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<document id="D9FACB90DF90145BE21E3FBA37EA3BE3" ID-DOI="10.5281/zenodo.5386711" ID-ISSN="2345-7600" ID-Zenodo-Dep="5386711" ID-ZooBank="BAA493D3-8195-417B-ABD8-A36D3FD90533" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630303402995" checkinUser="valdenar" docAuthor="Grinang, Jongkar &amp; Ng, Peter K. L." docDate="2015" docId="039387F0C66EFF9BAD91FB210A67F7AC" docLanguage="en" docName="63rbz564-582.pdf" docOrigin="Raffles Bulletin of Zoology 63" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Lepidothelphusa loi Grinang &amp; Ng 2015, new species" docType="treatment" docUuid="06A0A263-62C8-4A1D-AA3A-9F4FFACF5271" docUuidSource="ZooBank" docVersion="5" lastPageNumber="572" masterDocId="FFAAFF88C666FF93AD69FFE40E06FFB1" masterDocTitle="Taxonomy of the semiterrestrial crab Lepidothelphusa cognettii (Nobili, 1903) (Crustacea: Decapoda: Brachyura: Gecarcinucidae), with descriptions of five new species from Sarawak, Malaysia, Borneo" masterLastPageNumber="582" masterPageNumber="564" pageNumber="572" updateTime="1699221322658" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="0C285BB2D3BB24760AEF51DAEBEDCF91">Taxonomy of the semiterrestrial crab Lepidothelphusa cognettii (Nobili, 1903) (Crustacea: Decapoda: Brachyura: Gecarcinucidae), with descriptions of five new species from Sarawak, Malaysia, Borneo</mods:title>
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<mods:namePart id="6519F5CB039A9FBCA739CABDD11F7BCE">Grinang, Jongkar</mods:namePart>
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<mods:namePart id="2770982E5B5EFE12C56ECAB029DFDD11">Ng, Peter K. L.</mods:namePart>
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<mods:title id="1D975F8321A418147860C0F188C14CCE">Raffles Bulletin of Zoology</mods:title>
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<mods:part id="97F397BBEF44E55E3393ED633EE16C7E">
<mods:date id="C4B4E6B8A3025EF9D267FE041A845883">2015</mods:date>
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<mods:number id="F4D20A12B513FD6388187DE35B7EE99D">2015-12-18</mods:number>
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<treatment id="039387F0C66EFF9BAD91FB210A67F7AC" ID-DOI="http://doi.org/10.5281/zenodo.5467452" ID-Zenodo-Dep="5467452" LSID="urn:lsid:zoobank.org:act:06A0A263-62C8-4A1D-AA3A-9F4FFACF5271" httpUri="http://treatment.plazi.org/id/039387F0C66EFF9BAD91FB210A67F7AC" lastPageNumber="572" pageId="8" pageNumber="572">
<subSubSection id="C320656DC66EFF9BAD91FB210C6CFB6C" box="[248,618,1221,1245]" pageId="8" pageNumber="572" type="nomenclature">
<paragraph id="8B8536E6C66EFF9BAD91FB210C6CFB6C" blockId="8.[248,618,1221,1277]" box="[248,618,1221,1245]" pageId="8" pageNumber="572">
<heading id="D0CD818AC66EFF9BAD91FB210C6CFB6C" bold="true" box="[248,618,1221,1245]" centered="true" fontSize="10" level="2" pageId="8" pageNumber="572" reason="2">
<emphasis id="B94EEAF4C66EFF9BAD91FB210C6CFB6C" bold="true" box="[248,618,1221,1245]" pageId="8" pageNumber="572">
<taxonomicName id="4C3A4D65C66EFF9BAD91FB210FD3FB6C" ID-CoL="3TDTJ" authority="Grinang &amp; Ng, 2015" authorityName="Grinang &amp; Ng" authorityYear="2015" box="[248,469,1221,1245]" class="Malacostraca" family="Gecarcinucidae" genus="Lepidothelphusa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="572" phylum="Arthropoda" rank="species" species="loi" status="sp. nov.">
<emphasis id="B94EEAF4C66EFF9BAD91FB210FD3FB6C" bold="true" box="[248,469,1221,1245]" italics="true" pageId="8" pageNumber="572">Lepidothelphusa loi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A27D578FC66EFF9BAC8DFB210C6CFB6C" box="[484,618,1221,1245]" pageId="8" pageNumber="572" rank="species">new species</taxonomicNameLabel>
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<subSubSection id="C320656DC66EFF9BAC71FB010C4FFB4C" box="[280,585,1253,1277]" pageId="8" pageNumber="572" type="description">
<paragraph id="8B8536E6C66EFF9BAC71FB010C4FFB4C" blockId="8.[248,618,1221,1277]" box="[280,585,1253,1277]" pageId="8" pageNumber="572">
(
<figureCitation id="13012A63C66EFF9BAC49FB010F91FB4C" box="[288,407,1253,1277]" captionStart="Fig" captionStartId="9.[136,171,2006,2028]" captionTargetBox="[235,1376,189,1991]" captionTargetId="figure-91@9.[235,1376,189,1991]" captionTargetPageId="9" captionText="Fig. 7. Lepidothelphusa loi, new species, holotype male (10.3 × 9.6 mm) (ZRC 2015.0607), Gunung Ampungan, Sadong River Basin, Serian, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, left cheliped showing serrated inner margin of merus; H, outer view of left chela; I, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.5386737" httpUri="https://zenodo.org/record/5386737/files/figure.png" pageId="8" pageNumber="572">Figs. 7AI</figureCitation>
,
<figureCitation id="13012A63C66EFF9BACCDFB010FE2FB4C" box="[420,484,1253,1277]" captionStart="Fig" captionStartId="10.[113,148,1287,1309]" captionTargetBox="[115,1450,189,1271]" captionTargetId="figure-12@10.[95,1468,178,1283]" captionTargetPageId="10" captionText="Fig. 8. Lepidothelphusa loi, new species, holotype male (10.3 × 9.6 mm) (ZRC 2015.0607), Gunung Ampungan, Sadong River Basin, Serian, Sarawak. A, dorsal view of left G1; B, dorsal view of distal part of left G1; C, dorsal view of right G2; D, ventral view of left G1; E, ventral view of distal part of left G1. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5386739" httpUri="https://zenodo.org/record/5386739/files/figure.png" pageId="8" pageNumber="572">8AE</figureCitation>
,
<figureCitation id="13012A63C66EFF9BAC9BFB010C47FB4C" box="[498,577,1253,1277]" captionStart="Fig" captionStartId="16.[113,148,1308,1330]" captionTargetBox="[113,1452,189,1293]" captionTargetId="figure-491@16.[113,1452,188,1294]" captionTargetPageId="16" captionText="Fig. 14. Live colours of three Lepidothelphusa species from Kuching Division, southwestern Sarawak. A, B, Lepidothelphusa loi, holotype male (10.3 × 9.6 mm) (ZRC 2015.0607), Gunung Ampungan, Sadong River Basin, Serian, Sarawak; C, D, Lepidothelphusa padawan, new species, paratype male (15.8 × 13.2 mm) (SBC.C.004404), Bung Brungu, Padawan, Sarawak; E, F, Lepidothelphusa sangon, new species, paratype male (8.0 × 7.0 mm) (SBC.C.00476), Nawang Waterfall, Bung Bratak, Bau, Sarawak." figureDoi="http://doi.org/10.5281/zenodo.5386759" httpUri="https://zenodo.org/record/5386759/files/figure.png" pageId="8" pageNumber="572">14A, B</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C320656DC66EFF9BAD18FAC10CF7F9CC" pageId="8" pageNumber="572" type="materials_examined">
<paragraph id="8B8536E6C66EFF9BAD18FAC10CF7F9CC" blockId="8.[113,753,1317,1662]" pageId="8" pageNumber="572">
<emphasis id="B94EEAF4C66EFF9BAD18FAC10F49FA8C" bold="true" box="[113,335,1317,1341]" pageId="8" pageNumber="572">Material examined.</emphasis>
<materialsCitation id="3B523CBBC66EFF9BAC3CFAC10CEBFACC" collectingDate="2015-02-08" collectionCode="ZRC" collectorName="J Grinang" country="Malaysia" location="Sadong River" municipality="River Basin" pageId="8" pageNumber="572" specimenCode="ZRC 2015.0607" specimenCount="1" typeStatus="holotype">
<typeStatus id="54818844C66EFF9BAC3CFAC10FBAFA8C" box="[341,444,1317,1341]" pageId="8" pageNumber="572" type="holotype">Holotype</typeStatus>
: male (10.3 ×
<quantity id="4CC29B03C66EFF9BAF3AFAC10CA0FA8C" box="[595,678,1317,1341]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" pageId="8" pageNumber="572" unit="mm" value="9.6">9.6 mm</quantity>
) (
<specimenCode id="DB9C9E9DC66EFF9BAFD2FAC10EE9FAEC" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="8" pageNumber="572">ZRC 2015.0607</specimenCode>
), Gunung Ampungan,
<location id="8EE5603DC66EFF9BAC97FAA10C9BFAEC" LSID="urn:lsid:plazi:treatment:039387F0C66EFF9BAD91FB210A67F7AC:8EE5603DC66EFF9BAC97FAA10C9BFAEC" box="[510,669,1349,1373]" country="Malaysia" municipality="River Basin" name="Sadong River" pageId="8" pageNumber="572">
Sadong
<collectingMunicipality id="6BE1AC9CC66EFF9BAF37FAA10C9BFAEC" box="[606,669,1349,1373]" pageId="8" pageNumber="572">River</collectingMunicipality>
</location>
Basin, Serian,
<collectingCountry id="F32D7676C66EFF9BADA1FA810F2EFACC" box="[200,296,1381,1405]" name="Malaysia" pageId="8" pageNumber="572">Sarawak</collectingCountry>
, coll.
<collectorName id="26CF5330C66EFF9BAC19FA820FDBFACC" box="[368,477,1381,1405]" pageId="8" pageNumber="572">J Grinang</collectorName>
et al.,
<date id="FF841026C66EFF9BAF58FA810CEBFACC" box="[561,749,1381,1405]" pageId="8" pageNumber="572" value="2015-02-08">
<collectingDate id="EFC0E9CEC66EFF9BAF58FA810CEBFACC" box="[561,749,1381,1405]" pageId="8" pageNumber="572" value="2015-02-08">8 February 2015</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B523CBBC66EFF9BAD1BFA620F37FA6C" collectingDate="2015-02-08" collectionCode="ZRC" collectorName="J Grinang" country="Malaysia" location="Sadong River" municipality="River Basin" pageId="8" pageNumber="572" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<typeStatus id="54818844C66EFF9BAD1BFA620EE6FA2F" box="[114,224,1414,1438]" pageId="8" pageNumber="572" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D3CFD6FC66EFF9BAD84FA610F5FFA2C" box="[237,345,1413,1437]" pageId="8" pageNumber="572" type="female">2 females</specimenCount>
(larger 9.8 ×
<quantity id="4CC29B03C66EFF9BAC94FA610C51FA2C" box="[509,599,1413,1437]" metricMagnitude="-3" metricUnit="m" metricValue="9.2" pageId="8" pageNumber="572" unit="mm" value="9.2">9.2 mm</quantity>
), 2 ovigerous females (larger 10.3 ×
<quantity id="4CC29B03C66EFF9BAC1EFA410FC9FA0C" box="[375,463,1445,1469]" metricMagnitude="-3" metricUnit="m" metricValue="9.8" pageId="8" pageNumber="572" unit="mm" value="9.8">9.8 mm</quantity>
) (SBC.C.0043539), data same as holotype
</materialsCitation>
;
<specimenCount id="9D3CFD6FC66EFF9BAC57FA210FA6FA6C" box="[318,416,1477,1501]" pageId="8" pageNumber="572" type="male">14 males</specimenCount>
(largest 11.3 ×
<quantity id="4CC29B03C66EFF9BAF38FA210CB3FA6C" box="[593,693,1477,1501]" metricMagnitude="-2" metricUnit="m" metricValue="1.06" pageId="8" pageNumber="572" unit="mm" value="10.6">10.6 mm</quantity>
, 10.5 ×
<quantity id="4CC29B03C66EFF9BADE0FA010ED9FA4C" box="[137,223,1509,1533]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" pageId="8" pageNumber="572" unit="mm" value="9.6">9.6 mm</quantity>
, 11.8 ×
<quantity id="4CC29B03C66EFF9BAC55FA010FA6FA4C" box="[316,416,1509,1533]" metricMagnitude="-2" metricUnit="m" metricValue="1.09" pageId="8" pageNumber="572" unit="mm" value="10.9">10.9 mm</quantity>
),
<materialsCitation id="3B523CBBC66EFF9BACDCFA010CEBF9CC" collectingDate="2015-06-18" collectingDateMax="2015-06-20" collectingDateMin="2015-06-18" collectionCode="ZRC" collectorName="J Grinang" country="Malaysia" location="Sadong River" municipality="River Basin" pageId="8" pageNumber="572" specimenCode="ZRC 2015.0292" specimenCount="8" specimenCount-female="8" typeStatus="paratype">
<specimenCount id="9D3CFD6FC66EFF9BACDCFA010C18FA4C" box="[437,542,1509,1533]" pageId="8" pageNumber="572" type="female">8 females</specimenCount>
(largest 11.2 ×
<quantity id="4CC29B03C66EFF9BAFA6FA010E9AF9AC" metricMagnitude="-3" metricUnit="m" metricValue="9.8" pageId="8" pageNumber="572" unit="mm" value="9.8">9.8 mm</quantity>
) (
<specimenCode id="DB9C9E9DC66EFF9BADDBF9E10F63F9AC" box="[178,357,1541,1565]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="8" pageNumber="572">ZRC 2015.0292</specimenCode>
), on sides of small stream, mudstone and rocks substrate, densely forest covered slope of hill Gunung Ampungan,
<location id="8EE5603DC66EFF9BAC3CF9A10FEEF9EC" LSID="urn:lsid:plazi:treatment:039387F0C66EFF9BAD91FB210A67F7AC:8EE5603DC66EFF9BAC3CF9A10FEEF9EC" box="[341,488,1605,1629]" country="Malaysia" municipality="River Basin" name="Sadong River" pageId="8" pageNumber="572">
Sadong
<collectingMunicipality id="6BE1AC9CC66EFF9BACC4F9A10FEEF9EC" box="[429,488,1605,1629]" pageId="8" pageNumber="572">River</collectingMunicipality>
</location>
Basin, Serian,
<collectingCountry id="F32D7676C66EFF9BAFE7F9A10CEBF9EC" box="[654,749,1605,1629]" name="Malaysia" pageId="8" pageNumber="572">Sarawak</collectingCountry>
, coll. PKL Ng,
<collectorName id="26CF5330C66EFF9BAC78F9820F7CF9CC" box="[273,378,1637,1661]" pageId="8" pageNumber="572">J Grinang</collectorName>
&amp; PYC Ng,
<date id="FF841026C66EFF9BAF60F9810CEBF9CC" box="[521,749,1637,1661]" pageId="8" pageNumber="572" value="2015-06-18" valueMax="2015-06-20" valueMin="2015-06-18">
<collectingDate id="EFC0E9CEC66EFF9BAF60F9810CEBF9CC" box="[521,749,1637,1661]" pageId="8" pageNumber="572" value="2015-06-18" valueMax="2015-06-20" valueMin="2015-06-18">18 and 20 June 2015</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C320656DC66EFF9BAD18F9410ACFFA0C" pageId="8" pageNumber="572" type="diagnosis">
<paragraph id="8B8536E6C66EFF9BAD18F9410A3EFA8C" blockId="8.[113,753,1701,2109]" lastBlockId="8.[813,1452,1093,1342]" pageId="8" pageNumber="572">
<emphasis id="B94EEAF4C66EFF9BAD18F9410EE2F90C" bold="true" box="[113,228,1701,1725]" pageId="8" pageNumber="572">Diagnosis.</emphasis>
Carapace quadrate, surface relatively flat, smooth, anterolateral margins smooth; antero- and posterolateral regions rugose; epibranchial tooth undiscernible; external orbital tooth with outer margin concave, smooth; serrated projection on frontal region distinct; epigastric and postorbital cristae distinct, high, margins serrated; cervical and H-grooves relatively deep, not confluent (
<figureCitation id="13012A63C66EFF9BAF18F8810E81F82C" captionStart="Fig" captionStartId="9.[136,171,2006,2028]" captionTargetBox="[235,1376,189,1991]" captionTargetId="figure-91@9.[235,1376,189,1991]" captionTargetPageId="9" captionText="Fig. 7. Lepidothelphusa loi, new species, holotype male (10.3 × 9.6 mm) (ZRC 2015.0607), Gunung Ampungan, Sadong River Basin, Serian, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, left cheliped showing serrated inner margin of merus; H, outer view of left chela; I, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.5386737" httpUri="https://zenodo.org/record/5386737/files/figure.png" pageId="8" pageNumber="572">Fig. 7A, B, D</figureCitation>
); ischium of third maxilliped rhomboidal, much longer than broad, with incomplete median sulcus (
<figureCitation id="13012A63C66EFF9BAF3BF8410CA2F80C" box="[594,676,1957,1981]" captionStart="Fig" captionStartId="9.[136,171,2006,2028]" captionTargetBox="[235,1376,189,1991]" captionTargetId="figure-91@9.[235,1376,189,1991]" captionTargetPageId="9" captionText="Fig. 7. Lepidothelphusa loi, new species, holotype male (10.3 × 9.6 mm) (ZRC 2015.0607), Gunung Ampungan, Sadong River Basin, Serian, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, left cheliped showing serrated inner margin of merus; H, outer view of left chela; I, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.5386737" httpUri="https://zenodo.org/record/5386737/files/figure.png" pageId="8" pageNumber="572">Fig. 7F</figureCitation>
); inner margin of merus of chelipeds with relatively lower serrated projection (
<figureCitation id="13012A63C66EFF9BAD82F8010F47F84C" box="[235,321,2021,2045]" captionStart="Fig" captionStartId="9.[136,171,2006,2028]" captionTargetBox="[235,1376,189,1991]" captionTargetId="figure-91@9.[235,1376,189,1991]" captionTargetPageId="9" captionText="Fig. 7. Lepidothelphusa loi, new species, holotype male (10.3 × 9.6 mm) (ZRC 2015.0607), Gunung Ampungan, Sadong River Basin, Serian, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, left cheliped showing serrated inner margin of merus; H, outer view of left chela; I, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.5386737" httpUri="https://zenodo.org/record/5386737/files/figure.png" pageId="8" pageNumber="572">Fig. 7G</figureCitation>
); carpus of chelipeds rugose, inner angle with broad, low triangular tooth (
<figureCitation id="13012A63C66EFF9BACB7F7E10C33F7AC" box="[478,565,2053,2077]" captionStart="Fig" captionStartId="9.[136,171,2006,2028]" captionTargetBox="[235,1376,189,1991]" captionTargetId="figure-91@9.[235,1376,189,1991]" captionTargetPageId="9" captionText="Fig. 7. Lepidothelphusa loi, new species, holotype male (10.3 × 9.6 mm) (ZRC 2015.0607), Gunung Ampungan, Sadong River Basin, Serian, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, left cheliped showing serrated inner margin of merus; H, outer view of left chela; I, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.5386737" httpUri="https://zenodo.org/record/5386737/files/figure.png" pageId="8" pageNumber="572">Fig. 7A</figureCitation>
); fingers of adult major male chela narrowly gaping (
<figureCitation id="13012A63C66EFF9BAC9DF7C10C67F78C" box="[500,609,2085,2109]" captionStart="Fig" captionStartId="9.[136,171,2006,2028]" captionTargetBox="[235,1376,189,1991]" captionTargetId="figure-91@9.[235,1376,189,1991]" captionTargetPageId="9" captionText="Fig. 7. Lepidothelphusa loi, new species, holotype male (10.3 × 9.6 mm) (ZRC 2015.0607), Gunung Ampungan, Sadong River Basin, Serian, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, left cheliped showing serrated inner margin of merus; H, outer view of left chela; I, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.5386737" httpUri="https://zenodo.org/record/5386737/files/figure.png" pageId="8" pageNumber="572">Fig. 7H, I</figureCitation>
); ambulatory legs short, merus of fourth ambulatory legs about half of length of carapace (
<figureCitation id="13012A63C66EFF9BA96FFB810A78FBCC" box="[1030,1150,1125,1149]" captionStart="Fig" captionStartId="9.[136,171,2006,2028]" captionTargetBox="[235,1376,189,1991]" captionTargetId="figure-91@9.[235,1376,189,1991]" captionTargetPageId="9" captionText="Fig. 7. Lepidothelphusa loi, new species, holotype male (10.3 × 9.6 mm) (ZRC 2015.0607), Gunung Ampungan, Sadong River Basin, Serian, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, left cheliped showing serrated inner margin of merus; H, outer view of left chela; I, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.5386737" httpUri="https://zenodo.org/record/5386737/files/figure.png" pageId="8" pageNumber="572">Fig. 7A, E</figureCitation>
); male abdomen triangular, somite 6 width and length subequal, subequal to length of telson (
<figureCitation id="13012A63C66EFF9BAE15FB410DD7FB0C" box="[892,977,1189,1213]" captionStart="Fig" captionStartId="9.[136,171,2006,2028]" captionTargetBox="[235,1376,189,1991]" captionTargetId="figure-91@9.[235,1376,189,1991]" captionTargetPageId="9" captionText="Fig. 7. Lepidothelphusa loi, new species, holotype male (10.3 × 9.6 mm) (ZRC 2015.0607), Gunung Ampungan, Sadong River Basin, Serian, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, left cheliped showing serrated inner margin of merus; H, outer view of left chela; I, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.5386737" httpUri="https://zenodo.org/record/5386737/files/figure.png" pageId="8" pageNumber="572">Fig. 7C</figureCitation>
); G1 stout, sinuous in ventral view, straight in dorsal view; terminal segment straight in dorsal view, gently curving outwards in ventral view (
<figureCitation id="13012A63C66EFF9BA980FB010B98FB4C" box="[1257,1438,1253,1277]" captionStart="Fig" captionStartId="10.[113,148,1287,1309]" captionTargetBox="[115,1450,189,1271]" captionTargetId="figure-12@10.[95,1468,178,1283]" captionTargetPageId="10" captionText="Fig. 8. Lepidothelphusa loi, new species, holotype male (10.3 × 9.6 mm) (ZRC 2015.0607), Gunung Ampungan, Sadong River Basin, Serian, Sarawak. A, dorsal view of left G1; B, dorsal view of distal part of left G1; C, dorsal view of right G2; D, ventral view of left G1; E, ventral view of distal part of left G1. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5386739" httpUri="https://zenodo.org/record/5386739/files/figure.png" pageId="8" pageNumber="572">Fig. 8A, B, D, E</figureCitation>
); G2 with long distal segment, about three-quarters length of basal segment (
<figureCitation id="13012A63C66EFF9BAEBFFAC10A2AFA8C" box="[982,1068,1317,1341]" captionStart="Fig" captionStartId="10.[113,148,1287,1309]" captionTargetBox="[115,1450,189,1271]" captionTargetId="figure-12@10.[95,1468,178,1283]" captionTargetPageId="10" captionText="Fig. 8. Lepidothelphusa loi, new species, holotype male (10.3 × 9.6 mm) (ZRC 2015.0607), Gunung Ampungan, Sadong River Basin, Serian, Sarawak. A, dorsal view of left G1; B, dorsal view of distal part of left G1; C, dorsal view of right G2; D, ventral view of left G1; E, ventral view of distal part of left G1. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5386739" httpUri="https://zenodo.org/record/5386739/files/figure.png" pageId="8" pageNumber="572">Fig. 8C</figureCitation>
).
</paragraph>
<paragraph id="8B8536E6C66EFF9BAE44FA810ACFFA0C" blockId="8.[813,1452,1381,1469]" pageId="8" pageNumber="572">
<emphasis id="B94EEAF4C66EFF9BAE44FA810D82FACC" bold="true" box="[813,900,1381,1405]" pageId="8" pageNumber="572">Colour.</emphasis>
In life, males are yellowish-red carapace, reddishbrown ambulatory legs and yellowish chelipeds. Females are light brown overall (
<figureCitation id="13012A63C66EFF9BA95EFA410ABAFA0C" box="[1079,1212,1445,1469]" captionStart="Fig" captionStartId="16.[113,148,1308,1330]" captionTargetBox="[113,1452,189,1293]" captionTargetId="figure-491@16.[113,1452,188,1294]" captionTargetPageId="16" captionText="Fig. 14. Live colours of three Lepidothelphusa species from Kuching Division, southwestern Sarawak. A, B, Lepidothelphusa loi, holotype male (10.3 × 9.6 mm) (ZRC 2015.0607), Gunung Ampungan, Sadong River Basin, Serian, Sarawak; C, D, Lepidothelphusa padawan, new species, paratype male (15.8 × 13.2 mm) (SBC.C.004404), Bung Brungu, Padawan, Sarawak; E, F, Lepidothelphusa sangon, new species, paratype male (8.0 × 7.0 mm) (SBC.C.00476), Nawang Waterfall, Bung Bratak, Bau, Sarawak." figureDoi="http://doi.org/10.5281/zenodo.5386759" httpUri="https://zenodo.org/record/5386759/files/figure.png" pageId="8" pageNumber="572">Fig. 14A, B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C320656DC66EFF9BAE44FA010D81F98C" pageId="8" pageNumber="572" type="etymology">
<paragraph id="8B8536E6C66EFF9BAE44FA010D81F98C" blockId="8.[813,1452,1509,1597]" pageId="8" pageNumber="572">
<emphasis id="B94EEAF4C66EFF9BAE44FA010DA9FA4C" bold="true" box="[813,943,1509,1533]" pageId="8" pageNumber="572">Etymology.</emphasis>
The name is after Michael Lo in recognition of his kind help in helping us search for these interesting animals.
</paragraph>
</subSubSection>
<subSubSection id="C320656DC66EFF9BAE44F9810A67F7AC" pageId="8" pageNumber="572" type="discussion">
<paragraph id="8B8536E6C66EFF9BAE44F9810B23F8EC" blockId="8.[813,1452,1637,1885]" pageId="8" pageNumber="572">
<emphasis id="B94EEAF4C66EFF9BAE44F9810D9DF9CC" bold="true" box="[813,923,1637,1661]" pageId="8" pageNumber="572">Remarks.</emphasis>
<taxonomicName id="4C3A4D65C66EFF9BAECDF9810A7DF9CC" authority="Grinang &amp; Ng, 2015" authorityName="Grinang &amp; Ng" authorityYear="2015" box="[932,1147,1637,1661]" class="Malacostraca" family="Gecarcinucidae" genus="Lepidothelphusa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="572" phylum="Arthropoda" rank="species" species="loi" status="sp. nov.">
<emphasis id="B94EEAF4C66EFF9BAECDF9810A7DF9CC" box="[932,1147,1637,1661]" italics="true" pageId="8" pageNumber="572">Lepidothelphusa loi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A27D578FC66EFF9BA9E2F9820B16F9CC" box="[1163,1296,1637,1661]" pageId="8" pageNumber="572" rank="species">new species</taxonomicNameLabel>
, can be easily distinguished from
<taxonomicName id="4C3A4D65C66EFF9BA96FF9620A83F92C" authorityName="Nobili" authorityYear="1903" box="[1030,1157,1670,1693]" class="Malacostraca" family="Potamidae" genus="Potamon" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="572" phylum="Arthropoda" rank="species" sensu="stricto" species="cognettii">
<emphasis id="B94EEAF4C66EFF9BA96FF9620A83F92C" box="[1030,1157,1670,1693]" italics="true" pageId="8" pageNumber="572">L. cognettii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27D578FC66EFF9BA9E6F9620AC9F92C" box="[1167,1231,1670,1693]" pageId="8" pageNumber="572" sensu="stricto">s. str.</taxonomicNameLabel>
,
<taxonomicName id="4C3A4D65C66EFF9BA9B7F9620B6BF92C" authority="Grinang &amp; Ng, 2015" authorityName="Grinang &amp; Ng" authorityYear="2015" box="[1246,1389,1669,1693]" class="Malacostraca" family="Gecarcinucidae" genus="Lepidothelphusa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="572" phylum="Arthropoda" rank="species" species="flavochela" status="sp. nov.">
<emphasis id="B94EEAF4C66EFF9BA9B7F9620B6BF92C" box="[1246,1389,1669,1693]" italics="true" pageId="8" pageNumber="572">L. flavochela</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A27D578FC66EFF9BA816F9620D7AF90C" pageId="8" pageNumber="572" rank="species">new species</taxonomicNameLabel>
,
<taxonomicName id="4C3A4D65C66EFF9BAEECF9420DDBF90C" authority="Grinang &amp; Ng, 2015" authorityName="Grinang &amp; Ng" authorityYear="2015" box="[901,989,1701,1725]" class="Malacostraca" family="Gecarcinucidae" genus="Lepidothelphusa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="572" phylum="Arthropoda" rank="species" species="limau" status="sp. nov.">
<emphasis id="B94EEAF4C66EFF9BAEECF9420DDBF90C" box="[901,989,1701,1725]" italics="true" pageId="8" pageNumber="572">L. limau</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A27D578FC66EFF9BAE81F9420A6FF90C" box="[1000,1129,1701,1725]" pageId="8" pageNumber="572" rank="species">new species</taxonomicNameLabel>
, and
<taxonomicName id="4C3A4D65C66EFF9BA9F7F9420B1CF90C" authority="Grinang &amp; Ng, 2015" authorityName="Grinang &amp; Ng" authorityYear="2015" box="[1182,1306,1701,1725]" class="Malacostraca" family="Gecarcinucidae" genus="Lepidothelphusa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="572" phylum="Arthropoda" rank="species" species="padawan" status="sp. nov.">
<emphasis id="B94EEAF4C66EFF9BA9F7F9420B1CF90C" box="[1182,1306,1701,1725]" italics="true" pageId="8" pageNumber="572">L. padawan</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A27D578FC66EFF9BA84FF9420BA1F90C" box="[1318,1447,1701,1725]" pageId="8" pageNumber="572" rank="species">new species</taxonomicNameLabel>
, in having relatively shorter ambulatory legs (
<figureCitation id="13012A63C66EFF9BA858F9210BADF96C" box="[1329,1451,1733,1757]" captionStart="Fig" captionStartId="9.[136,171,2006,2028]" captionTargetBox="[235,1376,189,1991]" captionTargetId="figure-91@9.[235,1376,189,1991]" captionTargetPageId="9" captionText="Fig. 7. Lepidothelphusa loi, new species, holotype male (10.3 × 9.6 mm) (ZRC 2015.0607), Gunung Ampungan, Sadong River Basin, Serian, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, left cheliped showing serrated inner margin of merus; H, outer view of left chela; I, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.5386737" httpUri="https://zenodo.org/record/5386737/files/figure.png" pageId="8" pageNumber="572">Fig. 7A, E</figureCitation>
versus
<figureCitation id="13012A63C66EFF9BAE17F9010A00F94C" box="[894,1030,1765,1789]" captionStart="Fig" captionStartId="2.[113,149,2006,2028]" captionTargetBox="[190,1374,189,1991]" captionTargetId="figure-93@2.[190,1374,189,1991]" captionTargetPageId="2" captionText="Fig. 1. Lepidothelphusa cognettii (Nobili, 1903), male (12.2 × 10.2 mm) (ZRC 2015.0601), Batu Panggah Trail, Gunung Penrissen, Kuching, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, left cheliped showing serrated inner margin of merus; H, outer view of left chela; I, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.5386715" httpUri="https://zenodo.org/record/5386715/files/figure.png" pageId="8" pageNumber="572">Figs. 1A, E</figureCitation>
,
<figureCitation id="13012A63C66EFF9BA97CF9010A5DF94C" box="[1045,1115,1765,1789]" captionStart="Fig" captionStartId="5.[136,171,2006,2028]" captionTargetBox="[256,1353,189,1991]" captionTargetId="figure-90@5.[256,1354,189,1991]" captionTargetPageId="5" captionText="Fig. 3. Lepidothelphusa flavochela, new species, holotype male (8.1 × 7.2 mm) (ZRC 2015.0604), Pagar Besi, Kampung Gumbang, Bau, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, outer view of right chela; H, outer view of left chela; I, left cheliped showing serrated inner margin of merus." figureDoi="http://doi.org/10.5281/zenodo.5386723" httpUri="https://zenodo.org/record/5386723/files/figure.png" pageId="8" pageNumber="572">3A, E</figureCitation>
,
<figureCitation id="13012A63C66EFF9BA903F9010AB6F94C" box="[1130,1200,1765,1789]" captionStart="Fig" captionStartId="7.[136,171,2006,2028]" captionTargetBox="[177,1434,189,1991]" captionTargetId="figure-90@7.[177,1434,189,1991]" captionTargetPageId="7" captionText="Fig. 5. Lepidothelphusa limau, new species, holotype male (12.1 × 10.8 mm) (ZRC 2015.0605), Kampung Peros, Krokong, Bau, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, left cheliped showing serrated inner margin of merus; H, outer view of left chela; I, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.5386729" httpUri="https://zenodo.org/record/5386729/files/figure.png" pageId="8" pageNumber="572">5A, E</figureCitation>
,
<figureCitation id="13012A63C66EFF9BA9A9F9010B00F94C" box="[1216,1286,1765,1789]" captionStart="Fig" captionStartId="11.[136,171,2006,2028]" captionTargetBox="[218,1391,189,1991]" captionTargetId="figure-88@11.[218,1393,189,1991]" captionTargetPageId="11" captionText="Fig. 9. Lepidothelphusa padawan, new species, holotype male (13.6 × 12.5 mm) (ZRC 2015.0608), Kampung Sentah, Padawan, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, left cheliped showing serrated inner margin of merus; H, outer view of left chela; I, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.5386743" httpUri="https://zenodo.org/record/5386743/files/figure.png" pageId="8" pageNumber="572">9A, E</figureCitation>
,
<figureCitation id="13012A63C66EFF9BA87CF9010B6CF94C" box="[1301,1386,1765,1789]" captionStart="Fig" captionStartId="13.[136,171,2006,2028]" captionTargetBox="[176,1431,189,1991]" captionTargetId="figure-90@13.[176,1431,189,1991]" captionTargetPageId="13" captionText="Fig. 11. Lepidothelphusa sangon, new species, holotype male (8.4 × 7.6 mm) (ZRC 2015.0611), Nawang Waterfall, Bung Bratak, Bau, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, left cheliped showing serrated inner margin of merus; H, outer view of left chela; I, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.5386749" httpUri="https://zenodo.org/record/5386749/files/figure.png" pageId="8" pageNumber="572">11A, E</figureCitation>
). The distal segment of the G2 of
<taxonomicName id="4C3A4D65C66EFF9BA90DF8E20A98F8AC" authorityName="Grinang &amp; Ng" authorityYear="2015" box="[1124,1182,1797,1821]" class="Malacostraca" family="Gecarcinucidae" genus="Lepidothelphusa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="572" phylum="Arthropoda" rank="species" species="loi">
<emphasis id="B94EEAF4C66EFF9BA90DF8E20A98F8AC" box="[1124,1182,1797,1821]" italics="true" pageId="8" pageNumber="572">L. loi</emphasis>
</taxonomicName>
is relatively longer than congeners (
<figureCitation id="13012A63C66EFF9BAECEF8C10DFEF88C" box="[935,1016,1829,1853]" captionStart="Fig" captionStartId="10.[113,148,1287,1309]" captionTargetBox="[115,1450,189,1271]" captionTargetId="figure-12@10.[95,1468,178,1283]" captionTargetPageId="10" captionText="Fig. 8. Lepidothelphusa loi, new species, holotype male (10.3 × 9.6 mm) (ZRC 2015.0607), Gunung Ampungan, Sadong River Basin, Serian, Sarawak. A, dorsal view of left G1; B, dorsal view of distal part of left G1; C, dorsal view of right G2; D, ventral view of left G1; E, ventral view of distal part of left G1. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5386739" httpUri="https://zenodo.org/record/5386739/files/figure.png" pageId="8" pageNumber="572">Fig. 8C</figureCitation>
versus
<figureCitation id="13012A63C66EFF9BA920F8C10AA3F88C" box="[1097,1189,1829,1853]" captionStart="Fig" captionStartId="3.[135,171,1092,1114]" captionTargetBox="[137,1474,190,1077]" captionTargetId="figure-8@3.[116,1492,174,1091]" captionTargetPageId="3" captionText="Fig. 2. Lepidothelphusa cognettii (Nobili, 1903), male (12.2 × 10.2 mm) (ZRC 2015.0601), Batu Panggah Trail, Gunung Penrissen, Kuching, Sarawak. A, dorsal view of left G1; B, dorsal view of distal part of left G1; C, dorsal view of left G2 (distal segment damaged at proximal half); D, ventral view of left G1; E, ventral view of distal part of left G1. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5386717" httpUri="https://zenodo.org/record/5386717/files/figure.png" pageId="8" pageNumber="572">Figs. 2C</figureCitation>
,
<figureCitation id="13012A63C66EFF9BA9D9F8C10AD4F88C" box="[1200,1234,1829,1853]" captionStart="Fig" captionStartId="6.[112,147,1068,1090]" captionTargetBox="[117,1453,190,1052]" captionTargetId="figure-12@6.[91,1473,171,1063]" captionTargetPageId="6" captionText="Fig. 4. Lepidothelphusa flavochela, new species, holotype male (8.1 × 7.2 mm) (ZRC 2015.0604), Pagar Besi, Kampung Gumbang, Bau, Sarawak. A, dorsal view of left G1; B, dorsal view of distal part of left G1; C, dorsal view of right G2; D, ventral view of left G1; E, ventral view of distal part of left G1. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5386727" httpUri="https://zenodo.org/record/5386727/files/figure.png" pageId="8" pageNumber="572">4C</figureCitation>
,
<figureCitation id="13012A63C66EFF9BA9B4F8C10AFBF88C" box="[1245,1277,1829,1853]" captionStart="Fig" captionStartId="8.[113,148,958,980]" captionTargetBox="[114,1452,188,944]" captionTargetId="figure-12@8.[100,1462,177,951]" captionTargetPageId="8" captionText="Fig. 6. Lepidothelphusa limau, new species, holotype male (12.1 × 10.8 mm) (ZRC 2015.0605), Kampung Peros, Krokong, Bau, Sarawak. A, dorsal view of left G1; B, dorsal view of distal part of left G1; C, dorsal view of right G2; D, ventral view of left G1; E, ventral view of distal part of left G1. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5386733" httpUri="https://zenodo.org/record/5386733/files/figure.png" pageId="8" pageNumber="572">6C</figureCitation>
,
<figureCitation id="13012A63C66EFF9BA860F8C10B31F88C" box="[1289,1335,1829,1853]" captionStart="Fig" captionStartId="12.[113,148,1333,1355]" captionTargetBox="[111,1451,190,1316]" captionTargetId="figure-12@12.[95,1467,176,1328]" captionTargetPageId="12" captionText="Fig. 10. Lepidothelphusa padawan, new species, holotype male (13.6 × 12.5 mm) (ZRC 2015.0608), Kampung Sentah, Padawan, Sarawak. A, dorsal view of left G1; B, dorsal view of distal part of left G1; C, dorsal view of left G2; D, ventral view of left G1; E, ventral view of distal part of left G1. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5386745" httpUri="https://zenodo.org/record/5386745/files/figure.png" pageId="8" pageNumber="572">10C</figureCitation>
,
<figureCitation id="13012A63C66EFF9BA82BF8C10B75F88C" box="[1346,1395,1829,1853]" captionStart="Fig" captionStartId="14.[113,148,1078,1100]" captionTargetBox="[116,1452,188,1063]" captionTargetId="figure-12@14.[92,1472,172,1070]" captionTargetPageId="14" captionText="Fig. 12. Lepidothelphusa sangon, new species, holotype male (8.4 × 7.6 mm) (ZRC 2015.0611), Nawang Waterfall, Bung Bratak, Bau, Sarawak. A, dorsal view of left G1; B, dorsal view of distal part of left G1; C, dorsal view of left G2; D, ventral view of left G1; E, ventral view of distal part of left G1. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5386751" httpUri="https://zenodo.org/record/5386751/files/figure.png" pageId="8" pageNumber="572">12C</figureCitation>
). See also discussion for
<taxonomicName id="4C3A4D65C66EFF9BA96BF8A20A88F8EC" authorityName="Grinang &amp; Ng" authorityYear="2015" box="[1026,1166,1861,1885]" class="Malacostraca" family="Gecarcinucidae" genus="Lepidothelphusa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="572" phylum="Arthropoda" rank="species" species="flavochela">
<emphasis id="B94EEAF4C66EFF9BA96BF8A20A88F8EC" box="[1026,1166,1861,1885]" italics="true" pageId="8" pageNumber="572">L. flavochela</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3A4D65C66EFF9BA9ACF8A20B19F8EC" authorityName="Grinang &amp; Ng" authorityYear="2015" box="[1221,1311,1861,1885]" class="Malacostraca" family="Gecarcinucidae" genus="Lepidothelphusa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="572" phylum="Arthropoda" rank="species" species="limau">
<emphasis id="B94EEAF4C66EFF9BA9ACF8A20B19F8EC" box="[1221,1311,1861,1885]" italics="true" pageId="8" pageNumber="572">L. limau</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B8536E6C66EFF9BAE44F8610A67F7AC" blockId="8.[813,1452,1925,2077]" pageId="8" pageNumber="572">
<emphasis id="B94EEAF4C66EFF9BAE44F8610D8AF82C" bold="true" box="[813,908,1925,1949]" pageId="8" pageNumber="572">Habitat.</emphasis>
The habitat is a gentle slope, with the substrate composed of moist mud and sand, with leaf litter. The site is shaded, has a spring, and is part of a disturbed dipterocarp forest. The base substrate is sandstone, and the site is more than
<quantity id="4CC29B03C66EFF9BAE0DF7E10DACF7AC" box="[868,938,2053,2077]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="572" unit="m" value="500.0">500 m</quantity>
above sea level.
</paragraph>
</subSubSection>
</treatment>
</document>