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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1044.62615" ID-GBIF-Dataset="2a9d2591-e034-4433-8be1-517c6921c937" ID-PMC="PMC9425446" ID-Pensoft-Pub="1313-2970-1044-449" ID-Pensoft-UUID="9FB3984FE63050AEBDD824E47EF5EF42" ID-PubMed="36187209" ID-ZooBank="E2CB7962B7904B95BD003650F11AE2D" ModsDocID="1313-2970-1044-449" checkinTime="1623875390689" checkinUser="pensoft" docAuthor="Assmann, Thorsten, Boutaud, Esteve, Buse, Joern, Drees, Claudia, Friedman, Ariel-Leib-Leonid, Harry, Ingmar, Khoury, Fares, Orbach, Eylon, Renan, Ittai, Schmidt, Constantin, Schmidt, Kilian, Wrase, David W. &amp; Zumstein, Pascale" docDate="2021" docId="03964FA4A59E5E6EBEBCD57B19835F74" docLanguage="en" docName="ZooKeys 1044: 449-478" docOrigin="ZooKeys 1044" docPubDate="2021-06-16" docSource="http://dx.doi.org/10.3897/zookeys.1044.62615" docTitle="Agonum monachum subsp. syriacum Schmidt &amp; Liebherr 2009" docType="treatment" docVersion="4" id="9FB3984FE63050AEBDD824E47EF5EF42" lastPageNumber="449" masterDocId="9FB3984FE63050AEBDD824E47EF5EF42" masterDocTitle="The ground beetle tribe Platynini Bonelli, 1810 (Coleoptera, Carabidae) in the southern Levant: dichotomous and interactive identification tools, ecological traits, and distribution" masterLastPageNumber="478" masterPageNumber="449" pageNumber="449" updateTime="1668150476930" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The ground beetle tribe Platynini Bonelli, 1810 (Coleoptera, Carabidae) in the southern Levant: dichotomous and interactive identification tools, ecological traits, and distribution</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Assmann, Thorsten</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-9203-769X</mods:nameIdentifier>
<mods:affiliation>Institute of Ecology, Leuphana University Lueneburg, Universitaetsallee 1, D- 21335 Lueneburg, Germany</mods:affiliation>
<mods:nameIdentifier type="email">assmann@uni.leuphana.de</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Boutaud, Esteve</mods:namePart>
<mods:affiliation>Institute of Ecology, Leuphana University Lueneburg, Universitaetsallee 1, D- 21335 Lueneburg, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Buse, Joern</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-8226-1893</mods:nameIdentifier>
<mods:affiliation>Ecosystem Monitoring, Research and Wildlife Conservation (SB 23 Invertebrates and Biodiversity), Black Forest National Park, Kniebisstrasse 67, D- 72250 Freudenstadt, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Drees, Claudia</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-2743-395X</mods:nameIdentifier>
<mods:affiliation>School of Life Sciences, University of Sussex, Brighton, BN 1 9 QG, United Kingdom</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Friedman, Ariel-Leib-Leonid</mods:namePart>
<mods:affiliation>Steinhardt Museum of Natural History, Tel Aviv University, Klauzner 12, Tel Aviv, IL- 69978, Israel</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Harry, Ingmar</mods:namePart>
<mods:affiliation>Office for Conservation Biology ABL, Egonstrasse 55, D- 79106 Freiburg, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Khoury, Fares</mods:namePart>
<mods:affiliation>Department of Biology and Biotechnology, American University of Madaba, P. O. Box 2882, Amman, JO- 11821, Jordan</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Orbach, Eylon</mods:namePart>
<mods:affiliation>Remez St. 49, IL- 36044 Qiryat Tiv'on, Israel</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Renan, Ittai</mods:namePart>
<mods:affiliation>Steinhardt Museum of Natural History, Tel Aviv University, Klauzner 12, Tel Aviv, IL- 69978, Israel</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Schmidt, Constantin</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-3892-4255</mods:nameIdentifier>
<mods:affiliation>Gartenstr. 8, 21354 Bleckede, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Schmidt, Kilian</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-4854-1282</mods:nameIdentifier>
<mods:affiliation>Gartenstr. 8, 21354 Bleckede, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wrase, David W.</mods:namePart>
<mods:affiliation>Oderstrasse 2, D- 15306 Gusow-Platkow, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zumstein, Pascale</mods:namePart>
<mods:affiliation>Institute of Ecology, Leuphana University Lueneburg, Universitaetsallee 1, D- 21335 Lueneburg, Germany</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-06-16</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1044</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>449</mods:start>
<mods:end>478</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1044.62615</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1044.62615</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1044-449</mods:identifier>
<mods:identifier type="ZooBank">E2CB7962B7904B95BD003650F11AE2D</mods:identifier>
<mods:identifier type="Pensoft-UUID">9FB3984FE63050AEBDD824E47EF5EF42</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="182754001" LSID="urn:lsid:plazi:treatment:03964FA4A59E5E6EBEBCD57B19835F74" httpUri="http://treatment.plazi.org/id/03964FA4A59E5E6EBEBCD57B19835F74" lastPageNumber="449" pageId="0" pageNumber="449">
<subSubSection pageId="0" pageNumber="449" type="nomenclature">
<paragraph pageId="0" pageNumber="449">
<taxonomicName LSID="03964FA4-A59E-5E6E-BEBC-D57B19835F74" authority="Schmidt &amp; Liebherr, 2009" authorityName="Schmidt &amp; Liebherr" authorityYear="2009" class="Insecta" family="Carabidae" genus="Agonum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agonum monachum subsp. syriacum" order="Coleoptera" pageId="0" pageNumber="449" phylum="Arthropoda" rank="subSpecies" species="monachum" subSpecies="syriacum">Agonum (s. str.) monachum syriacum Schmidt &amp; Liebherr, 2009</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="449" type="dispersal power">
<paragraph pageId="0" pageNumber="449">Dispersal power.</paragraph>
<paragraph pageId="0" pageNumber="449">Fully winged and flight-active (pers. obs.).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="449" type="habitat">
<paragraph pageId="0" pageNumber="449">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="449">
Known only from winter ponds (Fig.
<figureCitation captionStart="Figures 3134" captionStartId="F10" captionText="Figures 31 - 34. Habitats of Platynini species in the southern Levant 31 Merom Golan (Golan Heights): water reservoir, habitat of Agonum rugicolle, A. sordidum, A. marginatum, A. viridicupreum, A. nigrum, and Anchomenus dorsalis infuscatus 32 Ein Akev (Central Negev): headwater stream in desert, habitat of Agonum nigrum 33 Breikhat Ya'ar (Coastal Plain, east of Hadera): winter pond, habitat of Agonum monachum syriacum, A. viridicupreum, A. nigrum, and Anchomenus dorsalis infuscatus 34 Nahal Bezet (Upper Galilee): softwood floodplain woodland with Oleander (Nerium oleander), habitat of Atranus ruficollis." figureDoi="10.3897/zookeys.1044.62615.figures31-34" httpUri="https://binary.pensoft.net/fig/554582" pageId="0" pageNumber="449">31</figureCitation>
) and intact large wetlands (e.g., swamps north of the Sea of Galilee).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="449" type="phenology">
<paragraph pageId="0" pageNumber="449">Phenology.</paragraph>
<paragraph pageId="0" pageNumber="449">Spring breeder, adults hibernate (pers. obs.).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="449" type="distribution">
<paragraph pageId="0" pageNumber="449">Distribution range.</paragraph>
<paragraph pageId="0" pageNumber="449">
The species is distributed from Southwest Europe to Central Asia, southwards to Israel and Iran (
<bibRefCitation author="Schmidt, J" editor="Loebl, I" journalOrPublisher="Brill, Leiden, Boston" pageId="0" pageNumber="449" pagination="642 - 675" refId="B71" refString="Schmidt, J, 2017. Tribe Platynini Bonelli, 1810. In: Loebl, I, Loebl, D, Eds., Catalogue of Palaearctic Coleoptera, Volume 1 (revised and updated edition, Aechostemata - Myxophaga - Adephaga). Brill, Leiden, Boston: 642 - 675" title="Tribe Platynini Bonelli, 1810." volumeTitle="Catalogue of Palaearctic Coleoptera, Volume 1 (revised and updated edition, Aechostemata - Myxophaga - Adephaga)." year="2017">Schmidt 2017</bibRefCitation>
). The subspecies
<taxonomicName authorityName="Schmidt &amp; Liebherr" authorityYear="2009" class="Insecta" family="Carabidae" genus="Agonum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agonum monachum subsp. syriacum" order="Coleoptera" pageId="0" pageNumber="449" phylum="Arthropoda" rank="subSpecies" species="monachum" subSpecies="syriacum">Agonum monachum syriacum</taxonomicName>
is known from Southeast Turkey to Israel (
<bibRefCitation author="Schmidt, J" journalOrPublisher="VERNATE" pageId="0" pageNumber="449" pagination="225 - 257" refId="B72" refString="Schmidt, J, Liebherr, JK, 2009. Beitraege zur Systematik und Verbreitung palaearktischer Arten der Platynini (Insecta: Coleoptera: Carabidae). VERNATE 28: 225 - 257" title="Beitraege zur Systematik und Verbreitung palaearktischer Arten der Platynini (Insecta: Coleoptera: Carabidae)." volume="28" year="2009">Schmidt and Liebherr 2009</bibRefCitation>
) and from Iran (
<bibRefCitation author="Schmidt, J" editor="Loebl, I" journalOrPublisher="Brill, Leiden, Boston" pageId="0" pageNumber="449" pagination="642 - 675" refId="B71" refString="Schmidt, J, 2017. Tribe Platynini Bonelli, 1810. In: Loebl, I, Loebl, D, Eds., Catalogue of Palaearctic Coleoptera, Volume 1 (revised and updated edition, Aechostemata - Myxophaga - Adephaga). Brill, Leiden, Boston: 642 - 675" title="Tribe Platynini Bonelli, 1810." volumeTitle="Catalogue of Palaearctic Coleoptera, Volume 1 (revised and updated edition, Aechostemata - Myxophaga - Adephaga)." year="2017">Schmidt 2017</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="449" type="distribution">
<paragraph pageId="0" pageNumber="449">Distribution in the southern Levant.</paragraph>
<paragraph pageId="0" pageNumber="449">
In the southern Levant recorded exclusively from four sites in Israel, three of them from winter pools in the Central Coastal Plain (Berekhat
<normalizedToken originalValue="Yaar">Ya'ar</normalizedToken>
(sometimes also written Breikhat
<normalizedToken originalValue="Yaar">Ya'ar</normalizedToken>
), Dora Pool and
<normalizedToken originalValue="Gaash">Ga'ash</normalizedToken>
Pool; from the latter two sites only single specimens). A large population exists also in the swamps north of the Sea of Galilee.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="449" type="taxonomic notes">
<paragraph pageId="0" pageNumber="449">Taxonomic notes.</paragraph>
<paragraph pageId="0" pageNumber="449">
<bibRefCitation author="Schmidt, J" editor="Loebl, I" journalOrPublisher="Brill, Leiden, Boston" pageId="0" pageNumber="449" pagination="642 - 675" refId="B71" refString="Schmidt, J, 2017. Tribe Platynini Bonelli, 1810. In: Loebl, I, Loebl, D, Eds., Catalogue of Palaearctic Coleoptera, Volume 1 (revised and updated edition, Aechostemata - Myxophaga - Adephaga). Brill, Leiden, Boston: 642 - 675" title="Tribe Platynini Bonelli, 1810." volumeTitle="Catalogue of Palaearctic Coleoptera, Volume 1 (revised and updated edition, Aechostemata - Myxophaga - Adephaga)." year="2017">Schmidt (2017)</bibRefCitation>
listed the nominate subspecies from Israel, but we know only the subspecies subspecies
<taxonomicName authorityName="Schmidt &amp; Liebherr" authorityYear="2009" class="Insecta" family="Carabidae" genus="Agonum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agonum monachum subsp. syriacum" order="Coleoptera" pageId="0" pageNumber="449" phylum="Arthropoda" rank="subSpecies" species="monachum" subSpecies="syriacum">Agonum monachum syriacum</taxonomicName>
from the southern Levant. All specimens we know from the southern Levant show a greenish metallic luster on the upper side, despite the indication in the original description.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="449" type="conservation">
<paragraph pageId="0" pageNumber="449">Conservation.</paragraph>
<paragraph pageId="0" pageNumber="449">
In contrast to
<taxonomicName lsidName="A. nigrum" pageId="0" pageNumber="449" rank="species" species="nigrum">
<emphasis italics="true" pageId="0" pageNumber="449">A. nigrum</emphasis>
</taxonomicName>
a rarely recorded species; we know only two larger populations (Sea of Galilee, Berekhat
<normalizedToken originalValue="Yaar">Ya'ar</normalizedToken>
). Threatened by habitat loss, drainage, and water table drawdown.
</paragraph>
</subSubSection>
</treatment>
</document>