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<mods:title id="836FBF01C9BD66A15E53BC45AEF4661C">On two new species of the shrimp genus Salmoneus Holthuis, 1955 (Decapoda, Caridea, Alpheidae) from the tropical eastern Pacific</mods:title>
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<emphasis id="B931EA9B2147FFC748A89360A3CEFCCD" bold="true" box="[151,386,780,806]" italics="true" pageId="1" pageNumber="521">Salmoneus alvarezi</emphasis>
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(
<figureCitation id="137E2A0C2147FFC748A09343A345FCA3" box="[159,265,815,840]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,250,1798,1820]" captionStartId-1="3.[151,250,1318,1340]" captionStartId-2="5.[151,250,1952,1974]" captionTargetBox-0="[162,1411,208,1768]" captionTargetBox-1="[162,1428,202,1288]" captionTargetBox-2="[151,1434,183,1854]" captionTargetId-0="figure@2.[151,1436,193,1777]" captionTargetId-1="figure@3.[151,1436,193,1297]" captionTargetId-2="figure@5.[151,1436,183,1942]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="5" captionText-0="FIGURE 1. Salmoneus alvarezi sp. nov., holotype from Bahía Málaga, Colombia (INV CRU 8365): A—frontal region, dorsal; B—same, lateral; C—posterior abdominal somites, lateral; D—telson, dorsal; E—same, posterior margin, dorsal; F—antennal scaphocerite, dorsal; G—third maxilliped, lateral; H—same, tip of ultimate article, dorsomesial; I—second pereiopod, lateral; J—third pereiopod, lateral; K—fourth pereiopod, lateral; L—fifth pereiopod, lateral; M—second pleopod, detail of appendices masculina and interna, mesial; N—uropod, dorsal; O—same, detail of distolateral area of exopod, dorsal. H drawn without scale." captionText-1="FIGURE 2. Salmoneus alvarezi sp. nov., holotype from Bahía Málaga, Colombia (INV CRU 8365): A—major cheliped, lateral (mastigobranch detached); B—same, mesial; C—same, unfolded, detail of merus and carpus, lateral; D—same, chela, lateral; E—minor cheliped, lateral; F—same, chela, mesial." captionText-2="FIGURE 3. Salmoneus alvarezi sp. nov., living specimens: A, B—paratype, ov. specimen from Bahía Málaga, Colombia (OUMNH. ZC. 2015.01.089); C, D—non-type, ov. specimen from Punta Morales, Costa Rica (OUMNH. ZC. 2015.01.090); A, C—dorsal view; B, D—lateral view. Photographs by A. Anker." httpUri-0="https://zenodo.org/record/243882/files/figure.png" httpUri-1="https://zenodo.org/record/243883/files/figure.png" httpUri-2="https://zenodo.org/record/243884/files/figure.png" pageId="1" pageNumber="521">Figs. 13</figureCitation>
)
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<taxonomicName id="4C454D0A2147FFC748A8931EA324FC63" ID-CoL="85Y2K" box="[151,360,882,904]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="521" phylum="Arthropoda" rank="species" species="ortmanni">
<emphasis id="B931EA9B2147FFC748A8931EA324FC63" box="[151,360,882,904]" italics="true" pageId="1" pageNumber="521">Salmoneus ortmanni</emphasis>
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<treatmentCitation id="0AE410982147FFC749BD931EA0EEFC63" author="Rios" box="[386,674,882,904]" page="462" pageId="1" pageNumber="521" year="1983">
<bibRefCitation id="EFD44B782147FFC749BD931EA022FC63" author="Rios" box="[386,622,882,904]" pageId="1" pageNumber="521" refString="Rios, R. &amp; Carvacho, A. (1983) Caridean shrimps of the Gulf of California. I. New records, with some remarks on amphi- American distribution. Pacific Science, 36, 459 - 465." type="journal article" year="1983">Ríos &amp; Carvacho 1983</bibRefCitation>
: 462
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;
<treatmentCitation id="0AE410982147FFC74A93931EA64BFC63" author="Christoffersen" box="[684,1031,882,904]" page="63" pageId="1" pageNumber="521" year="1988">
<bibRefCitation id="EFD44B782147FFC74A93931EA1ACFC63" author="Christoffersen" box="[684,992,882,904]" pageId="1" pageNumber="521" refString="Christoffersen, M. L. &amp; Ramos, G. E. (1988) A new species of Alpheus (Caridea, Alpheidae) from the Pacific coast of Colombia. Revista Nordestina de Biologia, 6, 61 - 65." type="journal article" year="1988">Christoffersen &amp; Ramos 1988</bibRefCitation>
: 63
</treatmentCitation>
;
<treatmentCitation id="0AE410982147FFC74C2D931EA72EFC63" author="Villalobos-Hiriart" box="[1042,1378,882,904]" page="16" pageId="1" pageNumber="521" year="1989">
<bibRefCitation id="EFD44B782147FFC74C2D931EA770FC63" author="Villalobos-Hiriart" box="[1042,1340,882,904]" pageId="1" pageNumber="521" refString="Villalobos-Hiriart, J. L., Nates-Rodriguez, J. C., Diaz-Barriga, A. C., Valle Martinez, M. D., Flores-Hernandez, P., Lira- Fernandez, E. &amp; Schmidtsdorf-Valencia, P. (1989) Listados faunisticos de Mexico. I. Crustaceos estomatopodos y decapodos intermareales de las islas del Golfo de California. Instituto de Biologia, Universidad Nacional Autonoma de Mexico, ix + 101 pp." type="book" year="1989">
Villalobos-Hiriart
<emphasis id="B931EA9B2147FFC74CF3931FA74CFC63" box="[1228,1280,882,904]" italics="true" pageId="1" pageNumber="521">et al.</emphasis>
1989
</bibRefCitation>
: 16
</treatmentCitation>
;
<treatmentCitation id="0AE410982147FFC74D52931EA37FFC4C" author="Rios" page="154" pageId="1" pageNumber="521" year="1989">
<bibRefCitation id="EFD44B782147FFC74D52931EA2B1FC4C" author="Rios" pageId="1" pageNumber="521" refString="Rios, R. G. (1989) Un catalogo de camarones carideos de Mulege y Bahia Concepcion, B. C. S. con anotaciones acerca de su biologia, ecologia, distribucion geografica y taxonomia. Tesis profesional, Ensenada, B. C., Mexico, 208 pp." type="book" year="1989">Ríos 1989</bibRefCitation>
: 154
</treatmentCitation>
, pl. 30;
<treatmentCitation id="0AE410982147FFC749AF93FDA029FC4C" author="Wicksten" box="[400,613,913,935]" page="151" pageId="1" pageNumber="521" year="1991">
<bibRefCitation id="EFD44B782147FFC749AF93FDA063FC4C" author="Wicksten" box="[400,559,913,935]" pageId="1" pageNumber="521" refString="Wicksten, M. K. (1991) Caridean and stenopodid shrimp of the Galapagos Islands. In: James, M. J. (Ed.), Galapagos marine invertebrates. Plenum, New York, pp. 147 - 156. http: // dx. doi. org / 10.1007 / 978 - 1 - 4899 - 0646 - 5 _ 7" type="book chapter" year="1991">Wicksten 1991</bibRefCitation>
: 151
</treatmentCitation>
;
<treatmentCitation id="0AE410982147FFC74A4B93FDA0B3FC4C" author="Rios" box="[628,767,913,935]" page="7" pageId="1" pageNumber="521" year="1992">
<bibRefCitation id="EFD44B782147FFC74A4B93FDA0AEFC4C" author="Rios" box="[628,738,913,935]" pageId="1" pageNumber="521" refString="Rios, R. (1992) Camarones carideos del Golfo de California. VI. Alpheidae del estuario de Mulege y de Bahia Concepcion, Baja California Sur, Mexico (Crustacea: Caridea). Proceedings of San Diego Society of Natural History, 14, 1 - 13." type="journal article" year="1992">Ríos 1992</bibRefCitation>
: 7
</treatmentCitation>
; Villalobos-Hiriart
<emphasis id="B931EA9B2147FFC74BF393FEA648FC4C" box="[972,1028,913,935]" italics="true" pageId="1" pageNumber="521">et al.</emphasis>
1992: 6;
<treatmentCitation id="0AE410982147FFC74C5093FDA70CFC4C" author="Villalobos" box="[1135,1344,913,935]" page="74" pageId="1" pageNumber="521" year="2000">
<bibRefCitation id="EFD44B782147FFC74C5093FDA754FC4C" author="Villalobos" box="[1135,1304,913,935]" pageId="1" pageNumber="521" refString="Villalobos, J. L. (2000) Estudio monografico de los crustaceos decapodos no braquiuros de la zona intermareal de las islas del Golfo de California, Mexico. Tesis de Maestria. Facultad de Ciencias, Universidad Nacional Autonoma de Mexico, 312 pp." type="book" year="2000">Villalobos 2000</bibRefCitation>
: 74
</treatmentCitation>
, fig. 37;
<treatmentCitation id="0AE410982147FFC748F893DCA0DEFC2D" author="Lazarus-Agudelo" box="[199,658,944,966]" page="228" pageId="1" pageNumber="521" year="2007">
<bibRefCitation id="EFD44B782147FFC748F893DCA02EFC2D" author="Lazarus-Agudelo" box="[199,610,944,966]" pageId="1" pageNumber="521" refString="Lazarus-Agudelo, J. F. &amp; Cantera-Kintz, J. R. (2007) Crustaceos (Crustacea: Sessilia, Stomatopoda, Isopoda, Amphipoda, Decapoda) de Bahia Malaga, Valle del Cauca (Pacifico colombiano). Biota Colombiana, 8, 221 - 239." type="journal article" year="2007">Lazarus-Agudelo &amp; Cantera-Kintz 2007</bibRefCitation>
: 228
</treatmentCitation>
[not
<taxonomicName id="4C454D0A2147FFC74AF793DDA1AFFC2D" ID-CoL="85Y2K" authority="Rankin, 1898" authorityName="Rankin" authorityYear="1898" box="[712,995,944,966]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="521" phylum="Arthropoda" rank="species" species="ortmanni">
<emphasis id="B931EA9B2147FFC74AF793DDA172FC2D" box="[712,830,945,966]" italics="true" pageId="1" pageNumber="521">S. ortmanni</emphasis>
(
<bibRefCitation id="EFD44B782147FFC74B7493DCA197FC2D" author="Rankin" box="[843,987,944,966]" pageId="1" pageNumber="521" refString="Rankin, W. M. (1898) The Northrop collection of Crustacea from the Bahamas. Annals of the New York Academy of Sciences, 11, 225 - 258. http: // dx. doi. org / 10.1111 / j. 1749 - 6632.1898. tb 54971. x" type="journal article" year="1898">Rankin, 1898</bibRefCitation>
)
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(?)
<taxonomicName id="4C454D0A2147FFC7488593A2A3C7FC0E" ID-CoL="85Y2K" box="[186,395,974,997]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="521" phylum="Arthropoda" rank="species" species="ortmanni">
<emphasis id="B931EA9B2147FFC7488593A2A3C7FC0E" box="[186,395,974,997]" italics="true" pageId="1" pageNumber="521">Salmoneus ortmanni</emphasis>
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<bibRefCitation id="EFD44B782147FFC7499A93A2A0DBFC0F" author="Flores-Hernandez" box="[421,663,974,996]" pageId="1" pageNumber="521" refString="Flores-Hernandez, P. (1991) Taxonomia y distribucion geografica de tres familias de camarones carideos (Crustacea: Decapoda) intermareales de las islas del Golfo de California, Mexico. Tesis profesional. Facultad de Ciencias, Universidad Nacional Autonoma de Mexico, 225 pp. [not seen]" type="book" year="1991">Flores-Hernandez 1991</bibRefCitation>
: 132;
<bibRefCitation id="EFD44B782147FFC74AEA93A2A1B2FC0F" author="Wicksten" box="[725,1022,974,996]" pageId="1" pageNumber="521" refString="Wicksten, M. K. &amp; Hendrickx, M. E. (1992) Checklist of penaeoid and caridean shrimps (Decapoda: Penaeoidea, Caridea) from the eastern tropical Pacific. Proceedings of the San Diego Society of Natural History, 9, 1 - 11." type="journal article" year="1992">Wicksten &amp; Hendrickx 1992</bibRefCitation>
: 6;
<bibRefCitation id="EFD44B782147FFC74C1C93A2A687FC0F" author="Hendrickx" box="[1059,1227,974,996]" pageId="1" pageNumber="521" refString="Hendrickx, M. E. (1993) Crustaceos decapodos bentonicos del Sur de Sinaloa, Mexico. Anales del Instituto de Biologia, Universidad Nacional Autonoma de Mexico, Ser. Zool., 64, 1 - 16." type="journal article" year="1993">Hendrickx 1993</bibRefCitation>
: 6;
<bibRefCitation id="EFD44B782147FFC74CCF93A2A7D5FC0F" author="Hendrickx" box="[1264,1433,974,996]" pageId="1" pageNumber="521" refString="Hendrickx, M. E. (1995) Camarones. In: Fischer, W., Krupp, F., Schneider, W., Sommer, C., Carpenter, K. E. &amp; Niem, V. H. (Eds.), Guia FAO para la identificacion de especies para los fines de la pesca. Pacifico Centro-Oriental. Vol. 1 Plantas e Invertebrados, pp. 417 - 537. FAO, Rome." type="journal article" year="1995">Hendrickx 1995</bibRefCitation>
: 433;
<bibRefCitation id="EFD44B782147FFC748C59381A06CFBE8" author="Wicksten" box="[250,544,1005,1027]" pageId="1" pageNumber="521" refString="Wicksten, M. K. &amp; Hendrickx, M. E. (2003) An updated checklist of benthic marine and brackish water shrimps (Decapoda: Penaeoidea, Stenopodidea, Caridea) from the eastern tropical Pacific. In Hendrickx, M. E. (Ed.), Contributions to the Study of East Pacific Crustaceans. Vol. 2. Instituto de Ciencias del Mar y Limnologia, UNAM, Mexico, pp. 49 - 76." type="book" year="2003">Wicksten &amp; Hendrickx 2003</bibRefCitation>
: 66 [not
<taxonomicName id="4C454D0A2147FFC74A469382A1D8FBE8" ID-CoL="85Y2K" authority="Rankin, 1898" authorityName="Rankin" authorityYear="1898" box="[633,916,1005,1027]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="521" phylum="Arthropoda" rank="species" species="ortmanni">
<emphasis id="B931EA9B2147FFC74A469382A0A3FBE8" box="[633,751,1006,1027]" italics="true" pageId="1" pageNumber="521">S. ortmanni</emphasis>
(
<bibRefCitation id="EFD44B782147FFC74AC39381A1C7FBE8" author="Rankin" box="[764,907,1005,1027]" pageId="1" pageNumber="521" refString="Rankin, W. M. (1898) The Northrop collection of Crustacea from the Bahamas. Annals of the New York Academy of Sciences, 11, 225 - 258. http: // dx. doi. org / 10.1111 / j. 1749 - 6632.1898. tb 54971. x" type="journal article" year="1898">Rankin, 1898</bibRefCitation>
)
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].
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<taxonomicName id="4C454D0A2147FFC748A89460A3C4FBC9" ID-CoL="85Y2K" box="[151,392,1036,1058]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="521" phylum="Arthropoda" rank="species" species="ortmanni">
<emphasis id="B931EA9B2147FFC748A89460A349FBC9" box="[151,261,1036,1058]" italics="true" pageId="1" pageNumber="521">Salmoneus</emphasis>
cf.
<emphasis id="B931EA9B2147FFC749149461A3C4FBC9" box="[299,392,1037,1058]" italics="true" pageId="1" pageNumber="521">ortmanni</emphasis>
</taxonomicName>
<treatmentCitation id="0AE410982147FFC7499E9460A001FBC9" ID-DOI="http://doi.org/10.5281/zenodo.5314472" ID-Zenodo-Dep="5314472" author="Anker" box="[417,589,1036,1058]" httpUri="http://treatment.plazi.org/id/2554563EFF91FB00A780FF4B03DEFCE1" page="201" pageId="1" pageNumber="521" year="2010">
<bibRefCitation id="EFD44B782147FFC7499E9460A052FBC9" author="Anker" box="[417,542,1036,1058]" pageId="1" pageNumber="521" refString="Anker, A. (2010) The shrimp genus Salmoneus Holthuis, 1955 (Crustacea, Decapoda, Alpheidae) in the tropical western Atlantic, with description of five new species. Zootaxa, 2372, 177 - 205." type="journal article" year="2010">Anker 2010</bibRefCitation>
: 201
</treatmentCitation>
[not
<taxonomicName id="4C454D0A2147FFC74ABD9460A1D1FBC9" ID-CoL="85Y2K" authority="Rankin, 1898" authorityName="Rankin" authorityYear="1898" box="[642,925,1036,1058]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="521" phylum="Arthropoda" rank="species" species="ortmanni">
<emphasis id="B931EA9B2147FFC74ABD9460A0B5FBC9" box="[642,761,1036,1058]" italics="true" pageId="1" pageNumber="521">S. ortmanni</emphasis>
(
<bibRefCitation id="EFD44B782147FFC74B399460A1D9FBC9" author="Rankin" box="[774,917,1036,1058]" pageId="1" pageNumber="521" refString="Rankin, W. M. (1898) The Northrop collection of Crustacea from the Bahamas. Annals of the New York Academy of Sciences, 11, 225 - 258. http: // dx. doi. org / 10.1111 / j. 1749 - 6632.1898. tb 54971. x" type="journal article" year="1898">Rankin, 1898</bibRefCitation>
)
</taxonomicName>
].
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C35F65022147FFC748A89425A1D9FA0E" pageId="1" pageNumber="521" type="materials_examined">
<paragraph id="8BFA36892147FFC748A89425A6FAFA90" blockId="1.[151,1437,1097,2037]" pageId="1" pageNumber="521">
<emphasis id="B931EA9B2147FFC748A89425A308FB89" bold="true" box="[151,324,1097,1122]" pageId="1" pageNumber="521">
<typeStatus id="54FE882B2147FFC748A89425A29EFB89" box="[151,210,1097,1122]" pageId="1" pageNumber="521">Type</typeStatus>
material
</emphasis>
.
<collectingCountry id="F35276192147FFC749679425A386FB89" box="[344,458,1097,1122]" name="Colombia" pageId="1" pageNumber="521">Colombia</collectingCountry>
.
<typeStatus id="54FE882B2147FFC749E49425A004FB89" box="[475,584,1097,1122]" pageId="1" pageNumber="521" type="holotype">Holotype</typeStatus>
: ov. specimen (cl
<quantity id="4CBD9B6C2147FFC74B169426A1C5FB89" box="[809,905,1098,1122]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="1" pageNumber="521" unit="mm" value="4.2">4.2 mm</quantity>
), INV CRU8365, Bahía Málaga, Curichichi,
<geoCoordinate id="EE71504E2147FFC748A89400A379FB6E" box="[151,309,1132,1157]" direction="north" orientation="latitude" pageId="1" pageNumber="521" precision="1" value="3.9938333">03°5937.8”N</geoCoordinate>
<geoCoordinate id="EE71504E2147FFC749029401A3AEFB6E" box="[317,482,1133,1157]" direction="west" orientation="longitude" pageId="1" pageNumber="521" precision="1" value="-77.31775">77°1903.9”W</geoCoordinate>
, intertidal mudflat exposed at low tide, under rocks, coll. A. Anker,
<date id="FFFB10492147FFC74CCE9400A72BFB6E" box="[1265,1383,1132,1157]" pageId="1" pageNumber="521" value="2009-04-26">26.iv.2009</date>
[fcn COL-00176].
<typeStatus id="54FE882B2147FFC7490794FCA3E6FB43" box="[312,426,1168,1192]" pageId="1" pageNumber="521" type="paratype">Paratypes</typeStatus>
: 1 non-ov. specimen (cl 3.0 mm), INV CRU8366, Bahía Málaga, Los Negros,
<geoCoordinate id="EE71504E2147FFC74D0494E3A7D0FB43" box="[1339,1436,1167,1192]" direction="north" orientation="latitude" pageId="1" pageNumber="521" precision="925" value="3.9833333">03°59N</geoCoordinate>
<geoCoordinate id="EE71504E2147FFC748A894DFA34DFB20" box="[151,257,1203,1227]" direction="west" orientation="longitude" pageId="1" pageNumber="521" precision="925" value="-77.28333">77°17”W</geoCoordinate>
, rocky intertidal, under large rocks on muddy sand, coll. A. Anker,
<date id="FFFB10492147FFC74C3294DFA6CFFB27" box="[1037,1155,1203,1228]" pageId="1" pageNumber="521" value="2009-04-25">25.iv.2009</date>
[fcn COL-00146]; 1 ov. specimen (cl 3.0 mm),
<collectionCode id="ED54AE4C2147FFC7499594BBA04BFB04" box="[426,519,1239,1263]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="521">MZUSP</collectionCode>
33198, Bahía Málaga, Naidizal,
<geoCoordinate id="EE71504E2147FFC74BB094BBA661FB04" box="[911,1069,1239,1263]" direction="north" orientation="latitude" pageId="1" pageNumber="521" precision="1" value="4.042861">04°0234.3”N</geoCoordinate>
<geoCoordinate id="EE71504E2147FFC74C0794BBA691FB04" box="[1080,1245,1239,1263]" direction="west" orientation="longitude" pageId="1" pageNumber="521" precision="1" value="-77.23011">77°1348.4”W</geoCoordinate>
, rocky intertidal exposed at low tide, under rocks in muddy water (near water edge), hand sieve, coll. A. Anker,
<date id="FFFB10492147FFC74CCF9495A729FAF9" box="[1264,1381,1273,1298]" pageId="1" pageNumber="521" value="2009-04-23">23.iv.2009</date>
[fcn COL-00030]; 1 ov. specimen (cl
<quantity id="4CBD9B6C2147FFC74A349571A02AFADE" box="[523,614,1309,1333]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="1" pageNumber="521" unit="mm" value="4.1">4.1 mm</quantity>
),
<collectionCode id="ED54AE4C2147FFC74A469571A09AFADE" box="[633,726,1309,1333]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="521">MZUSP</collectionCode>
33199, same collection data [fcn COL-00017]; 1 ov. specimen (cl 4.0 mm), 1 non-ov. specimen (cl
<quantity id="4CBD9B6C2147FFC74A15952CA0C8FAB3" box="[554,644,1344,1368]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="1" pageNumber="521" unit="mm" value="3.1">3.1 mm</quantity>
),
<collectionCode id="ED54AE4C2147FFC74AA9952CA14AFAB3" box="[662,774,1344,1368]" pageId="1" pageNumber="521">OUMNH</collectionCode>
.ZC. 2015.01.0 88, same collection data [fcn COL-00012]; 1 ov. specimen (cl
<quantity id="4CBD9B6C2147FFC7496B950FA3E2FA90" box="[340,430,1379,1403]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="1" pageNumber="521" unit="mm" value="4.2">4.2 mm</quantity>
),
<collectionCode id="ED54AE4C2147FFC749FE950FA07DFA90" box="[449,561,1379,1403]" pageId="1" pageNumber="521">OUMNH</collectionCode>
.ZC. 2015.01.0 89, same collection data [fcn COL-00018].
</paragraph>
<paragraph id="8BFA36892147FFC748F895EAA1D9FA0E" blockId="1.[151,1437,1097,2037]" pageId="1" pageNumber="521">
<emphasis id="B931EA9B2147FFC748F895EAA3FFFA74" bold="true" box="[199,435,1414,1439]" pageId="1" pageNumber="521">Additional material</emphasis>
.
<collectingCountry id="F35276192147FFC749FC95EBA00EFA74" box="[451,578,1414,1439]" name="Costa Rica" pageId="1" pageNumber="521">Costa Rica</collectingCountry>
. 1 ov. specimen (cl 4.0 mm),
<collectionCode id="ED54AE4C2147FFC74BAC95EBA1BCFA74" box="[915,1008,1415,1439]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="521">MZUSP</collectionCode>
33200, Punta Morales, Playa Blanca, muddy-rocky intertidal near mangrove, under rocks, coll. A. Anker,
<date id="FFFB10492147FFC74BAE95C5A644FA29" box="[913,1032,1449,1474]" pageId="1" pageNumber="521" value="2005-11-22">22.xi.2005</date>
[fcn 05-112]; 1 ov. specimen (cl
<quantity id="4CBD9B6C2147FFC74D4695C6A28BFA0E" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="1" pageNumber="521" unit="mm" value="4.4">4.4 mm</quantity>
),
<collectionCode id="ED54AE4C2147FFC748E595A1A306FA0E" box="[218,330,1485,1509]" pageId="1" pageNumber="521">OUMNH</collectionCode>
.ZC. 2015.01.0 90, same collection data [fcn 05-113].
</paragraph>
</subSubSection>
<subSubSection id="C35F65022147FFC248F89583A72DFCD3" lastPageId="4" lastPageNumber="524" pageId="1" pageNumber="521" type="description">
<paragraph id="8BFA36892147FFC748F89583A14CF953" blockId="1.[151,1437,1097,2037]" pageId="1" pageNumber="521">
<emphasis id="B931EA9B2147FFC748F89583A31DF9E3" bold="true" box="[199,337,1519,1544]" pageId="1" pageNumber="521">Description</emphasis>
. Carapace with numerous scattered setae (
<figureCitation id="137E2A0C2147FFC74B149583A125F9E3" box="[811,873,1519,1544]" captionStart="FIGURE 1" captionStartId="2.[151,250,1798,1820]" captionTargetBox="[162,1411,208,1768]" captionTargetId="figure@2.[151,1436,193,1777]" captionTargetPageId="2" captionText="FIGURE 1. Salmoneus alvarezi sp. nov., holotype from Bahía Málaga, Colombia (INV CRU 8365): A—frontal region, dorsal; B—same, lateral; C—posterior abdominal somites, lateral; D—telson, dorsal; E—same, posterior margin, dorsal; F—antennal scaphocerite, dorsal; G—third maxilliped, lateral; H—same, tip of ultimate article, dorsomesial; I—second pereiopod, lateral; J—third pereiopod, lateral; K—fourth pereiopod, lateral; L—fifth pereiopod, lateral; M—second pleopod, detail of appendices masculina and interna, mesial; N—uropod, dorsal; O—same, detail of distolateral area of exopod, dorsal. H drawn without scale." httpUri="https://zenodo.org/record/243882/files/figure.png" pageId="1" pageNumber="521">Fig. 1</figureCitation>
B). Rostrum subtriangular in dorsal view, about as long as broad at base, reaching beyond distal margin of first article of antennular peduncle, typically not reaching mid-length of second article, with acute tip; lateral margins slightly convex proximally; ventral margin distally unarmed; rostral carina present as low keel extending beyond level of eyes posteriorly; areas lateral to rostral carina shallowly depressed (
<figureCitation id="137E2A0C2147FFC749AE9610A398F97E" box="[401,468,1660,1685]" captionStart="FIGURE 1" captionStartId="2.[151,250,1798,1820]" captionTargetBox="[162,1411,208,1768]" captionTargetId="figure@2.[151,1436,193,1777]" captionTargetPageId="2" captionText="FIGURE 1. Salmoneus alvarezi sp. nov., holotype from Bahía Málaga, Colombia (INV CRU 8365): A—frontal region, dorsal; B—same, lateral; C—posterior abdominal somites, lateral; D—telson, dorsal; E—same, posterior margin, dorsal; F—antennal scaphocerite, dorsal; G—third maxilliped, lateral; H—same, tip of ultimate article, dorsomesial; I—second pereiopod, lateral; J—third pereiopod, lateral; K—fourth pereiopod, lateral; L—fifth pereiopod, lateral; M—second pleopod, detail of appendices masculina and interna, mesial; N—uropod, dorsal; O—same, detail of distolateral area of exopod, dorsal. H drawn without scale." httpUri="https://zenodo.org/record/243882/files/figure.png" pageId="1" pageNumber="521">Fig. 1</figureCitation>
A, B). Orbital teeth strong, acute, directed anteriorly (
<figureCitation id="137E2A0C2147FFC74C689610A6D5F97E" box="[1111,1177,1660,1685]" captionStart="FIGURE 1" captionStartId="2.[151,250,1798,1820]" captionTargetBox="[162,1411,208,1768]" captionTargetId="figure@2.[151,1436,193,1777]" captionTargetPageId="2" captionText="FIGURE 1. Salmoneus alvarezi sp. nov., holotype from Bahía Málaga, Colombia (INV CRU 8365): A—frontal region, dorsal; B—same, lateral; C—posterior abdominal somites, lateral; D—telson, dorsal; E—same, posterior margin, dorsal; F—antennal scaphocerite, dorsal; G—third maxilliped, lateral; H—same, tip of ultimate article, dorsomesial; I—second pereiopod, lateral; J—third pereiopod, lateral; K—fourth pereiopod, lateral; L—fifth pereiopod, lateral; M—second pleopod, detail of appendices masculina and interna, mesial; N—uropod, dorsal; O—same, detail of distolateral area of exopod, dorsal. H drawn without scale." httpUri="https://zenodo.org/record/243882/files/figure.png" pageId="1" pageNumber="521">Fig. 1</figureCitation>
A, B). Pterygostomial margin feebly protruding anteriorly, rounded (
<figureCitation id="137E2A0C2147FFC74AA496F3A096F953" box="[667,730,1695,1720]" captionStart="FIGURE 1" captionStartId="2.[151,250,1798,1820]" captionTargetBox="[162,1411,208,1768]" captionTargetId="figure@2.[151,1436,193,1777]" captionTargetPageId="2" captionText="FIGURE 1. Salmoneus alvarezi sp. nov., holotype from Bahía Málaga, Colombia (INV CRU 8365): A—frontal region, dorsal; B—same, lateral; C—posterior abdominal somites, lateral; D—telson, dorsal; E—same, posterior margin, dorsal; F—antennal scaphocerite, dorsal; G—third maxilliped, lateral; H—same, tip of ultimate article, dorsomesial; I—second pereiopod, lateral; J—third pereiopod, lateral; K—fourth pereiopod, lateral; L—fifth pereiopod, lateral; M—second pleopod, detail of appendices masculina and interna, mesial; N—uropod, dorsal; O—same, detail of distolateral area of exopod, dorsal. H drawn without scale." httpUri="https://zenodo.org/record/243882/files/figure.png" pageId="1" pageNumber="521">Fig. 1</figureCitation>
B).
</paragraph>
<paragraph id="8BFA36892147FFC748F896A8A3D4F8C9" blockId="1.[151,1437,1097,2037]" pageId="1" pageNumber="521">
Eyes completely concealed in dorsal view, anterior-most portion often visible in lateral view; anteromesial margin of eyestalk not markedly projecting, rounded; cornea well developed (
<figureCitation id="137E2A0C2147FFC74BC5968AA674F914" box="[1018,1080,1766,1791]" captionStart="FIGURE 1" captionStartId="2.[151,250,1798,1820]" captionTargetBox="[162,1411,208,1768]" captionTargetId="figure@2.[151,1436,193,1777]" captionTargetPageId="2" captionText="FIGURE 1. Salmoneus alvarezi sp. nov., holotype from Bahía Málaga, Colombia (INV CRU 8365): A—frontal region, dorsal; B—same, lateral; C—posterior abdominal somites, lateral; D—telson, dorsal; E—same, posterior margin, dorsal; F—antennal scaphocerite, dorsal; G—third maxilliped, lateral; H—same, tip of ultimate article, dorsomesial; I—second pereiopod, lateral; J—third pereiopod, lateral; K—fourth pereiopod, lateral; L—fifth pereiopod, lateral; M—second pleopod, detail of appendices masculina and interna, mesial; N—uropod, dorsal; O—same, detail of distolateral area of exopod, dorsal. H drawn without scale." httpUri="https://zenodo.org/record/243882/files/figure.png" pageId="1" pageNumber="521">Fig. 1</figureCitation>
A, B). Epistomial sclerites each with small sharp tooth.
</paragraph>
<paragraph id="8BFA36892147FFC748F89740A701F81E" blockId="1.[151,1437,1097,2037]" pageId="1" pageNumber="521">
First to third pleura rounded posteroventrally; fourth pleuron with blunt posteroventral angle; fifth pleuron with posteroventral angle produced into small tooth; sixth somite without articulated plate (
<figureCitation id="137E2A0C2147FFC74CA19723A691F883" box="[1182,1245,1871,1896]" captionStart="FIGURE 1" captionStartId="2.[151,250,1798,1820]" captionTargetBox="[162,1411,208,1768]" captionTargetId="figure@2.[151,1436,193,1777]" captionTargetPageId="2" captionText="FIGURE 1. Salmoneus alvarezi sp. nov., holotype from Bahía Málaga, Colombia (INV CRU 8365): A—frontal region, dorsal; B—same, lateral; C—posterior abdominal somites, lateral; D—telson, dorsal; E—same, posterior margin, dorsal; F—antennal scaphocerite, dorsal; G—third maxilliped, lateral; H—same, tip of ultimate article, dorsomesial; I—second pereiopod, lateral; J—third pereiopod, lateral; K—fourth pereiopod, lateral; L—fifth pereiopod, lateral; M—second pleopod, detail of appendices masculina and interna, mesial; N—uropod, dorsal; O—same, detail of distolateral area of exopod, dorsal. H drawn without scale." httpUri="https://zenodo.org/record/243882/files/figure.png" pageId="1" pageNumber="521">Fig. 1</figureCitation>
C); preanal plate rounded distally. Telson about 2.22.3 times as long as wide proximally, tapering posteriorly, with two pairs of strong dorsal spiniform setae situated at about 0.5 and 0.75 of telson length, respectively; posterior margin nearly straight or shallowly emarginated centrally, with two pairs of stout spiniform setae, lateral shorter and noticeably more slender than mesial, and two pairs of long plumose setae between mesial spiniform setae (
<figureCitation id="137E2A0C2147FFC74CF897B0A74AF81E" box="[1223,1286,2012,2037]" captionStart="FIGURE 1" captionStartId="2.[151,250,1798,1820]" captionTargetBox="[162,1411,208,1768]" captionTargetId="figure@2.[151,1436,193,1777]" captionTargetPageId="2" captionText="FIGURE 1. Salmoneus alvarezi sp. nov., holotype from Bahía Málaga, Colombia (INV CRU 8365): A—frontal region, dorsal; B—same, lateral; C—posterior abdominal somites, lateral; D—telson, dorsal; E—same, posterior margin, dorsal; F—antennal scaphocerite, dorsal; G—third maxilliped, lateral; H—same, tip of ultimate article, dorsomesial; I—second pereiopod, lateral; J—third pereiopod, lateral; K—fourth pereiopod, lateral; L—fifth pereiopod, lateral; M—second pleopod, detail of appendices masculina and interna, mesial; N—uropod, dorsal; O—same, detail of distolateral area of exopod, dorsal. H drawn without scale." httpUri="https://zenodo.org/record/243882/files/figure.png" pageId="1" pageNumber="521">Fig. 1</figureCitation>
D, E).
</paragraph>
<caption id="DF3A66012144FFC448A8976AA29CF85E" httpUri="https://zenodo.org/record/243882/files/figure.png" pageId="2" pageNumber="522" targetBox="[162,1411,208,1768]" targetPageId="2">
<paragraph id="8BFA36892144FFC448A8976AA29CF85E" blockId="2.[151,1436,1798,1973]" pageId="2" pageNumber="522">
<emphasis id="B931EA9B2144FFC448A8976AA35FF8F0" bold="true" box="[151,275,1798,1820]" pageId="2" pageNumber="522">FIGURE 1.</emphasis>
<taxonomicName id="4C454D0A2144FFC44926976AA392F8F7" box="[281,478,1798,1820]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="522" phylum="Arthropoda" rank="species" species="alvarezi" status="sp. nov.">
<emphasis id="B931EA9B2144FFC44926976AA392F8F7" box="[281,478,1798,1820]" italics="true" pageId="2" pageNumber="522">Salmoneus alvarezi</emphasis>
</taxonomicName>
<emphasis id="B931EA9B2144FFC449DB976BA078F8F7" bold="true" box="[484,564,1799,1820]" pageId="2" pageNumber="522">
<taxonomicNameLabel id="A20257E02144FFC449DB976BA078F8F7" box="[484,564,1799,1820]" pageId="2" pageNumber="522" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype from Bahía Málaga, Colombia (INV CRU8365): A—frontal region, dorsal; B—same, lateral; C—posterior abdominal somites, lateral; D—telson, dorsal; E—same, posterior margin, dorsal; F—antennal scaphocerite, dorsal; G—third maxilliped, lateral; H—same, tip of ultimate article, dorsomesial; I—second pereiopod, lateral; J—third pereiopod, lateral; K—fourth pereiopod, lateral; L—fifth pereiopod, lateral; M—second pleopod, detail of appendices masculina and interna, mesial; N—uropod, dorsal; O—same, detail of distolateral area of exopod, dorsal. H drawn without scale.
</paragraph>
</caption>
<caption id="DF3A66012145FFC548A8954AA0E9FA92" httpUri="https://zenodo.org/record/243883/files/figure.png" pageId="3" pageNumber="523" targetBox="[162,1428,202,1288]" targetPageId="3">
<paragraph id="8BFA36892145FFC548A8954AA0E9FA92" blockId="3.[151,1436,1318,1401]" pageId="3" pageNumber="523">
<emphasis id="B931EA9B2145FFC548A8954AA35EFAD0" bold="true" box="[151,274,1318,1340]" pageId="3" pageNumber="523">FIGURE 2.</emphasis>
<taxonomicName id="4C454D0A2145FFC54927954AA391FAD7" box="[280,477,1318,1340]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="523" phylum="Arthropoda" rank="species" species="alvarezi" status="sp. nov.">
<emphasis id="B931EA9B2145FFC54927954AA391FAD7" box="[280,477,1318,1340]" italics="true" pageId="3" pageNumber="523">Salmoneus alvarezi</emphasis>
</taxonomicName>
<emphasis id="B931EA9B2145FFC549DD954BA07EFAD7" bold="true" box="[482,562,1319,1340]" pageId="3" pageNumber="523">
<taxonomicNameLabel id="A20257E02145FFC549DD954BA07EFAD7" box="[482,562,1319,1340]" pageId="3" pageNumber="523" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype from Bahía Málaga, Colombia (INV CRU8365): A—major cheliped, lateral (mastigobranch detached); B—same, mesial; C—same, unfolded, detail of merus and carpus, lateral; D—same, chela, lateral; E—minor cheliped, lateral; F—same, chela, mesial.
</paragraph>
</caption>
<paragraph id="8BFA36892145FFC548F895CDA2FBF97A" blockId="3.[151,1436,1441,2006]" pageId="3" pageNumber="523">
Antennule robust; stylocerite distinctly overreaching distal margin of second article, with acute tip; ventromesial carina of first article with sharp, anteriorly directed tooth; second article about as long as wide; lateral flagellum bifurcating at about third or fourth segment, secondary ramus well developed, with several (typically six) groups of aesthetascs (
<figureCitation id="137E2A0C2145FFC549A89660A39AF9CE" box="[407,470,1548,1573]" captionStart="FIGURE 1" captionStartId="2.[151,250,1798,1820]" captionTargetBox="[162,1411,208,1768]" captionTargetId="figure@2.[151,1436,193,1777]" captionTargetPageId="2" captionText="FIGURE 1. Salmoneus alvarezi sp. nov., holotype from Bahía Málaga, Colombia (INV CRU 8365): A—frontal region, dorsal; B—same, lateral; C—posterior abdominal somites, lateral; D—telson, dorsal; E—same, posterior margin, dorsal; F—antennal scaphocerite, dorsal; G—third maxilliped, lateral; H—same, tip of ultimate article, dorsomesial; I—second pereiopod, lateral; J—third pereiopod, lateral; K—fourth pereiopod, lateral; L—fifth pereiopod, lateral; M—second pleopod, detail of appendices masculina and interna, mesial; N—uropod, dorsal; O—same, detail of distolateral area of exopod, dorsal. H drawn without scale." httpUri="https://zenodo.org/record/243882/files/figure.png" pageId="3" pageNumber="523">Fig. 1</figureCitation>
A, B). Antenna with basicerite bearing sharp distoventral tooth; scaphocerite broadly ovate, with strong acute distolateral tooth and relatively broad blade, latter with convex anterior margin reaching slightly beyond distolateral tooth; carpocerite short, not reaching beyond mid-length of scaphocerite (
<figureCitation id="137E2A0C2145FFC54D249639A717F986" box="[1307,1371,1621,1646]" captionStart="FIGURE 1" captionStartId="2.[151,250,1798,1820]" captionTargetBox="[162,1411,208,1768]" captionTargetId="figure@2.[151,1436,193,1777]" captionTargetPageId="2" captionText="FIGURE 1. Salmoneus alvarezi sp. nov., holotype from Bahía Málaga, Colombia (INV CRU 8365): A—frontal region, dorsal; B—same, lateral; C—posterior abdominal somites, lateral; D—telson, dorsal; E—same, posterior margin, dorsal; F—antennal scaphocerite, dorsal; G—third maxilliped, lateral; H—same, tip of ultimate article, dorsomesial; I—second pereiopod, lateral; J—third pereiopod, lateral; K—fourth pereiopod, lateral; L—fifth pereiopod, lateral; M—second pleopod, detail of appendices masculina and interna, mesial; N—uropod, dorsal; O—same, detail of distolateral area of exopod, dorsal. H drawn without scale." httpUri="https://zenodo.org/record/243882/files/figure.png" pageId="3" pageNumber="523">Fig. 1</figureCitation>
A, B, F).
</paragraph>
<paragraph id="8BFA36892145FFC548F896F1A394F916" blockId="3.[151,1436,1441,2006]" pageId="3" pageNumber="523">
Mouthparts not specific, typical for genus. Third maxilliped with rounded lateral plate; ultimate article tapering distally, ending in small corneous, apparently immovable spine, distodorsal surface with several movable spiniform setae (
<figureCitation id="137E2A0C2145FFC5496D9689A3DCF916" box="[338,400,1765,1790]" captionStart="FIGURE 1" captionStartId="2.[151,250,1798,1820]" captionTargetBox="[162,1411,208,1768]" captionTargetId="figure@2.[151,1436,193,1777]" captionTargetPageId="2" captionText="FIGURE 1. Salmoneus alvarezi sp. nov., holotype from Bahía Málaga, Colombia (INV CRU 8365): A—frontal region, dorsal; B—same, lateral; C—posterior abdominal somites, lateral; D—telson, dorsal; E—same, posterior margin, dorsal; F—antennal scaphocerite, dorsal; G—third maxilliped, lateral; H—same, tip of ultimate article, dorsomesial; I—second pereiopod, lateral; J—third pereiopod, lateral; K—fourth pereiopod, lateral; L—fifth pereiopod, lateral; M—second pleopod, detail of appendices masculina and interna, mesial; N—uropod, dorsal; O—same, detail of distolateral area of exopod, dorsal. H drawn without scale." httpUri="https://zenodo.org/record/243882/files/figure.png" pageId="3" pageNumber="523">Fig. 1</figureCitation>
G, H).
</paragraph>
<paragraph id="8BFA36892145FFC248F89765A615FF13" blockId="3.[151,1436,1441,2006]" lastBlockId="4.[151,1437,151,2013]" lastPageId="4" lastPageNumber="524" pageId="3" pageNumber="523">
Chelipeds strongly asymmetrical in shape and unequal in size; major cheliped by far most conspicuous in general view (
<figureCitation id="137E2A0C2145FFC549019741A3DEF8AE" box="[318,402,1837,1862]" captionStart="FIGURE 2" captionStartId="3.[151,250,1318,1340]" captionTargetBox="[162,1428,202,1288]" captionTargetId="figure@3.[151,1436,193,1297]" captionTargetPageId="3" captionText="FIGURE 2. Salmoneus alvarezi sp. nov., holotype from Bahía Málaga, Colombia (INV CRU 8365): A—major cheliped, lateral (mastigobranch detached); B—same, mesial; C—same, unfolded, detail of merus and carpus, lateral; D—same, chela, lateral; E—minor cheliped, lateral; F—same, chela, mesial." httpUri="https://zenodo.org/record/243883/files/figure.png" pageId="3" pageNumber="523">Figs. 2</figureCitation>
,
<figureCitation id="137E2A0C2145FFC5499D9741A3FEF8AE" box="[418,434,1837,1861]" captionStart="FIGURE 3" captionStartId="5.[151,250,1952,1974]" captionTargetBox="[151,1434,183,1854]" captionTargetId="figure@5.[151,1436,183,1942]" captionTargetPageId="5" captionText="FIGURE 3. Salmoneus alvarezi sp. nov., living specimens: A, B—paratype, ov. specimen from Bahía Málaga, Colombia (OUMNH. ZC. 2015.01.089); C, D—non-type, ov. specimen from Punta Morales, Costa Rica (OUMNH. ZC. 2015.01.090); A, C—dorsal view; B, D—lateral view. Photographs by A. Anker." httpUri="https://zenodo.org/record/243884/files/figure.png" pageId="3" pageNumber="523">3</figureCitation>
). Major cheliped with ischium unarmed, depressed ventrally; merus strongly inflated, widening distally, excavated ventrally (especially distally), with shallow depression distodorsally; carpus moderately long, with depressed ventral margin; chela stout, with fingers about 0.5 length of palm; palm conspicuously flattened ventromesially, excavated ventrally, distoventral area forming short shoulder; fingers moderately stout, with strongly curved, crossing tips; cutting edges of both fingers with about 10 subtriangular or rounded teeth extending from finger base to 0.80.9 of finger length (
<figureCitation id="137E2A0C2142FFC24B8290FBA64EFF5B" box="[957,1026,151,176]" captionStart="FIGURE 2" captionStartId="3.[151,250,1318,1340]" captionTargetBox="[162,1428,202,1288]" captionTargetId="figure@3.[151,1436,193,1297]" captionTargetPageId="3" captionText="FIGURE 2. Salmoneus alvarezi sp. nov., holotype from Bahía Málaga, Colombia (INV CRU 8365): A—major cheliped, lateral (mastigobranch detached); B—same, mesial; C—same, unfolded, detail of merus and carpus, lateral; D—same, chela, lateral; E—minor cheliped, lateral; F—same, chela, mesial." httpUri="https://zenodo.org/record/243883/files/figure.png" pageId="4" pageNumber="524">Fig. 2</figureCitation>
CD). Minor cheliped with ischium shorter than merus, unarmed; merus equal to carpus; carpus cylindrical, widening distally; chela small, simple, with fingers equal to palm, slightly gaping; finger cutting edges unarmed (
<figureCitation id="137E2A0C2142FFC24BE790B3A657FF13" box="[984,1051,223,248]" captionStart="FIGURE 2" captionStartId="3.[151,250,1318,1340]" captionTargetBox="[162,1428,202,1288]" captionTargetId="figure@3.[151,1436,193,1297]" captionTargetPageId="3" captionText="FIGURE 2. Salmoneus alvarezi sp. nov., holotype from Bahía Málaga, Colombia (INV CRU 8365): A—major cheliped, lateral (mastigobranch detached); B—same, mesial; C—same, unfolded, detail of merus and carpus, lateral; D—same, chela, lateral; E—minor cheliped, lateral; F—same, chela, mesial." httpUri="https://zenodo.org/record/243883/files/figure.png" pageId="4" pageNumber="524">Fig. 2</figureCitation>
E, F).
</paragraph>
<paragraph id="8BFA36892142FFC248F89168A0C3FD8B" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="524">
Second pereiopod relatively short, slender; ischium unarmed; merus longer than ischium; carpus five-jointed, first joint slightly longer than sum of remaining joints (
<figureCitation id="137E2A0C2142FFC24B19914BA127FEAB" box="[806,875,295,320]" captionStart="FIGURE 1" captionStartId="2.[151,250,1798,1820]" captionTargetBox="[162,1411,208,1768]" captionTargetId="figure@2.[151,1436,193,1777]" captionTargetPageId="2" captionText="FIGURE 1. Salmoneus alvarezi sp. nov., holotype from Bahía Málaga, Colombia (INV CRU 8365): A—frontal region, dorsal; B—same, lateral; C—posterior abdominal somites, lateral; D—telson, dorsal; E—same, posterior margin, dorsal; F—antennal scaphocerite, dorsal; G—third maxilliped, lateral; H—same, tip of ultimate article, dorsomesial; I—second pereiopod, lateral; J—third pereiopod, lateral; K—fourth pereiopod, lateral; L—fifth pereiopod, lateral; M—second pleopod, detail of appendices masculina and interna, mesial; N—uropod, dorsal; O—same, detail of distolateral area of exopod, dorsal. H drawn without scale." httpUri="https://zenodo.org/record/243882/files/figure.png" pageId="4" pageNumber="524">
Fig.
<date id="FFFB10492142FFC24B609144A127FEAB" box="[863,875,296,320]" pageId="4" pageNumber="524">1</date>
</figureCitation>
I). Third pereiopod moderately slender; ischium typically with two spiniform setae on ventrolateral surface, sometimes with one spiniform seta; merus about five times as long as wide; carpus with slender distoventral spiniform seta; propodus with three slender spiniform setae on ventral margin and pair of longer and stouter spiniform setae adjacent to dactylus; dactylus slightly more than 0.5 length of propodus, moderately slender, conical, simple, slightly curving distally (
<figureCitation id="137E2A0C2142FFC24C5091DBA6FEFE3B" box="[1135,1202,439,464]" captionStart="FIGURE 1" captionStartId="2.[151,250,1798,1820]" captionTargetBox="[162,1411,208,1768]" captionTargetId="figure@2.[151,1436,193,1777]" captionTargetPageId="2" captionText="FIGURE 1. Salmoneus alvarezi sp. nov., holotype from Bahía Málaga, Colombia (INV CRU 8365): A—frontal region, dorsal; B—same, lateral; C—posterior abdominal somites, lateral; D—telson, dorsal; E—same, posterior margin, dorsal; F—antennal scaphocerite, dorsal; G—third maxilliped, lateral; H—same, tip of ultimate article, dorsomesial; I—second pereiopod, lateral; J—third pereiopod, lateral; K—fourth pereiopod, lateral; L—fifth pereiopod, lateral; M—second pleopod, detail of appendices masculina and interna, mesial; N—uropod, dorsal; O—same, detail of distolateral area of exopod, dorsal. H drawn without scale." httpUri="https://zenodo.org/record/243882/files/figure.png" pageId="4" pageNumber="524">Fig. 1</figureCitation>
J). Fourth pereiopod generally similar to third; ischium typically unarmed, sometimes with one small spiniform seta on ventrolateral surface (
<figureCitation id="137E2A0C2142FFC248C89193A37CFDF3" box="[247,304,511,536]" captionStart="FIGURE 1" captionStartId="2.[151,250,1798,1820]" captionTargetBox="[162,1411,208,1768]" captionTargetId="figure@2.[151,1436,193,1777]" captionTargetPageId="2" captionText="FIGURE 1. Salmoneus alvarezi sp. nov., holotype from Bahía Málaga, Colombia (INV CRU 8365): A—frontal region, dorsal; B—same, lateral; C—posterior abdominal somites, lateral; D—telson, dorsal; E—same, posterior margin, dorsal; F—antennal scaphocerite, dorsal; G—third maxilliped, lateral; H—same, tip of ultimate article, dorsomesial; I—second pereiopod, lateral; J—third pereiopod, lateral; K—fourth pereiopod, lateral; L—fifth pereiopod, lateral; M—second pleopod, detail of appendices masculina and interna, mesial; N—uropod, dorsal; O—same, detail of distolateral area of exopod, dorsal. H drawn without scale." httpUri="https://zenodo.org/record/243882/files/figure.png" pageId="4" pageNumber="524">Fig 1</figureCitation>
K). Fifth pereiopod longer and more slender than third; ischium shorter than that of third pereiopod, unarmed; merus subequal to carpus; propodus with at least three slender spiniform setae on ventral margin and well-developed setal brush distally (
<figureCitation id="137E2A0C2142FFC24A13922BA027FD8B" box="[556,619,583,608]" captionStart="FIGURE 1" captionStartId="2.[151,250,1798,1820]" captionTargetBox="[162,1411,208,1768]" captionTargetId="figure@2.[151,1436,193,1777]" captionTargetPageId="2" captionText="FIGURE 1. Salmoneus alvarezi sp. nov., holotype from Bahía Málaga, Colombia (INV CRU 8365): A—frontal region, dorsal; B—same, lateral; C—posterior abdominal somites, lateral; D—telson, dorsal; E—same, posterior margin, dorsal; F—antennal scaphocerite, dorsal; G—third maxilliped, lateral; H—same, tip of ultimate article, dorsomesial; I—second pereiopod, lateral; J—third pereiopod, lateral; K—fourth pereiopod, lateral; L—fifth pereiopod, lateral; M—second pleopod, detail of appendices masculina and interna, mesial; N—uropod, dorsal; O—same, detail of distolateral area of exopod, dorsal. H drawn without scale." httpUri="https://zenodo.org/record/243882/files/figure.png" pageId="4" pageNumber="524">Fig. 1</figureCitation>
L).
</paragraph>
<paragraph id="8BFA36892142FFC248F89200A199FD27" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="524">
Second pleopod with appendix masculina slightly longer than appendix interna, with stiff setae on and near apex (
<figureCitation id="137E2A0C2142FFC248DE92E3A369FD43" box="[225,293,655,680]" captionStart="FIGURE 1" captionStartId="2.[151,250,1798,1820]" captionTargetBox="[162,1411,208,1768]" captionTargetId="figure@2.[151,1436,193,1777]" captionTargetPageId="2" captionText="FIGURE 1. Salmoneus alvarezi sp. nov., holotype from Bahía Málaga, Colombia (INV CRU 8365): A—frontal region, dorsal; B—same, lateral; C—posterior abdominal somites, lateral; D—telson, dorsal; E—same, posterior margin, dorsal; F—antennal scaphocerite, dorsal; G—third maxilliped, lateral; H—same, tip of ultimate article, dorsomesial; I—second pereiopod, lateral; J—third pereiopod, lateral; K—fourth pereiopod, lateral; L—fifth pereiopod, lateral; M—second pleopod, detail of appendices masculina and interna, mesial; N—uropod, dorsal; O—same, detail of distolateral area of exopod, dorsal. H drawn without scale." httpUri="https://zenodo.org/record/243882/files/figure.png" pageId="4" pageNumber="524">Fig. 1</figureCitation>
M). Uropod with moderately narrow exopod and endopod; diaeresis sinuous, with subtriangular distolateral tooth adjacent to stout distolateral spiniform seta (
<figureCitation id="137E2A0C2142FFC24B7392D8A1C6FD27" box="[844,906,692,717]" captionStart="FIGURE 1" captionStartId="2.[151,250,1798,1820]" captionTargetBox="[162,1411,208,1768]" captionTargetId="figure@2.[151,1436,193,1777]" captionTargetPageId="2" captionText="FIGURE 1. Salmoneus alvarezi sp. nov., holotype from Bahía Málaga, Colombia (INV CRU 8365): A—frontal region, dorsal; B—same, lateral; C—posterior abdominal somites, lateral; D—telson, dorsal; E—same, posterior margin, dorsal; F—antennal scaphocerite, dorsal; G—third maxilliped, lateral; H—same, tip of ultimate article, dorsomesial; I—second pereiopod, lateral; J—third pereiopod, lateral; K—fourth pereiopod, lateral; L—fifth pereiopod, lateral; M—second pleopod, detail of appendices masculina and interna, mesial; N—uropod, dorsal; O—same, detail of distolateral area of exopod, dorsal. H drawn without scale." httpUri="https://zenodo.org/record/243882/files/figure.png" pageId="4" pageNumber="524">Fig. 1</figureCitation>
N,
<collectingCountry id="F35276192142FFC24B8F92D8A18BFD27" box="[944,967,692,716]" name="Iceland" pageId="4" pageNumber="524">O</collectingCountry>
).
</paragraph>
<paragraph id="8BFA36892142FFC248F892BBA10CFD1B" blockId="4.[151,1437,151,2013]" box="[199,832,727,752]" pageId="4" pageNumber="524">
Gill/exopod formula typical for genus (see
<bibRefCitation id="EFD44B782142FFC24A9492BBA178FD1B" author="Anker" box="[683,820,727,752]" pageId="4" pageNumber="524" refString="Anker, A. (2010) The shrimp genus Salmoneus Holthuis, 1955 (Crustacea, Decapoda, Alpheidae) in the tropical western Atlantic, with description of five new species. Zootaxa, 2372, 177 - 205." type="journal article" year="2010">Anker 2010</bibRefCitation>
).
</paragraph>
<paragraph id="8BFA36892142FFC248F89290A72DFCD3" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="524">
<emphasis id="B931EA9B2142FFC248F89290A314FCFE" bold="true" box="[199,344,764,789]" pageId="4" pageNumber="524">Colouration</emphasis>
. Uniform bright yolk-yellow to yellow-orange; antennular and antennal flagella, second to fifth pereiopods, and pleopods pale straw-yellow; major chela fingers hyaline-whitish; ovaries orange-red (
<figureCitation id="137E2A0C2142FFC24D309373A71FFCD3" box="[1295,1363,799,824]" captionStart="FIGURE 3" captionStartId="5.[151,250,1952,1974]" captionTargetBox="[151,1434,183,1854]" captionTargetId="figure@5.[151,1436,183,1942]" captionTargetPageId="5" captionText="FIGURE 3. Salmoneus alvarezi sp. nov., living specimens: A, B—paratype, ov. specimen from Bahía Málaga, Colombia (OUMNH. ZC. 2015.01.089); C, D—non-type, ov. specimen from Punta Morales, Costa Rica (OUMNH. ZC. 2015.01.090); A, C—dorsal view; B, D—lateral view. Photographs by A. Anker." httpUri="https://zenodo.org/record/243884/files/figure.png" pageId="4" pageNumber="524">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35F65022142FFC248F89328A2B3FC4E" pageId="4" pageNumber="524" type="etymology">
<paragraph id="8BFA36892142FFC248F89328A2B3FC4E" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="524">
<emphasis id="B931EA9B2142FFC248F89328A30AFCB6" bold="true" box="[199,326,836,861]" pageId="4" pageNumber="524">Etymology</emphasis>
. The new species is named after our colleague, Dr. Fernando Alvarez (Universidad Nacional Autónoma de
<collectingCountry id="F35276192142FFC2490A930BA3C3FC6B" box="[309,399,871,896]" name="Mexico" pageId="4" pageNumber="524">México</collectingCountry>
), in recognition of his numerous contributions to the crustacean fauna of
<collectingCountry id="F35276192142FFC24C81930BA75FFC6B" box="[1214,1299,871,896]" name="Mexico" pageId="4" pageNumber="524">Mexico</collectingCountry>
and Central
<collectingCountry id="F35276192142FFC248A893E0A2B7FC4E" box="[151,251,908,933]" name="United States of America" pageId="4" pageNumber="524">America</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C35F65022142FFC248F893C3A0C0FC23" box="[199,652,943,968]" pageId="4" pageNumber="524" type="materials_examined">
<paragraph id="8BFA36892142FFC248F893C3A0C0FC23" blockId="4.[151,1437,151,2013]" box="[199,652,943,968]" pageId="4" pageNumber="524">
<emphasis id="B931EA9B2142FFC248F893C3A311FC23" bold="true" box="[199,349,943,968]" pageId="4" pageNumber="524">
<typeStatus id="54FE882B2142FFC248F893C3A34EFC23" box="[199,258,943,968]" pageId="4" pageNumber="524">Type</typeStatus>
locality
</emphasis>
. Bahía Málaga,
<collectingCountry id="F35276192142FFC24A2993C3A0C4FC23" box="[534,648,943,968]" name="Colombia" pageId="4" pageNumber="524">Colombia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C35F65022142FFC248F893B8A359FADB" pageId="4" pageNumber="524" type="distribution">
<paragraph id="8BFA36892142FFC248F893B8A1F2FB2E" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="524">
<emphasis id="B931EA9B2142FFC248F893B8A315FC06" bold="true" box="[199,345,980,1005]" pageId="4" pageNumber="524">Distribution</emphasis>
. Tropical eastern Pacific from northern Gulf of California to
<collectingCountry id="F35276192142FFC24C2B93B8A6CFFC06" box="[1044,1155,980,1005]" name="Colombia" pageId="4" pageNumber="524">Colombia</collectingCountry>
and Galapagos. Specific localities include
<collectingCountry id="F35276192142FFC24956939BA38FFBFB" box="[361,451,1015,1040]" name="Mexico" pageId="4" pageNumber="524">Mexico</collectingCountry>
: Laguna Percebú (near San Felipe),
<collectingCountry id="F35276192142FFC24B41939BA1EAFBFB" box="[894,934,1015,1040]" name="Iceland" pageId="4" pageNumber="524">Isla</collectingCountry>
Tiburón, Río Mulegé, Bahía Concepción, southern Baja California (
<bibRefCitation id="EFD44B782142FFC249879470A0F2FBDF" author="Rios" box="[440,702,1052,1077]" pageId="4" pageNumber="524" refString="Rios, R. &amp; Carvacho, A. (1983) Caridean shrimps of the Gulf of California. I. New records, with some remarks on amphi- American distribution. Pacific Science, 36, 459 - 465." type="journal article" year="1983">Ríos &amp; Carvacho 1983</bibRefCitation>
;
<bibRefCitation id="EFD44B782142FFC24AF69470A65DFBDF" author="Villalobos-Hiriart" box="[713,1041,1052,1077]" pageId="4" pageNumber="524" refString="Villalobos-Hiriart, J. L., Nates-Rodriguez, J. C., Diaz-Barriga, A. C., Valle Martinez, M. D., Flores-Hernandez, P., Lira- Fernandez, E. &amp; Schmidtsdorf-Valencia, P. (1989) Listados faunisticos de Mexico. I. Crustaceos estomatopodos y decapodos intermareales de las islas del Golfo de California. Instituto de Biologia, Universidad Nacional Autonoma de Mexico, ix + 101 pp." type="book" year="1989">
Villalobos-Hiriart
<emphasis id="B931EA9B2142FFC24BA89471A183FBDF" box="[919,975,1052,1076]" italics="true" pageId="4" pageNumber="524">et al.</emphasis>
1989
</bibRefCitation>
, 1992;
<bibRefCitation id="EFD44B782142FFC24C5D9470A694FBDF" author="Rios" box="[1122,1240,1052,1076]" pageId="4" pageNumber="524" refString="Rios, R. G. (1989) Un catalogo de camarones carideos de Mulege y Bahia Concepcion, B. C. S. con anotaciones acerca de su biologia, ecologia, distribucion geografica y taxonomia. Tesis profesional, Ensenada, B. C., Mexico, 208 pp." type="book" year="1989">Ríos 1989</bibRefCitation>
,
<bibRefCitation id="EFD44B782142FFC24CDD9470A751FBDF" author="Rios" box="[1250,1309,1052,1076]" pageId="4" pageNumber="524" refString="Rios, R. (1992) Camarones carideos del Golfo de California. VI. Alpheidae del estuario de Mulege y de Bahia Concepcion, Baja California Sur, Mexico (Crustacea: Caridea). Proceedings of San Diego Society of Natural History, 14, 1 - 13." type="journal article" year="1992">1992</bibRefCitation>
;
<bibRefCitation id="EFD44B782142FFC24D169470A29EFBB3" author="Villalobos" pageId="4" pageNumber="524" refString="Villalobos, J. L. (2000) Estudio monografico de los crustaceos decapodos no braquiuros de la zona intermareal de las islas del Golfo de California, Mexico. Tesis de Maestria. Facultad de Ciencias, Universidad Nacional Autonoma de Mexico, 312 pp." type="book" year="2000">Villalobos 2000</bibRefCitation>
, all as
<taxonomicName id="4C454D0A2142FFC2491E942CA3E9FBBC" box="[289,421,1088,1111]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="524" phylum="Arthropoda" rank="species" species="ortmanni">
<emphasis id="B931EA9B2142FFC2491E942CA3E9FBBC" box="[289,421,1088,1111]" italics="true" pageId="4" pageNumber="524">S. ortmanni</emphasis>
</taxonomicName>
);
<collectingCountry id="F35276192142FFC24981942CA072FBB3" box="[446,574,1087,1112]" name="Costa Rica" pageId="4" pageNumber="524">Costa Rica</collectingCountry>
: Punta Morales (present study);
<collectingCountry id="F35276192142FFC24B929453A66CFBB3" box="[941,1056,1087,1112]" name="Colombia" pageId="4" pageNumber="524">Colombia</collectingCountry>
:
<collectingCountry id="F35276192142FFC24C139453A619FBB3" box="[1068,1109,1087,1112]" name="Iceland" pageId="4" pageNumber="524">Isla</collectingCountry>
Playa Blanca near Ensenada de Utria (Chocó) (
<bibRefCitation id="EFD44B782142FFC2494A9408A1E5FB97" author="Christoffersen" box="[373,937,1124,1149]" pageId="4" pageNumber="524" refString="Christoffersen, M. L. &amp; Ramos, G. E. (1988) A new species of Alpheus (Caridea, Alpheidae) from the Pacific coast of Colombia. Revista Nordestina de Biologia, 6, 61 - 65." type="journal article" year="1988">
Christoffersen &amp; Ramos 1988, as
<taxonomicName id="4C454D0A2142FFC24B2D9408A1D6FB97" box="[786,922,1124,1148]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="524" phylum="Arthropoda" rank="species" species="ortmanni">
<emphasis id="B931EA9B2142FFC24B2D9408A1D6FB97" box="[786,922,1124,1148]" italics="true" pageId="4" pageNumber="524">S. ortmanni</emphasis>
</taxonomicName>
*
</bibRefCitation>
), Bahía Málaga (present study);
<collectingCountry id="F35276192142FFC24D089408A7D5FB96" box="[1335,1433,1124,1149]" name="Ecuador" pageId="4" pageNumber="524">Ecuador</collectingCountry>
: Galapagos Archipelago: Gordon Rocks, Post Office Bay, Bahía Gardner (
<bibRefCitation id="EFD44B782142FFC24BE494EBA6C6FB4B" author="Wicksten" box="[987,1162,1159,1184]" pageId="4" pageNumber="524" refString="Wicksten, M. K. (1991) Caridean and stenopodid shrimp of the Galapagos Islands. In: James, M. J. (Ed.), Galapagos marine invertebrates. Plenum, New York, pp. 147 - 156. http: // dx. doi. org / 10.1007 / 978 - 1 - 4899 - 0646 - 5 _ 7" type="book chapter" year="1991">Wicksten 1991</bibRefCitation>
;
<bibRefCitation id="EFD44B782142FFC24CA894EBA29EFB2F" author="Wicksten" pageId="4" pageNumber="524" refString="Wicksten, M. K. &amp; Hendrickx, M. E. (2003) An updated checklist of benthic marine and brackish water shrimps (Decapoda: Penaeoidea, Stenopodidea, Caridea) from the eastern tropical Pacific. In Hendrickx, M. E. (Ed.), Contributions to the Study of East Pacific Crustaceans. Vol. 2. Instituto de Ciencias del Mar y Limnologia, UNAM, Mexico, pp. 49 - 76." type="book" year="2003">Wicksten &amp; Hendrickx 2003</bibRefCitation>
, as
<taxonomicName id="4C454D0A2142FFC248C494C0A332FB2F" box="[251,382,1196,1220]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="524" phylum="Arthropoda" rank="species" species="ortmanni">
<emphasis id="B931EA9B2142FFC248C494C0A332FB2F" box="[251,382,1196,1220]" italics="true" pageId="4" pageNumber="524">S. ortmanni</emphasis>
</taxonomicName>
*) [*tentative re-assignment of records, see below].
</paragraph>
<paragraph id="8BFA36892142FFC248F894A3A359FADB" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="524">
<emphasis id="B931EA9B2142FFC248F894A3A368FB03" bold="true" box="[199,292,1231,1256]" pageId="4" pageNumber="524">Ecology</emphasis>
. Rocky and rocky-sandy intertidal, sometimes with silt or mud component, occasionally also on rocky shores near mangroves or in estuarine conditions; typically found under rocks on coarse or fine sand, presumably free-living.
</paragraph>
</subSubSection>
<subSubSection id="C35F65022142FFC048F89550A745FCD3" lastPageId="6" lastPageNumber="526" pageId="4" pageNumber="524" type="discussion">
<paragraph id="8BFA36892142FFC248F89550A70AF8C3" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="524">
<emphasis id="B931EA9B2142FFC248F89550A378FABE" bold="true" box="[199,308,1340,1365]" pageId="4" pageNumber="524">Remarks</emphasis>
.
<taxonomicName id="4C454D0A2142FFC2497C9550A052FABF" box="[323,542,1340,1364]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="524" phylum="Arthropoda" rank="species" species="alvarezi" status="sp. nov.">
<emphasis id="B931EA9B2142FFC2497C9550A052FABF" box="[323,542,1340,1364]" italics="true" pageId="4" pageNumber="524">Salmoneus alvarezi</emphasis>
</taxonomicName>
<emphasis id="B931EA9B2142FFC24A189550A0CEFABE" bold="true" box="[551,642,1340,1365]" pageId="4" pageNumber="524">
<taxonomicNameLabel id="A20257E02142FFC24A189550A0CEFABE" box="[551,642,1340,1365]" pageId="4" pageNumber="524" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is morphologically closest to the western Atlantic
<taxonomicName id="4C454D0A2142FFC24CF69550A701FABF" box="[1225,1357,1340,1364]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="524" phylum="Arthropoda" rank="species" species="ortmanni">
<emphasis id="B931EA9B2142FFC24CF69550A701FABF" box="[1225,1357,1340,1364]" italics="true" pageId="4" pageNumber="524">S. ortmanni</emphasis>
</taxonomicName>
and
<taxonomicName id="4C454D0A2142FFC24DB89550A351FA93" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="524" phylum="Arthropoda" rank="species" species="wehrtmanni">
<emphasis id="B931EA9B2142FFC24DB89550A351FA93" italics="true" pageId="4" pageNumber="524">S. wehrtmanni</emphasis>
</taxonomicName>
. The new species differs from
<taxonomicName id="4C454D0A2142FFC24A9E950CA167FA9C" box="[673,811,1376,1399]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="524" phylum="Arthropoda" rank="species" species="ortmanni">
<emphasis id="B931EA9B2142FFC24A9E950CA167FA9C" box="[673,811,1376,1399]" italics="true" pageId="4" pageNumber="524">S. ortmanni</emphasis>
</taxonomicName>
by the noticeably depressed rostro-orbital area (not particularly depressed in
<taxonomicName id="4C454D0A2142FFC2498B95E8A07BFA77" box="[436,567,1412,1436]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="524" phylum="Arthropoda" rank="species" species="ortmanni">
<emphasis id="B931EA9B2142FFC2498B95E8A07BFA77" box="[436,567,1412,1436]" italics="true" pageId="4" pageNumber="524">S. ortmanni</emphasis>
</taxonomicName>
); the feebly emarginated posterior margin of the telson (more deeply incised in
<taxonomicName id="4C454D0A2142FFC2488C95C4A379FA54" box="[179,309,1448,1471]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="524" phylum="Arthropoda" rank="species" species="ortmanni">
<emphasis id="B931EA9B2142FFC2488C95C4A379FA54" box="[179,309,1448,1471]" italics="true" pageId="4" pageNumber="524">S. ortmanni</emphasis>
</taxonomicName>
); the longer carpus of the minor cheliped; the longer propodus and more slender dactylus of the third pereiopod; and the slightly longer stylocerite, distinctly overreaching the distal margin of the second article of the antennular peduncle (barely reaching or slightly overreaching this margin in
<taxonomicName id="4C454D0A2142FFC24BCA959CA634F9EC" box="[1013,1144,1520,1543]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="524" phylum="Arthropoda" rank="species" species="ortmanni">
<emphasis id="B931EA9B2142FFC24BCA959CA634F9EC" box="[1013,1144,1520,1543]" italics="true" pageId="4" pageNumber="524">S. ortmanni</emphasis>
</taxonomicName>
) (cf.
<figureCitation id="137E2A0C2142FFC24C899583A74EF9E3" box="[1206,1282,1519,1544]" captionStart="FIGURE 1" captionStartId="2.[151,250,1798,1820]" captionTargetBox="[162,1411,208,1768]" captionTargetId="figure@2.[151,1436,193,1777]" captionTargetPageId="2" captionText="FIGURE 1. Salmoneus alvarezi sp. nov., holotype from Bahía Málaga, Colombia (INV CRU 8365): A—frontal region, dorsal; B—same, lateral; C—posterior abdominal somites, lateral; D—telson, dorsal; E—same, posterior margin, dorsal; F—antennal scaphocerite, dorsal; G—third maxilliped, lateral; H—same, tip of ultimate article, dorsomesial; I—second pereiopod, lateral; J—third pereiopod, lateral; K—fourth pereiopod, lateral; L—fifth pereiopod, lateral; M—second pleopod, detail of appendices masculina and interna, mesial; N—uropod, dorsal; O—same, detail of distolateral area of exopod, dorsal. H drawn without scale." httpUri="https://zenodo.org/record/243882/files/figure.png" pageId="4" pageNumber="524">Figs. 1</figureCitation>
,
<figureCitation id="137E2A0C2142FFC24D2E959CA753F9E3" box="[1297,1311,1520,1544]" captionStart="FIGURE 2" captionStartId="3.[151,250,1318,1340]" captionTargetBox="[162,1428,202,1288]" captionTargetId="figure@3.[151,1436,193,1297]" captionTargetPageId="3" captionText="FIGURE 2. Salmoneus alvarezi sp. nov., holotype from Bahía Málaga, Colombia (INV CRU 8365): A—major cheliped, lateral (mastigobranch detached); B—same, mesial; C—same, unfolded, detail of merus and carpus, lateral; D—same, chela, lateral; E—minor cheliped, lateral; F—same, chela, mesial." httpUri="https://zenodo.org/record/243883/files/figure.png" pageId="4" pageNumber="524">2</figureCitation>
and
<bibRefCitation id="EFD44B782142FFC24D699583A298F9C7" author="Anker" pageId="4" pageNumber="524" refString="Anker, A. (2007) New species and records of alpheid shrimps, genera Salmoneus Holthuis and Parabetaeus Coutiere, from the tropical western Atlantic (Decapoda, Caridea). Zootaxa, 1653, 21 - 39." type="journal article" year="2007">Anker 2007</bibRefCitation>
: figs. 1, 2). The new species can be also separated from
<taxonomicName id="4C454D0A2142FFC24B679678A1B6F9C7" box="[856,1018,1556,1580]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="524" phylum="Arthropoda" rank="species" species="wehrtmanni">
<emphasis id="B931EA9B2142FFC24B679678A1B6F9C7" box="[856,1018,1556,1580]" italics="true" pageId="4" pageNumber="524">S. wehrtmanni</emphasis>
</taxonomicName>
, for instance, by the broadly convex lateral margins of the rostrum (slightly concave to nearly straight in
<taxonomicName id="4C454D0A2142FFC24BB09654A67DF9BB" box="[911,1073,1592,1616]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="524" phylum="Arthropoda" rank="species" species="wehrtmanni">
<emphasis id="B931EA9B2142FFC24BB09654A67DF9BB" box="[911,1073,1592,1616]" italics="true" pageId="4" pageNumber="524">S. wehrtmanni</emphasis>
</taxonomicName>
); the distally wider merus of the major cheliped; and in life, by the bright yellow-orange colour (white in
<taxonomicName id="4C454D0A2142FFC24BED9630A639F99F" box="[978,1141,1628,1652]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="524" phylum="Arthropoda" rank="species" species="wehrtmanni">
<emphasis id="B931EA9B2142FFC24BED9630A639F99F" box="[978,1141,1628,1652]" italics="true" pageId="4" pageNumber="524">S. wehrtmanni</emphasis>
</taxonomicName>
) (cf.
<figureCitation id="137E2A0C2142FFC24C8B9630A752F99F" box="[1204,1310,1628,1653]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,250,1798,1820]" captionStartId-1="3.[151,250,1318,1340]" captionStartId-2="5.[151,250,1952,1974]" captionTargetBox-0="[162,1411,208,1768]" captionTargetBox-1="[162,1428,202,1288]" captionTargetBox-2="[151,1434,183,1854]" captionTargetId-0="figure@2.[151,1436,193,1777]" captionTargetId-1="figure@3.[151,1436,193,1297]" captionTargetId-2="figure@5.[151,1436,183,1942]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="5" captionText-0="FIGURE 1. Salmoneus alvarezi sp. nov., holotype from Bahía Málaga, Colombia (INV CRU 8365): A—frontal region, dorsal; B—same, lateral; C—posterior abdominal somites, lateral; D—telson, dorsal; E—same, posterior margin, dorsal; F—antennal scaphocerite, dorsal; G—third maxilliped, lateral; H—same, tip of ultimate article, dorsomesial; I—second pereiopod, lateral; J—third pereiopod, lateral; K—fourth pereiopod, lateral; L—fifth pereiopod, lateral; M—second pleopod, detail of appendices masculina and interna, mesial; N—uropod, dorsal; O—same, detail of distolateral area of exopod, dorsal. H drawn without scale." captionText-1="FIGURE 2. Salmoneus alvarezi sp. nov., holotype from Bahía Málaga, Colombia (INV CRU 8365): A—major cheliped, lateral (mastigobranch detached); B—same, mesial; C—same, unfolded, detail of merus and carpus, lateral; D—same, chela, lateral; E—minor cheliped, lateral; F—same, chela, mesial." captionText-2="FIGURE 3. Salmoneus alvarezi sp. nov., living specimens: A, B—paratype, ov. specimen from Bahía Málaga, Colombia (OUMNH. ZC. 2015.01.089); C, D—non-type, ov. specimen from Punta Morales, Costa Rica (OUMNH. ZC. 2015.01.090); A, C—dorsal view; B, D—lateral view. Photographs by A. Anker." httpUri-0="https://zenodo.org/record/243882/files/figure.png" httpUri-1="https://zenodo.org/record/243883/files/figure.png" httpUri-2="https://zenodo.org/record/243884/files/figure.png" pageId="4" pageNumber="524">Figs. 13</figureCitation>
and
<bibRefCitation id="EFD44B782142FFC24D699630A298F973" author="Anker" pageId="4" pageNumber="524" refString="Anker, A. (2010) The shrimp genus Salmoneus Holthuis, 1955 (Crustacea, Decapoda, Alpheidae) in the tropical western Atlantic, with description of five new species. Zootaxa, 2372, 177 - 205." type="journal article" year="2010">Anker 2010</bibRefCitation>
: 10, 11, 14d). In
<taxonomicName id="4C454D0A2142FFC249AC96ECA079F973" box="[403,565,1664,1688]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="524" phylum="Arthropoda" rank="species" species="wehrtmanni">
<emphasis id="B931EA9B2142FFC249AC96ECA079F973" box="[403,565,1664,1688]" italics="true" pageId="4" pageNumber="524">S. wehrtmanni</emphasis>
</taxonomicName>
, the first carpal article of the second pereiopod may be either longer (as in
<taxonomicName id="4C454D0A2142FFC24DB896ECA2BCF957" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="524" phylum="Arthropoda" rank="species" species="alvarezi" status="sp. nov.">
<emphasis id="B931EA9B2142FFC24DB896ECA2BCF957" italics="true" pageId="4" pageNumber="524">S. alvarezi</emphasis>
</taxonomicName>
<emphasis id="B931EA9B2142FFC248C596C8A31AF956" bold="true" box="[250,342,1700,1725]" pageId="4" pageNumber="524">
<taxonomicNameLabel id="A20257E02142FFC248C596C8A31AF956" box="[250,342,1700,1725]" pageId="4" pageNumber="524" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
) or shorter than the sum of the remaining articles (
<bibRefCitation id="EFD44B782142FFC24B9596C8A674F957" author="Anker" box="[938,1080,1700,1725]" pageId="4" pageNumber="524" refString="Anker, A. (2010) The shrimp genus Salmoneus Holthuis, 1955 (Crustacea, Decapoda, Alpheidae) in the tropical western Atlantic, with description of five new species. Zootaxa, 2372, 177 - 205." type="journal article" year="2010">Anker 2010</bibRefCitation>
: figs. 10e, 13b); therefore, this character cannot be used in the separation of the two species.
<taxonomicName id="4C454D0A2142FFC24B7396A4A669F90B" box="[844,1061,1736,1760]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="524" phylum="Arthropoda" rank="species" species="alvarezi" status="sp. nov.">
<emphasis id="B931EA9B2142FFC24B7396A4A669F90B" box="[844,1061,1736,1760]" italics="true" pageId="4" pageNumber="524">Salmoneus alvarezi</emphasis>
</taxonomicName>
<emphasis id="B931EA9B2142FFC24C1396ABA6CAF90B" bold="true" box="[1068,1158,1735,1760]" pageId="4" pageNumber="524">
<taxonomicNameLabel id="A20257E02142FFC24C1396ABA6CAF90B" box="[1068,1158,1735,1760]" pageId="4" pageNumber="524" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is more distantly related to
<taxonomicName id="4C454D0A2142FFC248889680A30AF8EF" box="[183,326,1772,1796]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="524" phylum="Arthropoda" rank="species" species="carvachoi">
<emphasis id="B931EA9B2142FFC248889680A30AF8EF" box="[183,326,1772,1796]" italics="true" pageId="4" pageNumber="524">S. carvachoi</emphasis>
</taxonomicName>
from the western Atlantic and its below-described sister species from the eastern Pacific, both differing from the new species by several morphological characters, colour pattern and habitat (see below).
</paragraph>
<paragraph id="8BFA36892142FFC048F89758A108FE8F" blockId="4.[151,1437,151,2013]" lastBlockId="6.[151,1437,151,824]" lastPageId="6" lastPageNumber="526" pageId="4" pageNumber="524">
All previous records of
<taxonomicName id="4C454D0A2142FFC249E59758A013F8A7" box="[474,607,1844,1868]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="524" phylum="Arthropoda" rank="species" species="ortmanni">
<emphasis id="B931EA9B2142FFC249E59758A013F8A7" box="[474,607,1844,1868]" italics="true" pageId="4" pageNumber="524">S. ortmanni</emphasis>
</taxonomicName>
from the eastern Pacific (e.g.,
<bibRefCitation id="EFD44B782142FFC24BFB9758A683F8A7" author="Rios" box="[964,1231,1844,1869]" pageId="4" pageNumber="524" refString="Rios, R. &amp; Carvacho, A. (1983) Caridean shrimps of the Gulf of California. I. New records, with some remarks on amphi- American distribution. Pacific Science, 36, 459 - 465." type="journal article" year="1983">Ríos &amp; Carvacho 1983</bibRefCitation>
;
<bibRefCitation id="EFD44B782142FFC24CE29758A36AF89B" author="Christoffersen" pageId="4" pageNumber="524" refString="Christoffersen, M. L. &amp; Ramos, G. E. (1988) A new species of Alpheus (Caridea, Alpheidae) from the Pacific coast of Colombia. Revista Nordestina de Biologia, 6, 61 - 65." type="journal article" year="1988">Christoffersen &amp; Ramos 1988</bibRefCitation>
;
<bibRefCitation id="EFD44B782142FFC2490E973BA03BF89B" author="Villalobos-Hiriart" box="[305,631,1879,1904]" pageId="4" pageNumber="524" refString="Villalobos-Hiriart, J. L., Nates-Rodriguez, J. C., Diaz-Barriga, A. C., Valle Martinez, M. D., Flores-Hernandez, P., Lira- Fernandez, E. &amp; Schmidtsdorf-Valencia, P. (1989) Listados faunisticos de Mexico. I. Crustaceos estomatopodos y decapodos intermareales de las islas del Golfo de California. Instituto de Biologia, Universidad Nacional Autonoma de Mexico, ix + 101 pp." type="book" year="1989">
Villalobos-Hiriart
<emphasis id="B931EA9B2142FFC249C19735A07AF89B" box="[510,566,1880,1904]" italics="true" pageId="4" pageNumber="524">et al.</emphasis>
1989
</bibRefCitation>
;
<bibRefCitation id="EFD44B782142FFC24ABC973BA161F89B" author="Wicksten" box="[643,813,1879,1904]" pageId="4" pageNumber="524" refString="Wicksten, M. K. (1991) Caridean and stenopodid shrimp of the Galapagos Islands. In: James, M. J. (Ed.), Galapagos marine invertebrates. Plenum, New York, pp. 147 - 156. http: // dx. doi. org / 10.1007 / 978 - 1 - 4899 - 0646 - 5 _ 7" type="book chapter" year="1991">Wicksten 1991</bibRefCitation>
;
<bibRefCitation id="EFD44B782142FFC24B079734A1E7F89B" author="Rios" box="[824,939,1880,1904]" pageId="4" pageNumber="524" refString="Rios, R. (1992) Camarones carideos del Golfo de California. VI. Alpheidae del estuario de Mulege y de Bahia Concepcion, Baja California Sur, Mexico (Crustacea: Caridea). Proceedings of San Diego Society of Natural History, 14, 1 - 13." type="journal article" year="1992">Ríos 1992</bibRefCitation>
) are reassigned to
<taxonomicName id="4C454D0A2142FFC24C449734A6A3F89B" box="[1147,1263,1880,1904]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="524" phylum="Arthropoda" rank="species" species="alvarezi" status="sp. nov.">
<emphasis id="B931EA9B2142FFC24C449734A6A3F89B" box="[1147,1263,1880,1904]" italics="true" pageId="4" pageNumber="524">S. alvarezi</emphasis>
</taxonomicName>
<emphasis id="B931EA9B2142FFC24CC9973BA703F89B" bold="true" box="[1270,1359,1879,1904]" pageId="4" pageNumber="524">
<taxonomicNameLabel id="A20257E02142FFC24CC9973BA703F89B" box="[1270,1359,1879,1904]" pageId="4" pageNumber="524" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, based on illustrations in
<bibRefCitation id="EFD44B782142FFC249589710A3A2F87F" author="Rios" box="[359,494,1916,1940]" pageId="4" pageNumber="524" refString="Rios, R. G. (1989) Un catalogo de camarones carideos de Mulege y Bahia Concepcion, B. C. S. con anotaciones acerca de su biologia, ecologia, distribucion geografica y taxonomia. Tesis profesional, Ensenada, B. C., Mexico, 208 pp." type="book" year="1989">Ríos (1989)</bibRefCitation>
and
<bibRefCitation id="EFD44B782142FFC24A169710A0BEF87F" author="Villalobos" box="[553,754,1916,1941]" pageId="4" pageNumber="524" refString="Villalobos, J. L. (2000) Estudio monografico de los crustaceos decapodos no braquiuros de la zona intermareal de las islas del Golfo de California, Mexico. Tesis de Maestria. Facultad de Ciencias, Universidad Nacional Autonoma de Mexico, 312 pp." type="book" year="2000">Villalobos (2000)</bibRefCitation>
, morphological notes and colour description in
<bibRefCitation id="EFD44B782142FFC24D2A9710A7D0F87F" author="Rios" box="[1301,1436,1916,1940]" pageId="4" pageNumber="524" refString="Rios, R. (1992) Camarones carideos del Golfo de California. VI. Alpheidae del estuario de Mulege y de Bahia Concepcion, Baja California Sur, Mexico (Crustacea: Caridea). Proceedings of San Diego Society of Natural History, 14, 1 - 13." type="journal article" year="1992">Ríos (1992)</bibRefCitation>
and some ecological data in
<bibRefCitation id="EFD44B782142FFC249E897CCA176F853" author="Christoffersen" box="[471,826,1952,1977]" pageId="4" pageNumber="524" refString="Christoffersen, M. L. &amp; Ramos, G. E. (1988) A new species of Alpheus (Caridea, Alpheidae) from the Pacific coast of Colombia. Revista Nordestina de Biologia, 6, 61 - 65." type="journal article" year="1988">Christoffersen &amp; Ramos (1988)</bibRefCitation>
and
<bibRefCitation id="EFD44B782142FFC24B4E97CCA67CF853" author="Wicksten" box="[881,1072,1952,1977]" pageId="4" pageNumber="524" refString="Wicksten, M. K. (1991) Caridean and stenopodid shrimp of the Galapagos Islands. In: James, M. J. (Ed.), Galapagos marine invertebrates. Plenum, New York, pp. 147 - 156. http: // dx. doi. org / 10.1007 / 978 - 1 - 4899 - 0646 - 5 _ 7" type="book chapter" year="1991">Wicksten (1991)</bibRefCitation>
.
<bibRefCitation id="EFD44B782142FFC24C0597CCA6F3F853" author="Rios" box="[1082,1215,1952,1976]" pageId="4" pageNumber="524" refString="Rios, R. (1992) Camarones carideos del Golfo de California. VI. Alpheidae del estuario de Mulege y de Bahia Concepcion, Baja California Sur, Mexico (Crustacea: Caridea). Proceedings of San Diego Society of Natural History, 14, 1 - 13." type="journal article" year="1992">Ríos (1992)</bibRefCitation>
stated that the only consistent difference between his specimens from the northern and central Gulf of California (Laguna Percebú, Río Mulegé and Bahía Concepción) and the Caribbean (
<collectingCountry id="F35276192140FFC04AC490FBA1CBFF5B" box="[763,903,151,176]" name="Guadeloupe" pageId="6" pageNumber="526">Guadeloupe</collectingCountry>
, most likely Carvachos (1979) material) and Brazilian specimens (
<emphasis id="B931EA9B2140FFC049B290D0A3F4FF3F" box="[397,440,188,212]" italics="true" pageId="6" pageNumber="526">fide</emphasis>
<bibRefCitation id="EFD44B782140FFC0498090D0A0EDFF3F" author="Christoffersen" box="[447,673,188,213]" pageId="6" pageNumber="526" refString="Christoffersen, M. L. (1982) Distribution of warm water alpheoid shrimp (Crustacea, Caridea) on the continental shelf of eastern South America between 23 and 35 ° lat. S. Boletim do Instituto Oceanografico, Sao Paulo, 31, 93 - 112. http: // dx. doi. org / 10.1590 / S 0373 - 55241982000100007" type="journal article" year="1982">Christoffersen 1982</bibRefCitation>
) was the absence of a spiniform seta on the ischium of the second pereiopod in the former specimens. However, the material from
<collectingCountry id="F35276192140FFC04B4B90B3A1B7FF13" box="[884,1019,223,248]" name="Guadeloupe" pageId="6" pageNumber="526">Guadeloupe</collectingCountry>
reported by Carvachos (1979) as
<taxonomicName id="4C454D0A2140FFC04DB8908CA2B2FEF7" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="526" phylum="Arthropoda" rank="species" species="ortmanni">
<emphasis id="B931EA9B2140FFC04DB8908CA2B2FEF7" italics="true" pageId="6" pageNumber="526">S. ortmanni</emphasis>
</taxonomicName>
, as well as Christoffersens (1982) material of
<taxonomicName id="4C454D0A2140FFC04B109168A1F9FEF7" box="[815,949,260,284]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="526" phylum="Arthropoda" rank="species" species="ortmanni">
<emphasis id="B931EA9B2140FFC04B109168A1F9FEF7" box="[815,949,260,284]" italics="true" pageId="6" pageNumber="526">S. ortmanni</emphasis>
</taxonomicName>
from
<collectingCountry id="F35276192140FFC04C3C9168A606FEF6" box="[1027,1098,260,285]" name="Brazil" pageId="6" pageNumber="526">Brazil</collectingCountry>
, was later re-identified as
<taxonomicName id="4C454D0A2140FFC04DB89168A34BFEAB" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="526" phylum="Arthropoda" rank="species" species="carvachoi">
<emphasis id="B931EA9B2140FFC04DB89168A34BFEAB" italics="true" pageId="6" pageNumber="526">S. carvachoi</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFD44B782140FFC04909914BA398FEAB" author="Anker" box="[310,468,295,320]" pageId="6" pageNumber="526" refString="Anker, A. (2007) New species and records of alpheid shrimps, genera Salmoneus Holthuis and Parabetaeus Coutiere, from the tropical western Atlantic (Decapoda, Caridea). Zootaxa, 1653, 21 - 39." type="journal article" year="2007">Anker (2007)</bibRefCitation>
. Thus
<bibRefCitation id="EFD44B782140FFC04A1C9144A0E7FEAB" author="Rios" box="[547,683,296,320]" pageId="6" pageNumber="526" refString="Rios, R. (1992) Camarones carideos del Golfo de California. VI. Alpheidae del estuario de Mulege y de Bahia Concepcion, Baja California Sur, Mexico (Crustacea: Caridea). Proceedings of San Diego Society of Natural History, 14, 1 - 13." type="journal article" year="1992">Ríos (1992)</bibRefCitation>
comparative material of
<taxonomicName id="4C454D0A2140FFC04BEF9144A618FED4" box="[976,1108,296,319]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="526" phylum="Arthropoda" rank="species" species="ortmanni">
<emphasis id="B931EA9B2140FFC04BEF9144A618FED4" box="[976,1108,296,319]" italics="true" pageId="6" pageNumber="526">S. ortmanni</emphasis>
</taxonomicName>
from
<collectingCountry id="F35276192140FFC04CA0914BA76AFEAB" box="[1183,1318,295,320]" name="Guadeloupe" pageId="6" pageNumber="526">Guadeloupe</collectingCountry>
was most likely
<taxonomicName id="4C454D0A2140FFC048E29120A326FE8F" box="[221,362,332,356]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="526" phylum="Arthropoda" rank="species" species="carvachoi">
<emphasis id="B931EA9B2140FFC048E29120A326FE8F" box="[221,362,332,356]" italics="true" pageId="6" pageNumber="526">S. carvachoi</emphasis>
</taxonomicName>
and not
<taxonomicName id="4C454D0A2140FFC049F49120A10CFE8F" authority="Rankin (1898)" authorityName="Rankin" authorityYear="1898" box="[459,832,332,357]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="526" phylum="Arthropoda" rank="species" species="ortmanni">
<emphasis id="B931EA9B2140FFC049F49120A0DEFE8F" box="[459,658,332,356]" italics="true" pageId="6" pageNumber="526">S. ortmanni sensu</emphasis>
<bibRefCitation id="EFD44B782140FFC04AA59120A10CFE8F" author="Rankin" box="[666,832,332,357]" pageId="6" pageNumber="526" refString="Rankin, W. M. (1898) The Northrop collection of Crustacea from the Bahamas. Annals of the New York Academy of Sciences, 11, 225 - 258. http: // dx. doi. org / 10.1111 / j. 1749 - 6632.1898. tb 54971. x" type="journal article" year="1898">Rankin (1898)</bibRefCitation>
</taxonomicName>
.
</paragraph>
<caption id="DF3A66012143FFC348A897CCA15BF81F" httpUri="https://zenodo.org/record/243884/files/figure.png" pageId="5" pageNumber="525" targetBox="[151,1434,183,1854]" targetPageId="5">
<paragraph id="8BFA36892143FFC348A897CCA15BF81F" blockId="5.[151,1436,1952,2036]" pageId="5" pageNumber="525">
<emphasis id="B931EA9B2143FFC348A897CCA35AF85E" bold="true" box="[151,278,1952,1974]" pageId="5" pageNumber="525">FIGURE 3.</emphasis>
<taxonomicName id="4C454D0A2143FFC3491F97CCA3A5F85D" box="[288,489,1952,1974]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="525" phylum="Arthropoda" rank="species" species="alvarezi" status="sp. nov.">
<emphasis id="B931EA9B2143FFC3491F97CCA3A5F85D" box="[288,489,1952,1974]" italics="true" pageId="5" pageNumber="525">Salmoneus alvarezi</emphasis>
</taxonomicName>
<emphasis id="B931EA9B2143FFC349CD97CDA00AF85D" bold="true" box="[498,582,1953,1974]" pageId="5" pageNumber="525">
<taxonomicNameLabel id="A20257E02143FFC349CD97CDA00AF85D" box="[498,582,1953,1974]" pageId="5" pageNumber="525" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, living specimens: A, B—paratype, ov. specimen from Bahía Málaga, Colombia (OUMNH.ZC. 2015.01.089); C, D—non-type, ov. specimen from Punta Morales, Costa Rica (OUMNH.ZC. 2015.01.090); A, C—dorsal view; B, D—lateral view. Photographs by A. Anker.
</paragraph>
</caption>
<paragraph id="8BFA36892140FFC048F8911CA1C5FDD6" blockId="6.[151,1437,151,824]" pageId="6" pageNumber="526">
<bibRefCitation id="EFD44B782140FFC048F8911CA301FE63" author="Rios" box="[199,333,368,392]" pageId="6" pageNumber="526" refString="Rios, R. (1992) Camarones carideos del Golfo de California. VI. Alpheidae del estuario de Mulege y de Bahia Concepcion, Baja California Sur, Mexico (Crustacea: Caridea). Proceedings of San Diego Society of Natural History, 14, 1 - 13." type="journal article" year="1992">Ríos (1992)</bibRefCitation>
noted that the colour in life of the Bahía Concepción specimens was “brilliant greenish yellow”. Since the unarmed ischium of the second pereiopod and the uniform bright yellow colour represent two key characters separating
<taxonomicName id="4C454D0A2140FFC049B591D4A3B3FE3B" box="[394,511,440,464]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="526" phylum="Arthropoda" rank="species" species="alvarezi" status="sp. nov.">
<emphasis id="B931EA9B2140FFC049B591D4A3B3FE3B" box="[394,511,440,464]" italics="true" pageId="6" pageNumber="526">S. alvarezi</emphasis>
</taxonomicName>
<emphasis id="B931EA9B2140FFC04A3991DBA02CFE3B" bold="true" box="[518,608,439,464]" pageId="6" pageNumber="526">
<taxonomicNameLabel id="A20257E02140FFC04A3991DBA02CFE3B" box="[518,608,439,464]" pageId="6" pageNumber="526" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from the below described
<taxonomicName id="4C454D0A2140FFC04BB291D4A66BFE3B" box="[909,1063,440,464]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="526" phylum="Arthropoda" rank="species" species="malagensis" status="sp. nov.">
<emphasis id="B931EA9B2140FFC04BB291D4A66BFE3B" box="[909,1063,440,464]" italics="true" pageId="6" pageNumber="526">S. malagensis</emphasis>
</taxonomicName>
<emphasis id="B931EA9B2140FFC04C1191DBA6C4FE3B" bold="true" box="[1070,1160,439,464]" pageId="6" pageNumber="526">
<taxonomicNameLabel id="A20257E02140FFC04C1191DBA6C4FE3B" box="[1070,1160,439,464]" pageId="6" pageNumber="526" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, the specimens from the Gulf of California reported as
<taxonomicName id="4C454D0A2140FFC049C791B0A031FE1F" box="[504,637,476,500]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="526" phylum="Arthropoda" rank="species" species="ortmanni">
<emphasis id="B931EA9B2140FFC049C791B0A031FE1F" box="[504,637,476,500]" italics="true" pageId="6" pageNumber="526">S. ortmanni</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFD44B782140FFC04A9391B0A162FE1F" author="Rios" box="[684,814,476,500]" pageId="6" pageNumber="526" refString="Rios, R. G. (1989) Un catalogo de camarones carideos de Mulege y Bahia Concepcion, B. C. S. con anotaciones acerca de su biologia, ecologia, distribucion geografica y taxonomia. Tesis profesional, Ensenada, B. C., Mexico, 208 pp." type="book" year="1989">Ríos (1989</bibRefCitation>
,
<bibRefCitation id="EFD44B782140FFC04B0391B0A134FE1F" author="Rios" box="[828,888,476,500]" pageId="6" pageNumber="526" refString="Rios, R. (1992) Camarones carideos del Golfo de California. VI. Alpheidae del estuario de Mulege y de Bahia Concepcion, Baja California Sur, Mexico (Crustacea: Caridea). Proceedings of San Diego Society of Natural History, 14, 1 - 13." type="journal article" year="1992">1992</bibRefCitation>
),
<bibRefCitation id="EFD44B782140FFC04BB291B0A6A1FE1F" box="[909,1261,476,501]" pageId="6" pageNumber="526" refString="Villalobos-Hiriart, J. L., Nates-Rodriguez, J. C., Diaz-Barriga, A. C., Valle Martinez, M. D., Flores-Hernandez, P., Lira- Fernandez, E. &amp; Schmidtsdorf-Valencia, P. (1989) Listados faunisticos de Mexico. I. Crustaceos estomatopodos y decapodos intermareales de las islas del Golfo de California. Instituto de Biologia, Universidad Nacional Autonoma de Mexico, ix + 101 pp." type="book">
Villalobos-Hiriart
<emphasis id="B931EA9B2140FFC04C6191B1A6D5FE1F" box="[1118,1177,476,500]" italics="true" pageId="6" pageNumber="526">et al.</emphasis>
(1989)
</bibRefCitation>
and
<bibRefCitation id="EFD44B782140FFC04D1691B0A2ADFDF3" author="Villalobos" pageId="6" pageNumber="526" refString="Villalobos, J. L. (2000) Estudio monografico de los crustaceos decapodos no braquiuros de la zona intermareal de las islas del Golfo de California, Mexico. Tesis de Maestria. Facultad de Ciencias, Universidad Nacional Autonoma de Mexico, 312 pp." type="book" year="2000">Villalobos (2000)</bibRefCitation>
are reassigned to
<taxonomicName id="4C454D0A2140FFC04993926CA06CFDF3" box="[428,544,512,536]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="526" phylum="Arthropoda" rank="species" species="alvarezi" status="sp. nov.">
<emphasis id="B931EA9B2140FFC04993926CA06CFDF3" box="[428,544,512,536]" italics="true" pageId="6" pageNumber="526">S. alvarezi</emphasis>
</taxonomicName>
<emphasis id="B931EA9B2140FFC04A179193A0CDFDF3" bold="true" box="[552,641,511,536]" pageId="6" pageNumber="526">
<taxonomicNameLabel id="A20257E02140FFC04A179193A0CDFDF3" box="[552,641,511,536]" pageId="6" pageNumber="526" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Records of
<taxonomicName id="4C454D0A2140FFC04B36926CA1C7FDFC" box="[777,907,512,535]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="526" phylum="Arthropoda" rank="species" species="ortmanni">
<emphasis id="B931EA9B2140FFC04B36926CA1C7FDFC" box="[777,907,512,535]" italics="true" pageId="6" pageNumber="526">S. ortmanni</emphasis>
</taxonomicName>
in several regional checklists (e.g.
<bibRefCitation id="EFD44B782140FFC04D289193A303FDD7" author="Wicksten" pageId="6" pageNumber="526" refString="Wicksten, M. K. &amp; Hendrickx, M. E. (1992) Checklist of penaeoid and caridean shrimps (Decapoda: Penaeoidea, Caridea) from the eastern tropical Pacific. Proceedings of the San Diego Society of Natural History, 9, 1 - 11." type="journal article" year="1992">Wicksten &amp; Hendrickx 1992</bibRefCitation>
,
<bibRefCitation id="EFD44B782140FFC049659248A3D8FDD7" author="Hendrickx" box="[346,404,548,572]" pageId="6" pageNumber="526" refString="Wicksten, M. K. &amp; Hendrickx, M. E. (2003) An updated checklist of benthic marine and brackish water shrimps (Decapoda: Penaeoidea, Stenopodidea, Caridea) from the eastern tropical Pacific. In Hendrickx, M. E. (Ed.), Contributions to the Study of East Pacific Crustaceans. Vol. 2. Instituto de Ciencias del Mar y Limnologia, UNAM, Mexico, pp. 49 - 76." type="book" year="2003">2003</bibRefCitation>
) most likely also refer to
<taxonomicName id="4C454D0A2140FFC04A8C9248A164FDD7" box="[691,808,548,572]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="526" phylum="Arthropoda" rank="species" species="alvarezi" status="sp. nov.">
<emphasis id="B931EA9B2140FFC04A8C9248A164FDD7" box="[691,808,548,572]" italics="true" pageId="6" pageNumber="526">S. alvarezi</emphasis>
</taxonomicName>
<emphasis id="B931EA9B2140FFC04B109248A1C5FDD6" bold="true" box="[815,905,548,573]" pageId="6" pageNumber="526">
<taxonomicNameLabel id="A20257E02140FFC04B109248A1C5FDD6" box="[815,905,548,573]" pageId="6" pageNumber="526" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BFA36892140FFC048F8922BA745FCD3" blockId="6.[151,1437,151,824]" pageId="6" pageNumber="526">
Christoffersen &amp; Ramos (1988) material of
<taxonomicName id="4C454D0A2140FFC04AFA9224A105FDB4" box="[709,841,584,607]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="526" phylum="Arthropoda" rank="species" species="ortmanni">
<emphasis id="B931EA9B2140FFC04AFA9224A105FDB4" box="[709,841,584,607]" italics="true" pageId="6" pageNumber="526">S. ortmanni</emphasis>
</taxonomicName>
from
<collectingCountry id="F35276192140FFC04BAE922BA1F5FD8B" box="[913,953,583,608]" name="Iceland" pageId="6" pageNumber="526">Isla</collectingCountry>
Playa Blanca,
<collectingCountry id="F35276192140FFC04C59922BA69BFD8B" box="[1126,1239,583,608]" name="Colombia" pageId="6" pageNumber="526">Colombia</collectingCountry>
, was collected in the intertidal coralline sand, adjacent to large and small rocks, a habitat more typical of
<taxonomicName id="4C454D0A2140FFC04C4E9200A6ABFD6F" box="[1137,1255,620,644]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="526" phylum="Arthropoda" rank="species" species="alvarezi" status="sp. nov.">
<emphasis id="B931EA9B2140FFC04C4E9200A6ABFD6F" box="[1137,1255,620,644]" italics="true" pageId="6" pageNumber="526">S. alvarezi</emphasis>
</taxonomicName>
<emphasis id="B931EA9B2140FFC04CD19200A704FD6E" bold="true" box="[1262,1352,620,645]" pageId="6" pageNumber="526">
<taxonomicNameLabel id="A20257E02140FFC04CD19200A704FD6E" box="[1262,1352,620,645]" pageId="6" pageNumber="526" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
than
<taxonomicName id="4C454D0A2140FFC04DB89200A359FD43" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="526" phylum="Arthropoda" rank="species" species="malagensis" status="sp. nov.">
<emphasis id="B931EA9B2140FFC04DB89200A359FD43" italics="true" pageId="6" pageNumber="526">S. malagensis</emphasis>
</taxonomicName>
<emphasis id="B931EA9B2140FFC0492192E3A335FD43" bold="true" box="[286,377,655,680]" pageId="6" pageNumber="526">
<taxonomicNameLabel id="A20257E02140FFC0492192E3A335FD43" box="[286,377,655,680]" pageId="6" pageNumber="526" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, which was collected from burrows on a tidal mudflat (see below).
<bibRefCitation id="EFD44B782140FFC04CB992E3A70EFD43" author="Wicksten" box="[1158,1346,655,680]" pageId="6" pageNumber="526" refString="Wicksten, M. K. (1991) Caridean and stenopodid shrimp of the Galapagos Islands. In: James, M. J. (Ed.), Galapagos marine invertebrates. Plenum, New York, pp. 147 - 156. http: // dx. doi. org / 10.1007 / 978 - 1 - 4899 - 0646 - 5 _ 7" type="book chapter" year="1991">Wicksten (1991)</bibRefCitation>
did not provide direct notes on the habitat of
<taxonomicName id="4C454D0A2140FFC04A0192D8A08EFD27" box="[574,706,692,716]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="526" phylum="Arthropoda" rank="species" species="ortmanni">
<emphasis id="B931EA9B2140FFC04A0192D8A08EFD27" box="[574,706,692,716]" italics="true" pageId="6" pageNumber="526">S. ortmanni</emphasis>
</taxonomicName>
in the Galapagos localities, but a brief analysis of the syntopical decapod fauna at the collection stations suggests an intertidal or shallow subtidal rocky-sand bottom, which is much more typical of
<taxonomicName id="4C454D0A2140FFC0499F9290A056FCFF" box="[416,538,764,788]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="526" phylum="Arthropoda" rank="species" species="alvarezi" status="sp. nov.">
<emphasis id="B931EA9B2140FFC0499F9290A056FCFF" box="[416,538,764,788]" italics="true" pageId="6" pageNumber="526">S. alvarezi</emphasis>
</taxonomicName>
<emphasis id="B931EA9B2140FFC04A199290A0C8FCFE" bold="true" box="[550,644,764,789]" pageId="6" pageNumber="526">
<taxonomicNameLabel id="A20257E02140FFC04A199290A0C8FCFE" box="[550,644,764,789]" pageId="6" pageNumber="526" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Therefore, Christoffersen &amp; Ramos (1988) and Wickstens (1991) records from
<collectingCountry id="F35276192140FFC049119373A3D0FCD3" box="[302,412,799,824]" name="Colombia" pageId="6" pageNumber="526">Colombia</collectingCountry>
and Galapagos, respectively, are tentatively re-assigned to
<taxonomicName id="4C454D0A2140FFC04C0B934CA6E5FCD3" box="[1076,1193,800,824]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="526" phylum="Arthropoda" rank="species" species="alvarezi" status="sp. nov.">
<emphasis id="B931EA9B2140FFC04C0B934CA6E5FCD3" box="[1076,1193,800,824]" italics="true" pageId="6" pageNumber="526">S. alvarezi</emphasis>
</taxonomicName>
<emphasis id="B931EA9B2140FFC04C8F9373A745FCD3" bold="true" box="[1200,1289,799,824]" pageId="6" pageNumber="526">
<taxonomicNameLabel id="A20257E02140FFC04C8F9373A745FCD3" box="[1200,1289,799,824]" pageId="6" pageNumber="526" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>