pageId="4" pageNumber="64" refString="Bechly, G., Nel, A. & Martinez-Delclos, X. (1996) Redescription of Nannogomphus bavaricus Handlirsch from the Upper Jurassic of Germany, with an analysis of its phylogenetic position (Odonata, Anisoptera: Gomphidae or Libellulidae). Archaeopteryx, 14, 51 - 66." type="journal article" year="1996">Bechly, 1996</bibRefCitation>
|
||
(clade comprising the Mesozoic families Valdicorduliidae
|
||
<bibRefCitation id="EFC64B5AF16FFF99644BDAE951BFA938" author="Bechly" box="[260,424,1512,1538]" pageId="4" pageNumber="64" refString="Bechly, G., Nel, A. & Martinez-Delclos, X. (1996) Redescription of Nannogomphus bavaricus Handlirsch from the Upper Jurassic of Germany, with an analysis of its phylogenetic position (Odonata, Anisoptera: Gomphidae or Libellulidae). Archaeopteryx, 14, 51 - 66." type="journal article" year="1996">Bechly, 1996</bibRefCitation>
|
||
, Araripephlebiidae Bechly, 1998, Eocorduliidae
|
||
<bibRefCitation id="EFC64B5AF16FFF996148DAE954BCA938" author="Bechly" box="[1031,1195,1512,1538]" pageId="4" pageNumber="64" refString="Bechly, G., Nel, A. & Martinez-Delclos, X. (1996) Redescription of Nannogomphus bavaricus Handlirsch from the Upper Jurassic of Germany, with an analysis of its phylogenetic position (Odonata, Anisoptera: Gomphidae or Libellulidae). Archaeopteryx, 14, 51 - 66." type="journal article" year="1996">Bechly, 1996</bibRefCitation>
|
||
, Araripelibellulidae
|
||
<bibRefCitation id="EFC64B5AF16FFF9965D8D9115123A910" author="Bechly" box="[151,308,1552,1578]" pageId="4" pageNumber="64" refString="Bechly, G., Nel, A. & Martinez-Delclos, X. (1996) Redescription of Nannogomphus bavaricus Handlirsch from the Upper Jurassic of Germany, with an analysis of its phylogenetic position (Odonata, Anisoptera: Gomphidae or Libellulidae). Archaeopteryx, 14, 51 - 66." type="journal article" year="1996">Bechly, 1996</bibRefCitation>
|
||
, and the fossil and modern Eurypalpida
|
||
<bibRefCitation id="EFC64B5AF16FFF99665FD91153B8A910" author="Bechly" box="[784,943,1552,1578]" pageId="4" pageNumber="64" refString="Bechly, G., Nel, A. & Martinez-Delclos, X. (1996) Redescription of Nannogomphus bavaricus Handlirsch from the Upper Jurassic of Germany, with an analysis of its phylogenetic position (Odonata, Anisoptera: Gomphidae or Libellulidae). Archaeopteryx, 14, 51 - 66." type="journal article" year="1996">Bechly, 1996</bibRefCitation>
|
||
), viz. ‘in the hind wing the elongated gaff (= basal CuA before its branching) sigmoidally curved, CuAb shifted very far distally, and CuAa with only one or two posterior branches’ (
|
||
<bibRefCitation id="EFC64B5AF16FFF996741D96152DEA940" author="Pritykina" box="[526,713,1632,1658]" pageId="4" pageNumber="64" refString="Pritykina, L. N. (1986) Two new dragonflies from the Lower Cretaceous deposits of west Mongolia (Anisoptera: Sonidae fam. nov., Corduliidae). Odonatologica, 15 (2), 169 - 184." type="journal article" year="1986">Pritykina, 1986</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFC64B5AF16FFF99679AD96153E5A940" author="Nel" box="[725,1010,1632,1658]" pageId="4" pageNumber="64" refString="Nel, A. & Paicheler, J. - C. (1994) Les Libelluloidea fossiles autres que Libellulidae. Un inventaire critique (Odonata, Corduliidae, Macromiidae, Synthemistidae, Chlorogomphidae et Mesophlebiidae). Nouvelle Revue d'Entomologie, (N. S.), 11, 321 - 334." type="journal article" year="1994">Nel and Paicheler, 1994</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFC64B5AF16FFF9966B0D96154F0A940" author="Ren" box="[1023,1255,1632,1658]" pageId="4" pageNumber="64" refString="Ren, D. & Guo, Z. - G. (1996) Three new genera and three new species of dragonflies from the Late Jurassic of Northeast China (Anisoptera: Aeshnidae, Gomphidae, Corduliidae). Entomologia Sinica, 3 (2), 95 - 105." type="journal article" year="1996">Ren and Guo, 1996</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFC64B5AF16FFF9961BBD9615128A998" author="Jarzembowski" pageId="4" pageNumber="64" refString="Jarzembowski, E. A. & Nel, A. (1996) New fossil dragonflies from the Lower Cretaceous of SE England and the phylogeny of the superfamily Libelluloidea (Insecta: Odonata). Cretaceous Research, 17 (1), 67 - 85." type="journal article" year="1996">Jarzembowski and Nel, 1996</bibRefCitation>
|
||
; Bechly, 1998).
|
||
</paragraph>
|
||
<paragraph id="8BE836ABF16FFF996588D9B151A7A878" blockId="4.[151,1437,152,2018]" pageId="4" pageNumber="63">
|
||
In conclusion,
|
||
<taxonomicName id="4C574D28F16FFF99643CD9B151F7A9F3" box="[371,480,1712,1737]" class="Insecta" family="Nodalulaidae" genus="Nodalula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="62" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B923EAB9F16FFF99643CD9B151F7A9F3" box="[371,480,1712,1737]" italics="true" pageId="4" pageNumber="63">Nodalula</emphasis>
|
||
</taxonomicName>
|
||
can be considered as a Neobrachystigmata that does not belong to the clade Chlorogomphida and shares some synapomorphies with the Paneurypalpidomorpha, but which is neither a Juracorduliidae nor a Eurypalpidomorpha. It could well be the sister group of the whole clade Paneurypalpidomorpha.
|
||
</paragraph>
|
||
<paragraph id="8BE836ABF16FFF986588D85150F2AE38" blockId="4.[151,1437,152,2018]" lastBlockId="5.[151,1436,151,258]" lastPageId="5" lastPageNumber="64" pageId="4" pageNumber="63">
|
||
A larval dragonfly from the same strata described by
|
||
<bibRefCitation id="EFC64B5AF16FFF996670D851545FA850" author="Huang" box="[831,1096,1872,1898]" pageId="4" pageNumber="64" refString="Huang, D. - Y. & Nel. A. (2004) Two new Lower Cretaceous dragonfly larvae from Northeastern China (Anisoptera). Odonatologica, 33, 199 - 205." type="journal article" year="2004">Huang and Nel (2004)</bibRefCitation>
|
||
has been placed in the clade Cavilabiata. This specimen shows some strong similarities with the ultimate larval instars of
|
||
<emphasis id="B923EAB9F16FFF996043D879558CA8AB" box="[1292,1435,1912,1937]" italics="true" pageId="4" pageNumber="63">Sona nectes</emphasis>
|
||
described from western
|
||
<collectingCountry id="F340763BF16FFF9964E0D8A15236A880" box="[431,545,1952,1978]" name="Mongolia" pageId="4" pageNumber="63">Mongolia</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFC64B5AF16FFF996760D8A152F5A880" author="Pritykina" box="[559,738,1952,1978]" pageId="4" pageNumber="64" refString="Pritykina, L. N. (1986) Two new dragonflies from the Lower Cretaceous deposits of west Mongolia (Anisoptera: Sonidae fam. nov., Corduliidae). Odonatologica, 15 (2), 169 - 184." type="journal article" year="1986">Pritykina 1986</bibRefCitation>
|
||
). The ultimate larval instars always display few comparable features to the corresponding adult in
|
||
<taxonomicName id="4C574D28F16FFF99671FD8C952A0A8D8" box="[592,695,1992,2018]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="63" phylum="Arthropoda" rank="order">Odonata</taxonomicName>
|
||
. This ultimate larval instars of the ‘
|
||
<emphasis id="B923EAB9F16FFF99611BD8C954C9A8DB" box="[1108,1246,1992,2017]" italics="true" pageId="4" pageNumber="63">Sona nectes</emphasis>
|
||
type’ larva bear strongly developed spines rows on legs that resemble those of
|
||
<taxonomicName id="4C574D28F16EFF9866CCDF99549BAF8B" box="[899,1164,152,177]" class="Insecta" family="Nodalulaidae" genus="Nodalula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="64" phylum="Arthropoda" rank="species" species="dalinghensis">
|
||
<emphasis id="B923EAB9F16EFF9866CCDF99549BAF8B" box="[899,1164,152,177]" italics="true" pageId="5" pageNumber="64">Nodalula dalinghensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B923EAB9F16EFF9861DADF965547AF8B" bold="true" box="[1173,1360,151,177]" pageId="5" pageNumber="64">gen. et sp. nov.</emphasis>
|
||
However, their correspondence still remains uncertain due to the strong morphological gap between larvae and adults.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |