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<document id="B01CFB6AE86B3BDE3684B705CB4D3FD4" ID-CLB-Dataset="7124" ID-DOI="10.12782/sd.20.1.037" ID-GBIF-Dataset="7c173c23-22cf-44c9-936c-7f9aedbb9a87" ID-ISSN="2189-7301" ID-Zenodo-Dep="4584966" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1614946735199" checkinUser="felipe" docAuthor="Saito, Nobuhiro &amp; Shimomura, Michitaka" docDate="2015" docId="03EC87FB3029FFE9FC0A139B7075F8FF" docLanguage="en" docName="SpeciesDiversity.20.1.37-43.pdf" docOrigin="Species Diversity 20 (1)" docSource="http://dx.doi.org/10.12782/sd.20.1.037" docStyle="DocumentStyle:27FECD2C3CD153DA405029F2D33C9514.7:SpeciesDiversity.2012-.journal_article" docStyleId="27FECD2C3CD153DA405029F2D33C9514" docStyleName="SpeciesDiversity.2012-.journal_article" docStyleVersion="7" docTitle="Anisarthrus okunoi Saito &amp; Shimomura 2015, sp. nov." docType="treatment" docUuid="72957321-67AE-41C8-9560-B5CCBA22C97E" docUuidSource="ZooBank" docVersion="5" lastPageNumber="39" masterDocId="FFD5FF833029FFEBFFD214197431FF85" masterDocTitle="A New Abdominally Parasitizing Bopyrid, Anisarthrus okunoi sp. nov. (Crustacea: Isopoda), Infesting the Hinge-Beak Shrimp Rhynchocinetes uritai Kubo, 1942 (Crustacea: Decapoda: Rhynchocinetidae)" masterLastPageNumber="43" masterPageNumber="37" pageNumber="37" updateTime="1698932737753" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="16C91AC7A3A8EAB780CE5BCF4C2F165E">A New Abdominally Parasitizing Bopyrid, Anisarthrus okunoi sp. nov. (Crustacea: Isopoda), Infesting the Hinge-Beak Shrimp Rhynchocinetes uritai Kubo, 1942 (Crustacea: Decapoda: Rhynchocinetidae)</mods:title>
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<mods:namePart id="E7F788552F3266CAAD7A126393925377">Saito, Nobuhiro</mods:namePart>
<mods:affiliation id="81C9BEB04ED8E1B5E7BC98A4F86C14FD">Suido-sha Co. Ltd., Ikuta 8 - 11 - 11, Tama-ku, Kawasaki, Kanagawa 214 - 0038, Japan E-mail: nsaitoh @ suidosha. co. jp &amp; Corresponding author</mods:affiliation>
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<mods:namePart id="C24D1A61479F814D890904C5D1F6BE79">Shimomura, Michitaka</mods:namePart>
<mods:affiliation id="4A7493DC5B860F10162C5C7567CF48E5">Kitakyushu Museum of Natural History and Human History, Higashida 2 - 4 - 1, Yahatahigashi-ku, Kitakyushu, Fukuoka 805 - 0071, Japan E-mail: shimomura @ kmnh. jp</mods:affiliation>
<mods:nameIdentifier id="3DFCF7392AC729F1124789905FFB6C65" type="email">shimomura@kmnh.jp</mods:nameIdentifier>
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<mods:title id="A06753A8AE2B6B12550F5F42173DEC51">Species Diversity</mods:title>
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<mods:date id="696D72876F116A61DC70D35DC7E112ED">2015</mods:date>
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<treatment id="03EC87FB3029FFE9FC0A139B7075F8FF" ID-DOI="http://doi.org/10.5281/zenodo.4733922" ID-GBIF-Taxon="181850835" ID-Zenodo-Dep="4733922" LSID="urn:lsid:zoobank.org:act:72957321-67AE-41C8-9560-B5CCBA22C97E" httpUri="http://treatment.plazi.org/id/03EC87FB3029FFE9FC0A139B7075F8FF" lastPageId="2" lastPageNumber="39" pageId="0" pageNumber="37">
<subSubSection id="C35F65663029FFEBFC0A139B70A8F859" pageId="0" pageNumber="37" type="nomenclature">
<subSubSection id="C35F65663029FFEBFC0A139B70C5F81F" box="[984,1268,1922,1946]" pageId="0" pageNumber="37" type="nomenclature">
<paragraph id="8BFA36ED3029FFEBFC0A139B70C5F81F" blockId="0.[910,1342,1857,2012]" box="[984,1268,1922,1946]" pageId="0" pageNumber="37">
<heading id="D0B281813029FFEBFC0A139B70C5F81F" box="[984,1268,1922,1946]" centered="true" fontSize="10" level="2" pageId="0" pageNumber="37" reason="2">
<taxonomicName id="4C454D6E3029FFEBFC0A139B7092F81F" ID-CoL="66RQ3" authority="Saito &amp; Shimomura, 2015" authorityName="Saito &amp; Shimomura" authorityYear="2015" box="[984,1187,1922,1946]" class="Malacostraca" family="Bopyridae" genus="Anisarthrus" kingdom="Animalia" order="Isopoda" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="okunoi" status="sp. nov.">
<emphasis id="B931EAFF3029FFEBFC0A139B7092F81F" bold="true" box="[984,1187,1922,1946]" italics="true" pageId="0" pageNumber="37">Anisarthrus okunoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20257843029FFEBFB7A139A70C5F81F" box="[1192,1268,1923,1946]" pageId="0" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="C35F65663029FFEBFC7213BD711CF83E" box="[928,1325,1955,1979]" pageId="0" pageNumber="37" type="vernacular_names">
<paragraph id="8BFA36ED3029FFEBFC7213BD711CF83E" blockId="0.[910,1342,1857,2012]" box="[928,1325,1955,1979]" pageId="0" pageNumber="37">[Japanese name: Sarasa-no-harayadori]</paragraph>
</subSubSection>
<subSubSection id="C35F65663029FFEBFBE113DD70A8F859" box="[1075,1177,1988,2012]" pageId="0" pageNumber="37" type="description">
<paragraph id="8BFA36ED3029FFEBFBE113DD70A8F859" blockId="0.[910,1342,1857,2012]" box="[1075,1177,1988,2012]" pageId="0" pageNumber="37">
(
<figureCitation id="137E2A683029FFEBFBE813DD70A0F859" box="[1082,1169,1988,2012]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="1.[156,187,1229,1251]" captionStartId-1="2.[156,187,1321,1343]" captionStartId-2="3.[156,187,1219,1241]" captionTargetBox-0="[318,1271,206,1188]" captionTargetBox-1="[312,1275,206,1281]" captionTargetBox-2="[284,1303,206,1179]" captionTargetId-0="figure-417@1.[314,1273,205,1190]" captionTargetId-1="figure-319@2.[311,1277,205,1283]" captionTargetId-2="figure-444@3.[283,1305,205,1181]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="Fig. 1. Anisarthrus okunoi sp. nov., holotype female, 3.88mm BL, KMNH IvR 500,727, Yura Harbor, Sumoto, Awaji Island, central Japan, 6m depth, originally attached to ventral surface of pleon of hinge-beak shrimp Rhynchocinetes uritai Kubo, 1942. A, dorsal view; B, ventral view, with partly hidden male (Ƌ; 1.12mm BL, KMNH IvR 500,728); C, pleon, ventral. Abbreviations: lp-14, lateral plates 14; oo-25, oostegites 25; p-34, pereopods 34; plm-5, pleomere 5 (=pleotelson); plp-14, pleopods 14." captionText-1="Fig. 2. Anisarthrus okunoi sp. nov., holotype female, 3.88mm BL, KMNH IvR 500,727. A, right antenna 1, dorsal view; B, right antenna 2, dorsal view; C, left maxilliped, ventral view; D, right maxilliped, ventral view; E, barbula, ventral view; F, left oostegite 1, ventral view; G, right oostegite 1, ventral view; H, left pereopod 1, lateral view; I, left pereopods 47, with pleopod 1 removed, ventral view." captionText-2="Fig. 3. Anisarthrus okunoi sp. nov., allotype male, 1.12mm BL, KMNH IvR 500,728, originally attached to ventral surface of pleon of holotype female. A, dorsal view; B, ventral view; C, left lateral view; D, anteroventral view of head with right antennae 1 and 2; E, left pereopod 1, lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.4584969" figureDoi-1="http://doi.org/10.5281/zenodo.4584973" figureDoi-2="http://doi.org/10.5281/zenodo.4584975" httpUri-0="https://zenodo.org/record/4584969/files/figure.png" httpUri-1="https://zenodo.org/record/4584973/files/figure.png" httpUri-2="https://zenodo.org/record/4584975/files/figure.png" pageId="0" pageNumber="37">Figs 13</figureCitation>
)
</paragraph>
</subSubSection>
</subSubSection>
<subSubSection id="C35F65663029FFEAFC9C1C1D751BF9D0" lastPageId="1" lastPageNumber="38" pageId="0" pageNumber="37" type="materials_examined">
<paragraph id="8BFA36ED3029FFEAFC9C1C1D751BF9D0" blockId="0.[846,1432,2052,2077]" lastBlockId="1.[156,768,1369,2077]" lastPageId="1" lastPageNumber="38" pageId="0" pageNumber="37">
<emphasis id="B931EAFF3029FFEBFC9C1C1D7012F799" bold="true" box="[846,1059,2052,2076]" pageId="0" pageNumber="37">Material examined.</emphasis>
<typeStatus id="54FE884F3029FFEBFBEF1C1C70AFF798" box="[1085,1182,2053,2077]" pageId="0" pageNumber="37" type="holotype">Holotype</typeStatus>
: ovigerous female (BL
<materialsCitation id="3B2D3CB03028FFEAFF4E1143761EF990" ID-GBIF-Occurrence="3097560302" collectingDate="2008-10-22" collectionCode="T" collectorName="J. Okuno &amp; Okuno, K &amp; Tamego." country="Japan" county="Awaji Island" elevation="6" latitude="34.291668" location="Yura Harbor" longLatPrecision="19" longitude="134.95" municipality="Sumoto" pageId="1" pageNumber="38" specimenCode="CMNH-ZC 02342" specimenCount="2" stateProvince="Hyogo" typeStatus="allotype">
3.88 mm), KMNH IvR 500,727, removed from ventral surface of pleon of hinge-beak shrimp
<taxonomicName id="4C454D6E3028FFEAFDF6116376CEFA17" authorityName="Kubo" authorityYear="1942" box="[548,767,1402,1426]" class="Malacostraca" family="Rhynchocinetidae" genus="Rhynchocinetes" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="38" phylum="Arthropoda" rank="species" species="uritai">
<emphasis id="B931EAFF3028FFEAFDF6116376CEFA17" box="[548,767,1402,1426]" italics="true" pageId="1" pageNumber="38">Rhynchocinetes uritai</emphasis>
</taxonomicName>
(female, CL 6.98 mm),
<specimenCode id="DBE39E963028FFEAFE4D11827652FA36" box="[415,611,1435,1459]" collectionCode="CMNH-ZC" pageId="1" pageNumber="38">CMNH-ZC 02342</specimenCode>
,
<location id="8E9A60363028FFEAFDA4118276CEFA36" LSID="urn:lsid:plazi:treatment:03EC87FB3029FFE9FC0A139B7075F8FF:8E9A60363028FFEAFDA4118276CEFA36" box="[630,767,1435,1459]" country="Japan" county="Awaji Island" latitude="34.291668" longLatPrecision="19" longitude="134.95" municipality="Sumoto" name="Yura Harbor" pageId="1" pageNumber="38" stateProvince="Hyogo">Yura Harbor</location>
(
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,
<geoCoordinate id="EE71502A3028FFEAFEF611A27590FA56" box="[292,417,1466,1491]" degrees="134" direction="east" minutes="57" orientation="longitude" pageId="1" pageNumber="38" precision="15" seconds="00" value="134.95">134°5700″E</geoCoordinate>
),
<collectingMunicipality id="6B9EAC973028FFEAFE6111A27630FA56" box="[435,513,1467,1491]" pageId="1" pageNumber="38">Sumoto</collectingMunicipality>
city,
<collectingCounty id="629B4E613028FFEAFDE711A27683FA56" box="[565,690,1467,1491]" pageId="1" pageNumber="38">Awaji Island</collectingCounty>
,
<collectingRegion id="4981F80F3028FFEAFD6E11A57537FA71" country="Japan" name="Hyogo" pageId="1" pageNumber="38">Hyogo Prefecture</collectingRegion>
, central
<collectingCountry id="F352767D3028FFEAFE8D11C575ABFA76" box="[351,410,1500,1523]" name="Japan" pageId="1" pageNumber="38">Japan</collectingCountry>
,
<elevation id="0068D1DE3028FFEAFE7411C5763FFA71" box="[422,526,1500,1524]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="38" unit="m" value="6.0">
<quantity id="4CBD9B083028FFEAFE7411C575FBFA76" box="[422,458,1500,1524]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="38" unit="m" value="6.0">6 m</quantity>
depth
</elevation>
,
<date id="FFFB102D3028FFEAFDC811C576F8FA71" box="[538,713,1500,1524]" pageId="1" pageNumber="38" value="2008-10-22">
<collectingDate id="EFBFE9C53028FFEAFDC811C576F8FA71" box="[538,713,1500,1524]" pageId="1" pageNumber="38" value="2008-10-22">22 October 2008</collectingDate>
</date>
, coll.
<collectorName id="26B0533B3028FFEAFF4E11E474CAF991" box="[156,251,1532,1556]" pageId="1" pageNumber="38">J. Okuno</collectorName>
, K. Hanano, and
<collectionCode id="ED54AE283028FFEAFE6911E475FAF991" box="[443,459,1533,1556]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="38">T</collectionCode>
.
<collectorName id="26B0533B3028FFEAFE0A11E4761EF990" box="[472,559,1533,1557]" pageId="1" pageNumber="38">Tamego.</collectorName>
</materialsCitation>
<materialsCitation id="3B2D3CB03028FFEAFDEA11E5751BF9D0" ID-GBIF-Occurrence="3097560301" collectingDate="2008-10-22" collectionCode="T" collectorName="J. Okuno &amp; Okuno, K &amp; Tamego." country="Japan" county="Awaji Island" elevation="6" latitude="34.291668" location="Yura Harbor" longLatPrecision="19" longitude="134.95" municipality="Sumoto" pageId="1" pageNumber="38" specimenCode="CMNH-ZC 02342" specimenCount="2" stateProvince="Hyogo" typeStatus="allotype">
<typeStatus id="54FE884F3028FFEAFDEA11E576A3F991" box="[568,658,1532,1556]" pageId="1" pageNumber="38" type="allotype">Allotype</typeStatus>
: male (BL 1.12 mm), KMNH IvR 500,728, removed from pleon of holotype female.
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="DF3A66653028FFEAFF4E10D477D2FAB3" ID-DOI="http://doi.org/10.5281/zenodo.4584969" ID-Zenodo-Dep="4584969" httpUri="https://zenodo.org/record/4584969/files/figure.png" pageId="1" pageNumber="38" startId="1.[156,187,1229,1251]" targetBox="[318,1271,206,1188]" targetPageId="1">
<paragraph id="8BFA36ED3028FFEAFF4E10D477D2FAB3" blockId="1.[156,1432,1229,1335]" pageId="1" pageNumber="38">
Fig. 1.
<taxonomicName id="4C454D6E3028FFEAFF2210D475ACFB67" authority="Saito &amp; Shimomura, 2015" authorityName="Saito &amp; Shimomura" authorityYear="2015" box="[240,413,1229,1250]" class="Malacostraca" family="Bopyridae" genus="Anisarthrus" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="38" phylum="Arthropoda" rank="species" species="okunoi" status="sp. nov.">
<emphasis id="B931EAFF3028FFEAFF2210D475ACFB67" box="[240,413,1229,1250]" italics="true" pageId="1" pageNumber="38">Anisarthrus okunoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20257843028FFEAFE7610D475DAFB67" box="[420,491,1229,1250]" pageId="1" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
, holotype female, 3.88mm BL, KMNH IvR 500,727, Yura Harbor, Sumoto, Awaji Island, central Japan, 6 m depth, originally attached to ventral surface of pleon of hinge-beak shrimp
<taxonomicName id="4C454D6E3028FFEAFCA410F07096FB7B" authority="Kubo, 1942" authorityName="Kubo" authorityYear="1942" box="[886,1191,1257,1279]" class="Malacostraca" family="Rhynchocinetidae" genus="Rhynchocinetes" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="38" phylum="Arthropoda" rank="species" species="uritai">
<emphasis id="B931EAFF3028FFEAFCA410F07007FB7B" box="[886,1078,1257,1278]" italics="true" pageId="1" pageNumber="38">Rhynchocinetes uritai</emphasis>
Kubo, 1942
</taxonomicName>
. A, dorsal view; B, ventral view, with partly hidden male (
<emphasis id="B931EAFF3028FFEAFE6A111F75F8FA99" box="[440,457,1286,1308]" italics="true" pageId="1" pageNumber="38">Ƌ</emphasis>
; 1.12 mm BL, KMNH IvR 500,728); C, pleon, ventral. Abbreviations: lp-14, lateral plates 14; oo-25, oostegites 25; p-34, pereopods 34; plm-5, pleomere 5 (=pleotelson); plp-14, pleopods 14.
</paragraph>
</caption>
<subSubSection id="C35F65663028FFE9FF67124777E6F89D" lastPageId="2" lastPageNumber="39" pageId="1" pageNumber="38" type="description">
<paragraph id="8BFA36ED3028FFEAFF67124776CBF97D" blockId="1.[156,768,1369,2077]" pageId="1" pageNumber="38">
<emphasis id="B931EAFF3028FFEAFF671247763BF9F3" bold="true" box="[181,522,1630,1654]" pageId="1" pageNumber="38">
Description of
<typeStatus id="54FE884F3028FFEAFE8E12477588F9F3" box="[348,441,1630,1654]" pageId="1" pageNumber="38" type="holotype">holotype</typeStatus>
female
</emphasis>
(KMNH IvR 500,727). Body (
<figureCitation id="137E2A683028FFEAFF3212677578F913" box="[224,329,1662,1686]" captionStart="Fig" captionStartId="1.[156,187,1229,1251]" captionTargetBox="[318,1271,206,1188]" captionTargetId="figure-417@1.[314,1273,205,1190]" captionTargetPageId="1" captionText="Fig. 1. Anisarthrus okunoi sp. nov., holotype female, 3.88mm BL, KMNH IvR 500,727, Yura Harbor, Sumoto, Awaji Island, central Japan, 6m depth, originally attached to ventral surface of pleon of hinge-beak shrimp Rhynchocinetes uritai Kubo, 1942. A, dorsal view; B, ventral view, with partly hidden male (Ƌ; 1.12mm BL, KMNH IvR 500,728); C, pleon, ventral. Abbreviations: lp-14, lateral plates 14; oo-25, oostegites 25; p-34, pereopods 34; plm-5, pleomere 5 (=pleotelson); plp-14, pleopods 14." figureDoi="http://doi.org/10.5281/zenodo.4584969" httpUri="https://zenodo.org/record/4584969/files/figure.png" pageId="1" pageNumber="38">Fig. 1A, B</figureCitation>
) markedly asymmetrical, greatly expanded on convex (left) side, approximating an inverted equilateral triangle in general outline; body axis distortion about 40º; maximal body width about 1.2 times as long as body length.
</paragraph>
<paragraph id="8BFA36ED3028FFEAFF671319700BFA56" blockId="1.[156,768,1369,2077]" lastBlockId="1.[821,1432,1369,2077]" pageId="1" pageNumber="38">
Head (
<figureCitation id="137E2A683028FFEAFF2B13197574F89C" box="[249,325,1792,1817]" captionStart="Fig" captionStartId="1.[156,187,1229,1251]" captionTargetBox="[318,1271,206,1188]" captionTargetId="figure-417@1.[314,1273,205,1190]" captionTargetPageId="1" captionText="Fig. 1. Anisarthrus okunoi sp. nov., holotype female, 3.88mm BL, KMNH IvR 500,727, Yura Harbor, Sumoto, Awaji Island, central Japan, 6m depth, originally attached to ventral surface of pleon of hinge-beak shrimp Rhynchocinetes uritai Kubo, 1942. A, dorsal view; B, ventral view, with partly hidden male (Ƌ; 1.12mm BL, KMNH IvR 500,728); C, pleon, ventral. Abbreviations: lp-14, lateral plates 14; oo-25, oostegites 25; p-34, pereopods 34; plm-5, pleomere 5 (=pleotelson); plp-14, pleopods 14." figureDoi="http://doi.org/10.5281/zenodo.4584969" httpUri="https://zenodo.org/record/4584969/files/figure.png" pageId="1" pageNumber="38">Fig. 1A</figureCitation>
) quadrilateral, completely embedded in pereon, representing about 35% of body length; frontal margin concave and deeply cleft. Eyes absent. Right antenna 1 (
<figureCitation id="137E2A683028FFEAFD0A1358748AF8FF" captionStart="Fig" captionStartId="2.[156,187,1321,1343]" captionTargetBox="[312,1275,206,1281]" captionTargetId="figure-319@2.[311,1277,205,1283]" captionTargetPageId="2" captionText="Fig. 2. Anisarthrus okunoi sp. nov., holotype female, 3.88mm BL, KMNH IvR 500,727. A, right antenna 1, dorsal view; B, right antenna 2, dorsal view; C, left maxilliped, ventral view; D, right maxilliped, ventral view; E, barbula, ventral view; F, left oostegite 1, ventral view; G, right oostegite 1, ventral view; H, left pereopod 1, lateral view; I, left pereopods 47, with pleopod 1 removed, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4584973" httpUri="https://zenodo.org/record/4584973/files/figure.png" pageId="1" pageNumber="38">Fig. 2A</figureCitation>
) of 3 articles, articles 2 and 3 with distal and terminal setae, respectively; right antenna 2 (
<figureCitation id="137E2A683028FFEAFDDE139A7666F81E" box="[524,599,1923,1947]" captionStart="Fig" captionStartId="2.[156,187,1321,1343]" captionTargetBox="[312,1275,206,1281]" captionTargetId="figure-319@2.[311,1277,205,1283]" captionTargetPageId="2" captionText="Fig. 2. Anisarthrus okunoi sp. nov., holotype female, 3.88mm BL, KMNH IvR 500,727. A, right antenna 1, dorsal view; B, right antenna 2, dorsal view; C, left maxilliped, ventral view; D, right maxilliped, ventral view; E, barbula, ventral view; F, left oostegite 1, ventral view; G, right oostegite 1, ventral view; H, left pereopod 1, lateral view; I, left pereopods 47, with pleopod 1 removed, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4584973" httpUri="https://zenodo.org/record/4584973/files/figure.png" pageId="1" pageNumber="38">Fig. 2B</figureCitation>
) of 5 articles, of which articles 2, 3, and 4 with marginal and distal setae, article 5 with terminal setae. Right and left maxillipeds (
<figureCitation id="137E2A683028FFEAFD0513DD74EAF879" captionStart="Fig" captionStartId="2.[156,187,1321,1343]" captionTargetBox="[312,1275,206,1281]" captionTargetId="figure-319@2.[311,1277,205,1283]" captionTargetPageId="2" captionText="Fig. 2. Anisarthrus okunoi sp. nov., holotype female, 3.88mm BL, KMNH IvR 500,727. A, right antenna 1, dorsal view; B, right antenna 2, dorsal view; C, left maxilliped, ventral view; D, right maxilliped, ventral view; E, barbula, ventral view; F, left oostegite 1, ventral view; G, right oostegite 1, ventral view; H, left pereopod 1, lateral view; I, left pereopods 47, with pleopod 1 removed, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4584973" httpUri="https://zenodo.org/record/4584973/files/figure.png" pageId="1" pageNumber="38">Fig. 2C, D</figureCitation>
) symmetrical, semi-ovoidal, nearly straight medially, roundly angled anteriorly, without palp, bearing short spur; anterior article about 2.4 times as long as posterior article. Barbula (
<figureCitation id="137E2A683028FFEAFC43116377EAFA17" box="[913,987,1402,1426]" captionStart="Fig" captionStartId="2.[156,187,1321,1343]" captionTargetBox="[312,1275,206,1281]" captionTargetId="figure-319@2.[311,1277,205,1283]" captionTargetPageId="2" captionText="Fig. 2. Anisarthrus okunoi sp. nov., holotype female, 3.88mm BL, KMNH IvR 500,727. A, right antenna 1, dorsal view; B, right antenna 2, dorsal view; C, left maxilliped, ventral view; D, right maxilliped, ventral view; E, barbula, ventral view; F, left oostegite 1, ventral view; G, right oostegite 1, ventral view; H, left pereopod 1, lateral view; I, left pereopods 47, with pleopod 1 removed, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4584973" httpUri="https://zenodo.org/record/4584973/files/figure.png" pageId="1" pageNumber="38">Fig. 2E</figureCitation>
) with pair of short, stout lateral projections, its lateral margins expanded and mid-region produced slightly into medial point.
</paragraph>
<paragraph id="8BFA36ED3028FFE9FC9C11C576EDFA36" blockId="1.[821,1432,1369,2077]" lastBlockId="2.[156,767,1435,2077]" lastPageId="2" lastPageNumber="39" pageId="1" pageNumber="38">
Pereon (
<figureCitation id="137E2A683028FFEAFC7111C577C3FA71" box="[931,1010,1500,1524]" captionStart="Fig" captionStartId="1.[156,187,1229,1251]" captionTargetBox="[318,1271,206,1188]" captionTargetId="figure-417@1.[314,1273,205,1190]" captionTargetPageId="1" captionText="Fig. 1. Anisarthrus okunoi sp. nov., holotype female, 3.88mm BL, KMNH IvR 500,727, Yura Harbor, Sumoto, Awaji Island, central Japan, 6m depth, originally attached to ventral surface of pleon of hinge-beak shrimp Rhynchocinetes uritai Kubo, 1942. A, dorsal view; B, ventral view, with partly hidden male (Ƌ; 1.12mm BL, KMNH IvR 500,728); C, pleon, ventral. Abbreviations: lp-14, lateral plates 14; oo-25, oostegites 25; p-34, pereopods 34; plm-5, pleomere 5 (=pleotelson); plp-14, pleopods 14." figureDoi="http://doi.org/10.5281/zenodo.4584969" httpUri="https://zenodo.org/record/4584969/files/figure.png" pageId="1" pageNumber="38">Fig. 1A</figureCitation>
): pereomeres 1 and 2 completely distinct dorsally; pereomeres 37 distinct on concave (right) side and medially but segmental boundaries not clearly expressed on convex (left) side. All pereomeres with dorsolateral bosses on concave side. Inflated closed brood pouch (
<figureCitation id="137E2A683028FFEAFCEE126777BBF913" box="[828,906,1662,1686]" captionStart="Fig" captionStartId="1.[156,187,1229,1251]" captionTargetBox="[318,1271,206,1188]" captionTargetId="figure-417@1.[314,1273,205,1190]" captionTargetPageId="1" captionText="Fig. 1. Anisarthrus okunoi sp. nov., holotype female, 3.88mm BL, KMNH IvR 500,727, Yura Harbor, Sumoto, Awaji Island, central Japan, 6m depth, originally attached to ventral surface of pleon of hinge-beak shrimp Rhynchocinetes uritai Kubo, 1942. A, dorsal view; B, ventral view, with partly hidden male (Ƌ; 1.12mm BL, KMNH IvR 500,728); C, pleon, ventral. Abbreviations: lp-14, lateral plates 14; oo-25, oostegites 25; p-34, pereopods 34; plm-5, pleomere 5 (=pleotelson); plp-14, pleopods 14." figureDoi="http://doi.org/10.5281/zenodo.4584969" httpUri="https://zenodo.org/record/4584969/files/figure.png" pageId="1" pageNumber="38">Fig. 1B</figureCitation>
) expanded far laterally, formed of fused convex oostegites 25. Oostegite 1 of convex side (
<figureCitation id="137E2A683028FFEAFB3F12867106F932" box="[1261,1335,1695,1719]" captionStart="Fig" captionStartId="2.[156,187,1321,1343]" captionTargetBox="[312,1275,206,1281]" captionTargetId="figure-319@2.[311,1277,205,1283]" captionTargetPageId="2" captionText="Fig. 2. Anisarthrus okunoi sp. nov., holotype female, 3.88mm BL, KMNH IvR 500,727. A, right antenna 1, dorsal view; B, right antenna 2, dorsal view; C, left maxilliped, ventral view; D, right maxilliped, ventral view; E, barbula, ventral view; F, left oostegite 1, ventral view; G, right oostegite 1, ventral view; H, left pereopod 1, lateral view; I, left pereopods 47, with pleopod 1 removed, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4584973" httpUri="https://zenodo.org/record/4584973/files/figure.png" pageId="1" pageNumber="38">Fig. 2F</figureCitation>
) bilobed; that of concave side (
<figureCitation id="137E2A683028FFEAFBC012A67050F952" box="[1042,1121,1727,1751]" captionStart="Fig" captionStartId="2.[156,187,1321,1343]" captionTargetBox="[312,1275,206,1281]" captionTargetId="figure-319@2.[311,1277,205,1283]" captionTargetPageId="2" captionText="Fig. 2. Anisarthrus okunoi sp. nov., holotype female, 3.88mm BL, KMNH IvR 500,727. A, right antenna 1, dorsal view; B, right antenna 2, dorsal view; C, left maxilliped, ventral view; D, right maxilliped, ventral view; E, barbula, ventral view; F, left oostegite 1, ventral view; G, right oostegite 1, ventral view; H, left pereopod 1, lateral view; I, left pereopods 47, with pleopod 1 removed, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4584973" httpUri="https://zenodo.org/record/4584973/files/figure.png" pageId="1" pageNumber="38">Fig. 2G</figureCitation>
) oval, not segmented, lacking posterolateral point. Oostegites 25 of concave side all separated and aligned. Seven pereopods on each side of body; on both sides, first two pairs of pereopods adjacent to head: both pairs large and with all articles distinct, basis rectangular, ischium trapezoidal and shorter than basis, merus trapezoidal with short spines on inferior margin, carpus triangular, propodus ovate, dactylus stout (
<figureCitation id="137E2A683028FFEAFB2213BA7173F83E" box="[1264,1346,1955,1979]" captionStart="Fig" captionStartId="2.[156,187,1321,1343]" captionTargetBox="[312,1275,206,1281]" captionTargetId="figure-319@2.[311,1277,205,1283]" captionTargetPageId="2" captionText="Fig. 2. Anisarthrus okunoi sp. nov., holotype female, 3.88mm BL, KMNH IvR 500,727. A, right antenna 1, dorsal view; B, right antenna 2, dorsal view; C, left maxilliped, ventral view; D, right maxilliped, ventral view; E, barbula, ventral view; F, left oostegite 1, ventral view; G, right oostegite 1, ventral view; H, left pereopod 1, lateral view; I, left pereopods 47, with pleopod 1 removed, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4584973" httpUri="https://zenodo.org/record/4584973/files/figure.png" pageId="1" pageNumber="38">Fig. 2H</figureCitation>
). Pereopods 3 and 4 of convex side widely separated by expanded part of pereomere (
<figureCitation id="137E2A683028FFEAFC2813FD7078F879" box="[1018,1097,2020,2044]" captionStart="Fig" captionStartId="1.[156,187,1229,1251]" captionTargetBox="[318,1271,206,1188]" captionTargetId="figure-417@1.[314,1273,205,1190]" captionTargetPageId="1" captionText="Fig. 1. Anisarthrus okunoi sp. nov., holotype female, 3.88mm BL, KMNH IvR 500,727, Yura Harbor, Sumoto, Awaji Island, central Japan, 6m depth, originally attached to ventral surface of pleon of hinge-beak shrimp Rhynchocinetes uritai Kubo, 1942. A, dorsal view; B, ventral view, with partly hidden male (Ƌ; 1.12mm BL, KMNH IvR 500,728); C, pleon, ventral. Abbreviations: lp-14, lateral plates 14; oo-25, oostegites 25; p-34, pereopods 34; plm-5, pleomere 5 (=pleotelson); plp-14, pleopods 14." figureDoi="http://doi.org/10.5281/zenodo.4584969" httpUri="https://zenodo.org/record/4584969/files/figure.png" pageId="1" pageNumber="38">Fig. 1A</figureCitation>
). Pereopods 57 present on posterior margin of pleon, covered by left pleopod 1 (
<figureCitation id="137E2A683028FFEAFA961C1C71BAF798" box="[1348,1419,2053,2077]" captionStart="Fig" captionStartId="2.[156,187,1321,1343]" captionTargetBox="[312,1275,206,1281]" captionTargetId="figure-319@2.[311,1277,205,1283]" captionTargetPageId="2" captionText="Fig. 2. Anisarthrus okunoi sp. nov., holotype female, 3.88mm BL, KMNH IvR 500,727. A, right antenna 1, dorsal view; B, right antenna 2, dorsal view; C, left maxilliped, ventral view; D, right maxilliped, ventral view; E, barbula, ventral view; F, left oostegite 1, ventral view; G, right oostegite 1, ventral view; H, left pereopod 1, lateral view; I, left pereopods 47, with pleopod 1 removed, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4584973" httpUri="https://zenodo.org/record/4584973/files/figure.png" pageId="1" pageNumber="38">Fig. 2I</figureCitation>
). Pereopods 37 of concave side tightly clumped (
<figureCitation id="137E2A68302BFFE9FD50118276FEFA36" box="[642,719,1435,1459]" captionStart="Fig" captionStartId="1.[156,187,1229,1251]" captionTargetBox="[318,1271,206,1188]" captionTargetId="figure-417@1.[314,1273,205,1190]" captionTargetPageId="1" captionText="Fig. 1. Anisarthrus okunoi sp. nov., holotype female, 3.88mm BL, KMNH IvR 500,727, Yura Harbor, Sumoto, Awaji Island, central Japan, 6m depth, originally attached to ventral surface of pleon of hinge-beak shrimp Rhynchocinetes uritai Kubo, 1942. A, dorsal view; B, ventral view, with partly hidden male (Ƌ; 1.12mm BL, KMNH IvR 500,728); C, pleon, ventral. Abbreviations: lp-14, lateral plates 14; oo-25, oostegites 25; p-34, pereopods 34; plm-5, pleomere 5 (=pleotelson); plp-14, pleopods 14." figureDoi="http://doi.org/10.5281/zenodo.4584969" httpUri="https://zenodo.org/record/4584969/files/figure.png" pageId="2" pageNumber="39">Fig. 1A</figureCitation>
).
</paragraph>
<caption id="DF3A6665302BFFE9FF4E113070C1FAF2" ID-DOI="http://doi.org/10.5281/zenodo.4584973" ID-Zenodo-Dep="4584973" httpUri="https://zenodo.org/record/4584973/files/figure.png" pageId="2" pageNumber="39" startId="2.[156,187,1321,1343]" targetBox="[312,1275,206,1281]" targetPageId="2">
<paragraph id="8BFA36ED302BFFE9FF4E113070C1FAF2" blockId="2.[156,1432,1321,1399]" pageId="2" pageNumber="39">
Fig. 2.
<taxonomicName id="4C454D6E302BFFE9FF22113075AFFABB" authority="Saito &amp; Shimomura, 2015" authorityName="Saito &amp; Shimomura" authorityYear="2015" box="[240,414,1321,1342]" class="Malacostraca" family="Bopyridae" genus="Anisarthrus" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="39" phylum="Arthropoda" rank="species" species="okunoi" status="sp. nov.">
<emphasis id="B931EAFF302BFFE9FF22113075AFFABB" box="[240,414,1321,1342]" italics="true" pageId="2" pageNumber="39">Anisarthrus okunoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2025784302BFFE9FE77113375DCFABA" box="[421,493,1322,1343]" pageId="2" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
, holotype female, 3.88 mm BL, KMNH IvR 500,727. A, right antenna 1, dorsal view; B, right antenna 2, dorsal view; C, left maxilliped, ventral view; D, right maxilliped, ventral view; E, barbula, ventral view; F, left oostegite 1, ventral view; G, right oostegite 1, ventral view; H, left pereopod 1, lateral view; I, left pereopods 47, with pleopod 1 removed, ventral view.
</paragraph>
</caption>
<paragraph id="8BFA36ED302BFFE9FF6711A2764CF89D" blockId="2.[156,767,1435,2077]" pageId="2" pageNumber="39">
Pleon (
<figureCitation id="137E2A68302BFFE9FED111A2750EFA56" box="[259,319,1467,1491]" captionStart="Fig" captionStartId="1.[156,187,1229,1251]" captionTargetBox="[318,1271,206,1188]" captionTargetId="figure-417@1.[314,1273,205,1190]" captionTargetPageId="1" captionText="Fig. 1. Anisarthrus okunoi sp. nov., holotype female, 3.88mm BL, KMNH IvR 500,727, Yura Harbor, Sumoto, Awaji Island, central Japan, 6m depth, originally attached to ventral surface of pleon of hinge-beak shrimp Rhynchocinetes uritai Kubo, 1942. A, dorsal view; B, ventral view, with partly hidden male (Ƌ; 1.12mm BL, KMNH IvR 500,728); C, pleon, ventral. Abbreviations: lp-14, lateral plates 14; oo-25, oostegites 25; p-34, pereopods 34; plm-5, pleomere 5 (=pleotelson); plp-14, pleopods 14." figureDoi="http://doi.org/10.5281/zenodo.4584969" httpUri="https://zenodo.org/record/4584969/files/figure.png" pageId="2" pageNumber="39">Fig. 1</figureCitation>
AC): all pleomeres fused dorsally; pleomeres 14 distinct laterally and ventrally; pleomere 1 bearing single midventral tubercle, pleomere 2 with two similar ventral tubercles; pleomere 5 (= pleotelson) in form of simple bulb visible only in ventral view. Four pairs of large lamellar lateral plates and four pairs of pleopods present, all well-developed; lateral plates (
<figureCitation id="137E2A68302BFFE9FDE2126776B3F913" box="[560,642,1662,1686]" captionStart="Fig" captionStartId="1.[156,187,1229,1251]" captionTargetBox="[318,1271,206,1188]" captionTargetId="figure-417@1.[314,1273,205,1190]" captionTargetPageId="1" captionText="Fig. 1. Anisarthrus okunoi sp. nov., holotype female, 3.88mm BL, KMNH IvR 500,727, Yura Harbor, Sumoto, Awaji Island, central Japan, 6m depth, originally attached to ventral surface of pleon of hinge-beak shrimp Rhynchocinetes uritai Kubo, 1942. A, dorsal view; B, ventral view, with partly hidden male (Ƌ; 1.12mm BL, KMNH IvR 500,728); C, pleon, ventral. Abbreviations: lp-14, lateral plates 14; oo-25, oostegites 25; p-34, pereopods 34; plm-5, pleomere 5 (=pleotelson); plp-14, pleopods 14." figureDoi="http://doi.org/10.5281/zenodo.4584969" httpUri="https://zenodo.org/record/4584969/files/figure.png" pageId="2" pageNumber="39">Fig. 1A</figureCitation>
) irregularly oval, shorter posteriorly; those on concave side smaller than those opposite. Pleopods (
<figureCitation id="137E2A68302BFFE9FE2312A67673F95D" box="[497,578,1727,1752]" captionStart="Fig" captionStartId="1.[156,187,1229,1251]" captionTargetBox="[318,1271,206,1188]" captionTargetId="figure-417@1.[314,1273,205,1190]" captionTargetPageId="1" captionText="Fig. 1. Anisarthrus okunoi sp. nov., holotype female, 3.88mm BL, KMNH IvR 500,727, Yura Harbor, Sumoto, Awaji Island, central Japan, 6m depth, originally attached to ventral surface of pleon of hinge-beak shrimp Rhynchocinetes uritai Kubo, 1942. A, dorsal view; B, ventral view, with partly hidden male (Ƌ; 1.12mm BL, KMNH IvR 500,728); C, pleon, ventral. Abbreviations: lp-14, lateral plates 14; oo-25, oostegites 25; p-34, pereopods 34; plm-5, pleomere 5 (=pleotelson); plp-14, pleopods 14." figureDoi="http://doi.org/10.5281/zenodo.4584969" httpUri="https://zenodo.org/record/4584969/files/figure.png" pageId="2" pageNumber="39">Fig. 1C</figureCitation>
) uniramous, first two large, lanceolate; pleopods 3 and 4 slightly bilobed and smaller than pleopods 1 and 2. Uropods absent.
</paragraph>
<paragraph id="8BFA36ED302BFFE9FF67133875E6F81F" blockId="2.[156,767,1435,2077]" pageId="2" pageNumber="39">
<emphasis id="B931EAFF302BFFE9FF67133875CDF8BC" bold="true" box="[181,508,1825,1849]" pageId="2" pageNumber="39">
Description of
<typeStatus id="54FE884F302BFFE9FEB71338758BF8BC" box="[357,442,1825,1849]" pageId="2" pageNumber="39" type="allotype">allotype</typeStatus>
male
</emphasis>
(KMNH IvR 500,728). Body (
<figureCitation id="137E2A68302BFFE9FF0C13587527F8DF" box="[222,278,1857,1882]" captionStart="Fig" captionStartId="3.[156,187,1219,1241]" captionTargetBox="[284,1303,206,1179]" captionTargetId="figure-444@3.[283,1305,205,1181]" captionTargetPageId="3" captionText="Fig. 3. Anisarthrus okunoi sp. nov., allotype male, 1.12mm BL, KMNH IvR 500,728, originally attached to ventral surface of pleon of holotype female. A, dorsal view; B, ventral view; C, left lateral view; D, anteroventral view of head with right antennae 1 and 2; E, left pereopod 1, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4584975" httpUri="https://zenodo.org/record/4584975/files/figure.png" pageId="2" pageNumber="39">Fig. 3</figureCitation>
AC) small, about 29% as long as female, elliptical in general outline; body width 54% of body length, with maximal width at pereomere 5.
</paragraph>
<paragraph id="8BFA36ED302BFFE9FF6713BA77E3FA56" blockId="2.[156,767,1435,2077]" lastBlockId="2.[820,1432,1434,1914]" pageId="2" pageNumber="39">
Head (
<figureCitation id="137E2A68302BFFE9FF2E13BA757DF83E" box="[252,332,1955,1979]" captionStart="Fig" captionStartId="3.[156,187,1219,1241]" captionTargetBox="[284,1303,206,1179]" captionTargetId="figure-444@3.[283,1305,205,1181]" captionTargetPageId="3" captionText="Fig. 3. Anisarthrus okunoi sp. nov., allotype male, 1.12mm BL, KMNH IvR 500,728, originally attached to ventral surface of pleon of holotype female. A, dorsal view; B, ventral view; C, left lateral view; D, anteroventral view of head with right antennae 1 and 2; E, left pereopod 1, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4584975" httpUri="https://zenodo.org/record/4584975/files/figure.png" pageId="2" pageNumber="39">Fig. 3A</figureCitation>
) distinct from pereomere 1, semicircular, with tiny medial notch on anterior margin, slightly longer than pereomere 1. Eyes absent. Antenna 1 (
<figureCitation id="137E2A68302BFFE9FDB613FD7685F879" box="[612,692,2020,2044]" captionStart="Fig" captionStartId="3.[156,187,1219,1241]" captionTargetBox="[284,1303,206,1179]" captionTargetId="figure-444@3.[283,1305,205,1181]" captionTargetPageId="3" captionText="Fig. 3. Anisarthrus okunoi sp. nov., allotype male, 1.12mm BL, KMNH IvR 500,728, originally attached to ventral surface of pleon of holotype female. A, dorsal view; B, ventral view; C, left lateral view; D, anteroventral view of head with right antennae 1 and 2; E, left pereopod 1, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4584975" httpUri="https://zenodo.org/record/4584975/files/figure.png" pageId="2" pageNumber="39">Fig. 3D</figureCitation>
) short, of 3 articles; articles 2 and 3 each with tuft of long seta distally. Antenna 2 (
<figureCitation id="137E2A68302BFFE9FC3711837002FA36" box="[997,1075,1434,1459]" captionStart="Fig" captionStartId="3.[156,187,1219,1241]" captionTargetBox="[284,1303,206,1179]" captionTargetId="figure-444@3.[283,1305,205,1181]" captionTargetPageId="3" captionText="Fig. 3. Anisarthrus okunoi sp. nov., allotype male, 1.12mm BL, KMNH IvR 500,728, originally attached to ventral surface of pleon of holotype female. A, dorsal view; B, ventral view; C, left lateral view; D, anteroventral view of head with right antennae 1 and 2; E, left pereopod 1, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4584975" httpUri="https://zenodo.org/record/4584975/files/figure.png" pageId="2" pageNumber="39">Fig. 3D</figureCitation>
) of 7 articles, surpassing midpoint of pereomere 1.
</paragraph>
<paragraph id="8BFA36ED302BFFE9FC9C11C570A9F913" blockId="2.[820,1432,1434,1914]" pageId="2" pageNumber="39">
All pereomeres (
<figureCitation id="137E2A68302BFFE9FBC411C5706CFA71" box="[1046,1117,1500,1524]" captionStart="Fig" captionStartId="3.[156,187,1219,1241]" captionTargetBox="[284,1303,206,1179]" captionTargetId="figure-444@3.[283,1305,205,1181]" captionTargetPageId="3" captionText="Fig. 3. Anisarthrus okunoi sp. nov., allotype male, 1.12mm BL, KMNH IvR 500,728, originally attached to ventral surface of pleon of holotype female. A, dorsal view; B, ventral view; C, left lateral view; D, anteroventral view of head with right antennae 1 and 2; E, left pereopod 1, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4584975" httpUri="https://zenodo.org/record/4584975/files/figure.png" pageId="2" pageNumber="39">Fig. 3</figureCitation>
AC) distinctly demarcated dorsally, lacking tubercles ventrally; lateral margins convex; pereomere 7 and pleon distinct laterally on both sides, fused in central region both dorsally and ventrally. Pereopods (
<figureCitation id="137E2A68302BFFE9FCEE124777B6F9F3" box="[828,903,1630,1654]" captionStart="Fig" captionStartId="3.[156,187,1219,1241]" captionTargetBox="[284,1303,206,1179]" captionTargetId="figure-444@3.[283,1305,205,1181]" captionTargetPageId="3" captionText="Fig. 3. Anisarthrus okunoi sp. nov., allotype male, 1.12mm BL, KMNH IvR 500,728, originally attached to ventral surface of pleon of holotype female. A, dorsal view; B, ventral view; C, left lateral view; D, anteroventral view of head with right antennae 1 and 2; E, left pereopod 1, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4584975" httpUri="https://zenodo.org/record/4584975/files/figure.png" pageId="2" pageNumber="39">Fig. 3E</figureCitation>
) all of similar size and structure; all articles similar to those of female in segmentation.
</paragraph>
<paragraph id="8BFA36ED302BFFE9FC9C128677E6F89D" blockId="2.[820,1432,1434,1914]" pageId="2" pageNumber="39">
Pleon (
<figureCitation id="137E2A68302BFFE9FC4412867033F932" box="[918,1026,1695,1719]" captionStart="Fig" captionStartId="3.[156,187,1219,1241]" captionTargetBox="[284,1303,206,1179]" captionTargetId="figure-444@3.[283,1305,205,1181]" captionTargetPageId="3" captionText="Fig. 3. Anisarthrus okunoi sp. nov., allotype male, 1.12mm BL, KMNH IvR 500,728, originally attached to ventral surface of pleon of holotype female. A, dorsal view; B, ventral view; C, left lateral view; D, anteroventral view of head with right antennae 1 and 2; E, left pereopod 1, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4584975" httpUri="https://zenodo.org/record/4584975/files/figure.png" pageId="2" pageNumber="39">Fig. 3A, C</figureCitation>
) completely fused, semi-ovoidal, representing 27% of whole body length; posterior margin rounded, smooth, without anal tube or long taper. Pleopods and uropods absent.
</paragraph>
</subSubSection>
<subSubSection id="C35F6566302BFFE9FC9C13387075F8FF" pageId="2" pageNumber="39" type="etymology">
<paragraph id="8BFA36ED302BFFE9FC9C13387075F8FF" blockId="2.[820,1432,1434,1914]" pageId="2" pageNumber="39">
<emphasis id="B931EAFF302BFFE9FC9C133877F4F8BC" bold="true" box="[846,965,1825,1849]" pageId="2" pageNumber="39">Etymology.</emphasis>
Named in honour of the collector, Dr. Junji Okuno, in recognition of his contribution to the knowledge of the hinge-beak shrimps.
</paragraph>
</subSubSection>
</treatment>
</document>