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<document id="C7516E0D717AE853AF381389610481D1" ID-CLB-Dataset="24654" ID-DOI="10.11646/zootaxa.4706.4.4" ID-GBIF-Dataset="a5168d5b-f2f3-4ca9-be27-cf0a4b3421df" ID-ISSN="1175-5326" ID-Zenodo-Dep="3616903" ID-ZooBank="D5917547-D026-42E4-AE59-A891C30E756C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1576046767784" checkinUser="plazi" docAuthor="Hugel, Sylvain" docDate="2019" docId="03EC87C4FFFF540AFF49D5884C9DFB83" docLanguage="en" docName="zootaxa.4706.4.4.pdf" docOrigin="Zootaxa 4706 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Toliaridectes Hugel 2019, n. gen." docType="treatment" docVersion="10" lastPageNumber="548" masterDocId="FFD5FFBCFFFE5408FFDED7754D06FF90" masterDocTitle="First record of shield-backed katydids in Madagascar with the description of a new genus and three new species (Orthoptera: Ensifera: Tettigoniidae: Tettigoniinae Arytropteridini)" masterLastPageNumber="560" masterPageNumber="546" pageNumber="547" updateTime="1698773708133" updateUser="ExternalLinkService">
<mods:mods id="E561D6191EA6863BC5BC0D873646D024" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E731EE108232331C0C94457D6F6C5048">
<mods:title id="E159F4C051CD96077B6169C919895D7C">First record of shield-backed katydids in Madagascar with the description of a new genus and three new species (Orthoptera: Ensifera: Tettigoniidae: Tettigoniinae Arytropteridini)</mods:title>
</mods:titleInfo>
<mods:name id="6ECCBCE83B10819B60D1A388E496D5FF" type="personal">
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<mods:namePart id="758D9FCB85CE0FAB4D66466E2641F753">Hugel, Sylvain</mods:namePart>
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<mods:typeOfResource id="669A76F2A67BD791AE70EAAE20434B39">text</mods:typeOfResource>
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<mods:title id="9DFA9A2F96D51508EA9DC185843F9418">Zootaxa</mods:title>
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<mods:part id="4243ED6406A9173DAF81DAEFD90DFD4E">
<mods:date id="1C9478363B0566E1C929CCFF9F66AF00">2019</mods:date>
<mods:detail id="9B980723219A3F49E59AF85DB1051CBE" type="pubDate">
<mods:number id="1B0F96F5D8E63CA463D325F09096425E">2019-12-11</mods:number>
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<mods:detail id="521C44AF8768CA34A1D14583CF8CD023" type="volume">
<mods:number id="14734EBFBFE62EDAEDCC36A8EC6B20A7">4706</mods:number>
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<mods:classification id="7FD0E297AD74B797B3F0DEAC41E6BB4A">journal article</mods:classification>
<mods:identifier id="E1BBB59DAB97CCCC791C0B9093352E75" type="CLB-Dataset">24654</mods:identifier>
<mods:identifier id="EF930E64CB4894CBF78A93B52047F8E4" type="DOI">10.11646/zootaxa.4706.4.4</mods:identifier>
<mods:identifier id="C8790440734D5C625CD2F7D6E84D7888" type="GBIF-Dataset">a5168d5b-f2f3-4ca9-be27-cf0a4b3421df</mods:identifier>
<mods:identifier id="9211E80623AFC282D0A8483932A804B1" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03EC87C4FFFF540AFF49D5884C9DFB83" ID-DOI="http://doi.org/10.5281/zenodo.5615491" ID-GBIF-Taxon="160689647" ID-Zenodo-Dep="5615491" LSID="urn:lsid:plazi:treatment:03EC87C4FFFF540AFF49D5884C9DFB83" httpUri="http://treatment.plazi.org/id/03EC87C4FFFF540AFF49D5884C9DFB83" lastPageId="2" lastPageNumber="548" pageId="1" pageNumber="547">
<subSubSection id="C35F6559FFFF5409FF49D5884CE7FC88" box="[151,481,765,792]" pageId="1" pageNumber="547" type="nomenclature">
<paragraph id="8BFA36D2FFFF5409FF49D5884CE7FC88" blockId="1.[151,481,765,792]" box="[151,481,765,792]" pageId="1" pageNumber="547">
<heading id="D0B281BEFFFF5409FF49D5884CE7FC88" bold="true" box="[151,481,765,792]" fontSize="11" level="1" pageId="1" pageNumber="547" reason="1">
<emphasis id="B931EAC0FFFF5409FF49D5884CE7FC88" bold="true" box="[151,481,765,792]" pageId="1" pageNumber="547">
Genus
<taxonomicName id="4C454D51FFFF5409FF30D5884C8EFC88" ID-CoL="8PVGX" authority="Hugel, 2019" authorityName="Hugel" authorityYear="2019" box="[238,392,765,792]" class="Insecta" family="Tettigoniidae" genus="Toliaridectes" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="547" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B931EAC0FFFF5409FF30D5884C8EFC88" bold="true" box="[238,392,765,792]" italics="true" pageId="1" pageNumber="547">Toliaridectes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20257BBFFFF5409FE51D58B4CE7FC88" box="[399,481,766,792]" pageId="1" pageNumber="547" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35F6559FFFF5409FF49D4334EA8FCCF" box="[151,942,837,864]" pageId="1" pageNumber="547" type="type_taxon">
<paragraph id="8BFA36D2FFFF5409FF49D4334EA8FCCF" blockId="1.[151,1437,837,2016]" box="[151,942,837,864]" pageId="1" pageNumber="547">
<emphasis id="B931EAC0FFFF5409FF49D4334C34FCF0" bold="true" box="[151,306,837,864]" pageId="1" pageNumber="547">
<typeStatus id="54FE8870FFFF5409FF49D4334DD4FCF0" box="[151,210,838,864]" pageId="1" pageNumber="547">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C454D51FFFF5409FEE7D4304F57FCCF" ID-CoL="8Q4HT" authority="Hugel, 2019" authorityName="Hugel" authorityYear="2019" box="[313,593,837,863]" class="Insecta" family="Tettigoniidae" genus="Toliaridectes" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="547" phylum="Arthropoda" rank="species" species="antsycurvis" status="gen. nov.">
<emphasis id="B931EAC0FFFF5409FEE7D4304F57FCCF" box="[313,593,837,863]" italics="true" pageId="1" pageNumber="547">Toliaridectes antsycurvis</emphasis>
</taxonomicName>
<emphasis id="B931EAC0FFFF5409FD86D4334FA0FCF0" bold="true" box="[600,678,838,864]" pageId="1" pageNumber="547">
<taxonomicNameLabel id="A20257BBFFFF5409FD86D4334FA0FCF0" box="[600,678,838,864]" pageId="1" pageNumber="547" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
, n. sp, here designated.
</paragraph>
</subSubSection>
<subSubSection id="C35F6559FFFF5409FF19D41C4F9DFC13" box="[199,667,873,900]" pageId="1" pageNumber="547" type="distribution">
<paragraph id="8BFA36D2FFFF5409FF19D41C4F9DFC13" blockId="1.[151,1437,837,2016]" box="[199,667,873,900]" pageId="1" pageNumber="547">
<emphasis id="B931EAC0FFFF5409FF19D41C4C67FC14" bold="true" box="[199,353,873,900]" pageId="1" pageNumber="547">Distribution.</emphasis>
South-Eastern
<collectingCountry id="F3527642FFFF5409FDD3D41C4F90FC13" box="[525,662,873,899]" name="Madagascar" pageId="1" pageNumber="547">Madagascar</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C35F6559FFFF5409FF19D4F84F50FB34" pageId="1" pageNumber="547" type="diagnosis">
<paragraph id="8BFA36D2FFFF5409FF19D4F84F50FB34" blockId="1.[151,1437,837,2016]" pageId="1" pageNumber="547">
<emphasis id="B931EAC0FFFF5409FF19D4F84C44FC38" bold="true" box="[199,322,909,936]" pageId="1" pageNumber="547">Diagnosis.</emphasis>
<taxonomicName id="4C454D51FFFF5409FE95D4F84CDAFC37" authority="Hugel, 2019" authorityName="Hugel" authorityYear="2019" box="[331,476,909,935]" class="Insecta" family="Tettigoniidae" genus="Toliaridectes" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="547" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B931EAC0FFFF5409FE95D4F84CDAFC37" box="[331,476,909,935]" italics="true" pageId="1" pageNumber="547">Toliaridectes</emphasis>
</taxonomicName>
<emphasis id="B931EAC0FFFF5409FE3BD4FB4F30FC38" bold="true" box="[485,566,910,936]" pageId="1" pageNumber="547">
<taxonomicNameLabel id="A20257BBFFFF5409FE3BD4FB4F30FC38" box="[485,566,910,936]" pageId="1" pageNumber="547" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
is characterized by: Fastigium of vertex narrower than antennal scape (
<figureCitation id="137E2A57FFFF5409FAB1D4F84DC2FC5B" captionStart="FIGURES 1–9" captionStartId="3.[151,264,1841,1866]" captionTargetBox="[158,1422,482,1819]" captionTargetId="figure@3.[151,1436,482,1821]" captionTargetPageId="3" captionText="FIGURES 1–9. Toliaridectes n. gen. male head and pronotum. 1–3: head and pronotum in dorsal view, before removing the forewings for examination. 4–6: head and pronotum in left side view. 7–9: face in frontal view. 1, 4, 7: Toliaridectes meridionalis n. gen. n. sp. 2, 5, 8: Toliaridectes wendenbaumi n. gen. n. sp. 3, 6, 9: Toliaridectes antsycurvis n. gen. n. sp. All pictured specimens are holotypes." figureDoi="http://doi.org/10.5281/zenodo.3616905" httpUri="https://zenodo.org/record/3616905/files/figure.png" pageId="1" pageNumber="547">Fig. 1–3</figureCitation>
; as broad or broader in
<taxonomicName id="4C454D51FFFF5409FE38D4C44F59FC5B" authorityName="Rentz" authorityYear="1988" box="[486,607,945,971]" class="Insecta" family="Tettigoniidae" genus="Zuludectes" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="547" phylum="Arthropoda" rank="genus">
<emphasis id="B931EAC0FFFF5409FE38D4C44F59FC5B" box="[486,607,945,971]" italics="true" pageId="1" pageNumber="547">Zuludectes</emphasis>
</taxonomicName>
,
<taxonomicName id="4C454D51FFFF5409FDAFD4C74E13FC5C" authorityName="Uvarov" authorityYear="1924" box="[625,789,946,972]" class="Insecta" family="Tettigoniidae" genus="Anarytropteris" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="547" phylum="Arthropoda" rank="genus">
<emphasis id="B931EAC0FFFF5409FDAFD4C74E13FC5C" box="[625,789,946,972]" italics="true" pageId="1" pageNumber="547">Anarytropteris</emphasis>
</taxonomicName>
,
<taxonomicName id="4C454D51FFFF5409FCF9D4C44EDCFC5B" authorityName="Rentz" authorityYear="1988" box="[807,986,945,971]" class="Insecta" family="Tettigoniidae" genus="Namaquadectes" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="547" phylum="Arthropoda" rank="genus">
<emphasis id="B931EAC0FFFF5409FCF9D4C44EDCFC5B" box="[807,986,945,971]" italics="true" pageId="1" pageNumber="547">Namaquadectes</emphasis>
</taxonomicName>
,
<taxonomicName id="4C454D51FFFF5409FC32D4C44976FC5B" authorityName="Pictet" authorityYear="1888" box="[1004,1136,945,971]" class="Insecta" family="Tettigoniidae" genus="Thoracistus" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="547" phylum="Arthropoda" rank="genus">
<emphasis id="B931EAC0FFFF5409FC32D4C44976FC5B" box="[1004,1136,945,971]" italics="true" pageId="1" pageNumber="547">Thoracistus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C454D51FFFF5409FB5CD4C449F9FC5B" authorityName="Rentz" authorityYear="1988" box="[1154,1279,945,971]" class="Insecta" family="Tettigoniidae" genus="Alfredectes" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="547" phylum="Arthropoda" rank="genus">
<emphasis id="B931EAC0FFFF5409FB5CD4C449F9FC5B" box="[1154,1279,945,971]" italics="true" pageId="1" pageNumber="547">Alfredectes</emphasis>
</taxonomicName>
), no fastigial gap (
<figureCitation id="137E2A57FFFF5409FF0FD4A04C33FC7F" box="[209,309,981,1007]" captionStart="FIGURES 1–9" captionStartId="3.[151,264,1841,1866]" captionTargetBox="[158,1422,482,1819]" captionTargetId="figure@3.[151,1436,482,1821]" captionTargetPageId="3" captionText="FIGURES 1–9. Toliaridectes n. gen. male head and pronotum. 1–3: head and pronotum in dorsal view, before removing the forewings for examination. 4–6: head and pronotum in left side view. 7–9: face in frontal view. 1, 4, 7: Toliaridectes meridionalis n. gen. n. sp. 2, 5, 8: Toliaridectes wendenbaumi n. gen. n. sp. 3, 6, 9: Toliaridectes antsycurvis n. gen. n. sp. All pictured specimens are holotypes." figureDoi="http://doi.org/10.5281/zenodo.3616905" httpUri="https://zenodo.org/record/3616905/files/figure.png" pageId="1" pageNumber="547">Fig. 4–6</figureCitation>
; with a gap in
<taxonomicName id="4C454D51FFFF5409FE3DD4A34F6DFC60" authorityName="Herman" authorityYear="1874" box="[483,619,982,1008]" class="Insecta" family="Tettigoniidae" genus="Arytropteris" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="547" phylum="Arthropoda" rank="genus">
<emphasis id="B931EAC0FFFF5409FE3DD4A34F6DFC60" box="[483,619,982,1008]" italics="true" pageId="1" pageNumber="547">Arytropteris</emphasis>
</taxonomicName>
). Pronotum little modified (
<figureCitation id="137E2A57FFFF5409FC70D4A04917FC7F" box="[942,1041,981,1007]" captionStart="FIGURES 1–9" captionStartId="3.[151,264,1841,1866]" captionTargetBox="[158,1422,482,1819]" captionTargetId="figure@3.[151,1436,482,1821]" captionTargetPageId="3" captionText="FIGURES 1–9. Toliaridectes n. gen. male head and pronotum. 1–3: head and pronotum in dorsal view, before removing the forewings for examination. 4–6: head and pronotum in left side view. 7–9: face in frontal view. 1, 4, 7: Toliaridectes meridionalis n. gen. n. sp. 2, 5, 8: Toliaridectes wendenbaumi n. gen. n. sp. 3, 6, 9: Toliaridectes antsycurvis n. gen. n. sp. All pictured specimens are holotypes." figureDoi="http://doi.org/10.5281/zenodo.3616905" httpUri="https://zenodo.org/record/3616905/files/figure.png" pageId="1" pageNumber="547">Fig. 1–6</figureCitation>
; elongate in males of
<taxonomicName id="4C454D51FFFF5409FACFD4A04893FC7F" authorityName="Pictet" authorityYear="1888" box="[1297,1429,981,1007]" class="Insecta" family="Tettigoniidae" genus="Thoracistus" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="547" phylum="Arthropoda" rank="genus">
<emphasis id="B931EAC0FFFF5409FACFD4A04893FC7F" box="[1297,1429,981,1007]" italics="true" pageId="1" pageNumber="547">Thoracistus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C454D51FFFF5409FF49D48C4C12FB83" authorityName="Rentz" authorityYear="1988" box="[151,276,1017,1043]" class="Insecta" family="Tettigoniidae" genus="Alfredectes" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="547" phylum="Arthropoda" rank="genus">
<emphasis id="B931EAC0FFFF5409FF49D48C4C12FB83" box="[151,276,1017,1043]" italics="true" pageId="1" pageNumber="547">Alfredectes</emphasis>
</taxonomicName>
, some
<taxonomicName id="4C454D51FFFF5409FEB4D48F4CF4FB84" authorityName="Herman" authorityYear="1874" box="[362,498,1018,1044]" class="Insecta" family="Tettigoniidae" genus="Arytropteris" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="547" phylum="Arthropoda" rank="genus">
<emphasis id="B931EAC0FFFF5409FEB4D48F4CF4FB84" box="[362,498,1018,1044]" italics="true" pageId="1" pageNumber="547">Arytropteris</emphasis>
</taxonomicName>
), not exceeding first abdominal segment. Ventral surface of fore and mid femora armed with minute teeth (
<figureCitation id="137E2A57FFFF5409FE6CD3684CF2FBA7" box="[434,500,1053,1079]" captionStart="FIGURES 1–9" captionStartId="3.[151,264,1841,1866]" captionTargetBox="[158,1422,482,1819]" captionTargetId="figure@3.[151,1436,482,1821]" captionTargetPageId="3" captionText="FIGURES 1–9. Toliaridectes n. gen. male head and pronotum. 1–3: head and pronotum in dorsal view, before removing the forewings for examination. 4–6: head and pronotum in left side view. 7–9: face in frontal view. 1, 4, 7: Toliaridectes meridionalis n. gen. n. sp. 2, 5, 8: Toliaridectes wendenbaumi n. gen. n. sp. 3, 6, 9: Toliaridectes antsycurvis n. gen. n. sp. All pictured specimens are holotypes." figureDoi="http://doi.org/10.5281/zenodo.3616905" httpUri="https://zenodo.org/record/3616905/files/figure.png" pageId="1" pageNumber="547">Fig. 1</figureCitation>
; unarmed in
<taxonomicName id="4C454D51FFFF5409FD5AD3684FFBFBA7" authorityName="Rentz" authorityYear="1988" box="[644,765,1053,1079]" class="Insecta" family="Tettigoniidae" genus="Zuludectes" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="547" phylum="Arthropoda" rank="genus">
<emphasis id="B931EAC0FFFF5409FD5AD3684FFBFBA7" box="[644,765,1053,1079]" italics="true" pageId="1" pageNumber="547">Zuludectes</emphasis>
</taxonomicName>
). Fore tibia unarmed dorsally (
<figureCitation id="137E2A57FFFF5409FB8DD36849D6FBA8" box="[1107,1232,1053,1080]" captionStart="FIGURES 10–24" captionStartId="4.[151,264,1676,1701]" captionTargetBox="[156,1432,167,1652]" captionTargetId="figure@4.[151,1436,166,1657]" captionTargetPageId="4" captionText="FIGURES 10–24. Toliaridectes n. gen. legs. 10–12: left fore tibia inner view. 13–15: left fore tibia outer view. 16–18: left mid tibia anterior view. 19–21: left mid tibia posterior view. 22–24: left hind femur outer view. 10, 13, 16, 19, 22: Toliaridectes meridionalis n. gen. n. sp. 11, 14, 17, 20, 23: Toliaridectes wendenbaumi n. gen. n. sp. 12, 15, 18, 21, 24: Toliaridectes antsycurvis n. gen. n. sp. All pictured specimens are holotypes." figureDoi="http://doi.org/10.5281/zenodo.3616909" httpUri="https://zenodo.org/record/3616909/files/figure.png" pageId="1" pageNumber="547">Fig. 10–15</figureCitation>
; armed in all other
<taxonomicName id="4C454D51FFFF5409FF49D3344C46FBCB" authorityName="Caudell" authorityYear="1908" box="[151,320,1089,1115]" class="Insecta" family="Tettigoniidae" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="547" phylum="Arthropoda" rank="tribe" tribe="Arytropteridini">
<emphasis id="B931EAC0FFFF5409FF49D3344C46FBCB" box="[151,320,1089,1115]" italics="true" pageId="1" pageNumber="547">Arytropteridini</emphasis>
</taxonomicName>
). Mid tibia armed with 3 dorsal posterior spurs, not armed on dorsal anterior surface (
<figureCitation id="137E2A57FFFF5409FAC7D3344891FBCB" box="[1305,1431,1089,1115]" captionStart="FIGURES 10–24" captionStartId="4.[151,264,1676,1701]" captionTargetBox="[156,1432,167,1652]" captionTargetId="figure@4.[151,1436,166,1657]" captionTargetPageId="4" captionText="FIGURES 10–24. Toliaridectes n. gen. legs. 10–12: left fore tibia inner view. 13–15: left fore tibia outer view. 16–18: left mid tibia anterior view. 19–21: left mid tibia posterior view. 22–24: left hind femur outer view. 10, 13, 16, 19, 22: Toliaridectes meridionalis n. gen. n. sp. 11, 14, 17, 20, 23: Toliaridectes wendenbaumi n. gen. n. sp. 12, 15, 18, 21, 24: Toliaridectes antsycurvis n. gen. n. sp. All pictured specimens are holotypes." figureDoi="http://doi.org/10.5281/zenodo.3616909" httpUri="https://zenodo.org/record/3616909/files/figure.png" pageId="1" pageNumber="547">Fig. 16–21</figureCitation>
; armed in all other
<taxonomicName id="4C454D51FFFF5409FEB8D3104F09FBEF" authorityName="Caudell" authorityYear="1908" box="[358,527,1125,1151]" class="Insecta" family="Tettigoniidae" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="547" phylum="Arthropoda" rank="tribe" tribe="Arytropteridini">
<emphasis id="B931EAC0FFFF5409FEB8D3104F09FBEF" box="[358,527,1125,1151]" italics="true" pageId="1" pageNumber="547">Arytropteridini</emphasis>
</taxonomicName>
). Male forewings with fewer than 45 teeth (
<figureCitation id="137E2A57FFFF5409FBDED31049AEFBEF" box="[1024,1192,1125,1151]" captionStart="FIGURES 25–33" captionStartId="6.[151,264,1907,1932]" captionTargetBox="[446,1113,175,1881]" captionTargetId="figure@6.[445,1116,166,1887]" captionTargetPageId="6" captionText="FIGURES 25–33. Toliaridectes n. gen. male forewings. 25, 28, 31: left forewing in dorsal view (removed from specimen). 26, 29, 32: right forewing in dorsal view (removed from specimen). 27, 30, 33: file in ventral view. 25–27: Toliaridectes meridionalis n. gen. n. sp. 28–30: Toliaridectes wendenbaumi n. gen. n. sp. 31–33: Toliaridectes antsycurvis n. gen. n. sp. All pictured specimens are holotypes." figureDoi="http://doi.org/10.5281/zenodo.3616911" httpUri="https://zenodo.org/record/3616911/files/figure.png" pageId="1" pageNumber="547">Fig. 27, 30, 33</figureCitation>
; when known,&gt;
<quantity id="4CBD9B37FFFF5409FABDD310489AFBEF" box="[1379,1436,1125,1151]" metricMagnitude="0" metricUnit="m" metricValue="1.2446" pageId="1" pageNumber="547" unit="in" value="49.0">49 in</quantity>
all
<taxonomicName id="4C454D51FFFF5409FF64D3FC4C6DFB33" authorityName="Caudell" authorityYear="1908" box="[186,363,1161,1187]" class="Insecta" family="Tettigoniidae" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="547" phylum="Arthropoda" rank="tribe" tribe="Arytropteridini">
<emphasis id="B931EAC0FFFF5409FF64D3FC4C6DFB33" box="[186,363,1161,1187]" italics="true" pageId="1" pageNumber="547">Arytropteridini</emphasis>
</taxonomicName>
except
<taxonomicName id="4C454D51FFFF5409FE18D3FC4F43FB33" authorityName="Rentz" authorityYear="1988" box="[454,581,1161,1187]" class="Insecta" family="Tettigoniidae" genus="Zuludectes" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="547" phylum="Arthropoda" rank="genus">
<emphasis id="B931EAC0FFFF5409FE18D3FC4F43FB33" box="[454,581,1161,1187]" italics="true" pageId="1" pageNumber="547">Zuludectes</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C35F6559FFFF540AFF19D3D84C9DFB83" lastPageId="2" lastPageNumber="548" pageId="1" pageNumber="547" type="description">
<paragraph id="8BFA36D2FFFF5409FF19D3D84E57FB57" blockId="1.[151,1437,837,2016]" box="[199,849,1197,1224]" pageId="1" pageNumber="547">
<emphasis id="B931EAC0FFFF5409FF19D3D84C5FFB58" bold="true" box="[199,345,1197,1224]" pageId="1" pageNumber="547">Description.</emphasis>
Body size average for the tribe, form robust.
</paragraph>
<paragraph id="8BFA36D2FFFF5409FF19D3A44885FAEC" blockId="1.[151,1437,837,2016]" pageId="1" pageNumber="547">
<emphasis id="B931EAC0FFFF5409FF19D3A44C16FB7C" bold="true" box="[199,272,1233,1260]" pageId="1" pageNumber="547">Color.</emphasis>
General coloration brown with dark and light patterns (
<figureCitation id="137E2A57FFFF5409FCA9D3A44EF2FB7C" box="[887,1012,1233,1260]" captionStart="FIGURES 68–71" captionStartId="11.[151,264,1792,1817]" captionTargetBox="[151,1436,166,1773]" captionTargetId="figure@11.[151,1436,166,1773]" captionTargetPageId="11" captionText="FIGURES 68–71. Toliaridectes n. gen. in natura. 68–69: Toliaridectes wendenbaumi n. gen. n. sp., male in resting position (68); female eating a caterpillar (69). 70–71: Toliaridectes antsycurvis n. gen. n. sp., male calling (70; specimen 2019MA- DASH042, see figures 64–67); female in oviposition (71)." figureDoi="http://doi.org/10.5281/zenodo.3616931" httpUri="https://zenodo.org/record/3616931/files/figure.png" pageId="1" pageNumber="547">Fig. 68–71</figureCitation>
). Face with sagittal black pattern from antennal sockets to labrum tip. Black retro-ocular patterns often edged with dorsal light pattern. Scapus and pedicel with black patterns. Pronotum folds (see
<bibRefCitation id="EFD44B23FFFF5409FDB6D26F4FECFAA3" author="Rentz, D. C. F." box="[616,746,1305,1332]" pageId="1" pageNumber="547" pagination="223 - 335" refId="ref7226" refString="Rentz, D. C. F. (1988) The shield-backed katylids of Southern Africa: their taxonomy, ecology and relationships to the faunas of Australia and South America (Orthoptera: Tettigoniidae: Tettigoniinae). Invertebrate Systematics, 2 (2), 223 - 335. https: // doi. org / 10.1071 / IT 9880223" type="journal article" year="1988">Rentz 1988</bibRefCitation>
) with black pattern in line with retroocular black lines, often edged dorsally with light stripes; ventral and posterior rims of lateral lobes black. Abdomen with black patterns laterally, often edged dorsally with light stripes. Hind femur outer field with a dark or black longitudinal pattern.
</paragraph>
<paragraph id="8BFA36D2FFFF5409FF19D2F34E95F974" blockId="1.[151,1437,837,2016]" pageId="1" pageNumber="547">
<emphasis id="B931EAC0FFFF5409FF19D2F34C08FA30" bold="true" box="[199,270,1414,1440]" pageId="1" pageNumber="547">Head.</emphasis>
Head in frontal view 1.4–1.8 times higher than wide, narrower than pronotum in dorsal view (
<figureCitation id="137E2A57FFFF5409FAF0D2F04889FA0F" box="[1326,1423,1413,1439]" captionStart="FIGURES 1–9" captionStartId="3.[151,264,1841,1866]" captionTargetBox="[158,1422,482,1819]" captionTargetId="figure@3.[151,1436,482,1821]" captionTargetPageId="3" captionText="FIGURES 1–9. Toliaridectes n. gen. male head and pronotum. 1–3: head and pronotum in dorsal view, before removing the forewings for examination. 4–6: head and pronotum in left side view. 7–9: face in frontal view. 1, 4, 7: Toliaridectes meridionalis n. gen. n. sp. 2, 5, 8: Toliaridectes wendenbaumi n. gen. n. sp. 3, 6, 9: Toliaridectes antsycurvis n. gen. n. sp. All pictured specimens are holotypes." figureDoi="http://doi.org/10.5281/zenodo.3616905" httpUri="https://zenodo.org/record/3616905/files/figure.png" pageId="1" pageNumber="547">Fig. 1–3</figureCitation>
). Fastigium verticis narrower than scapus (
<figureCitation id="137E2A57FFFF5409FDB5D2DC4FCAFA53" box="[619,716,1449,1475]" captionStart="FIGURES 1–9" captionStartId="3.[151,264,1841,1866]" captionTargetBox="[158,1422,482,1819]" captionTargetId="figure@3.[151,1436,482,1821]" captionTargetPageId="3" captionText="FIGURES 1–9. Toliaridectes n. gen. male head and pronotum. 1–3: head and pronotum in dorsal view, before removing the forewings for examination. 4–6: head and pronotum in left side view. 7–9: face in frontal view. 1, 4, 7: Toliaridectes meridionalis n. gen. n. sp. 2, 5, 8: Toliaridectes wendenbaumi n. gen. n. sp. 3, 6, 9: Toliaridectes antsycurvis n. gen. n. sp. All pictured specimens are holotypes." figureDoi="http://doi.org/10.5281/zenodo.3616905" httpUri="https://zenodo.org/record/3616905/files/figure.png" pageId="1" pageNumber="547">Fig. 1–3</figureCitation>
), triangular in frontal view (
<figureCitation id="137E2A57FFFF5409FBCED2DC4974FA53" box="[1040,1138,1449,1475]" captionStart="FIGURES 1–9" captionStartId="3.[151,264,1841,1866]" captionTargetBox="[158,1422,482,1819]" captionTargetId="figure@3.[151,1436,482,1821]" captionTargetPageId="3" captionText="FIGURES 1–9. Toliaridectes n. gen. male head and pronotum. 1–3: head and pronotum in dorsal view, before removing the forewings for examination. 4–6: head and pronotum in left side view. 7–9: face in frontal view. 1, 4, 7: Toliaridectes meridionalis n. gen. n. sp. 2, 5, 8: Toliaridectes wendenbaumi n. gen. n. sp. 3, 6, 9: Toliaridectes antsycurvis n. gen. n. sp. All pictured specimens are holotypes." figureDoi="http://doi.org/10.5281/zenodo.3616905" httpUri="https://zenodo.org/record/3616905/files/figure.png" pageId="1" pageNumber="547">Fig. 7–9</figureCitation>
), no fastigial gap between fastigium verticis and fastigium frontis, fastigium frontis contacting the fastigium verticis in a narrow ridge (
<figureCitation id="137E2A57FFFF5409FAB1D2B84DC3F99B" captionStart="FIGURES 1–9" captionStartId="3.[151,264,1841,1866]" captionTargetBox="[158,1422,482,1819]" captionTargetId="figure@3.[151,1436,482,1821]" captionTargetPageId="3" captionText="FIGURES 1–9. Toliaridectes n. gen. male head and pronotum. 1–3: head and pronotum in dorsal view, before removing the forewings for examination. 4–6: head and pronotum in left side view. 7–9: face in frontal view. 1, 4, 7: Toliaridectes meridionalis n. gen. n. sp. 2, 5, 8: Toliaridectes wendenbaumi n. gen. n. sp. 3, 6, 9: Toliaridectes antsycurvis n. gen. n. sp. All pictured specimens are holotypes." figureDoi="http://doi.org/10.5281/zenodo.3616905" httpUri="https://zenodo.org/record/3616905/files/figure.png" pageId="1" pageNumber="547">Fig. 4–9</figureCitation>
). Median ocellus very distinct, in line with ventral part of antennal scrobus (
<figureCitation id="137E2A57FFFF5409FBFCD2844983F99B" box="[1058,1157,1521,1547]" captionStart="FIGURES 1–9" captionStartId="3.[151,264,1841,1866]" captionTargetBox="[158,1422,482,1819]" captionTargetId="figure@3.[151,1436,482,1821]" captionTargetPageId="3" captionText="FIGURES 1–9. Toliaridectes n. gen. male head and pronotum. 1–3: head and pronotum in dorsal view, before removing the forewings for examination. 4–6: head and pronotum in left side view. 7–9: face in frontal view. 1, 4, 7: Toliaridectes meridionalis n. gen. n. sp. 2, 5, 8: Toliaridectes wendenbaumi n. gen. n. sp. 3, 6, 9: Toliaridectes antsycurvis n. gen. n. sp. All pictured specimens are holotypes." figureDoi="http://doi.org/10.5281/zenodo.3616905" httpUri="https://zenodo.org/record/3616905/files/figure.png" pageId="1" pageNumber="547">Fig. 7–9</figureCitation>
). Genae with no carinae. Scapus unarmed.
<emphasis id="B931EAC0FFFF5409FEBED1634CC6F9A0" bold="true" box="[352,448,1558,1584]" pageId="1" pageNumber="547">Thorax.</emphasis>
Pronotum little modified, slightly different in males and females, in both cases not exceeding first abdominal segment, not producing posteriorly, without median carina, without lateral carinae, rims of lateral lobe and disk not enlarged; disk flat in side view, arched in cross section; anterior margin inconspicuously concave; posterior margin broadly rounded; surface of disk smooth, faintly shining; disk prozona traversed by a transverse sulcus shaped as a curly bracket (
<figureCitation id="137E2A57FFFF5409FDA0D1D04FD8F92F" box="[638,734,1701,1727]" captionStart="FIGURES 1–9" captionStartId="3.[151,264,1841,1866]" captionTargetBox="[158,1422,482,1819]" captionTargetId="figure@3.[151,1436,482,1821]" captionTargetPageId="3" captionText="FIGURES 1–9. Toliaridectes n. gen. male head and pronotum. 1–3: head and pronotum in dorsal view, before removing the forewings for examination. 4–6: head and pronotum in left side view. 7–9: face in frontal view. 1, 4, 7: Toliaridectes meridionalis n. gen. n. sp. 2, 5, 8: Toliaridectes wendenbaumi n. gen. n. sp. 3, 6, 9: Toliaridectes antsycurvis n. gen. n. sp. All pictured specimens are holotypes." figureDoi="http://doi.org/10.5281/zenodo.3616905" httpUri="https://zenodo.org/record/3616905/files/figure.png" pageId="1" pageNumber="547">Fig. 1–3</figureCitation>
). Prosternum armed with a pair of broadly spaced blunt erected spines; meso and metasternum with a pair of more robust processes.
</paragraph>
<paragraph id="8BFA36D2FFFF5409FF19D1984E2AF897" blockId="1.[151,1437,837,2016]" box="[199,812,1773,1800]" pageId="1" pageNumber="547">
<emphasis id="B931EAC0FFFF5409FF19D1984C1CF898" bold="true" box="[199,282,1773,1800]" pageId="1" pageNumber="547">Wings.</emphasis>
Forewings present in males, absent in females.
</paragraph>
<paragraph id="8BFA36D2FFFF540AFF19D0674CD5FEF7" blockId="1.[151,1437,837,2016]" lastBlockId="2.[151,1437,152,1043]" lastPageId="2" lastPageNumber="548" pageId="1" pageNumber="547">
<emphasis id="B931EAC0FFFF5409FF19D0674C00F8BC" bold="true" box="[199,262,1810,1836]" pageId="1" pageNumber="547">Legs.</emphasis>
Leg surface covered with numerous minute imprints, often pigmented and bearing small setae. Fore coxa with one median antero lateral spine, relatively flattened (
<figureCitation id="137E2A57FFFF5409FCB3D0404ED6F8DF" box="[877,976,1845,1871]" captionStart="FIGURES 1–9" captionStartId="3.[151,264,1841,1866]" captionTargetBox="[158,1422,482,1819]" captionTargetId="figure@3.[151,1436,482,1821]" captionTargetPageId="3" captionText="FIGURES 1–9. Toliaridectes n. gen. male head and pronotum. 1–3: head and pronotum in dorsal view, before removing the forewings for examination. 4–6: head and pronotum in left side view. 7–9: face in frontal view. 1, 4, 7: Toliaridectes meridionalis n. gen. n. sp. 2, 5, 8: Toliaridectes wendenbaumi n. gen. n. sp. 3, 6, 9: Toliaridectes antsycurvis n. gen. n. sp. All pictured specimens are holotypes." figureDoi="http://doi.org/10.5281/zenodo.3616905" httpUri="https://zenodo.org/record/3616905/files/figure.png" pageId="1" pageNumber="547">Fig. 1–3</figureCitation>
). Fore femora with 1–4 minute spinules in anterior and posterior ventral margins (rarely none on posterior margin of
<taxonomicName id="4C454D51FFFF5409FBCFD02F49A9F8E3" authority="Hugel, 2019" authorityName="Hugel" authorityYear="2019" box="[1041,1199,1882,1907]" class="Insecta" family="Tettigoniidae" genus="Toliaridectes" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="547" phylum="Arthropoda" rank="species" species="antsycurvis" status="gen. nov.">
<emphasis id="B931EAC0FFFF5409FBCFD02F49A9F8E3" box="[1041,1199,1882,1907]" italics="true" pageId="1" pageNumber="547">T. antsycurvis</emphasis>
</taxonomicName>
<emphasis id="B931EAC0FFFF5409FB66D02F4852F8E4" bold="true" box="[1208,1364,1882,1908]" pageId="1" pageNumber="547">
<taxonomicNameLabel id="A20257BBFFFF5409FB66D02F480FF8E4" box="[1208,1289,1882,1908]" pageId="1" pageNumber="547" rank="genus">n. gen.</taxonomicNameLabel>
n. sp.
</emphasis>
); fore femora genicular lobes with 2 very small spinules on both sides. Fore tibia (
<figureCitation id="137E2A57FFFF5409FC35D008496CF808" box="[1003,1130,1917,1944]" captionStart="FIGURES 10–24" captionStartId="4.[151,264,1676,1701]" captionTargetBox="[156,1432,167,1652]" captionTargetId="figure@4.[151,1436,166,1657]" captionTargetPageId="4" captionText="FIGURES 10–24. Toliaridectes n. gen. legs. 10–12: left fore tibia inner view. 13–15: left fore tibia outer view. 16–18: left mid tibia anterior view. 19–21: left mid tibia posterior view. 22–24: left hind femur outer view. 10, 13, 16, 19, 22: Toliaridectes meridionalis n. gen. n. sp. 11, 14, 17, 20, 23: Toliaridectes wendenbaumi n. gen. n. sp. 12, 15, 18, 21, 24: Toliaridectes antsycurvis n. gen. n. sp. All pictured specimens are holotypes." figureDoi="http://doi.org/10.5281/zenodo.3616909" httpUri="https://zenodo.org/record/3616909/files/figure.png" pageId="1" pageNumber="547">Fig. 10–15</figureCitation>
): with 6 ventral inner spurs (including apical spur); with 6 ventral outer spurs (including apical spur); without dorsal apical and subapical spurs. Auditory foramen narrow, slit shaped, visible in side view (
<figureCitation id="137E2A57FFFF5409FCF9D0B04EA2F870" box="[807,932,1989,2016]" captionStart="FIGURES 10–24" captionStartId="4.[151,264,1676,1701]" captionTargetBox="[156,1432,167,1652]" captionTargetId="figure@4.[151,1436,166,1657]" captionTargetPageId="4" captionText="FIGURES 10–24. Toliaridectes n. gen. legs. 10–12: left fore tibia inner view. 13–15: left fore tibia outer view. 16–18: left mid tibia anterior view. 19–21: left mid tibia posterior view. 22–24: left hind femur outer view. 10, 13, 16, 19, 22: Toliaridectes meridionalis n. gen. n. sp. 11, 14, 17, 20, 23: Toliaridectes wendenbaumi n. gen. n. sp. 12, 15, 18, 21, 24: Toliaridectes antsycurvis n. gen. n. sp. All pictured specimens are holotypes." figureDoi="http://doi.org/10.5281/zenodo.3616909" httpUri="https://zenodo.org/record/3616909/files/figure.png" pageId="1" pageNumber="547">Fig. 10–15</figureCitation>
). Mid coxa with a short ventral blunt process. Mid femora with 2–5 minute spinules in anterior and posterior ventral margins (rarely absent on posterior margin of
<taxonomicName id="4C454D51FFFC540AFF49D7CB4C35FF47" authority="Hugel, 2019" authorityName="Hugel" authorityYear="2019" box="[151,307,190,215]" class="Insecta" family="Tettigoniidae" genus="Toliaridectes" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="548" phylum="Arthropoda" rank="species" species="antsycurvis" status="gen. nov.">
<emphasis id="B931EAC0FFFC540AFF49D7CB4C35FF47" box="[151,307,190,215]" italics="true" pageId="2" pageNumber="548">T. antsycurvis</emphasis>
</taxonomicName>
<emphasis id="B931EAC0FFFC540AFEE4D7C84CD6FF47" bold="true" box="[314,464,189,215]" pageId="2" pageNumber="548">
<taxonomicNameLabel id="A20257BBFFFC540AFEE4D7C84C8FFF47" box="[314,393,189,215]" pageId="2" pageNumber="548" rank="genus">n. gen.</taxonomicNameLabel>
n. sp.
</emphasis>
); mid femora genicular lobes with 2 very small spinules on both sides. Mid tibia (
<figureCitation id="137E2A57FFFC540AFAB1D7C84DE4FF6C" captionStart="FIGURES 10–24" captionStartId="4.[151,264,1676,1701]" captionTargetBox="[156,1432,167,1652]" captionTargetId="figure@4.[151,1436,166,1657]" captionTargetPageId="4" captionText="FIGURES 10–24. Toliaridectes n. gen. legs. 10–12: left fore tibia inner view. 13–15: left fore tibia outer view. 16–18: left mid tibia anterior view. 19–21: left mid tibia posterior view. 22–24: left hind femur outer view. 10, 13, 16, 19, 22: Toliaridectes meridionalis n. gen. n. sp. 11, 14, 17, 20, 23: Toliaridectes wendenbaumi n. gen. n. sp. 12, 15, 18, 21, 24: Toliaridectes antsycurvis n. gen. n. sp. All pictured specimens are holotypes." figureDoi="http://doi.org/10.5281/zenodo.3616909" httpUri="https://zenodo.org/record/3616909/files/figure.png" pageId="2" pageNumber="548">Fig. 16–21</figureCitation>
): with 3 dorsal posterior subapical spurs; without dorsal apical spurs; with 6 ventral inner spurs (including apical spur); with 6 ventral outer spurs (including apical spur). Hind coxa with a short ventral blunt process. Hind femora with 6–10 spines on ventral inner and ventral outer carinae (
<figureCitation id="137E2A57FFFC540AFC4BD65C4912FED3" box="[917,1044,297,323]" captionStart="FIGURES 10–24" captionStartId="4.[151,264,1676,1701]" captionTargetBox="[156,1432,167,1652]" captionTargetId="figure@4.[151,1436,166,1657]" captionTargetPageId="4" captionText="FIGURES 10–24. Toliaridectes n. gen. legs. 10–12: left fore tibia inner view. 13–15: left fore tibia outer view. 16–18: left mid tibia anterior view. 19–21: left mid tibia posterior view. 22–24: left hind femur outer view. 10, 13, 16, 19, 22: Toliaridectes meridionalis n. gen. n. sp. 11, 14, 17, 20, 23: Toliaridectes wendenbaumi n. gen. n. sp. 12, 15, 18, 21, 24: Toliaridectes antsycurvis n. gen. n. sp. All pictured specimens are holotypes." figureDoi="http://doi.org/10.5281/zenodo.3616909" httpUri="https://zenodo.org/record/3616909/files/figure.png" pageId="2" pageNumber="548">Fig. 22–24</figureCitation>
); hind femora genicular lobes with a minute spine on each side.
</paragraph>
<paragraph id="8BFA36D2FFFC540AFF19D6054EDCFE1B" blockId="2.[151,1437,152,1043]" box="[199,986,368,395]" pageId="2" pageNumber="548">
<emphasis id="B931EAC0FFFC540AFF19D6054C45FE1B" bold="true" box="[199,323,368,395]" pageId="2" pageNumber="548">Abdomen.</emphasis>
Tergites with inconspicuous longitudinal carina (
<figureCitation id="137E2A57FFFC540AFCB2D6044ECBFE1B" box="[876,973,369,395]" captionStart="FIGURES 1–9" captionStartId="3.[151,264,1841,1866]" captionTargetBox="[158,1422,482,1819]" captionTargetId="figure@3.[151,1436,482,1821]" captionTargetPageId="3" captionText="FIGURES 1–9. Toliaridectes n. gen. male head and pronotum. 1–3: head and pronotum in dorsal view, before removing the forewings for examination. 4–6: head and pronotum in left side view. 7–9: face in frontal view. 1, 4, 7: Toliaridectes meridionalis n. gen. n. sp. 2, 5, 8: Toliaridectes wendenbaumi n. gen. n. sp. 3, 6, 9: Toliaridectes antsycurvis n. gen. n. sp. All pictured specimens are holotypes." figureDoi="http://doi.org/10.5281/zenodo.3616905" httpUri="https://zenodo.org/record/3616905/files/figure.png" pageId="2" pageNumber="548">Fig. 1–3</figureCitation>
).
</paragraph>
<paragraph id="8BFA36D2FFFC540AFF19D6E04CFBFC87" blockId="2.[151,1437,152,1043]" pageId="2" pageNumber="548">
<emphasis id="B931EAC0FFFC540AFF19D6E04C0DFE3F" bold="true" box="[199,267,405,431]" pageId="2" pageNumber="548">Male.</emphasis>
Pronotum (
<figureCitation id="137E2A57FFFC540AFE48D6E04CFEFE20" box="[406,504,405,432]" captionStart="FIGURES 1–9" captionStartId="3.[151,264,1841,1866]" captionTargetBox="[158,1422,482,1819]" captionTargetId="figure@3.[151,1436,482,1821]" captionTargetPageId="3" captionText="FIGURES 1–9. Toliaridectes n. gen. male head and pronotum. 1–3: head and pronotum in dorsal view, before removing the forewings for examination. 4–6: head and pronotum in left side view. 7–9: face in frontal view. 1, 4, 7: Toliaridectes meridionalis n. gen. n. sp. 2, 5, 8: Toliaridectes wendenbaumi n. gen. n. sp. 3, 6, 9: Toliaridectes antsycurvis n. gen. n. sp. All pictured specimens are holotypes." figureDoi="http://doi.org/10.5281/zenodo.3616905" httpUri="https://zenodo.org/record/3616905/files/figure.png" pageId="2" pageNumber="548">Fig. 1–6</figureCitation>
) reaching the middle of the first abdominal segment; disk more arched in cross section than in females, leaving room for the wings. Forewings (
<figureCitation id="137E2A57FFFC540AFCA9D6CC4EDEFE43" box="[887,984,441,467]" captionStart="FIGURES 1–9" captionStartId="3.[151,264,1841,1866]" captionTargetBox="[158,1422,482,1819]" captionTargetId="figure@3.[151,1436,482,1821]" captionTargetPageId="3" captionText="FIGURES 1–9. Toliaridectes n. gen. male head and pronotum. 1–3: head and pronotum in dorsal view, before removing the forewings for examination. 4–6: head and pronotum in left side view. 7–9: face in frontal view. 1, 4, 7: Toliaridectes meridionalis n. gen. n. sp. 2, 5, 8: Toliaridectes wendenbaumi n. gen. n. sp. 3, 6, 9: Toliaridectes antsycurvis n. gen. n. sp. All pictured specimens are holotypes." figureDoi="http://doi.org/10.5281/zenodo.3616905" httpUri="https://zenodo.org/record/3616905/files/figure.png" pageId="2" pageNumber="548">Fig. 1–3</figureCitation>
,
<figureCitation id="137E2A57FFFC540AFC38D6CD492BFE43" box="[998,1069,440,467]" captionStart="FIGURES 25–33" captionStartId="6.[151,264,1907,1932]" captionTargetBox="[446,1113,175,1881]" captionTargetId="figure@6.[445,1116,166,1887]" captionTargetPageId="6" captionText="FIGURES 25–33. Toliaridectes n. gen. male forewings. 25, 28, 31: left forewing in dorsal view (removed from specimen). 26, 29, 32: right forewing in dorsal view (removed from specimen). 27, 30, 33: file in ventral view. 25–27: Toliaridectes meridionalis n. gen. n. sp. 28–30: Toliaridectes wendenbaumi n. gen. n. sp. 31–33: Toliaridectes antsycurvis n. gen. n. sp. All pictured specimens are holotypes." figureDoi="http://doi.org/10.5281/zenodo.3616911" httpUri="https://zenodo.org/record/3616911/files/figure.png" pageId="2" pageNumber="548">25–33</figureCitation>
) slightly exceeding hind margin of pronotum, reaching the distal margin of the first abdominal segment; visible part of the forewing white or light green; file with 38–42 teeth, largest on the middle.
<superScript id="7C309B9AFFFC540AFD0BD5744FF0FD9E" attach="left" box="[725,758,512,539]" fontSize="6" pageId="2" pageNumber="548">Xth</superScript>
abdominal tergite posterior end with a median quadrangular notch of variable width separating two pointing lobes (
<figureCitation id="137E2A57FFFC540AFD2AD5504E9FFDAF" box="[756,921,548,575]" captionStart="FIGURES 34–54" captionStartId="7.[151,264,1792,1817]" captionTargetBox="[152,1436,176,1773]" captionTargetId="figure@7.[151,1436,166,1773]" captionTargetPageId="7" captionText="FIGURES 34–54. Toliaridectes n. gen. male terminalia. 34, 41, 48: subgenital plate (ventral view). 35, 42, 49: left paraproct (removed from specimen). 36, 43, 50: male suranal plate (dorsal view, the subgenital plate is lightened for better visualization). 37, 44, 51: left cercus (dorsal view). 38, 45, 52: left cercus (inner view). 39, 46, 53: left cercus (ventral view). 40, 47, 54: left cercus (outer view). 34–40: Toliaridectes meridionalis n. gen. n. sp. 41–47: Toliaridectes wendenbaumi n. gen. n. sp. 48–54: Toliaridectes antsycurvis n. gen. n. sp. All pictured specimens are holotypes." figureDoi="http://doi.org/10.5281/zenodo.3616915" httpUri="https://zenodo.org/record/3616915/files/figure.png" pageId="2" pageNumber="548">Fig. 36, 43, 50</figureCitation>
). Paraprocts very small, with a ventral-median projection variable in size (
<figureCitation id="137E2A57FFFC540AFE10D53C4F7DFDF3" box="[462,635,584,611]" captionStart="FIGURES 34–54" captionStartId="7.[151,264,1792,1817]" captionTargetBox="[152,1436,176,1773]" captionTargetId="figure@7.[151,1436,166,1773]" captionTargetPageId="7" captionText="FIGURES 34–54. Toliaridectes n. gen. male terminalia. 34, 41, 48: subgenital plate (ventral view). 35, 42, 49: left paraproct (removed from specimen). 36, 43, 50: male suranal plate (dorsal view, the subgenital plate is lightened for better visualization). 37, 44, 51: left cercus (dorsal view). 38, 45, 52: left cercus (inner view). 39, 46, 53: left cercus (ventral view). 40, 47, 54: left cercus (outer view). 34–40: Toliaridectes meridionalis n. gen. n. sp. 41–47: Toliaridectes wendenbaumi n. gen. n. sp. 48–54: Toliaridectes antsycurvis n. gen. n. sp. All pictured specimens are holotypes." figureDoi="http://doi.org/10.5281/zenodo.3616915" httpUri="https://zenodo.org/record/3616915/files/figure.png" pageId="2" pageNumber="548">Fig. 35, 42, 49</figureCitation>
). Subgenital plate (
<figureCitation id="137E2A57FFFC540AFC89D53C4902FDF3" box="[855,1028,585,611]" captionStart="FIGURES 34–54" captionStartId="7.[151,264,1792,1817]" captionTargetBox="[152,1436,176,1773]" captionTargetId="figure@7.[151,1436,166,1773]" captionTargetPageId="7" captionText="FIGURES 34–54. Toliaridectes n. gen. male terminalia. 34, 41, 48: subgenital plate (ventral view). 35, 42, 49: left paraproct (removed from specimen). 36, 43, 50: male suranal plate (dorsal view, the subgenital plate is lightened for better visualization). 37, 44, 51: left cercus (dorsal view). 38, 45, 52: left cercus (inner view). 39, 46, 53: left cercus (ventral view). 40, 47, 54: left cercus (outer view). 34–40: Toliaridectes meridionalis n. gen. n. sp. 41–47: Toliaridectes wendenbaumi n. gen. n. sp. 48–54: Toliaridectes antsycurvis n. gen. n. sp. All pictured specimens are holotypes." figureDoi="http://doi.org/10.5281/zenodo.3616915" httpUri="https://zenodo.org/record/3616915/files/figure.png" pageId="2" pageNumber="548">Fig. 34, 41, 48</figureCitation>
): broadly triangular in ventral view, with a shallow median notch between styli; with distinct styli. Cercus (
<figureCitation id="137E2A57FFFC540AFC68D51849DDFD17" box="[950,1243,620,647]" captionStart="FIGURES 34–54" captionStartId="7.[151,264,1792,1817]" captionTargetBox="[152,1436,176,1773]" captionTargetId="figure@7.[151,1436,166,1773]" captionTargetPageId="7" captionText="FIGURES 34–54. Toliaridectes n. gen. male terminalia. 34, 41, 48: subgenital plate (ventral view). 35, 42, 49: left paraproct (removed from specimen). 36, 43, 50: male suranal plate (dorsal view, the subgenital plate is lightened for better visualization). 37, 44, 51: left cercus (dorsal view). 38, 45, 52: left cercus (inner view). 39, 46, 53: left cercus (ventral view). 40, 47, 54: left cercus (outer view). 34–40: Toliaridectes meridionalis n. gen. n. sp. 41–47: Toliaridectes wendenbaumi n. gen. n. sp. 48–54: Toliaridectes antsycurvis n. gen. n. sp. All pictured specimens are holotypes." figureDoi="http://doi.org/10.5281/zenodo.3616915" httpUri="https://zenodo.org/record/3616915/files/figure.png" pageId="2" pageNumber="548">Fig. 37–40, 44–47, 51–54</figureCitation>
) elongate, proximal half broad, distal half narrow, with two projections on the middle: one dorsal lobe pointing toward midline, one median projection pointing toward midline. Titillators well sclerotized, external (distal) part pointing (
<figureCitation id="137E2A57FFFC540AFB20D5C04871FD5F" box="[1278,1399,692,719]" captionStart="FIGURES 55–57" captionStartId="8.[151,264,711,736]" captionTargetBox="[152,1430,173,682]" captionTargetId="figure@8.[151,1436,166,691]" captionTargetPageId="8" captionText="FIGURES 55–57. Toliaridectes n. gen. male genitalia (dorsal view, posterior side on top). 55: Toliaridectes meridionalis n. gen. n. sp. 56: Toliaridectes wendenbaumi n. gen. n. sp. 57: Toliaridectes antsycurvis n. gen. n. sp. All pictured specimens are holotypes." figureDoi="http://doi.org/10.5281/zenodo.3616919" httpUri="https://zenodo.org/record/3616919/files/figure.png" pageId="2" pageNumber="548">Fig. 55, 56</figureCitation>
) or enlarged (
<figureCitation id="137E2A57FFFC540AFED8D5AC4C50FD63" box="[262,342,728,755]" captionStart="FIGURES 55–57" captionStartId="8.[151,264,711,736]" captionTargetBox="[152,1430,173,682]" captionTargetId="figure@8.[151,1436,166,691]" captionTargetPageId="8" captionText="FIGURES 55–57. Toliaridectes n. gen. male genitalia (dorsal view, posterior side on top). 55: Toliaridectes meridionalis n. gen. n. sp. 56: Toliaridectes wendenbaumi n. gen. n. sp. 57: Toliaridectes antsycurvis n. gen. n. sp. All pictured specimens are holotypes." figureDoi="http://doi.org/10.5281/zenodo.3616919" httpUri="https://zenodo.org/record/3616919/files/figure.png" pageId="2" pageNumber="548">Fig. 57</figureCitation>
) distally in dorsal view, with minute denticles; internal (proximal) part with two branches forming an inverted Y or T (
<figureCitation id="137E2A57FFFC540AFEACD5884CF7FC87" box="[370,497,764,791]" captionStart="FIGURES 55–57" captionStartId="8.[151,264,711,736]" captionTargetBox="[152,1430,173,682]" captionTargetId="figure@8.[151,1436,166,691]" captionTargetPageId="8" captionText="FIGURES 55–57. Toliaridectes n. gen. male genitalia (dorsal view, posterior side on top). 55: Toliaridectes meridionalis n. gen. n. sp. 56: Toliaridectes wendenbaumi n. gen. n. sp. 57: Toliaridectes antsycurvis n. gen. n. sp. All pictured specimens are holotypes." figureDoi="http://doi.org/10.5281/zenodo.3616919" httpUri="https://zenodo.org/record/3616919/files/figure.png" pageId="2" pageNumber="548">Fig. 55–57</figureCitation>
).
</paragraph>
<paragraph id="8BFA36D2FFFC540AFF19D4544DE3FC5B" blockId="2.[151,1437,152,1043]" pageId="2" pageNumber="548">
<emphasis id="B931EAC0FFFC540AFF19D4544C23FCAB" bold="true" box="[199,293,801,827]" pageId="2" pageNumber="548">Female.</emphasis>
Pronotum hardly reaching metathoracic posterior margin, disk more flattened posteriorly than in males. Wings absent. Suranal plate small, triangular or rounded, with or without a longitudinal sulcus. Subgenital plate (
<figureCitation id="137E2A57FFFC540AFF7ED41C4C45FC14" box="[160,323,872,900]" captionStart="FIGURES 58–63" captionStartId="8.[151,264,1829,1854]" captionTargetBox="[152,1429,861,1810]" captionTargetId="figure@8.[151,1436,851,1810]" captionTargetPageId="8" captionText="FIGURES 58–63. Toliaridectes n. gen. female terminalia. 58, 60, 62: subgenital plate (ventral view, posterior side on top). 59, 61, 63: ovipositor (left side view). 58–59: Toliaridectes meridionalis n. gen. n. sp. 60–61: Toliaridectes wendenbaumi n. gen. n. sp. 62–63: Toliaridectes antsycurvis n. gen. n. sp." figureDoi="http://doi.org/10.5281/zenodo.3616923" httpUri="https://zenodo.org/record/3616923/files/figure.png" pageId="2" pageNumber="548">Fig. 58, 60, 62</figureCitation>
) of variable shape, posterior end with a median emargination of variable width and depth separating two pointing lobes. Ovipositor (
<figureCitation id="137E2A57FFFC540AFE26D4F84F9CFC38" box="[504,666,908,936]" captionStart="FIGURES 58–63" captionStartId="8.[151,264,1829,1854]" captionTargetBox="[152,1429,861,1810]" captionTargetId="figure@8.[151,1436,851,1810]" captionTargetPageId="8" captionText="FIGURES 58–63. Toliaridectes n. gen. female terminalia. 58, 60, 62: subgenital plate (ventral view, posterior side on top). 59, 61, 63: ovipositor (left side view). 58–59: Toliaridectes meridionalis n. gen. n. sp. 60–61: Toliaridectes wendenbaumi n. gen. n. sp. 62–63: Toliaridectes antsycurvis n. gen. n. sp." figureDoi="http://doi.org/10.5281/zenodo.3616923" httpUri="https://zenodo.org/record/3616923/files/figure.png" pageId="2" pageNumber="548">Fig. 59, 61, 63</figureCitation>
) flattened proximally, upcurved distally; the flattened part of variable length.
</paragraph>
<paragraph id="8BFA36D2FFFC540AFF19D4A14C9DFB83" blockId="2.[151,1437,152,1043]" pageId="2" pageNumber="548">
<emphasis id="B931EAC0FFFC540AFF19D4A14CBEFC7F" bold="true" box="[199,440,980,1007]" pageId="2" pageNumber="548">Derivation of name:</emphasis>
Named after the Malagasy province “
<collectingRegion id="4981F830FFFC540AFCBAD4A04EBFFC7F" box="[868,953,981,1007]" country="Madagascar" name="Toliara" pageId="2" pageNumber="548">Toliara</collectingRegion>
” where all known species of
<taxonomicName id="4C454D51FFFC540AFADBD4A0489AFC7F" authority="Hugel, 2019" authorityName="Hugel" authorityYear="2019" box="[1285,1436,981,1007]" class="Insecta" family="Tettigoniidae" genus="Toliaradectes" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="548" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B931EAC0FFFC540AFADBD4A0489AFC7F" box="[1285,1436,981,1007]" italics="true" pageId="2" pageNumber="548">Toliaradectes</emphasis>
</taxonomicName>
<emphasis id="B931EAC0FFFC540AFF49D48C4DE0FB83" bold="true" box="[151,230,1017,1043]" pageId="2" pageNumber="548">
<taxonomicNameLabel id="A20257BBFFFC540AFF49D48C4DE0FB83" box="[151,230,1017,1043]" pageId="2" pageNumber="548" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
occur (
<figureCitation id="137E2A57FFFC540AFEE4D48C4C89FB83" box="[314,399,1017,1043]" captionStart="FIGURE 72" captionStartId="12.[151,250,1594,1619]" captionTargetId="figure@12.[290,1297,166,1574]" captionTargetPageId="12" captionText="FIGURE 72. Distribution of Toliaridectes n. gen." figureDoi="http://doi.org/10.5281/zenodo.3616935" httpUri="https://zenodo.org/record/3616935/files/figure.png" pageId="2" pageNumber="548">Fig. 72</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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