treatments-xml/data/03/EC/87/03EC87CD7B18FFFBBDE7FA3FFF14FC17.xml
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<document id="5333FC220E6B734B421C909F93E0EC08" ID-DOI="10.11646/zootaxa.4748.2.11" ID-GBIF-Dataset="fd9edc0b-4e36-4fa6-b4e9-33599b22f984" ID-ISSN="1175-5326" ID-Zenodo-Dep="3698739" ID-ZooBank="B3F989CF-A760-46B2-8619-B0A633DAA0B6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1583482537524" checkinUser="plazi" docAuthor="Anichtchenko, Alexander &amp; Medina, Milton Norman" docDate="2020" docId="03EC87CD7B18FFFBBDE7FA3FFF14FC17" docLanguage="en" docName="zootaxa.4748.2.11.pdf" docOrigin="Zootaxa 4748 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Thopeutica wiesneri Anichtchenko &amp; Medina 2020, sp. nov." docType="treatment" docVersion="5" lastPageNumber="395" masterDocId="FFD5FFB57B1AFFFDBD70FF97FFD3FFF8" masterDocTitle="Contribution to the knowledge of three species of the genus Thopeutica Schaum, 1861 from the Philippines (Coleoptera: Carabidae: Cicindelinae)" masterLastPageNumber="395" masterPageNumber="389" pageNumber="391" updateTime="1698725122065" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="C0B13B9D0992252B2D14A94C07BC9CD9">Contribution to the knowledge of three species of the genus Thopeutica Schaum, 1861 from the Philippines (Coleoptera: Carabidae: Cicindelinae)</mods:title>
</mods:titleInfo>
<mods:name id="497CA0E5D91CC7FB2F2BB0B13CD51D78" type="personal">
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<mods:namePart id="04232409E76D493CE257655E005F9C8C">Anichtchenko, Alexander</mods:namePart>
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<mods:namePart id="70293DC12AA9B3B874C4555DAA82B141">Medina, Milton Norman</mods:namePart>
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<mods:title id="31DD8FE213C406EF1647F2D35109C1B6">Zootaxa</mods:title>
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<mods:part id="CE2A1EE0ECB7986D8FA02D517D50D768">
<mods:date id="3D75642F647B838B880ED56A4CC2516B">2020</mods:date>
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<mods:number id="9F9FD0BC04A816F054AE07C65C121DFE">2020-03-06</mods:number>
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<mods:number id="176114A883A6044DC4204F5EE1B66059">4748</mods:number>
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<treatment id="03EC87CD7B18FFFBBDE7FA3FFF14FC17" ID-DOI="http://doi.org/10.5281/zenodo.3705120" ID-GBIF-Taxon="162462215" ID-Zenodo-Dep="3705120" LSID="urn:lsid:plazi:treatment:03EC87CD7B18FFFBBDE7FA3FFF14FC17" httpUri="http://treatment.plazi.org/id/03EC87CD7B18FFFBBDE7FA3FFF14FC17" lastPageId="6" lastPageNumber="395" pageId="2" pageNumber="391">
<subSubSection id="C35F65507B18FFFFBDE7FA3FFD95FA3B" box="[151,582,1448,1475]" pageId="2" pageNumber="391" type="nomenclature">
<paragraph id="8BFA36DB7B18FFFFBDE7FA3FFD95FA3B" blockId="2.[151,582,1448,1475]" box="[151,582,1448,1475]" pageId="2" pageNumber="391">
<heading id="D0B281B77B18FFFFBDE7FA3FFD95FA3B" bold="true" box="[151,582,1448,1475]" fontSize="11" level="1" pageId="2" pageNumber="391" reason="1">
<emphasis id="B931EAC97B18FFFFBDE7FA3FFD95FA3B" bold="true" box="[151,582,1448,1475]" pageId="2" pageNumber="391">
<taxonomicName id="4C454D587B18FFFFBDE7FA3FFE32FA3B" ID-CoL="8MJ4B" authority="Anichtchenko &amp; Medina, 2020" authorityName="Anichtchenko &amp; Medina" authorityYear="2020" box="[151,481,1448,1475]" class="Insecta" family="Carabidae" genus="Thopeutica" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="391" phylum="Arthropoda" rank="species" species="wiesneri" status="sp. nov.">
<emphasis id="B931EAC97B18FFFFBDE7FA3FFEF3FA3B" bold="true" box="[151,288,1448,1475]" italics="true" pageId="2" pageNumber="391">Thopeutica</emphasis>
(s.str.)
<emphasis id="B931EAC97B18FFFFBC0CFA3EFE32FA3B" bold="true" box="[380,481,1449,1475]" italics="true" pageId="2" pageNumber="391">wiesneri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20257B27B18FFFFBC98FA3EFD95FA3B" box="[488,582,1449,1475]" pageId="2" pageNumber="391" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35F65507B18FFFFBDE7FA66FD1CF91B" pageId="2" pageNumber="391" type="materials_examined">
<paragraph id="8BFA36DB7B18FFFFBDE7FA66FD1CF91B" blockId="2.[151,1437,1520,2014]" pageId="2" pageNumber="391">
<emphasis id="B931EAC97B18FFFFBDE7FA66FE9AF9F3" bold="true" box="[151,329,1520,1547]" pageId="2" pageNumber="391">Type material.</emphasis>
<materialsCitation id="3B2D3C867B18FFFFBC24FA67FCB2F9D7" ID-GBIF-Occurrence="2574624210" collectingDate="2019-04-23" collectionCode="DUBC" collectorName="M. N. Medina" country="Philippines" county="Mindanao" elevation="1785" location="Cagan" municipality="New" pageId="2" pageNumber="391" specimenCount="1" stateProvince="Bataan" typeStatus="holotype">
<typeStatus id="54FE88797B18FFFFBC24FA67FE6CF9F2" box="[340,447,1520,1546]" pageId="2" pageNumber="391" type="holotype">Holotype</typeStatus>
, male: “
<collectingCountry id="F352764B7B18FFFFBF51FA67FD76F9F2" box="[545,677,1520,1546]" name="Philippines" pageId="2" pageNumber="391">Philippines</collectingCountry>
,
<collectingCounty id="629B4E577B18FFFFBFC4FA67FCFBF9F2" box="[692,808,1520,1546]" pageId="2" pageNumber="391">Mindanao</collectingCounty>
,
<collectingMunicipality id="6B9EACA17B18FFFFBE48FA66FCBEF9F3" box="[824,877,1521,1547]" pageId="2" pageNumber="391">New</collectingMunicipality>
<collectingRegion id="4981F8397B18FFFFBE08FA66FC1AF9F3" box="[888,969,1521,1547]" country="Philippines" name="Bataan" pageId="2" pageNumber="391">Bataan</collectingRegion>
,
<location id="8E9A60007B18FFFFBEA8FA66FBF1F9F3" LSID="urn:lsid:plazi:treatment:03EC87CD7B18FFFBBDE7FA3FFF14FC17:8E9A60007B18FFFFBEA8FA66FBF1F9F3" box="[984,1058,1521,1547]" country="Philippines" county="Mindanao" municipality="New" name="Cagan" pageId="2" pageNumber="391" stateProvince="Bataan">Cagan</location>
, 7˚2423.68”N, 126˚1334.81”E,
<quantity id="4CBD9B3E7B18FFFFBDE7F983FF3DF9D7" box="[151,238,1556,1583]" metricMagnitude="3" metricUnit="m" metricValue="1.785" pageId="2" pageNumber="391" unit="m" value="1785.0">
<elevation id="0068D1E87B18FFFFBDE7F983FF3DF9D7" box="[151,238,1556,1583]" metricMagnitude="3" metricUnit="m" metricValue="1.785" pageId="2" pageNumber="391" unit="m" value="1785.0">1785 m</elevation>
</quantity>
,
<date id="FFFB101B7B18FFFFBD89F983FE72F9D7" box="[249,417,1556,1583]" pageId="2" pageNumber="391" value="2019-04-23">
<collectingDate id="EFBFE9F37B18FFFFBD89F983FE72F9D7" box="[249,417,1556,1583]" pageId="2" pageNumber="391" value="2019-04-23">April 23, 2019</collectingDate>
</date>
,
<collectingMethod id="52044ECC7B18FFFFBCDCF983FDC6F9D7" box="[428,533,1556,1583]" pageId="2" pageNumber="391">light trap</collectingMethod>
,
<collectorName id="26B0530D7B18FFFFBF50F982FD6AF9D6" box="[544,697,1556,1583]" pageId="2" pageNumber="391">M.N. Medina</collectorName>
leg.” (
<collectionCode id="ED54AE1E7B18FFFFBE74F982FC8BF9D7" box="[772,856,1557,1583]" pageId="2" pageNumber="391">DUBC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B2D3C867B18FFFFBE1BF982FD18F91B" ID-GBIF-Occurrence="2574624204" collectingDate="2014-06" collectingDateMax="2019-04" collectingDateMin="2014-06" collectionCode="UMCRC" collectorName="M. Apo" country="Philippines" location="City" municipality="Mindanao" pageId="2" pageNumber="391" specimenCount="11" specimenCount-female="7" specimenCount-male="4" stateProvince="Davao" typeStatus="paratype">
<typeStatus id="54FE88797B18FFFFBE1BF982FC0FF9D7" box="[875,988,1557,1583]" pageId="2" pageNumber="391" type="paratype">Paratypes</typeStatus>
: same labeled as the
<typeStatus id="54FE88797B18FFFFB9B8F983FAF9F9D6" box="[1224,1322,1556,1582]" pageId="2" pageNumber="391" type="holotype">holotype</typeStatus>
(
<specimenCount id="9D43FD527B18FFFFB84DF982FA4FF9D6" box="[1341,1436,1556,1583]" pageId="2" pageNumber="391" type="female">1 female</specimenCount>
in cJW,
<specimenCount id="9D43FD527B18FFFFBD85F9AEFE91F9AA" box="[245,322,1592,1619]" pageId="2" pageNumber="391" type="male">1 male</specimenCount>
and
<specimenCount id="9D43FD527B18FFFFBC0CF9AEFE0CF9AA" box="[380,479,1592,1619]" pageId="2" pageNumber="391" type="female">1 female</specimenCount>
in
<collectionCode id="ED54AE1E7B18FFFFBF76F9AEFDA3F9AB" box="[518,624,1593,1619]" pageId="2" pageNumber="391">UMCRC</collectionCode>
); “
<collectingMunicipality id="6B9EACA17B18FFFFBFFFF9AFFCD5F9AA" box="[655,774,1592,1618]" pageId="2" pageNumber="391">Mindanao</collectingMunicipality>
,
<collectingRegion id="4981F8397B18FFFFBE63F9AEFC8FF9AB" box="[787,860,1593,1619]" country="Philippines" name="Davao" pageId="2" pageNumber="391">Davao</collectingRegion>
<location id="8E9A60007B18FFFFBE15F9AFFC45F9AA" LSID="urn:lsid:plazi:treatment:03EC87CD7B18FFFBBDE7FA3FFF14FC17:8E9A60007B18FFFFBE15F9AFFC45F9AA" box="[869,918,1592,1618]" country="Philippines" municipality="Mindanao" name="City" pageId="2" pageNumber="391" stateProvince="Davao">City</location>
,
<date id="FFFB101B7B18FFFFBED3F9AEFC2CF9AB" box="[931,1023,1593,1619]" pageId="2" pageNumber="391" value="2019-04">
<collectingDate id="EFBFE9F37B18FFFFBED3F9AEFC2CF9AB" box="[931,1023,1593,1619]" pageId="2" pageNumber="391" value="2019-04">IV.2019</collectingDate>
</date>
” (
<specimenCount id="9D43FD527B18FFFFB96CF9AEFBB9F9AA" box="[1052,1130,1592,1619]" pageId="2" pageNumber="391" type="male">1 male</specimenCount>
,
<specimenCount id="9D43FD527B18FFFFB908F9AEFB09F9AA" box="[1144,1242,1592,1619]" pageId="2" pageNumber="391" type="female">1 female</specimenCount>
in cRS), “Mindanao, Panamokan,
<collectingRegion id="4981F8397B18FFFFBC1DF9CBFE0DF98E" box="[365,478,1628,1654]" country="Philippines" name="Bukidnon" pageId="2" pageNumber="391">Bukidnon</collectingRegion>
,
<date id="FFFB101B7B18FFFFBC9AF9CAFD8FF98F" box="[490,604,1629,1655]" pageId="2" pageNumber="391" value="2014-08">
<collectingDate id="EFBFE9F37B18FFFFBC9AF9CAFD8FF98F" box="[490,604,1629,1655]" pageId="2" pageNumber="391" value="2014-08">VIII.2014</collectingDate>
</date>
” (
<specimenCount id="9D43FD527B18FFFFBF09F9CAFD0AF98E" box="[633,729,1628,1655]" pageId="2" pageNumber="391" type="female">1 female</specimenCount>
in cRS); “Mindanao, Panamokan,
<collectingRegion id="4981F8397B18FFFFB916F9CBFB04F98E" box="[1126,1239,1628,1654]" country="Philippines" name="Bukidnon" pageId="2" pageNumber="391">Bukidnon</collectingRegion>
,
<date id="FFFB101B7B18FFFFB993F9CAFA9FF98F" box="[1251,1356,1629,1655]" pageId="2" pageNumber="391" value="2014-07">
<collectingDate id="EFBFE9F37B18FFFFB993F9CAFA9FF98F" box="[1251,1356,1629,1655]" pageId="2" pageNumber="391" value="2014-07">VII.2014</collectingDate>
</date>
” (
<specimenCount id="9D43FD527B18FFFFB819F9CAFF1EF962" pageId="2" pageNumber="391" type="female">1 female</specimenCount>
in cRS); “Mindanao, Cabanglasan,
<collectingRegion id="4981F8397B18FFFFBF12F917FD00F962" box="[610,723,1664,1690]" country="Philippines" name="Bukidnon" pageId="2" pageNumber="391">Bukidnon</collectingRegion>
,
<date id="FFFB101B7B18FFFFBFAEF916FC95F963" box="[734,838,1665,1691]" pageId="2" pageNumber="391" value="2014-07">
<collectingDate id="EFBFE9F37B18FFFFBFAEF916FC95F963" box="[734,838,1665,1691]" pageId="2" pageNumber="391" value="2014-07">VII.2014</collectingDate>
</date>
” (
<specimenCount id="9D43FD527B18FFFFBE12F916FC11F962" box="[866,962,1664,1691]" pageId="2" pageNumber="391" type="female">1 female</specimenCount>
in cRS); “Mindanao,
<collectingRegion id="4981F8397B18FFFFB9C9F916FA8BF963" box="[1209,1368,1664,1691]" country="Philippines" name="Lanao del Sur" pageId="2" pageNumber="391">Lanao del Sur</collectingRegion>
, Wao,
<date id="FFFB101B7B18FFFFBDE7F932FED3F947" box="[151,256,1701,1727]" pageId="2" pageNumber="391" value="2018-07">
<collectingDate id="EFBFE9F37B18FFFFBDE7F932FED3F947" box="[151,256,1701,1727]" pageId="2" pageNumber="391" value="2018-07">VII.2018</collectingDate>
</date>
” (
<specimenCount id="9D43FD527B18FFFFBC6CF932FEB6F946" box="[284,357,1700,1727]" pageId="2" pageNumber="391" type="male">1 male</specimenCount>
in cRS); “Mindanao,
<collectingRegion id="4981F8397B18FFFFBF2DF933FCDEF947" box="[605,781,1700,1727]" country="Philippines" name="Surigao del Sur" pageId="2" pageNumber="391">Surigao del Sur</collectingRegion>
, Bugak, Lingig,
<date id="FFFB101B7B18FFFFBEB8F932FBE2F947" box="[968,1073,1701,1727]" pageId="2" pageNumber="391" value="2018-07">
<collectingDate id="EFBFE9F37B18FFFFBEB8F932FBE2F947" box="[968,1073,1701,1727]" pageId="2" pageNumber="391" value="2018-07">VII.2018</collectingDate>
</date>
” (
<specimenCount id="9D43FD527B18FFFFB93DF932FB7FF946" box="[1101,1196,1700,1727]" pageId="2" pageNumber="391" type="female">1 female</specimenCount>
in cRS); “Mindanao,
<collectingRegion id="4981F8397B18FFFFBDE7F95EFEEAF91B" box="[151,313,1736,1763]" country="Philippines" name="Davao del Sur" pageId="2" pageNumber="391">Davao del Sur</collectingRegion>
,
<collectorName id="26B0530D7B18FFFFBC34F95EFE4EF91B" box="[324,413,1737,1763]" pageId="2" pageNumber="391">M. Apo</collectorName>
,
<date id="FFFB101B7B18FFFFBCD7F95EFDD5F91B" box="[423,518,1737,1763]" pageId="2" pageNumber="391" value="2014-06">
<collectingDate id="EFBFE9F37B18FFFFBCD7F95EFDD5F91B" box="[423,518,1737,1763]" pageId="2" pageNumber="391" value="2014-06">VI.2014</collectingDate>
</date>
” (
<specimenCount id="9D43FD527B18FFFFBF52F95EFDBFF91A" box="[546,620,1736,1763]" pageId="2" pageNumber="391" type="male">1 male</specimenCount>
in cRS)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35F65507B18FFF9BDB7F97BFAF6FC3F" lastPageId="4" lastPageNumber="393" pageId="2" pageNumber="391" type="diagnosis">
<paragraph id="8BFA36DB7B18FFFFBDB7F97BFE9CF826" blockId="2.[151,1437,1520,2014]" pageId="2" pageNumber="391">
<emphasis id="B931EAC97B18FFFFBDB7F97BFE91F8FF" bold="true" box="[199,322,1772,1799]" pageId="2" pageNumber="391">Diagnosis.</emphasis>
The new species belongs, like
<taxonomicName id="4C454D587B18FFFFBFD2F97AFB8CF8FF" authority="Cassola, 2000" authorityName="Cassola" authorityYear="2000" box="[674,1119,1772,1799]" class="Insecta" family="Carabidae" genus="Thopeutica" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="391" phylum="Arthropoda" rank="species" species="rolandmuelleri">
<emphasis id="B931EAC97B18FFFFBFD2F97AFD64F8FE" box="[674,695,1773,1798]" italics="true" pageId="2" pageNumber="391">T.</emphasis>
(s. str.)
<emphasis id="B931EAC97B18FFFFBE63F97BFC69F8FE" box="[787,954,1772,1798]" italics="true" pageId="2" pageNumber="391">rolandmuelleri</emphasis>
Cassola, 2000
</taxonomicName>
to the stem 3 sensu Cassola &amp; Ward (2004). Similar to
<taxonomicName id="4C454D587B18FFFFBCBAF886FD48F8D2" authorityName="Des. F. Cassola" authorityYear="1997" baseAuthorityName="W. Horn" baseAuthorityYear="1929" box="[458,667,1808,1834]" class="Insecta" family="Carabidae" genus="Thopeutica" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="391" phylum="Arthropoda" rank="species" species="angulihumerosa">
<emphasis id="B931EAC97B18FFFFBCBAF886FD48F8D2" box="[458,667,1808,1834]" italics="true" pageId="2" pageNumber="391">T. angulihumerosa</emphasis>
</taxonomicName>
in body size and elytral pattern, but easily distinguished by the half sized discal mirror on elytra in females, by entirely metallic, dark violet tibiae and tarsi, and by relatively small aedeagus, dorsally regularly arcuate inflated from basal to apical part, with apex slightly bent down, while in
<taxonomicName id="4C454D587B18FFFFB8F8F8CEFE9FF86E" authorityName="Des. F. Cassola" authorityYear="1997" baseAuthorityName="W. Horn" baseAuthorityYear="1929" class="Insecta" family="Carabidae" genus="Thopeutica" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="391" phylum="Arthropoda" rank="species" species="angulihumerosa">
<emphasis id="B931EAC97B18FFFFB8F8F8CEFE9FF86E" italics="true" pageId="2" pageNumber="391">T. angulihumerosa</emphasis>
</taxonomicName>
the aedeagus is larger, narrow in basal part and with stronger downturned apex. Endophallus of the new species with almost indistinct basal shield (bs) (Figs. 56), while in
<taxonomicName id="4C454D587B18FFFFBECEF836FB5DF842" authorityName="Des. F. Cassola" authorityYear="1997" baseAuthorityName="W. Horn" baseAuthorityYear="1929" box="[958,1166,1952,1978]" class="Insecta" family="Carabidae" genus="Thopeutica" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="391" phylum="Arthropoda" rank="species" species="angulihumerosa">
<emphasis id="B931EAC97B18FFFFBECEF836FB5DF842" box="[958,1166,1952,1978]" italics="true" pageId="2" pageNumber="391">T. angulihumerosa</emphasis>
</taxonomicName>
basal shield is large and well sclerotized.
</paragraph>
<caption id="DF3A66537B19FFFEBDE7F80FFA4EF849" ID-DOI="http://doi.org/10.5281/zenodo.3698743" ID-Zenodo-Dep="3698743" box="[151,1437,1944,1969]" httpUri="https://zenodo.org/record/3698743/files/figure.png" pageId="3" pageNumber="392" startId="3.[151,264,1944,1969]" targetBox="[399,1187,190,1907]" targetPageId="3">
<paragraph id="8BFA36DB7B19FFFEBDE7F80FFA4EF849" blockId="3.[151,1437,1944,1969]" box="[151,1437,1944,1969]" pageId="3" pageNumber="392">
<emphasis id="B931EAC97B19FFFEBDE7F80FFEEFF849" bold="true" box="[151,316,1944,1969]" pageId="3" pageNumber="392">FIGURES 34.</emphasis>
Comparative length of antennae of males of
<taxonomicName id="4C454D587B19FFFEBE61F80EFC50F849" authorityName="Schaum" authorityYear="1861" box="[785,899,1945,1969]" class="Insecta" family="Carabidae" genus="Thopeutica" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="392" phylum="Arthropoda" rank="genus">
<emphasis id="B931EAC97B19FFFEBE61F80EFC50F849" box="[785,899,1945,1969]" italics="true" pageId="3" pageNumber="392">Thopeutica</emphasis>
</taxonomicName>
spp.: 3—holotype of
<taxonomicName id="4C454D587B19FFFEB916F80DFABAF849" authorityName="Schule" authorityYear="2017" box="[1126,1385,1945,1969]" class="Insecta" family="Carabidae" genus="Thopeutica" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="392" phylum="Arthropoda" rank="species" species="juergenwiesneri" status="syn.">
<emphasis id="B931EAC97B19FFFEB916F80DFBAAF849" box="[1126,1145,1946,1969]" italics="true" pageId="3" pageNumber="392">T.</emphasis>
(s.str.)
<emphasis id="B931EAC97B19FFFEB9B7F80DFABAF849" box="[1223,1385,1946,1969]" italics="true" pageId="3" pageNumber="392">juergenwiesneri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20257B27B19FFFEB801F80EFA4EF849" box="[1393,1437,1945,1969]" pageId="3" pageNumber="392">syn.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BFA36DB7B1EFFF9BDE7FCF1FD7AFC86" blockId="4.[151,1436,870,967]" box="[151,681,870,895]" pageId="4" pageNumber="393">
<emphasis id="B931EAC97B1EFFF9BDE7FCF1FF10FC86" bold="true" box="[151,195,870,894]" pageId="4" pageNumber="393">nov.</emphasis>
; 4—
<typeStatus id="54FE88797B1EFFF9BD87FCF1FE53FC87" box="[247,384,870,895]" pageId="4" pageNumber="393" type="paralectotype">paralectotype</typeStatus>
of
<taxonomicName id="4C454D587B1EFFF9BCD1FCF0FD77FC86" authorityName="Des. F. Cassola" authorityYear="1997" baseAuthorityName="W. Horn" baseAuthorityYear="1929" box="[417,676,870,894]" class="Insecta" family="Carabidae" genus="Thopeutica" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="393" phylum="Arthropoda" rank="species" species="angulihumerosa">
<emphasis id="B931EAC97B1EFFF9BCD1FCF0FE66FC86" box="[417,437,871,894]" italics="true" pageId="4" pageNumber="393">T.</emphasis>
(s.str.)
<emphasis id="B931EAC97B1EFFF9BF70FCF1FD77FC86" box="[512,676,870,894]" italics="true" pageId="4" pageNumber="393">angulihumerosa</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BFA36DB7B1EFFF9BDE7FC1DFAF6FC3F" blockId="4.[151,1436,870,967]" pageId="4" pageNumber="393">
<emphasis id="B931EAC97B1EFFF9BDE7FC1DFEE9FC5B" bold="true" box="[151,314,906,931]" pageId="4" pageNumber="393">FIGURES 59.</emphasis>
Structure of aedeagus of
<taxonomicName id="4C454D587B1EFFF9BF49FC1DFD78FC5A" authorityName="Schaum" authorityYear="1861" box="[569,683,906,930]" class="Insecta" family="Carabidae" genus="Thopeutica" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="393" phylum="Arthropoda" rank="genus">
<emphasis id="B931EAC97B1EFFF9BF49FC1DFD78FC5A" box="[569,683,906,930]" italics="true" pageId="4" pageNumber="393">Thopeutica</emphasis>
</taxonomicName>
spp.: 56—
<typeStatus id="54FE88797B1EFFF9BE55FC1DFC56FC5B" box="[805,901,906,931]" pageId="4" pageNumber="393" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="4C454D587B1EFFF9BED5FC1CFB8BFC5A" authority="Anichtchenko &amp; Medina, 2020" authorityName="Anichtchenko &amp; Medina" authorityYear="2020" box="[933,1112,906,930]" class="Insecta" family="Carabidae" genus="Thopeutica" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="393" phylum="Arthropoda" rank="species" species="wiesneri" status="sp. nov.">
<emphasis id="B931EAC97B1EFFF9BED5FC1CFC6BFC5A" box="[933,952,907,930]" italics="true" pageId="4" pageNumber="393">T.</emphasis>
(s.str.)
<emphasis id="B931EAC97B1EFFF9B972FC1CFB8BFC5A" box="[1026,1112,907,930]" italics="true" pageId="4" pageNumber="393">wiesneri</emphasis>
</taxonomicName>
<emphasis id="B931EAC97B1EFFF9B92DFC1DFB7EFC5A" bold="true" box="[1117,1197,906,930]" pageId="4" pageNumber="393">
<taxonomicNameLabel id="A20257B27B1EFFF9B92DFC1DFB7EFC5A" box="[1117,1197,906,930]" pageId="4" pageNumber="393" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; 78—
<typeStatus id="54FE88797B1EFFF9B987FC1DFA51FC5B" box="[1271,1410,906,931]" pageId="4" pageNumber="393" type="paralectotype">paralectotype</typeStatus>
of
<taxonomicName id="4C454D587B1EFFF9BDE7FC38FE49FC3E" authorityName="Des. F. Cassola" authorityYear="1997" baseAuthorityName="W. Horn" baseAuthorityYear="1929" box="[151,410,942,966]" class="Insecta" family="Carabidae" genus="Thopeutica" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="393" phylum="Arthropoda" rank="species" species="angulihumerosa">
<emphasis id="B931EAC97B1EFFF9BDE7FC38FF78FC3E" box="[151,171,943,966]" italics="true" pageId="4" pageNumber="393">T.</emphasis>
(s.str.)
<emphasis id="B931EAC97B1EFFF9BD86FC39FE49FC3E" box="[246,410,942,966]" italics="true" pageId="4" pageNumber="393">angulihumerosa</emphasis>
</taxonomicName>
; 9—
<typeStatus id="54FE88797B1EFFF9BCBCFC39FDF6FC3F" box="[460,549,942,967]" pageId="4" pageNumber="393" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C454D587B1EFFF9BF37FC38FC9BFC3E" authorityName="Schule" authorityYear="2017" box="[583,840,942,966]" class="Insecta" family="Carabidae" genus="Thopeutica" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="393" phylum="Arthropoda" rank="species" species="juergenwiesneri">
<emphasis id="B931EAC97B1EFFF9BF37FC38FD88FC3E" box="[583,603,943,966]" italics="true" pageId="4" pageNumber="393">T.</emphasis>
(s.str.)
<emphasis id="B931EAC97B1EFFF9BFD6FC38FC9BFC3E" box="[678,840,943,966]" italics="true" pageId="4" pageNumber="393">juergenwiesneri</emphasis>
</taxonomicName>
<emphasis id="B931EAC97B1EFFF9BE3EFC39FC7FFC3E" bold="true" box="[846,940,942,966]" pageId="4" pageNumber="393">
<taxonomicNameLabel id="A20257B27B1EFFF9BE3EFC39FC7FFC3E" box="[846,940,942,966]" pageId="4" pageNumber="393">syn. nov.</taxonomicNameLabel>
</emphasis>
; 5, 7, 9—left view; 6, 8—right view.
</paragraph>
</subSubSection>
<caption id="DF3A66537B1EFFF9BDE7F8A8FB16F884" ID-DOI="http://doi.org/10.5281/zenodo.3698747" ID-Zenodo-Dep="3698747" httpUri="https://zenodo.org/record/3698747/files/figure.png" pageId="4" pageNumber="393" startId="4.[151,264,1855,1880]" targetBox="[170,1418,992,1825]" targetPageId="4">
<paragraph id="8BFA36DB7B1EFFF9BDE7F8A8FB16F884" blockId="4.[151,1436,1855,1916]" pageId="4" pageNumber="393">
<emphasis id="B931EAC97B1EFFF9BDE7F8A8FE86F8A0" bold="true" box="[151,341,1855,1880]" pageId="4" pageNumber="393">FIGURES 1012.</emphasis>
Labrum of
<taxonomicName id="4C454D587B1EFFF9BCBFF8D7FD92F8A0" authorityName="Schaum" authorityYear="1861" box="[463,577,1856,1880]" class="Insecta" family="Carabidae" genus="Thopeutica" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="393" phylum="Arthropoda" rank="genus">
<emphasis id="B931EAC97B1EFFF9BCBFF8D7FD92F8A0" box="[463,577,1856,1880]" italics="true" pageId="4" pageNumber="393">Thopeutica</emphasis>
</taxonomicName>
spp. (a—males, b—females): 10—paralectotype of
<taxonomicName id="4C454D587B1EFFF9B928F8D6FA8EF8A0" authorityName="Des. F. Cassola" authorityYear="1997" baseAuthorityName="W. Horn" baseAuthorityYear="1929" box="[1112,1373,1856,1880]" class="Insecta" family="Carabidae" genus="Thopeutica" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="393" phylum="Arthropoda" rank="species" species="angulihumerosa">
<emphasis id="B931EAC97B1EFFF9B928F8D6FBBFF8A0" box="[1112,1132,1857,1880]" italics="true" pageId="4" pageNumber="393">T.</emphasis>
(s.str.)
<emphasis id="B931EAC97B1EFFF9B9C9F8D7FA8EF8A0" box="[1209,1373,1856,1880]" italics="true" pageId="4" pageNumber="393">angulihumerosa</emphasis>
</taxonomicName>
; 11— holotype of
<taxonomicName id="4C454D587B1EFFF9BC62F8F2FDC1F884" authorityName="Schule" authorityYear="2017" box="[274,530,1892,1916]" class="Insecta" family="Carabidae" genus="Thopeutica" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="393" phylum="Arthropoda" rank="species" species="juergenwiesneri">
<emphasis id="B931EAC97B1EFFF9BC62F8F2FEF6F884" box="[274,293,1893,1916]" italics="true" pageId="4" pageNumber="393">T.</emphasis>
(s.str.)
<emphasis id="B931EAC97B1EFFF9BC00F8F2FDC1F884" box="[368,530,1893,1916]" italics="true" pageId="4" pageNumber="393">juergenwiesneri</emphasis>
</taxonomicName>
<emphasis id="B931EAC97B1EFFF9BF69F8F3FDA5F884" bold="true" box="[537,630,1892,1916]" pageId="4" pageNumber="393">
<taxonomicNameLabel id="A20257B27B1EFFF9BF69F8F3FDA5F884" box="[537,630,1892,1916]" pageId="4" pageNumber="393">syn. nov.</taxonomicNameLabel>
</emphasis>
; 12—holotype and paratype of
<taxonomicName id="4C454D587B1EFFF9BEC9F8F2FBBDF884" authority="Anichtchenko &amp; Medina, 2020" authorityName="Anichtchenko &amp; Medina" authorityYear="2020" box="[953,1134,1892,1916]" class="Insecta" family="Carabidae" genus="Thopeutica" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="393" phylum="Arthropoda" rank="species" species="wiesneri" status="sp. nov.">
<emphasis id="B931EAC97B1EFFF9BEC9F8F2FC1EF884" box="[953,973,1893,1916]" italics="true" pageId="4" pageNumber="393">T.</emphasis>
(s.str.)
<emphasis id="B931EAC97B1EFFF9B968F8F2FBBDF884" box="[1048,1134,1893,1916]" italics="true" pageId="4" pageNumber="393">wiesneri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20257B27B1EFFF9B904F8F3FB16F884" box="[1140,1221,1892,1916]" pageId="4" pageNumber="393" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<subSubSection id="C35F65507B1FFFFBBDB7FF0FFDA2FCC3" lastPageId="6" lastPageNumber="395" pageId="5" pageNumber="394" type="description">
<paragraph id="8BFA36DB7B1FFFF8BDB7FF0FFB83FF03" blockId="5.[151,1437,152,611]" pageId="5" pageNumber="394">
<emphasis id="B931EAC97B1FFFF8BDB7FF0FFE8AFF4B" bold="true" box="[199,345,152,179]" pageId="5" pageNumber="394">Description.</emphasis>
Body length 7.58.5 mm. Basic coloration of dorsal side dark bronze, with green, blue, and violet hue on head and pronotum. Elytra dark bronze, exterior sides and all elytral pores greenish, dorsal maculation of elytra white, humera of female shiny green around white humeral spot (
<figureCitation id="137E2A5E7B1FFFF8BECAFF77FB90FF03" box="[954,1091,224,251]" captionStart="FIGURES 1314" captionStartId="5.[151,264,1883,1908]" captionTargetBox="[158,1425,637,1857]" captionTargetId="figure@5.[151,1436,632,1857]" captionTargetPageId="5" captionText="FIGURES 1314. Habitus of Thopeutica (s.str.) wiesneri sp. n.: 13—holotype, male; 14—paratype, female." figureDoi="http://doi.org/10.5281/zenodo.3698749" httpUri="https://zenodo.org/record/3698749/files/figure.png" pageId="5" pageNumber="394">Figs. 1314</figureCitation>
).
</paragraph>
<paragraph id="8BFA36DB7B1FFFF8BDB7FE93FD92FD9A" blockId="5.[151,1437,152,611]" pageId="5" pageNumber="394">
<emphasis id="B931EAC97B1FFFF8BDB7FE93FED8FEE6" box="[199,267,260,286]" italics="true" pageId="5" pageNumber="394">Head.</emphasis>
Glabrous, dorsal surface dark bronze, dull, with very fine microreticulation,; lateral areas, genae, clypeus and area above antennal insertion with stronger metallic green and blue lustre; surface of frons with minute semicircular, concentrically arranged rugae, vertex strongly reticulated, sublateral areas and juxta-orbital areas striaterugose; two orbital setae on each side of the head. Labrum from goldish green to dark blue and violet coloration; transverse, lateral margins subparallel, median portion broadly protruding in males and females; with 610 setae in apical margin (
<figureCitation id="137E2A5E7B1FFFF8BC2DFE2FFE6AFE2B" box="[349,441,440,467]" captionStart="FIGURES 1012" captionStartId="4.[151,264,1855,1880]" captionTargetBox="[170,1418,992,1825]" captionTargetId="figure@4.[163,1425,988,1830]" captionTargetPageId="4" captionText="FIGURES 1012. Labrum of Thopeutica spp. (a—males, b—females): 10—paralectotype of T. (s.str.) angulihumerosa; 11— holotype of T. (s.str.) juergenwiesneri syn. nov.; 12—holotype and paratype of T. (s.str.) wiesneri sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3698747" httpUri="https://zenodo.org/record/3698747/files/figure.png" pageId="5" pageNumber="394">Figs. 12</figureCitation>
ab). Mandibles dark brown, with green metallic luster and with white baso-lateral spot; narrow, long, with arcuate lateral margins. Palpi elongate and slender, apical joints moderately and gradually dilated towards apex, light yellow except green apical palpomeres. Antennae long, in male reaching the apical fourth of elytra; scape with coppery-green lustre; pedicellum concolorous, antennomeres three and four black with violet lustre, the rest of antennomeres black.
</paragraph>
<caption id="DF3A66537B1FFFF8BDE7F8CCFB35F88C" ID-DOI="http://doi.org/10.5281/zenodo.3698749" ID-Zenodo-Dep="3698749" box="[151,1254,1883,1908]" httpUri="https://zenodo.org/record/3698749/files/figure.png" pageId="5" pageNumber="394" startId="5.[151,264,1883,1908]" targetBox="[158,1425,637,1857]" targetPageId="5">
<paragraph id="8BFA36DB7B1FFFF8BDE7F8CCFB35F88C" blockId="5.[151,1254,1883,1908]" box="[151,1254,1883,1908]" pageId="5" pageNumber="394">
<emphasis id="B931EAC97B1FFFF8BDE7F8CCFE87F88C" bold="true" box="[151,340,1883,1908]" pageId="5" pageNumber="394">FIGURES 1314.</emphasis>
Habitus of
<taxonomicName id="4C454D587B1FFFF8BCBBF8CCFD0CF88B" authority="Anichtchenko &amp; Medina, 2020" authorityName="Anichtchenko &amp; Medina" authorityYear="2020" box="[459,735,1883,1907]" class="Insecta" family="Carabidae" genus="Thopeutica" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="394" phylum="Arthropoda" rank="species" species="wiesneri" status="sp. nov.">
<emphasis id="B931EAC97B1FFFF8BCBBF8CCFDEEF88B" box="[459,573,1883,1907]" italics="true" pageId="5" pageNumber="394">Thopeutica</emphasis>
(s.str.)
<emphasis id="B931EAC97B1FFFF8BFF9F8CBFD0CF88B" box="[649,735,1884,1907]" italics="true" pageId="5" pageNumber="394">wiesneri</emphasis>
</taxonomicName>
<emphasis id="B931EAC97B1FFFF8BF95F8CBFCF4F88C" bold="true" box="[741,807,1884,1908]" pageId="5" pageNumber="394">
<taxonomicNameLabel id="A20257B27B1FFFF8BF95F8CBFCF2F88C" box="[741,801,1884,1908]" pageId="5" pageNumber="394" rank="species">sp. n.</taxonomicNameLabel>
:
</emphasis>
13—holotype, male; 14—paratype, female.
</paragraph>
</caption>
<paragraph id="8BFA36DB7B1FFFFBBDB7F835FCEAFEBB" blockId="5.[151,1436,1954,2016]" lastBlockId="6.[151,1437,153,1007]" lastPageId="6" lastPageNumber="395" pageId="5" pageNumber="394">
<emphasis id="B931EAC97B1FFFF8BDB7F835FECBF844" box="[199,280,1954,1980]" italics="true" pageId="5" pageNumber="394">Thorax</emphasis>
. Pronotum as long as wide in females, and slightly narrower in males; anterior and posterior sulcus well pronounced, with metallic-green lustre, posterior sulcus doubly bowed each side curved towards the fine median line of pronotum; disc of pronotum dull, with strong microreticulation and shallow wrinkles; lateral edge of pronotum coppery, anterior and posterior angles with a group of white curved setae; proepisterna coppery-green, with fine vermiculate wrinkles, ventral half, anterior and posterior margin with long white setae, prosternum setose. Mesepisterna slightly rugose on ventral half, with metallic green lustre; thoracic segments entirely setose with dense, white setae, except mesosternum and central area of metasternum.
</paragraph>
<paragraph id="8BFA36DB7B1CFFFBBDB7FEDAFF68FD9B" blockId="6.[151,1437,153,1007]" pageId="6" pageNumber="395">
<emphasis id="B931EAC97B1CFFFBBDB7FEDAFEDEFE9F" box="[199,269,333,359]" italics="true" pageId="6" pageNumber="395">Elytra</emphasis>
. Elongate, subparallel with well-pronounced humeri in males, and with strongly hooked humeri in females; slightly dilated laterally with apical angles rounded towards and a very small and blunt apical sutural tooth; apical margin with microserrulation; dorsal surface with fine microreticulation, dull, finely and shallowly punctate of green punctures, slightly greenish iridescent towards lateral margin; elytral maculation consisting of a yellow humeral dot, white roundish posthumeral, sublateral-median, slightly smaller discal and anteapical dots and a curved apical lunule not reaching suture nor subapical angle (
<figureCitation id="137E2A5E7B1CFFFBBF8EFD96FC5BFDE3" box="[766,904,513,539]" captionStart="FIGURES 1314" captionStartId="5.[151,264,1883,1908]" captionTargetBox="[158,1425,637,1857]" captionTargetId="figure@5.[151,1436,632,1857]" captionTargetPageId="5" captionText="FIGURES 1314. Habitus of Thopeutica (s.str.) wiesneri sp. n.: 13—holotype, male; 14—paratype, female." figureDoi="http://doi.org/10.5281/zenodo.3698749" httpUri="https://zenodo.org/record/3698749/files/figure.png" pageId="6" pageNumber="395">Figs. 1314</figureCitation>
). Females with large green posthumeral mirror spot, and with relatively small, black diagonal mirror on the disc of elytra. Scutellum metallic green with gold luster.
</paragraph>
<paragraph id="8BFA36DB7B1CFFFBBDB7FDF9FE21FD53" blockId="6.[151,1437,153,1007]" pageId="6" pageNumber="395">
<emphasis id="B931EAC97B1CFFFBBDB7FDF9FF2FFD7F" box="[199,252,622,647]" italics="true" pageId="6" pageNumber="395">Legs</emphasis>
. Trochanters yellow; femora with bronze and green lustre, apices rufescent, with violet lustre; tibiae violet, basally green; tarsi dark violet.
</paragraph>
<paragraph id="8BFA36DB7B1CFFFBBDB7FD22FBF1FD37" blockId="6.[151,1437,153,1007]" box="[199,1058,693,719]" pageId="6" pageNumber="395">
<emphasis id="B931EAC97B1CFFFBBDB7FD22FEE2FD37" box="[199,305,693,719]" italics="true" pageId="6" pageNumber="395">Abdomen</emphasis>
. Ventrites black with coppery-green lustre, covered by white setae.
</paragraph>
<paragraph id="8BFA36DB7B1CFFFBBDB7FD4EFDA2FCC3" blockId="6.[151,1437,153,1007]" pageId="6" pageNumber="395">
<emphasis id="B931EAC97B1CFFFBBDB7FD4EFEE7FD0B" box="[199,308,729,755]" italics="true" pageId="6" pageNumber="395">Aedeagus</emphasis>
. (Figs. 56) short, dorsally regularly arcuate inflated from basal to apical part, apex slightly bent ventrally. The endophallus structure is typical for
<taxonomicName id="4C454D587B1CFFFBBFA5FD6AFC80FCEF" authorityName="Schaum" authorityYear="1861" box="[725,851,765,791]" class="Insecta" family="Carabidae" genus="Thopeutica" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="395" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B931EAC97B1CFFFBBFA5FD6AFC80FCEF" box="[725,851,765,791]" italics="true" pageId="6" pageNumber="395">Thopeutica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20257B27B1CFFFBBE2CFD6AFC42FCEF" box="[860,913,765,791]" pageId="6" pageNumber="395" sensu="stricto">s.str.</taxonomicNameLabel>
, with a long coiled flagellum; basal shield of endophallus small and weakly sclerotized.
</paragraph>
</subSubSection>
<subSubSection id="C35F65507B1CFFFBBDB7FCD2FF14FC17" pageId="6" pageNumber="395" type="etymology">
<paragraph id="8BFA36DB7B1CFFFBBDB7FCD2FAF1FCA7" blockId="6.[151,1437,153,1007]" box="[199,1314,837,863]" pageId="6" pageNumber="395">
<emphasis id="B931EAC97B1CFFFBBDB7FCD2FE9CFCA7" bold="true" box="[199,335,837,863]" pageId="6" pageNumber="395">Etymology.</emphasis>
Named after the famous German Tiger-beetle specialist and our friend Jürgen Wiesner.
</paragraph>
<paragraph id="8BFA36DB7B1CFFFBBDB7FCFEFF14FC17" blockId="6.[151,1437,153,1007]" pageId="6" pageNumber="395">
<emphasis id="B931EAC97B1CFFFBBDB7FCFEFE44FC7B" bold="true" box="[199,407,873,899]" pageId="6" pageNumber="395">Notes on ecology.</emphasis>
The recently described species was collected near the slow moving fluvial system covered with secondary Dipterocarp species between 800 and
<quantity id="4CBD9B3E7B1CFFFBBF8BFC1AFCA0FC5F" box="[763,883,909,935]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="6" pageNumber="395" unit="m" value="1300.0">1300 masl</quantity>
. They tend to rest among moist bryophytes fully or partially shaded. This species is very hard to catch opportunistically but attracted to light between 6:00 to 8:00 PM.
</paragraph>
</subSubSection>
</treatment>
</document>