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<document id="7517E02D9FF6B8CAD41943B5C4E47431" ID-DOI="10.5281/zenodo.5740224" ID-ISSN="0374-1036" ID-Zenodo-Dep="5740224" ID-ZooBank="84292869-A733-4F7F-B2B8-B65A0356127B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638254309187" checkinUser="marcus" docAuthor="Lackner, Tomáš" docDate="2013" docId="03C4C81DFFBAFFE3244CE331A0B6FE66" docLanguage="en" docName="ActaEntMusNatPra.2013.53.2.623-632.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 53 (2)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Pilisaprinus verschureni Lackner 2013, comb. nov." docType="treatment" docVersion="3" lastPageNumber="630" masterDocId="FFFDB065FFB8FFE42514E12AA363FF95" masterDocTitle="Elevation of Pilisaprinus to full generic status with notes on the evolution of termitoxeny in the Saprininae (Coleoptera: Histeridae)" masterLastPageNumber="632" masterPageNumber="623" pageNumber="625" updateTime="1699435727309" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="450E0157A37F1556D99F9C256F468024" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B652C762F93B9C7F0B15F57B7DD1DD75">
<mods:title id="4A235DB51D83DE577DDFB99975F4D903">Elevation of Pilisaprinus to full generic status with notes on the evolution of termitoxeny in the Saprininae (Coleoptera: Histeridae)</mods:title>
</mods:titleInfo>
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<mods:roleTerm id="24F7FF61D10630C0C8E9660CB24C1C25">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5307D2D7395C9831658A3F65E70B4C5D">Lackner, Tomáš</mods:namePart>
</mods:name>
<mods:typeOfResource id="7F6CCA064F30BF499933B2F663CC5CD8">text</mods:typeOfResource>
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<mods:title id="D266078755A5B8ED85CEF5B95A2ABD48">Acta Entomologica Musei Nationalis Pragae</mods:title>
</mods:titleInfo>
<mods:part id="459EB96018F79FD8C3F4ADCFD59ED0B9">
<mods:date id="5F8BA09D01B1366D6D11C7DEFB5D81C1">2013</mods:date>
<mods:detail id="96ECF0BA5A323778A745DF4EF156528E" type="pubDate">
<mods:number id="EE1232815087CB117DA2F7C615256C89">2013-11-15</mods:number>
</mods:detail>
<mods:detail id="31B71D395973271C8A05196B573BC763" type="volume">
<mods:number id="DEF2006A3BD3D530114D3AF7D2CC48DD">53</mods:number>
</mods:detail>
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<mods:number id="B82F45DCB84730AABFA37F9C25277894">2</mods:number>
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<mods:start id="935ADE6F1D8F8D26B88B5D82A7833658">623</mods:start>
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<mods:classification id="70B79F62BD96FC3329B70226E4251B14">journal article</mods:classification>
<mods:identifier id="376710E103F54A7371EDCE20C92F045D" type="DOI">10.5281/zenodo.5740224</mods:identifier>
<mods:identifier id="E008800103FD9C6B300284E2E1381B9A" type="ISSN">0374-1036</mods:identifier>
<mods:identifier id="3532C020A676474739E4983A80429519" type="Zenodo-Dep">5740224</mods:identifier>
<mods:identifier id="AFFBEDE9B6E8868115ADD0201F527375" type="ZooBank">84292869-A733-4F7F-B2B8-B65A0356127B</mods:identifier>
</mods:mods>
<treatment id="03C4C81DFFBAFFE3244CE331A0B6FE66" LSID="urn:lsid:plazi:treatment:03C4C81DFFBAFFE3244CE331A0B6FE66" httpUri="http://treatment.plazi.org/id/03C4C81DFFBAFFE3244CE331A0B6FE66" lastPageId="7" lastPageNumber="630" pageId="2" pageNumber="625">
<subSubSection id="C3772A80FFBAFFE6244CE331A0C5FDA1" box="[344,934,539,564]" pageId="2" pageNumber="625" type="nomenclature">
<paragraph id="8BD2790BFFBAFFE6244CE331A0C5FDA1" blockId="2.[344,934,539,564]" box="[344,934,539,564]" pageId="2" pageNumber="625">
<heading id="D09ACE67FFBAFFE6244CE331A0C5FDA1" bold="true" box="[344,934,539,564]" centered="true" fontSize="8" level="2" pageId="2" pageNumber="625" reason="3">
<emphasis id="B919A519FFBAFFE6244CE331A0C5FDA1" bold="true" box="[344,934,539,564]" pageId="2" pageNumber="625">
<taxonomicName id="4C6D0288FFBAFFE6244CE331A047FDA6" ID-CoL="4HTRW" authority="(Therond, 1959)" authorityName="Lackner" authorityYear="2013" baseAuthorityName="Therond" baseAuthorityYear="1959" box="[344,804,539,564]" class="Insecta" family="Histeridae" genus="Pilisaprinus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="625" phylum="Arthropoda" rank="species" species="verschureni" status="comb. nov.">
<emphasis id="B919A519FFBAFFE6244CE331A105FDA1" bold="true" box="[344,614,539,564]" italics="true" pageId="2" pageNumber="625">Pilisaprinus verschureni</emphasis>
(
<bibRefCitation id="EFFC04FAFFBAFFE62761E331A07EFDA6" author="THEROND J." box="[629,797,539,564]" pageId="2" pageNumber="625" pagination="1 - 39" refId="ref4728" refString="THEROND J. 1959: Histeridae (Coleoptera, Staphylinoidea). Exploration de la Parc Naturel du Garamba, Mission de Saeger 15 (1): 1 - 39." type="journal article" year="1959">Thérond, 1959</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A22A1862FFBAFFE6263FE336A0C5FDA1" box="[811,934,540,564]" pageId="2" pageNumber="625" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3772A80FFBAFFE62759E310A1D2FDD8" box="[589,689,570,589]" pageId="2" pageNumber="625" type="description">
<paragraph id="8BD2790BFFBAFFE62759E310A1D2FDD8" blockId="2.[151,1029,570,677]" box="[589,689,570,589]" pageId="2" pageNumber="625">
(
<figureCitation id="1356658EFFBAFFE62747E310A1CAFDD8" box="[595,681,570,589]" captionStart-0="Figs 1–9" captionStart-1="Figs 10–12" captionStart-2="Figs 13–19" captionStartId-0="3.[113,150,1513,1532]" captionStartId-1="4.[151,187,1104,1123]" captionStartId-2="5.[113,150,1122,1141]" captionTargetBox-0="[211,991,219,1492]" captionTargetBox-1="[247,1028,220,1075]" captionTargetBox-2="[252,944,222,1095]" captionTargetId-0="figure-12@3.[211,991,219,1492]" captionTargetId-1="figure-153@4.[246,1029,219,1075]" captionTargetId-2="figure-149@5.[251,950,219,1107]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="Figs 1–9. External morphology of Pilisaprinus verschureni (Thérond, 1959). 1 – habitus, dorsal view; 2 – same, ventral view; 3 – antennal club, latero-ventral view; 4 – head, dorsal view; 5 – mentum, ventral view; 6 – prosternum; 7 – propygidium and pygidium; 8 – lateral disc of metaventrite + metepisternum; 9 – protibia, dorsal view." captionText-1="Figs 10–12. Mouthparts and sensory structures of the antenna of Pilisaprinus verschureni (Thérond, 1959).10 – mandibles, dorsal view; 11 – labrum, left half showing dorsal view; right half showing ventral view; 12 – antennal club showing sensory structures of the antenna." captionText-2="Figs 13–19. Male terminalia of Pilisaprinus verschureni (Thérond, 1959). 13 – eighth sternite and tergite, ventral view; 14 – same, ventral view; 15 – same, lateral view; 16 – ninth and tenth tergites, dorsal view; spiculum gastrale, ventral view; 17 – ninth, tenth tergites and spiculum gastrale, lateral view; 18 – aedeagus, dorsal view; 19 – same, lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.5740226" figureDoi-1="http://doi.org/10.5281/zenodo.5740230" figureDoi-2="http://doi.org/10.5281/zenodo.5740232" httpUri-0="https://zenodo.org/record/5740226/files/figure.png" httpUri-1="https://zenodo.org/record/5740230/files/figure.png" httpUri-2="https://zenodo.org/record/5740232/files/figure.png" pageId="2" pageNumber="625">Figs 1–19</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3772A80FFBAFFE62583E377A11DFD30" pageId="2" pageNumber="625" type="reference_group">
<paragraph id="8BD2790BFFBAFFE62583E377A11DFD30" blockId="2.[151,1029,570,677]" pageId="2" pageNumber="625">
<treatmentCitationGroup id="AB7D5E25FFBAFFE62583E377A11DFD30" pageId="2" pageNumber="625">
<treatmentCitation id="0ACC5F1AFFBAFFE62583E377A292FDFA" author="THEROND J." box="[151,497,604,623]" page="17" pageId="2" pageNumber="625" year="1959">
<taxonomicName id="4C6D0288FFBAFFE62583E377A292FDFA" ID-CoL="79KMQ" authority="Therond, 1959: 17" authorityName="Therond" authorityPageNumber="17" authorityYear="1959" box="[151,497,604,623]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="625" phylum="Arthropoda" rank="species" species="verschureni">
<emphasis id="B919A519FFBAFFE62583E377A22FFDFA" box="[151,332,605,623]" italics="true" pageId="2" pageNumber="625">Saprinus verschureni</emphasis>
<bibRefCitation id="EFFC04FAFFBAFFE62445E376A292FDFA" author="THEROND J." box="[337,497,604,623]" pageId="2" pageNumber="625" pagination="1 - 39" refId="ref4728" refString="THEROND J. 1959: Histeridae (Coleoptera, Staphylinoidea). Exploration de la Parc Naturel du Garamba, Mission de Saeger 15 (1): 1 - 39." type="journal article" year="1959">Thérond, 1959: 17</bibRefCitation>
</taxonomicName>
</treatmentCitation>
; MAZUR (1984: 62);
<treatmentCitation id="0ACC5F1AFFBAFFE627B4E377A02BFDFA" author="MAZUR S." box="[672,840,604,624]" page="231" pageId="2" pageNumber="625" year="1997">
<bibRefCitation id="EFFC04FAFFBAFFE627B4E377A02BFDFA" author="MAZUR S." box="[672,840,604,624]" pageId="2" pageNumber="625" pagination="1 - 373" refId="ref4643" refString="MAZUR S. 1997: A world catalogue of the Histeridae (Coleoptera: Histeridae). Genus, Supplement, 1 - 373." type="book chapter" year="1997">MAZUR (1997: 231)</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0ACC5F1AFFBAFFE62645E377A761FDFA" author="KANAAR P." box="[849,1026,604,624]" page="130" pageId="2" pageNumber="625" year="1996">
<bibRefCitation id="EFFC04FAFFBAFFE62645E377A761FDFA" author="KANAAR P." box="[849,1026,604,624]" pageId="2" pageNumber="625" pagination="130" refId="ref4271" refString="KANAAR P. 1996: Styphrus termitophilus Kanaar, 1990 = Saprinus verschureni Therond, 1959. nov. syn., type d'une nouveau subgenus Pilisaprinus dans le genre Saprinus Er. Nouvelle Revue d'Entomologie (N. S.) 13 (2): 130." type="journal article" year="1996">KANAAR (1996: 130)</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation id="0ACC5F1AFFBAFFE62583E352A29DFD1F" author="KANAAR P." box="[151,510,631,650]" page="145" pageId="2" pageNumber="625" year="1990">
<taxonomicName id="4C6D0288FFBAFFE62583E352A29DFD1F" ID-CoL="9TNHL" authority="Kanaar, 1990: 145" authorityName="Kanaar" authorityPageNumber="145" authorityYear="1990" box="[151,510,631,650]" class="Insecta" family="Histeridae" genus="Styphrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="625" phylum="Arthropoda" rank="species" species="termitophilus">
<emphasis id="B919A519FFBAFFE62583E352A23AFD1F" box="[151,345,632,650]" italics="true" pageId="2" pageNumber="625">Styphrus termitophilus</emphasis>
<bibRefCitation id="EFFC04FAFFBAFFE6244AE35DA29DFD1F" author="KANAAR P." box="[350,510,631,650]" pageId="2" pageNumber="625" pagination="145 - 147" refId="ref4229" refString="KANAAR P. 1990: L'Entomofaune des termitieres mortes de Macrotermes. Un Styphrus nouveau de l'Afrique Occidentale (Coleoptera, Histeridae). Revue Francaise d'Entomologie (N. S.) 12 (3): 145 - 147." type="journal article" year="1990">Kanaar, 1990: 145</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. Synonymized by
<treatmentCitation id="0ACC5F1AFFBAFFE62788E35DA02EFD1F" author="KANAAR P." box="[668,845,631,650]" page="130" pageId="2" pageNumber="625" year="1996">
<bibRefCitation id="EFFC04FAFFBAFFE62788E35DA02EFD1F" author="KANAAR P." box="[668,845,631,650]" pageId="2" pageNumber="625" pagination="130" refId="ref4271" refString="KANAAR P. 1996: Styphrus termitophilus Kanaar, 1990 = Saprinus verschureni Therond, 1959. nov. syn., type d'une nouveau subgenus Pilisaprinus dans le genre Saprinus Er. Nouvelle Revue d'Entomologie (N. S.) 13 (2): 130." type="journal article" year="1996">KANAAR (1996: 130)</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation id="0ACC5F1AFFBAFFE62583E3B8A118FD30" author="MAZUR S." box="[151,635,658,677]" page=")" pageId="2" pageNumber="625" year="2011">
<taxonomicName id="4C6D0288FFBAFFE62583E3B8A118FD30" ID-CoL="79KMQ" authority=": MAZUR (2011: 179)" authorityName=": MAZUR" authorityPageNumber="179" authorityYear="2011" box="[151,635,658,677]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="625" phylum="Arthropoda" rank="species" species="verschureni" subGenus="Pilisaprinus">
<emphasis id="B919A519FFBAFFE62583E3B8A380FD31" box="[151,227,658,676]" italics="true" pageId="2" pageNumber="625">Saprinus</emphasis>
(
<emphasis id="B919A519FFBAFFE625E4E3B8A23BFD31" box="[240,344,658,676]" italics="true" pageId="2" pageNumber="625">Pilisaprinus</emphasis>
)
<emphasis id="B919A519FFBAFFE62470E3B8A2ABFD31" box="[356,456,658,676]" italics="true" pageId="2" pageNumber="625">verschureni</emphasis>
:
<bibRefCitation id="EFFC04FAFFBAFFE624C7E3B8A118FD30" author="MAZUR S." box="[467,635,658,677]" pageId="2" pageNumber="625" refId="ref4668" refString="MAZUR S. 2011: A concise catalogue of the Histeridae (Coleoptera). Warsaw University of Life Sciences, SGGW Press, Warszawa, 332 pp." type="book" year="2011">MAZUR (2011: 179)</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3772A80FFBAFFE62583E391A035FC4C" pageId="2" pageNumber="625" type="materials_examined">
<paragraph id="8BD2790BFFBAFFE62583E391A1C4FD5B" blockId="2.[151,1127,698,985]" box="[151,679,698,718]" pageId="2" pageNumber="625">
<emphasis id="B919A519FFBAFFE62583E391A273FD5B" bold="true" box="[151,272,698,718]" pageId="2" pageNumber="625">
<typeStatus id="54D6C7A9FFBAFFE62583E391A3A7FD5B" box="[151,196,699,718]" pageId="2" pageNumber="625">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F37A399BFFBAFFE62401E391A143FD5B" box="[277,544,699,718]" name="Democratic Republic of the Congo" pageId="2" pageNumber="625">Democratic Republic of Congo</collectingCountry>
(former
<collectingCountry id="F37A399BFFBAFFE62778E391A1FEFD5B" box="[620,669,699,718]" name="Democratic Republic of the Congo" pageId="2" pageNumber="625">Zairë</collectingCountry>
).
</paragraph>
<paragraph id="8BD2790BFFBAFFE62583E3FCA2FFFCFB" blockId="2.[151,1127,698,985]" pageId="2" pageNumber="625">
<emphasis id="B919A519FFBAFFE62583E3FCA2E3FD7C" bold="true" box="[151,384,725,745]" pageId="2" pageNumber="625">Type specimen examined.</emphasis>
<taxonomicName id="4C6D0288FFBAFFE62491E3FDA159FD7C" authorityName=": MAZUR" authorityYear="2011" box="[389,570,727,745]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="625" phylum="Arthropoda" rank="species" species="verschureni">
<emphasis id="B919A519FFBAFFE62491E3FDA159FD7C" box="[389,570,727,745]" italics="true" pageId="2" pageNumber="625">Saprinus verschureni</emphasis>
</taxonomicName>
.
<materialsCitation id="3B057356FFBAFFE62751E3FCA2FFFCFB" collectingDate="1952-05-29" collectionCode="RMCA" collectorName="Miss. H. De Saeger &amp; J. Verschuren." country="Democratic Republic of the Congo" determinerName="J. Therond" location="Congo Belge" pageId="2" pageNumber="625" specimenCount="2" typeStatus="holotype">
<typeStatus id="54D6C7A9FFBAFFE62751E3FCA1FBFD7D" box="[581,664,726,745]" pageId="2" pageNumber="625" type="holotype">HOLOTYPE</typeStatus>
: “
<collectingCountry id="F37A399BFFBAFFE627B8E3FCA07CFD7C" box="[684,799,726,745]" name="Democratic Republic of the Congo" pageId="2" pageNumber="625">Congo Belge</collectingCountry>
, P.N.G. /
<collectorName id="26981CDDFFBAFFE62664E3FCA775FD7C" box="[880,1046,726,745]" pageId="2" pageNumber="625">Miss. H. De Saeger</collectorName>
/ II/gd/4,
<date id="FFD35FCBFFBAFFE62583E3DAA393FC96" box="[151,240,752,771]" pageId="2" pageNumber="625" value="1952-05-29">
<collectingDate id="EF97A623FFBAFFE62583E3DAA393FC96" box="[151,240,752,771]" pageId="2" pageNumber="625" value="1952-05-29">29-v-1952</collectingDate>
</date>
/
<collectorName id="26981CDDFFBAFFE62410E3DBA21CFC91" box="[260,383,753,772]" pageId="2" pageNumber="625">J. Verschuren.</collectorName>
3545” (printed); “
<typeStatus id="54D6C7A9FFBAFFE62734E3DBA1C2FC91" box="[544,673,753,772]" pageId="2" pageNumber="625" type="holotype">HOLOTYPUS</typeStatus>
” (orange, red-margined label, printed); “
<taxonomicName id="4C6D0288FFBAFFE6211AE3D8A739FC91" authorityName="Erichson" authorityYear="1834" box="[1038,1114,754,772]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="625" phylum="Arthropoda" rank="genus">
<emphasis id="B919A519FFBAFFE6211AE3D8A739FC91" box="[1038,1114,754,772]" italics="true" pageId="2" pageNumber="625">Saprinus</emphasis>
</taxonomicName>
/
<taxonomicName id="4C6D0288FFBAFFE62583E226A398FC8B" authority="Lackner, 2013" authorityName="Lackner" authorityYear="2013" box="[151,251,780,798]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="625" phylum="Arthropoda" rank="species" species="verschureni" status="sp. nov.">
<emphasis id="B919A519FFBAFFE62583E226A398FC8B" box="[151,251,780,798]" italics="true" pageId="2" pageNumber="625">verschureni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22A1862FFBAFFE62414E226A221FC8A" box="[256,322,780,799]" pageId="2" pageNumber="625" rank="species">nov. sp.</taxonomicNameLabel>
/
<determinerName id="7E0E95CFFFBAFFE62445E226A2C8FC8B" box="[337,427,779,799]" pageId="2" pageNumber="625">J. Thérond</determinerName>
det., 1958” (printed-written); “TYPE” (purple label, printed) (
<collectionCode id="ED7CE1CEFFBAFFE626A9E221A762FC8B" box="[957,1025,779,798]" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="2" pageNumber="625" type="Museum">RMCA</collectionCode>
). The specimen is of unidentified sex: genitalia lost, sixth abdominal tergite broken in two, left antenna missing, both protibiae missing, right mesotibia broken off glued to the same triangular point with specimen, both mesotarsi missing sidemounted on a triangular point.
</materialsCitation>
</paragraph>
<paragraph id="8BD2790BFFBAFFE62583E25FA035FC4C" blockId="2.[151,1127,698,985]" pageId="2" pageNumber="625">
<emphasis id="B919A519FFBAFFE62583E25FA2CFFC1C" bold="true" box="[151,428,885,905]" pageId="2" pageNumber="625">Additional material examined.</emphasis>
<materialsCitation id="3B057356FFBAFFE624A5E25CA25AFC31" collectingDate="2002-02" collectionCode="TLAN" collectorName="G. Goergen" country="Benin" location="Niaouli" pageId="2" pageNumber="625" specimenCount="1">
<emphasis id="B919A519FFBAFFE624A5E25CA29FFC1C" bold="true" box="[433,508,886,905]" pageId="2" pageNumber="625">
<collectingCountry id="F37A399BFFBAFFE624A5E25CA29BFC1C" box="[433,504,886,905]" name="Benin" pageId="2" pageNumber="625">BENIN</collectingCountry>
:
</emphasis>
1 J, “BENIN –
<location id="8EB22FD0FFBAFFE6279CE25CA1ABFC1C" LSID="urn:lsid:plazi:treatment:03C4C81DFFBAFFE3244CE331A0B6FE66:8EB22FD0FFBAFFE6279CE25CA1ABFC1C" box="[648,712,886,905]" country="Benin" name="Niaouli" pageId="2" pageNumber="625">Niaouli</location>
II /
<date id="FFD35FCBFFBAFFE627FFE25CA004FC1C" box="[747,871,886,905]" pageId="2" pageNumber="625" value="2002-02">
<collectingDate id="EF97A623FFBAFFE627FFE25CA004FC1C" box="[747,871,886,905]" pageId="2" pageNumber="625" value="2002-02">February 2002</collectingDate>
</date>
/ in
<taxonomicName id="4C6D0288FFBAFFE62698E25DA09EFC1C" box="[908,1021,887,905]" class="Insecta" family="Termitidae" genus="Macrotermes" kingdom="Animalia" order="Blattodea" pageId="2" pageNumber="625" phylum="Arthropoda" rank="genus">
<emphasis id="B919A519FFBAFFE62698E25DA09EFC1C" box="[908,1021,887,905]" italics="true" pageId="2" pageNumber="625">Macrotermes</emphasis>
</taxonomicName>
hill / leg.
<collectorName id="26981CDDFFBAFFE62146E25CA382FC31" pageId="2" pageNumber="625">G. Goergen</collectorName>
” (
<collectionCode id="ED7CE1CEFFBAFFE625E1E2BBA250FC31" box="[245,307,913,932]" pageId="2" pageNumber="625">TLAN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B057356FFBAFFE62456E2BBA108FC31" box="[322,619,907,933]" collectionCode="RMNH" country="Benin" pageId="2" pageNumber="625" specimenCount="1" specimenCount-female="1">
1 J
<specimenCount id="9D6BB282FFBAFFE62473E2BBA2E4FC31" box="[359,391,907,932]" pageId="2" pageNumber="625" type="female">1 ♀</specimenCount>
, same label data (
<collectionCode id="ED7CE1CEFFBAFFE62734E2BBA106FC31" box="[544,613,913,932]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="2" pageNumber="625" type="Museum">RMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B057356FFBAFFE62767E2BBA21CFC2B" collectingDate="2002-01" collectionCode="RMNH" collectorName="G. Goergen" country="Benin" location="Benin" municipality="Toffo" pageId="2" pageNumber="625" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D6BB282FFBAFFE62767E2BBA1F7FC31" box="[627,660,907,932]" pageId="2" pageNumber="625" type="female">1 ♀</specimenCount>
, “
<location id="8EB22FD0FFBAFFE627B8E2BBA183FC31" LSID="urn:lsid:plazi:treatment:03C4C81DFFBAFFE3244CE331A0B6FE66:8EB22FD0FFBAFFE627B8E2BBA183FC31" box="[684,736,913,932]" country="Benin" municipality="Toffo" name="Benin" pageId="2" pageNumber="625">Benin</location>
/
<collectingMunicipality id="6BB6E371FFBAFFE627E4E2BBA07CFC31" box="[752,799,913,932]" pageId="2" pageNumber="625">Toffo</collectingMunicipality>
/
<date id="FFD35FCBFFBAFFE6263BE2BBA0C1FC31" box="[815,930,913,932]" pageId="2" pageNumber="625" value="2002-01">
<collectingDate id="EF97A623FFBAFFE6263BE2BBA0C1FC31" box="[815,930,913,932]" pageId="2" pageNumber="625" value="2002-01">January 2002</collectingDate>
</date>
/ in
<taxonomicName id="4C6D0288FFBAFFE626DCE2B8A75AFC31" box="[968,1081,914,932]" class="Insecta" family="Termitidae" genus="Macrotermes" kingdom="Animalia" order="Blattodea" pageId="2" pageNumber="625" phylum="Arthropoda" rank="genus">
<emphasis id="B919A519FFBAFFE626DCE2B8A75AFC31" box="[968,1081,914,932]" italics="true" pageId="2" pageNumber="625">Macrotermes</emphasis>
</taxonomicName>
hill / leg.
<collectorName id="26981CDDFFBAFFE625A8E281A243FC2B" box="[188,288,939,958]" pageId="2" pageNumber="625">G. Goergen</collectorName>
” (
<collectionCode id="ED7CE1CEFFBAFFE62420E281A21AFC2B" box="[308,377,939,958]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="2" pageNumber="625" type="Museum">RMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B057356FFBAFFE62493E280A031FC4C" collectingDate="1994-04-15" collectionCode="RMNH" collectorName="Taabo" country="Ivory Coast" location="Lamto" municipality="Termitiere" pageId="2" pageNumber="625" specimenCount="1">
<emphasis id="B919A519FFBAFFE62493E280A142FC2B" bold="true" box="[391,545,938,958]" pageId="2" pageNumber="625">
<collectingCountry id="F37A399BFFBAFFE62493E280A17EFC2B" box="[391,541,938,958]" name="Ivory Coast" pageId="2" pageNumber="625">IVORY COAST</collectingCountry>
:
</emphasis>
1 J, “
<location id="8EB22FD0FFBAFFE6274DE281A1F1FC2B" LSID="urn:lsid:plazi:treatment:03C4C81DFFBAFFE3244CE331A0B6FE66:8EB22FD0FFBAFFE6274DE281A1F1FC2B" box="[601,658,939,958]" country="Ivory Coast" municipality="Termitiere" name="Lamto" pageId="2" pageNumber="625">Lamto</location>
/
<date id="FFD35FCBFFBAFFE627B7E281A19FFC2B" box="[675,764,939,958]" pageId="2" pageNumber="625" value="1994-04-15">
<collectingDate id="EF97A623FFBAFFE627B7E281A19FFC2B" box="[675,764,939,958]" pageId="2" pageNumber="625" value="1994-04-15">15.iv.1994</collectingDate>
</date>
/
<collectingMunicipality id="6BB6E371FFBAFFE62619E281A005FC2B" box="[781,870,939,958]" pageId="2" pageNumber="625">Termitière</collectingMunicipality>
morte en [= dead termitarium in] / forêt dégradée [= degraded forest]
<quantity id="4C95D4EEFFBAFFE624FCE2ECA170FC4C" box="[488,531,966,985]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="625" unit="km" value="2.0">2 km</quantity>
de /
<collectorName id="26981CDDFFBAFFE62729E2ECA112FC4C" box="[573,625,966,985]" pageId="2" pageNumber="625">Taabo</collectorName>
leg. Cl. Girard” (
<collectionCode id="ED7CE1CEFFBAFFE62613E2ECA02FFC4C" box="[775,844,966,985]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="2" pageNumber="625" type="Museum">RMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3772A80FFBAFFE32583E2DBA0B6FE66" lastPageId="7" lastPageNumber="630" pageId="2" pageNumber="625" type="description">
<paragraph id="8BD2790BFFBAFFE12583E2DBA1B3FA66" blockId="2.[151,1127,1009,1577]" lastBlockId="5.[113,1089,1275,1587]" lastPageId="5" lastPageNumber="628" pageId="2" pageNumber="625">
<emphasis id="B919A519FFBAFFE62583E2DBA25EFB9C" bold="true" box="[151,317,1009,1033]" pageId="2" pageNumber="625">Redescription.</emphasis>
Body moderately convex, PEL:
<quantity id="4C95D4EEFFBAFFE62781E2DBA050FB9C" box="[661,819,1009,1033]" metricMagnitude="-3" metricUnit="m" metricValue="4.625" metricValueMax="4.75" metricValueMin="4.5" pageId="2" pageNumber="625" unit="mm" value="4.625" valueMax="4.75" valueMin="4.5">4.50–4.75 mm</quantity>
; APW: 1.85–2.00 mm; PPW: 3.75–4.00 mm; EL: 2.75–3.00 mm; EW: 4.00–
<quantity id="4C95D4EEFFBAFFE6279CE53BA18BFBBC" box="[648,744,1041,1065]" metricMagnitude="-3" metricUnit="m" metricValue="4.25" pageId="2" pageNumber="625" unit="mm" value="4.25">4.25 mm</quantity>
, dorsally with various kinds of punctuation, cuticle black, not metallic; legs, mouthparts and antennae rufopiceous. Antennal scape (
<figureCitation id="1356658EFFBAFFE625F6E57BA247FBFC" box="[226,292,1105,1129]" captionStart="Figs 1–9" captionStartId="3.[113,150,1513,1532]" captionTargetBox="[211,991,219,1492]" captionTargetId="figure-12@3.[211,991,219,1492]" captionTargetPageId="3" captionText="Figs 1–9. External morphology of Pilisaprinus verschureni (Thérond, 1959). 1 – habitus, dorsal view; 2 – same, ventral view; 3 – antennal club, latero-ventral view; 4 – head, dorsal view; 5 – mentum, ventral view; 6 – prosternum; 7 – propygidium and pygidium; 8 – lateral disc of metaventrite + metepisternum; 9 – protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5740226" httpUri="https://zenodo.org/record/5740226/files/figure.png" pageId="2" pageNumber="625">Fig. 4</figureCitation>
) slightly dilated and thickened, densely punctate, with numerous long amber setae; antennal club (
<figureCitation id="1356658EFFBAFFE62462E55BA2D8FB1C" box="[374,443,1137,1161]" captionStart="Figs 1–9" captionStartId="3.[113,150,1513,1532]" captionTargetBox="[211,991,219,1492]" captionTargetId="figure-12@3.[211,991,219,1492]" captionTargetPageId="3" captionText="Figs 1–9. External morphology of Pilisaprinus verschureni (Thérond, 1959). 1 – habitus, dorsal view; 2 – same, ventral view; 3 – antennal club, latero-ventral view; 4 – head, dorsal view; 5 – mentum, ventral view; 6 – prosternum; 7 – propygidium and pygidium; 8 – lateral disc of metaventrite + metepisternum; 9 – protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5740226" httpUri="https://zenodo.org/record/5740226/files/figure.png" pageId="2" pageNumber="625">Figs 3</figureCitation>
,
<figureCitation id="1356658EFFBAFFE624D2E55BA282FB1C" box="[454,481,1137,1161]" captionStart="Figs 10–12" captionStartId="4.[151,187,1104,1123]" captionTargetBox="[247,1028,220,1075]" captionTargetId="figure-153@4.[246,1029,219,1075]" captionTargetPageId="4" captionText="Figs 10–12. Mouthparts and sensory structures of the antenna of Pilisaprinus verschureni (Thérond, 1959).10 – mandibles, dorsal view; 11 – labrum, left half showing dorsal view; right half showing ventral view; 12 – antennal club showing sensory structures of the antenna." figureDoi="http://doi.org/10.5281/zenodo.5740230" httpUri="https://zenodo.org/record/5740230/files/figure.png" pageId="2" pageNumber="625">12</figureCitation>
) ventrally with two slit-like pits almost encircling it entirely, dorsally with two complementary slit-like pits; lower half (approximately) glabrous or fringed only with few microscopic setae, upper half (approximately) with dense short setae; sensory structures of antennal club (
<figureCitation id="1356658EFFBAFFE624D3E5FBA176FB7C" box="[455,533,1233,1257]" captionStart="Figs 10–12" captionStartId="4.[151,187,1104,1123]" captionTargetBox="[247,1028,220,1075]" captionTargetId="figure-153@4.[246,1029,219,1075]" captionTargetPageId="4" captionText="Figs 10–12. Mouthparts and sensory structures of the antenna of Pilisaprinus verschureni (Thérond, 1959).10 – mandibles, dorsal view; 11 – labrum, left half showing dorsal view; right half showing ventral view; 12 – antennal club showing sensory structures of the antenna." figureDoi="http://doi.org/10.5281/zenodo.5740230" httpUri="https://zenodo.org/record/5740230/files/figure.png" pageId="2" pageNumber="625">Fig. 12</figureCitation>
) in form of six vesicles, four of them situated in pairs under ventral slit-like pits and another two of them situated under each-other under dorsal surface medially. Mandibles (
<figureCitation id="1356658EFFBAFFE624CDE43BA143FABC" box="[473,544,1297,1321]" captionStart="Figs 1–9" captionStartId="3.[113,150,1513,1532]" captionTargetBox="[211,991,219,1492]" captionTargetId="figure-12@3.[211,991,219,1492]" captionTargetPageId="3" captionText="Figs 1–9. External morphology of Pilisaprinus verschureni (Thérond, 1959). 1 – habitus, dorsal view; 2 – same, ventral view; 3 – antennal club, latero-ventral view; 4 – head, dorsal view; 5 – mentum, ventral view; 6 – prosternum; 7 – propygidium and pygidium; 8 – lateral disc of metaventrite + metepisternum; 9 – protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5740226" httpUri="https://zenodo.org/record/5740226/files/figure.png" pageId="2" pageNumber="625">Figs 4</figureCitation>
,
<figureCitation id="1356658EFFBAFFE62739E43BA12BFABC" box="[557,584,1297,1321]" captionStart="Figs 10–12" captionStartId="4.[151,187,1104,1123]" captionTargetBox="[247,1028,220,1075]" captionTargetId="figure-153@4.[246,1029,219,1075]" captionTargetPageId="4" captionText="Figs 10–12. Mouthparts and sensory structures of the antenna of Pilisaprinus verschureni (Thérond, 1959).10 – mandibles, dorsal view; 11 – labrum, left half showing dorsal view; right half showing ventral view; 12 – antennal club showing sensory structures of the antenna." figureDoi="http://doi.org/10.5281/zenodo.5740230" httpUri="https://zenodo.org/record/5740230/files/figure.png" pageId="2" pageNumber="625">10</figureCitation>
) stout, moderately punctate, with evenly rounded outer margin strongly curved inwardly; acutely pointed; sub-apical tooth on inner margin of left mandible moderately large, almost perpendicular; labrum (
<figureCitation id="1356658EFFBAFFE62649E47BA0C9FAFC" box="[861,938,1361,1385]" captionStart="Figs 10–12" captionStartId="4.[151,187,1104,1123]" captionTargetBox="[247,1028,220,1075]" captionTargetId="figure-153@4.[246,1029,219,1075]" captionTargetPageId="4" captionText="Figs 10–12. Mouthparts and sensory structures of the antenna of Pilisaprinus verschureni (Thérond, 1959).10 – mandibles, dorsal view; 11 – labrum, left half showing dorsal view; right half showing ventral view; 12 – antennal club showing sensory structures of the antenna." figureDoi="http://doi.org/10.5281/zenodo.5740230" httpUri="https://zenodo.org/record/5740230/files/figure.png" pageId="2" pageNumber="625">Fig. 11</figureCitation>
) slightly convex, slightly depressed medially, finely punctate; labral pits rather deep and large with two short amber setae arising from each; terminal labial palpomere elongate, its width about one-fourth its length; mentum (
<figureCitation id="1356658EFFBAFFE6247EE49BA2CAFA5C" box="[362,425,1457,1481]" captionStart="Figs 1–9" captionStartId="3.[113,150,1513,1532]" captionTargetBox="[211,991,219,1492]" captionTargetId="figure-12@3.[211,991,219,1492]" captionTargetPageId="3" captionText="Figs 1–9. External morphology of Pilisaprinus verschureni (Thérond, 1959). 1 – habitus, dorsal view; 2 – same, ventral view; 3 – antennal club, latero-ventral view; 4 – head, dorsal view; 5 – mentum, ventral view; 6 – prosternum; 7 – propygidium and pygidium; 8 – lateral disc of metaventrite + metepisternum; 9 – protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5740226" httpUri="https://zenodo.org/record/5740226/files/figure.png" pageId="2" pageNumber="625">Fig. 5</figureCitation>
) square-shaped, anterior angles slightly produced, anterior margin with a shallow notch; disc of mentum covered with numerous short setae that are almost absent along posterior margin; cardo of maxilla laterally with numerous short setae; stipes triangular, with four short setae; terminal maxillary palpomere elongate, its width about one-fifth its length, about three times as long as penultimate; labial palpomeres with dense brush of long setae. Clypeus and frons (
<figureCitation id="1356658EFFBCFFE024BCE5D3A284FA87" box="[424,487,1273,1298]" captionStart="Figs 1–9" captionStartId="3.[113,150,1513,1532]" captionTargetBox="[211,991,219,1492]" captionTargetId="figure-12@3.[211,991,219,1492]" captionTargetPageId="3" captionText="Figs 1–9. External morphology of Pilisaprinus verschureni (Thérond, 1959). 1 – habitus, dorsal view; 2 – same, ventral view; 3 – antennal club, latero-ventral view; 4 – head, dorsal view; 5 – mentum, ventral view; 6 – prosternum; 7 – propygidium and pygidium; 8 – lateral disc of metaventrite + metepisternum; 9 – protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5740226" httpUri="https://zenodo.org/record/5740226/files/figure.png" pageId="4" pageNumber="627">Fig. 4</figureCitation>
) rectangular, broad, with moderate punctures and fine transverse wrinkles; frontal and supraorbital striae absent; eyes slightly flattened, well visible from above. Pronotal sides (
<figureCitation id="1356658EFFBCFFE0249EE413A2ABFAC7" box="[394,456,1337,1362]" captionStart="Figs 1–9" captionStartId="3.[113,150,1513,1532]" captionTargetBox="[211,991,219,1492]" captionTargetId="figure-12@3.[211,991,219,1492]" captionTargetPageId="3" captionText="Figs 1–9. External morphology of Pilisaprinus verschureni (Thérond, 1959). 1 – habitus, dorsal view; 2 – same, ventral view; 3 – antennal club, latero-ventral view; 4 – head, dorsal view; 5 – mentum, ventral view; 6 – prosternum; 7 – propygidium and pygidium; 8 – lateral disc of metaventrite + metepisternum; 9 – protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5740226" httpUri="https://zenodo.org/record/5740226/files/figure.png" pageId="4" pageNumber="627">Fig. 1</figureCitation>
) strongly narrowing anteriorly; anterior emargination for head deep; apical angles acute and conspicuous, marginal pronotal stria slightly carinate, complete, slightly weakened behind head; disc entirely covered with coarse round punctuation, punctures separated by about their diameter, laterally coarser than medially; pronotal depressions rather deep and large; scutellum very small; pronotal hypomeron with dense long amber setae (
<figureCitation id="1356658EFFBCFFE0212FE493A3C4FA67" captionStart="Figs 1–9" captionStartId="3.[113,150,1513,1532]" captionTargetBox="[211,991,219,1492]" captionTargetId="figure-12@3.[211,991,219,1492]" captionTargetPageId="3" captionText="Figs 1–9. External morphology of Pilisaprinus verschureni (Thérond, 1959). 1 – habitus, dorsal view; 2 – same, ventral view; 3 – antennal club, latero-ventral view; 4 – head, dorsal view; 5 – mentum, ventral view; 6 – prosternum; 7 – propygidium and pygidium; 8 – lateral disc of metaventrite + metepisternum; 9 – protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5740226" httpUri="https://zenodo.org/record/5740226/files/figure.png" pageId="4" pageNumber="627">Fig. 2</figureCitation>
). Elytral epipleuron (
<figureCitation id="1356658EFFBCFFE02492E4F3A2A0FA67" box="[390,451,1497,1522]" captionStart="Figs 1–9" captionStartId="3.[113,150,1513,1532]" captionTargetBox="[211,991,219,1492]" captionTargetId="figure-12@3.[211,991,219,1492]" captionTargetPageId="3" captionText="Figs 1–9. External morphology of Pilisaprinus verschureni (Thérond, 1959). 1 – habitus, dorsal view; 2 – same, ventral view; 3 – antennal club, latero-ventral view; 4 – head, dorsal view; 5 – mentum, ventral view; 6 – prosternum; 7 – propygidium and pygidium; 8 – lateral disc of metaventrite + metepisternum; 9 – protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5740226" httpUri="https://zenodo.org/record/5740226/files/figure.png" pageId="4" pageNumber="627">Fig. 8</figureCitation>
) with long amber setae; marginal epipleural stria thin, complete, continued as complete apical stria; marginal elytral stria well impressed, complete, slightly carinate, apically attaining marginal epipleural stria. Humeral elytral stria deeply impressed on basal third, joined with inner subhumeral stria forming a supplementary dorsal elytral stria parallel to first dorsal elytral, reaching about half of elytral length apically; elytral disc with four deeply impressed dorsal elytral striae 1–4, in shallow punctures, all about the same length, reaching about half of elytral length apically, fourth dorsal elytral stria basally connected with sutural elytral stria; sutural elytral stria well impressed, reaching about two-thirds of elytral length apically, otherwise obliterated. Elytral disc entirely with fine small dense punctuation (with the exception of fourth elytral interval and elytral flanks) intermingled with dense and coarse punctures separated by about half their diameters.
</paragraph>
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<paragraph id="8BD2790BFFBBFFE72565E4C3A763F9A4" blockId="3.[113,1090,1513,1586]" pageId="3" pageNumber="626">
Figs 1–9. External morphology of
<taxonomicName id="4C6D0288FFBBFFE7248AE4C0A064FA69" authority="(Therond, 1959)" authorityName="Lackner" authorityYear="2013" baseAuthorityName="Therond" baseAuthorityYear="1959" box="[414,775,1513,1532]" class="Insecta" family="Histeridae" genus="Pilisaprinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="626" phylum="Arthropoda" rank="species" species="verschureni">
<emphasis id="B919A519FFBBFFE7248AE4C0A112FA69" box="[414,625,1514,1532]" italics="true" pageId="3" pageNumber="626">Pilisaprinus verschureni</emphasis>
(
<bibRefCitation id="EFFC04FAFFBBFFE72769E4C3A062FA69" author="THEROND J." box="[637,769,1513,1532]" pageId="3" pageNumber="626" pagination="1 - 39" refId="ref4728" refString="THEROND J. 1959: Histeridae (Coleoptera, Staphylinoidea). Exploration de la Parc Naturel du Garamba, Mission de Saeger 15 (1): 1 - 39." type="journal article" year="1959">Thérond, 1959</bibRefCitation>
)
</taxonomicName>
. 1 – habitus, dorsal view; 2 – same, ventral view; 3 – antennal club, latero-ventral view; 4 – head, dorsal view; 5 – mentum, ventral view; 6 – prosternum; 7 – propygidium and pygidium; 8 – lateral disc of metaventrite + metepisternum; 9 – protibia, dorsal view.
</paragraph>
</caption>
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<paragraph id="8BD2790BFFBCFFE02583E57AA160FB0C" blockId="4.[151,1127,1104,1177]" pageId="4" pageNumber="627">
Figs 10–12. Mouthparts and sensory structures of the antenna of
<taxonomicName id="4C6D0288FFBCFFE027B8E57BA764FBF6" authority="(Therond, 1959)" authorityName="Lackner" authorityYear="2013" baseAuthorityName="Therond" baseAuthorityYear="1959" box="[684,1031,1104,1123]" class="Insecta" family="Histeridae" genus="Pilisaprinus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="627" phylum="Arthropoda" rank="species" species="verschureni">
<emphasis id="B919A519FFBCFFE027B8E57BA01AFBF6" box="[684,889,1105,1123]" italics="true" pageId="4" pageNumber="627">Pilisaprinus verschureni</emphasis>
(
<bibRefCitation id="EFFC04FAFFBCFFE02696E57AA762FBF6" author="THEROND J." box="[898,1025,1104,1123]" pageId="4" pageNumber="627" pagination="1 - 39" refId="ref4728" refString="THEROND J. 1959: Histeridae (Coleoptera, Staphylinoidea). Exploration de la Parc Naturel du Garamba, Mission de Saeger 15 (1): 1 - 39." type="journal article" year="1959">Thérond, 1959</bibRefCitation>
)
</taxonomicName>
.10 – mandibles, dorsal view; 11 – labrum, left half showing dorsal view; right half showing ventral view; 12 – antennal club showing sensory structures of the antenna.
</paragraph>
</caption>
<caption id="DF122983FFBDFFE12565E548A3B8FB50" ID-DOI="http://doi.org/10.5281/zenodo.5740232" ID-Zenodo-Dep="5740232" httpUri="https://zenodo.org/record/5740232/files/figure.png" pageId="5" pageNumber="628" startId="5.[113,150,1122,1141]" targetBox="[252,944,222,1095]" targetPageId="5">
<paragraph id="8BD2790BFFBDFFE12565E548A3B8FB50" blockId="5.[113,1094,1122,1221]" pageId="5" pageNumber="628">
Figs 13–19. Male terminalia of
<taxonomicName id="4C6D0288FFBDFFE12497E549A189FBE0" authority="(Therond, 1959)" authorityName="Lackner" authorityYear="2013" baseAuthorityName="Therond" baseAuthorityYear="1959" box="[387,746,1122,1141]" class="Insecta" family="Histeridae" genus="Pilisaprinus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="628" phylum="Arthropoda" rank="species" species="verschureni">
<emphasis id="B919A519FFBDFFE12497E549A136FBE0" box="[387,597,1123,1141]" italics="true" pageId="5" pageNumber="628">Pilisaprinus verschureni</emphasis>
(
<bibRefCitation id="EFFC04FAFFBDFFE12775E548A187FBE0" author="THEROND J." box="[609,740,1122,1141]" pageId="5" pageNumber="628" pagination="1 - 39" refId="ref4728" refString="THEROND J. 1959: Histeridae (Coleoptera, Staphylinoidea). Exploration de la Parc Naturel du Garamba, Mission de Saeger 15 (1): 1 - 39." type="journal article" year="1959">Thérond, 1959</bibRefCitation>
)
</taxonomicName>
. 13 – eighth sternite and tergite, ventral view; 14 – same, ventral view; 15 – same, lateral view; 16 – ninth and tenth tergites, dorsal view; spiculum gastrale, ventral view; 17 – ninth, tenth tergites and spiculum gastrale, lateral view; 18 – aedeagus, dorsal view; 19 – same, lateral view.
</paragraph>
</caption>
<paragraph id="8BD2790BFFBDFFE22581E4D1A770FE26" blockId="5.[113,1089,1275,1587]" lastBlockId="6.[151,1127,219,1587]" lastPageId="6" lastPageNumber="629" pageId="5" pageNumber="628">
Propygidium (
<figureCitation id="1356658EFFBDFFE12439E4D1A20FF986" box="[301,364,1531,1555]" captionStart="Figs 1–9" captionStartId="3.[113,150,1513,1532]" captionTargetBox="[211,991,219,1492]" captionTargetId="figure-12@3.[211,991,219,1492]" captionTargetPageId="3" captionText="Figs 1–9. External morphology of Pilisaprinus verschureni (Thérond, 1959). 1 – habitus, dorsal view; 2 – same, ventral view; 3 – antennal club, latero-ventral view; 4 – head, dorsal view; 5 – mentum, ventral view; 6 – prosternum; 7 – propygidium and pygidium; 8 – lateral disc of metaventrite + metepisternum; 9 – protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5740226" httpUri="https://zenodo.org/record/5740226/files/figure.png" pageId="5" pageNumber="628">Fig. 7</figureCitation>
) partly covered by elytra; with dense and coarse punctures becoming sparser and finer medially; pygidium (
<figureCitation id="1356658EFFBDFFE12712E731A125F9A6" box="[518,582,1563,1587]" captionStart="Figs 1–9" captionStartId="3.[113,150,1513,1532]" captionTargetBox="[211,991,219,1492]" captionTargetId="figure-12@3.[211,991,219,1492]" captionTargetPageId="3" captionText="Figs 1–9. External morphology of Pilisaprinus verschureni (Thérond, 1959). 1 – habitus, dorsal view; 2 – same, ventral view; 3 – antennal club, latero-ventral view; 4 – head, dorsal view; 5 – mentum, ventral view; 6 – prosternum; 7 – propygidium and pygidium; 8 – lateral disc of metaventrite + metepisternum; 9 – protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5740226" httpUri="https://zenodo.org/record/5740226/files/figure.png" pageId="5" pageNumber="628">Fig. 7</figureCitation>
) long, convex, with very coarse and dense punctuation, punctures separated by less than half of their diameter. Anterior margin of median portion of prosternum (
<figureCitation id="1356658EFFBEFFE22498E1D1A2A8FE86" box="[396,459,251,275]" captionStart="Figs 1–9" captionStartId="3.[113,150,1513,1532]" captionTargetBox="[211,991,219,1492]" captionTargetId="figure-12@3.[211,991,219,1492]" captionTargetPageId="3" captionText="Figs 1–9. External morphology of Pilisaprinus verschureni (Thérond, 1959). 1 – habitus, dorsal view; 2 – same, ventral view; 3 – antennal club, latero-ventral view; 4 – head, dorsal view; 5 – mentum, ventral view; 6 – prosternum; 7 – propygidium and pygidium; 8 – lateral disc of metaventrite + metepisternum; 9 – protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5740226" httpUri="https://zenodo.org/record/5740226/files/figure.png" pageId="6" pageNumber="629">Fig. 6</figureCitation>
) rounded; marginal prosternal stria very thin; prosternal foveae absent; prosternal process compressed, dorso-laterally with irregular coarse to fine punctures with long amber setae; carinal prosternal striae present as tiny fragments on prosternal apophysis, otherwise absent (in some specimens reaching approximately half the length of the prosternum apically, weakened and very approximate); lateral prosternal striae (
<figureCitation id="1356658EFFBEFFE226FEE051A749FE06" box="[1002,1066,379,403]" captionStart="Figs 1–9" captionStartId="3.[113,150,1513,1532]" captionTargetBox="[211,991,219,1492]" captionTargetId="figure-12@3.[211,991,219,1492]" captionTargetPageId="3" captionText="Figs 1–9. External morphology of Pilisaprinus verschureni (Thérond, 1959). 1 – habitus, dorsal view; 2 – same, ventral view; 3 – antennal club, latero-ventral view; 4 – head, dorsal view; 5 – mentum, ventral view; 6 – prosternum; 7 – propygidium and pygidium; 8 – lateral disc of metaventrite + metepisternum; 9 – protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5740226" httpUri="https://zenodo.org/record/5740226/files/figure.png" pageId="6" pageNumber="629">Fig. 6</figureCitation>
) well impressed, carinate, somewhat divergent anteriorly, united in front by anterior stria.
</paragraph>
<paragraph id="8BD2790BFFBEFFE225A1E091A26AFDA6" blockId="6.[151,1127,219,1587]" pageId="6" pageNumber="629">Anterior margin of mesoventrite medially emarginate; discal marginal mesoventral stria well impressed, carinate; disc of mesoventrite with dense shallow punctures separated by about their own to twice their diameters; meso-metaventral suture weakly impressed, curved outwardly.</paragraph>
<paragraph id="8BD2790BFFBEFFE225A1E311A0ECFC86" blockId="6.[151,1127,219,1587]" pageId="6" pageNumber="629">
Intercoxal disc of metaventrite with long median excavation (male), with punctuation similar to that of mesoventrite, punctures becoming sparser and finer medially; lateral metaventral stria well impressed, carinate, straight, curved outwardly, connected to metaventral-metepisternal suture; lateral disc of metaventrite (
<figureCitation id="1356658EFFBEFFE22701E3B1A137FD21" box="[533,596,667,692]" captionStart="Figs 1–9" captionStartId="3.[113,150,1513,1532]" captionTargetBox="[211,991,219,1492]" captionTargetId="figure-12@3.[211,991,219,1492]" captionTargetPageId="3" captionText="Figs 1–9. External morphology of Pilisaprinus verschureni (Thérond, 1959). 1 – habitus, dorsal view; 2 – same, ventral view; 3 – antennal club, latero-ventral view; 4 – head, dorsal view; 5 – mentum, ventral view; 6 – prosternum; 7 – propygidium and pygidium; 8 – lateral disc of metaventrite + metepisternum; 9 – protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5740226" httpUri="https://zenodo.org/record/5740226/files/figure.png" pageId="6" pageNumber="629">Fig. 8</figureCitation>
) excavated, with large deep setiferous punctures; metepisternum on basal two-thirds with even coarser and denser punctuation with long amber setae, punctuation on apical third becoming much sparser and finer, metepisternum (
<figureCitation id="1356658EFFBEFFE2210AE3F1A73EFD61" box="[1054,1117,731,756]" captionStart="Figs 1–9" captionStartId="3.[113,150,1513,1532]" captionTargetBox="[211,991,219,1492]" captionTargetId="figure-12@3.[211,991,219,1492]" captionTargetPageId="3" captionText="Figs 1–9. External morphology of Pilisaprinus verschureni (Thérond, 1959). 1 – habitus, dorsal view; 2 – same, ventral view; 3 – antennal club, latero-ventral view; 4 – head, dorsal view; 5 – mentum, ventral view; 6 – prosternum; 7 – propygidium and pygidium; 8 – lateral disc of metaventrite + metepisternum; 9 – protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5740226" httpUri="https://zenodo.org/record/5740226/files/figure.png" pageId="6" pageNumber="629">Fig. 8</figureCitation>
) laterally with an additional straight, well-impressed metepisternal stria.
</paragraph>
<paragraph id="8BD2790BFFBEFFE225A1E231A397FCE6" blockId="6.[151,1127,219,1587]" pageId="6" pageNumber="629">Intercoxal disc of first abdominal sternite almost completely striate laterally, with punctuation similar to that of metaventrite; lateral disc of all visible abdominal sternites setose laterally.</paragraph>
<paragraph id="8BD2790BFFBEFFE225A1E251A392FB46" blockId="6.[151,1127,219,1587]" pageId="6" pageNumber="629">
Protibia (
<figureCitation id="1356658EFFBEFFE2240DE251A23AFC01" box="[281,345,891,916]" captionStart="Figs 1–9" captionStartId="3.[113,150,1513,1532]" captionTargetBox="[211,991,219,1492]" captionTargetId="figure-12@3.[211,991,219,1492]" captionTargetPageId="3" captionText="Figs 1–9. External morphology of Pilisaprinus verschureni (Thérond, 1959). 1 – habitus, dorsal view; 2 – same, ventral view; 3 – antennal club, latero-ventral view; 4 – head, dorsal view; 5 – mentum, ventral view; 6 – prosternum; 7 – propygidium and pygidium; 8 – lateral disc of metaventrite + metepisternum; 9 – protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5740226" httpUri="https://zenodo.org/record/5740226/files/figure.png" pageId="6" pageNumber="629">Fig. 9</figureCitation>
) dilated, outer margin with two low triangular distal teeth topped by tiny denticle, followed by two tiny denticles rather distant from the two distal teeth and another two microscopic denticles visible only from ventral view; setae of outer row regular, diminishing in size apically; protarsal groove deep; anterior protibial stria present on basal two-thirds, otherwise obliterated; setae of median row similar in length to those of outer row, present only along the shortened anterior protibial stria; tarsal denticles absent; protibial spur well developed, bent, inserted near protarsal insertion; apical margin of protibia posteriorly with about nine minuscule denticles abutting each other; outer part of posterior surface obscurely variolate-punctate, delimitation of outer and median parts of posterior surface non-existent; posterior protibial stria absent; inner row of setae double, setae dense, increasing in length apically.
</paragraph>
<paragraph id="8BD2790BFFBEFFE225A1E5F1A07CF9A6" blockId="6.[151,1127,219,1587]" pageId="6" pageNumber="629">Mesotibia slender, outer margin with several closely-set thin denticles near mesotarsal insertion; setae of outer row shifted to approximately middle of posterior mesotibial surface, very regular and dense; setae of median row situated right below the setae of outer row, much shorter; posterior mesotibial stria present only on basal third, otherwise (or apically) obliterated; anterior surface of mesotibia with dense row of thin closely-set denticles near outer margin, rest of surface of anterior mesotibial stria imbricate-punctate; anterior mesotibial stria almost complete; inner anterior denticles absent; mesotibial spur short, thick; apical margin of mesotibia anteriorly with two widely-spaced short denticles; inner margin of mesotibia with sparse row of long thin setae; claws of apical tarsomere bent, shorter than half its length; each mesotarsomere with a single long seta ventrally; dorsally without setae; metatibia more slender and longer than mesotibia, in all aspects similar to it.</paragraph>
<paragraph id="8BD2790BFFBFFFE32584E1F1A129FE41" blockId="7.[113,1089,219,499]" pageId="7" pageNumber="630">
Male genitalia. Eighth sternite (
<figureCitation id="1356658EFFBFFFE324C8E1F1A136FF61" box="[476,597,219,244]" captionStart="Figs 13–19" captionStartId="5.[113,150,1122,1141]" captionTargetBox="[252,944,222,1095]" captionTargetId="figure-149@5.[251,950,219,1107]" captionTargetPageId="5" captionText="Figs 13–19. Male terminalia of Pilisaprinus verschureni (Thérond, 1959). 13 – eighth sternite and tergite, ventral view; 14 – same, ventral view; 15 – same, lateral view; 16 – ninth and tenth tergites, dorsal view; spiculum gastrale, ventral view; 17 – ninth, tenth tergites and spiculum gastrale, lateral view; 18 – aedeagus, dorsal view; 19 – same, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5740232" httpUri="https://zenodo.org/record/5740232/files/figure.png" pageId="7" pageNumber="630">Figs 13–14</figureCitation>
) separated longitudinally; vela absent; apex of eighth sternite laterally with dense brush of short setae (
<figureCitation id="1356658EFFBFFFE327A2E1D1A052FE86" box="[694,817,251,275]" captionStart="Figs 13–19" captionStartId="5.[113,150,1122,1141]" captionTargetBox="[252,944,222,1095]" captionTargetId="figure-149@5.[251,950,219,1107]" captionTargetPageId="5" captionText="Figs 13–19. Male terminalia of Pilisaprinus verschureni (Thérond, 1959). 13 – eighth sternite and tergite, ventral view; 14 – same, ventral view; 15 – same, lateral view; 16 – ninth and tenth tergites, dorsal view; spiculum gastrale, ventral view; 17 – ninth, tenth tergites and spiculum gastrale, lateral view; 18 – aedeagus, dorsal view; 19 – same, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5740232" httpUri="https://zenodo.org/record/5740232/files/figure.png" pageId="7" pageNumber="630">Figs 13–15</figureCitation>
); eighth tergite and eighth sternite fused laterally (
<figureCitation id="1356658EFFBFFFE32466E031A2A1FEA6" box="[370,450,283,307]" captionStart="Figs 13–19" captionStartId="5.[113,150,1122,1141]" captionTargetBox="[252,944,222,1095]" captionTargetId="figure-149@5.[251,950,219,1107]" captionTargetPageId="5" captionText="Figs 13–19. Male terminalia of Pilisaprinus verschureni (Thérond, 1959). 13 – eighth sternite and tergite, ventral view; 14 – same, ventral view; 15 – same, lateral view; 16 – ninth and tenth tergites, dorsal view; spiculum gastrale, ventral view; 17 – ninth, tenth tergites and spiculum gastrale, lateral view; 18 – aedeagus, dorsal view; 19 – same, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5740232" httpUri="https://zenodo.org/record/5740232/files/figure.png" pageId="7" pageNumber="630">Fig. 15</figureCitation>
). Ninth tergite (
<figureCitation id="1356658EFFBFFFE32764E031A18FFEA6" box="[624,748,283,307]" captionStart="Figs 13–19" captionStartId="5.[113,150,1122,1141]" captionTargetBox="[252,944,222,1095]" captionTargetId="figure-149@5.[251,950,219,1107]" captionTargetPageId="5" captionText="Figs 13–19. Male terminalia of Pilisaprinus verschureni (Thérond, 1959). 13 – eighth sternite and tergite, ventral view; 14 – same, ventral view; 15 – same, lateral view; 16 – ninth and tenth tergites, dorsal view; spiculum gastrale, ventral view; 17 – ninth, tenth tergites and spiculum gastrale, lateral view; 18 – aedeagus, dorsal view; 19 – same, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5740232" httpUri="https://zenodo.org/record/5740232/files/figure.png" pageId="7" pageNumber="630">Figs 16–17</figureCitation>
) longitudinally fused medially; spiculum gastrale (
<figureCitation id="1356658EFFBFFFE3242AE011A2E8FEC6" box="[318,395,315,339]" captionStart="Figs 13–19" captionStartId="5.[113,150,1122,1141]" captionTargetBox="[252,944,222,1095]" captionTargetId="figure-149@5.[251,950,219,1107]" captionTargetPageId="5" captionText="Figs 13–19. Male terminalia of Pilisaprinus verschureni (Thérond, 1959). 13 – eighth sternite and tergite, ventral view; 14 – same, ventral view; 15 – same, lateral view; 16 – ninth and tenth tergites, dorsal view; spiculum gastrale, ventral view; 17 – ninth, tenth tergites and spiculum gastrale, lateral view; 18 – aedeagus, dorsal view; 19 – same, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5740232" httpUri="https://zenodo.org/record/5740232/files/figure.png" pageId="7" pageNumber="630">Fig. 16</figureCitation>
) gradually dilated from middle in most of apical half, basal end only slightly triangularly dilated. Aedeagus (
<figureCitation id="1356658EFFBFFFE32750E071A1DFFEE1" box="[580,700,347,372]" captionStart="Figs 13–19" captionStartId="5.[113,150,1122,1141]" captionTargetBox="[252,944,222,1095]" captionTargetId="figure-149@5.[251,950,219,1107]" captionTargetPageId="5" captionText="Figs 13–19. Male terminalia of Pilisaprinus verschureni (Thérond, 1959). 13 – eighth sternite and tergite, ventral view; 14 – same, ventral view; 15 – same, lateral view; 16 – ninth and tenth tergites, dorsal view; spiculum gastrale, ventral view; 17 – ninth, tenth tergites and spiculum gastrale, lateral view; 18 – aedeagus, dorsal view; 19 – same, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5740232" httpUri="https://zenodo.org/record/5740232/files/figure.png" pageId="7" pageNumber="630">Figs 18–19</figureCitation>
) slender, almost parallel-sided; basal piece of aedeagus short, ratio of its length: length of parameres 1: 5; parameres of aedeagus fused almost along their basal three-fourths; aedeagus only slightly curved ventrad, apex of aedeagus flattened dorso-ventrally (
<figureCitation id="1356658EFFBFFFE324FAE091A15EFE41" box="[494,573,443,468]" captionStart="Figs 13–19" captionStartId="5.[113,150,1122,1141]" captionTargetBox="[252,944,222,1095]" captionTargetId="figure-149@5.[251,950,219,1107]" captionTargetPageId="5" captionText="Figs 13–19. Male terminalia of Pilisaprinus verschureni (Thérond, 1959). 13 – eighth sternite and tergite, ventral view; 14 – same, ventral view; 15 – same, lateral view; 16 – ninth and tenth tergites, dorsal view; spiculum gastrale, ventral view; 17 – ninth, tenth tergites and spiculum gastrale, lateral view; 18 – aedeagus, dorsal view; 19 – same, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5740232" httpUri="https://zenodo.org/record/5740232/files/figure.png" pageId="7" pageNumber="630">Fig. 19</figureCitation>
).
</paragraph>
<paragraph id="8BD2790BFFBFFFE32565E0F1A0B6FE66" blockId="7.[113,1089,219,499]" box="[113,981,475,499]" pageId="7" pageNumber="630">
<emphasis id="B919A519FFBFFFE32565E0F1A235FE66" bold="true" box="[113,342,475,499]" pageId="7" pageNumber="630">Sexual dimorphism.</emphasis>
Females are devoid of longitudinal metaventral depression.
</paragraph>
</subSubSection>
</treatment>
</document>
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