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<document id="6AA79D016F445B460F453A1CF11CB06F" ID-DOI="10.5281/zenodo.5386711" ID-ISSN="2345-7600" ID-Zenodo-Dep="5386711" ID-ZooBank="BAA493D3-8195-417B-ABD8-A36D3FD90533" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630303402995" checkinUser="valdenar" docAuthor="Grinang, Jongkar &amp; Ng, Peter K. L." docDate="2015" docId="039387F0C66CFF9FADBCFA690AF5F942" docLanguage="en" docName="63rbz564-582.pdf" docOrigin="Raffles Bulletin of Zoology 63" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Lepidothelphusa padawan Grinang &amp; Ng 2015, new species" docType="treatment" docVersion="3" lastPageNumber="576" masterDocId="FFAAFF88C666FF93AD69FFE40E06FFB1" masterDocTitle="Taxonomy of the semiterrestrial crab Lepidothelphusa cognettii (Nobili, 1903) (Crustacea: Decapoda: Brachyura: Gecarcinucidae), with descriptions of five new species from Sarawak, Malaysia, Borneo" masterLastPageNumber="582" masterPageNumber="564" pageNumber="574" updateTime="1699221322658" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="260510DB110E7C078B91A771AC48B036" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E99368061B86D4225D5AEE8A6BA9BCD6">
<mods:title id="7E0E214E03E87D61E7CF25E6ECD13251">Taxonomy of the semiterrestrial crab Lepidothelphusa cognettii (Nobili, 1903) (Crustacea: Decapoda: Brachyura: Gecarcinucidae), with descriptions of five new species from Sarawak, Malaysia, Borneo</mods:title>
</mods:titleInfo>
<mods:name id="3E341514248E1970E5A86BE7C1BDB6C2" type="personal">
<mods:role id="64B6BA009E01B20D6E4094FB977ED66C">
<mods:roleTerm id="5097CC035BFACE25CD439C9E047D1A3C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E6A4EECC648B858AC557938ADF52386E">Grinang, Jongkar</mods:namePart>
</mods:name>
<mods:name id="9411192E57104ADEC752FD9E925D9128" type="personal">
<mods:role id="3B66133067A099C580E492657B75C33A">
<mods:roleTerm id="8242149E3312B5ABE6BC1B302A71C75D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8BA19E6B7ED92B898795BA313C5EAA97">Ng, Peter K. L.</mods:namePart>
</mods:name>
<mods:typeOfResource id="27B1FD445749D77A86B63E1AA4A3B2A1">text</mods:typeOfResource>
<mods:relatedItem id="E758800D0609C52215C9460D7065717D" type="host">
<mods:titleInfo id="4EFE513709BD6021A71DE20573E90F88">
<mods:title id="99440E1E8F37101B5AB40BF1DCD3DDC7">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="17A5ABF0EBAB8A1359A9CE23576B1434">
<mods:date id="4444F5B83EC7D720ECDD2BE0505E7A46">2015</mods:date>
<mods:detail id="0554F391B2A34EB51CF0E91FDF979A01" type="pubDate">
<mods:number id="77DED730C6DB890892EC1ED2141F083D">2015-12-18</mods:number>
</mods:detail>
<mods:detail id="D4DA9E2AB3659C57423E2147AC333F0A" type="volume">
<mods:number id="A8D676BF36E0CE031DCA9E69BFF5813C">63</mods:number>
</mods:detail>
<mods:extent id="A3A51804136D3ED6BB450209885F409F" unit="page">
<mods:start id="BAD4AAB1145AFFE6511C1E6737688A5F">564</mods:start>
<mods:end id="36D593F4B62F6BDAEC9BCFA6C9850F92">582</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="8825701371D996A0170DD0E7D3ED79FA">journal article</mods:classification>
<mods:identifier id="6EE4CEAC3B1023BD6211422F9648DF72" type="DOI">10.5281/zenodo.5386711</mods:identifier>
<mods:identifier id="148ECEECF92ED9D57C686E5C01B6509A" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="34E58E11A10FD81681DA38C7FA5DB1A5" type="Zenodo-Dep">5386711</mods:identifier>
<mods:identifier id="582F76F61DC5689732133A58D3DF4677" type="ZooBank">BAA493D3-8195-417B-ABD8-A36D3FD90533</mods:identifier>
</mods:mods>
<treatment id="039387F0C66CFF9FADBCFA690AF5F942" LSID="urn:lsid:plazi:treatment:039387F0C66CFF9FADBCFA690AF5F942" httpUri="http://treatment.plazi.org/id/039387F0C66CFF9FADBCFA690AF5F942" lastPageId="12" lastPageNumber="576" pageId="10" pageNumber="574">
<subSubSection id="C320656DC66CFF99ADBCFA690C8BFA14" box="[213,653,1421,1446]" pageId="10" pageNumber="574" type="nomenclature">
<paragraph id="8B8536E6C66CFF99ADBCFA690C8BFA14" blockId="10.[213,653,1421,1478]" box="[213,653,1421,1446]" pageId="10" pageNumber="574">
<heading id="D0CD818AC66CFF99ADBCFA690C8BFA14" bold="true" box="[213,653,1421,1446]" centered="true" fontSize="10" level="2" pageId="10" pageNumber="574" reason="2">
<emphasis id="B94EEAF4C66CFF99ADBCFA690C8BFA14" bold="true" box="[213,653,1421,1446]" pageId="10" pageNumber="574">
<taxonomicName id="4C3A4D65C66CFF99ADBCFA690FFEFA14" ID-CoL="3TDTK" authority="Grinang &amp; Ng, 2015" authorityName="Grinang &amp; Ng" authorityYear="2015" box="[213,504,1421,1445]" class="Malacostraca" family="Gecarcinucidae" genus="Lepidothelphusa" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="574" phylum="Arthropoda" rank="species" species="padawan" status="sp. nov.">
<emphasis id="B94EEAF4C66CFF99ADBCFA690FFEFA14" bold="true" box="[213,504,1421,1445]" italics="true" pageId="10" pageNumber="574">Lepidothelphusa padawan</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A27D578FC66CFF99AF61FA6A0C8BFA14" box="[520,653,1421,1446]" pageId="10" pageNumber="574" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C320656DC66CFF99AD8DFA4A0C7BFA74" box="[228,637,1453,1478]" pageId="10" pageNumber="574" type="description">
<paragraph id="8B8536E6C66CFF99AD8DFA4A0C7BFA74" blockId="10.[213,653,1421,1478]" box="[228,637,1453,1478]" pageId="10" pageNumber="574">
(
<figureCitation id="13012A63C66CFF99AD85FA4A0F62FA77" box="[236,356,1454,1478]" captionStart="Fig" captionStartId="11.[136,171,2006,2028]" captionTargetBox="[218,1391,189,1991]" captionTargetId="figure-88@11.[218,1393,189,1991]" captionTargetPageId="11" captionText="Fig. 9. Lepidothelphusa padawan, new species, holotype male (13.6 × 12.5 mm) (ZRC 2015.0608), Kampung Sentah, Padawan, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, left cheliped showing serrated inner margin of merus; H, outer view of left chela; I, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.5386743" httpUri="https://zenodo.org/record/5386743/files/figure.png" pageId="10" pageNumber="574">Figs. 9A–I</figureCitation>
,
<figureCitation id="13012A63C66CFF99AC18FA4A0FB9FA77" box="[369,447,1454,1478]" captionStart="Fig" captionStartId="12.[113,148,1333,1355]" captionTargetBox="[111,1451,190,1316]" captionTargetId="figure-12@12.[95,1467,176,1328]" captionTargetPageId="12" captionText="Fig. 10. Lepidothelphusa padawan, new species, holotype male (13.6 × 12.5 mm) (ZRC 2015.0608), Kampung Sentah, Padawan, Sarawak. A, dorsal view of left G1; B, dorsal view of distal part of left G1; C, dorsal view of left G2; D, ventral view of left G1; E, ventral view of distal part of left G1. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5386745" httpUri="https://zenodo.org/record/5386745/files/figure.png" pageId="10" pageNumber="574">10A–E</figureCitation>
,
<figureCitation id="13012A63C66CFF99ACA5FA4A0C1AFA74" box="[460,540,1454,1478]" captionStart="Fig" captionStartId="16.[113,148,1308,1330]" captionTargetBox="[113,1452,189,1293]" captionTargetId="figure-491@16.[113,1452,188,1294]" captionTargetPageId="16" captionText="Fig. 14. Live colours of three Lepidothelphusa species from Kuching Division, southwestern Sarawak. A, B, Lepidothelphusa loi, holotype male (10.3 × 9.6 mm) (ZRC 2015.0607), Gunung Ampungan, Sadong River Basin, Serian, Sarawak; C, D, Lepidothelphusa padawan, new species, paratype male (15.8 × 13.2 mm) (SBC.C.00440–4), Bung Brungu, Padawan, Sarawak; E, F, Lepidothelphusa sangon, new species, paratype male (8.0 × 7.0 mm) (SBC.C.00476), Nawang Waterfall, Bung Bratak, Bau, Sarawak." figureDoi="http://doi.org/10.5281/zenodo.5386759" httpUri="https://zenodo.org/record/5386759/files/figure.png" pageId="10" pageNumber="574">14C, D</figureCitation>
,
<figureCitation id="13012A63C66CFF99AF43FA490C70FA74" box="[554,630,1453,1477]" captionStart="Fig" captionStartId="17.[135,170,1446,1468]" captionTargetBox="[135,1473,188,1430]" captionTargetId="figure-69@17.[134,1475,188,1430]" captionTargetPageId="17" captionText="Fig. 15. Live colours of Lepidothelphusa padawan, new species, paratypes (ZRC 2015.0609), showing variation. A, male (ca. 11.0 mm carapace width); B, male (ca. 11.0 mm carapace width); C, male (ca. 12.0 mm carapace width); D, male (ca. 12.0 mm carapace width); E, male (ca. 12.0 mm carapace width); F, female (ca. 9.0 mm carapace width). Kampung Sentah, Padawan, Sarawak" figureDoi="http://doi.org/10.5281/zenodo.5386761" httpUri="https://zenodo.org/record/5386761/files/figure.png" pageId="10" pageNumber="574">15A–F</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C320656DC66CFF99AD18FA090F8AF794" pageId="10" pageNumber="574" type="materials_examined">
<paragraph id="8B8536E6C66CFF99AD18FA090F8AF794" blockId="10.[113,753,1517,2086]" pageId="10" pageNumber="574">
<emphasis id="B94EEAF4C66CFF99AD18FA090F5BF9B4" bold="true" box="[113,349,1517,1541]" pageId="10" pageNumber="574">Material examined.</emphasis>
<materialsCitation id="3B523CBBC66CFF99AC0EFA0A0C72F9F7" collectingDate="2013-11-05" collectionCode="ZRC" collectorName="J Grinang &amp; HH Tan" country="Malaysia" location="Sarawak" pageId="10" pageNumber="574" specimenCode="ZRC 2015.0608" specimenCount="1" typeStatus="holotype">
<typeStatus id="54818844C66CFF99AC0EFA0A0FD5F9B7" box="[359,467,1518,1542]" pageId="10" pageNumber="574" type="holotype">Holotype</typeStatus>
: male (13.6 ×
<quantity id="4CC29B03C66CFF99AFE8FA090CEFF9B7" box="[641,745,1517,1542]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="10" pageNumber="574" unit="mm" value="12.5">12.5 mm</quantity>
) (
<specimenCode id="DB9C9E9DC66CFF99AD10F9EA0F34F994" box="[121,306,1549,1574]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="10" pageNumber="574">ZRC 2015.0608</specimenCode>
), Kampung Sentah, Padawan,
<collectingCountry id="F32D7676C66CFF99AFE5F9EA0CEAF997" box="[652,748,1550,1574]" name="Malaysia" pageId="10" pageNumber="574">Sarawak</collectingCountry>
, coll.
<collectorName id="26CF5330C66CFF99ADC3F9CA0F10F9F7" box="[170,278,1582,1606]" pageId="10" pageNumber="574">J Grinang</collectorName>
&amp;
<collectorName id="26CF5330C66CFF99AC56F9CA0F9DF9F4" box="[319,411,1582,1605]" pageId="10" pageNumber="574">HH Tan</collectorName>
,
<date id="FF841026C66CFF99ACC0F9C90C72F9F7" box="[425,628,1581,1606]" pageId="10" pageNumber="574" value="2013-11-05">
<collectingDate id="EFC0E9CEC66CFF99ACC0F9C90C72F9F7" box="[425,628,1581,1606]" pageId="10" pageNumber="574" value="2013-11-05">5 November 2013</collectingDate>
</date>
</materialsCitation>
.
<typeStatus id="54818844C66CFF99AFE8F9CA0CEBF9F7" box="[641,749,1582,1606]" pageId="10" pageNumber="574" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D3CFD6FC66CFF99AD18F9A90EC1F9D7" box="[113,199,1613,1638]" pageId="10" pageNumber="574" type="male">5 males</specimenCount>
(largest 14.2 ×
<quantity id="4CC29B03C66CFF99AC17F9AA0FE3F9D7" box="[382,485,1614,1638]" metricMagnitude="-2" metricUnit="m" metricValue="1.21" pageId="10" pageNumber="574" unit="mm" value="12.1">12.1 mm</quantity>
),
<materialsCitation id="3B523CBBC66CFF99AC92F9AA0FAEF917" collectingDate="2014-01-12" collectionCode="ZRC" collectorName="J Grinang" country="Malaysia" location="Sarawak" pageId="10" pageNumber="574" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<specimenCount id="9D3CFD6FC66CFF99AC92F9AA0C63F9D7" box="[507,613,1614,1638]" pageId="10" pageNumber="574" type="female">2 females</specimenCount>
(larger 12.6 ×
<quantity id="4CC29B03C66CFF99ADE0F98A0EEDF937" box="[137,235,1646,1670]" metricMagnitude="-2" metricUnit="m" metricValue="1.0699999999999998" pageId="10" pageNumber="574" unit="mm" value="10.7">10.7 mm</quantity>
) (SBC.C.00407–13), location as holotype, coll.
<collectorName id="26CF5330C66CFF99AD18F96A0EE6F917" box="[113,224,1678,1702]" pageId="10" pageNumber="574">J Grinang</collectorName>
,
<date id="FF841026C66CFF99AD87F96A0FAEF917" box="[238,424,1678,1702]" pageId="10" pageNumber="574" value="2014-01-12">
<collectingDate id="EFC0E9CEC66CFF99AD87F96A0FAEF917" box="[238,424,1678,1702]" pageId="10" pageNumber="574" value="2014-01-12">12 January 2014</collectingDate>
</date>
</materialsCitation>
;
<specimenCount id="9D3CFD6FC66CFF99ACDFF96A0C0AF917" box="[438,524,1678,1702]" pageId="10" pageNumber="574" type="male">7 males</specimenCount>
(largest 10.7 ×
<quantity id="4CC29B03C66CFF99AFA8F96A0E99F977" metricMagnitude="-2" metricUnit="m" metricValue="1.01" pageId="10" pageNumber="574" unit="mm" value="10.1">10.1 mm</quantity>
),
<materialsCitation id="3B523CBBC66CFF99ADDFF94A0F0EF8B7" collectingDate="2014-01-12" collectorName="J Grinang &amp; A Alek" country="Malaysia" county="Padawan" location="Sebarau" municipality="Kampung Bidak" pageId="10" pageNumber="574" specimenCount="3" specimenCount-juvenile="3" typeStatus="paratype">
<specimenCount id="9D3CFD6FC66CFF99ADDFF94A0F29F977" box="[182,303,1710,1734]" pageId="10" pageNumber="574" type="juvenile">3 juveniles</specimenCount>
(SBC.C.00414–23),
<location id="8EE5603DC66CFF99AF74F94A0C71F977" LSID="urn:lsid:plazi:treatment:039387F0C66CFF9FADBCFA690AF5F942:8EE5603DC66CFF99AF74F94A0C71F977" box="[541,631,1710,1734]" country="Malaysia" county="Padawan" municipality="Kampung Bidak" name="Sebarau" pageId="10" pageNumber="574">Sebarau</location>
,
<collectingMunicipality id="6BE1AC9CC66CFF99AFEFF94A0EB3F957" pageId="10" pageNumber="574">Kampung Bidak</collectingMunicipality>
,
<collectingCounty id="62E44E6AC66CFF99ADAAF92A0F21F957" box="[195,295,1742,1766]" pageId="10" pageNumber="574">Padawan</collectingCounty>
,
<collectingCountry id="F32D7676C66CFF99AC5CF92A0F92F957" box="[309,404,1742,1766]" name="Malaysia" pageId="10" pageNumber="574">Sarawak</collectingCountry>
, coll.
<collectorName id="26CF5330C66CFF99ACB5F92A0C4FF957" box="[476,585,1742,1766]" pageId="10" pageNumber="574">J Grinang</collectorName>
&amp;
<collectorName id="26CF5330C66CFF99AF1BF92A0CC1F957" box="[626,711,1742,1766]" pageId="10" pageNumber="574">A Alek</collectorName>
,
<date id="FF841026C66CFF99AFBCF92A0F0EF8B7" pageId="10" pageNumber="574" value="2014-01-12">
<collectingDate id="EFC0E9CEC66CFF99AFBCF92A0F0EF8B7" pageId="10" pageNumber="574" value="2014-01-12">12 January 2014</collectingDate>
</date>
</materialsCitation>
;
<specimenCount id="9D3CFD6FC66CFF99AC7FF9090F6BF8B7" box="[278,365,1773,1798]" pageId="10" pageNumber="574" type="male">5 males</specimenCount>
(largest 9.6 ×
<quantity id="4CC29B03C66CFF99AF7FF9090C76F8B7" box="[534,624,1773,1798]" metricMagnitude="-3" metricUnit="m" metricValue="8.6" pageId="10" pageNumber="574" unit="mm" value="8.6">8.6 mm</quantity>
),
<materialsCitation id="3B523CBBC66CFF99AFECF90A0CC1F8F7" collectingDate="2014-01-14" collectorName="Kampung Petag &amp; Padawan &amp; J Grinang" country="Malaysia" location="Sarawak" pageId="10" pageNumber="574" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<specimenCount id="9D3CFD6FC66CFF99AFECF90A0CF6F8B7" box="[645,752,1774,1798]" pageId="10" pageNumber="574" type="female">2 females</specimenCount>
(larger 9.3 ×
<quantity id="4CC29B03C66CFF99AD96F8EA0F55F897" box="[255,339,1806,1830]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="10" pageNumber="574" unit="mm" value="8.2">8.2 mm</quantity>
) (SBC.C.00424–30),
<collectorName id="26CF5330C66CFF99AF57F8EA0CEAF897" box="[574,748,1806,1830]" pageId="10" pageNumber="574">Kampung Petag</collectorName>
,
<collectorName id="26CF5330C66CFF99AD18F8CA0ED3F8F7" box="[113,213,1838,1862]" pageId="10" pageNumber="574">Padawan</collectorName>
,
<collectingCountry id="F32D7676C66CFF99AD8DF8CA0F42F8F7" box="[228,324,1838,1862]" name="Malaysia" pageId="10" pageNumber="574">Sarawak</collectingCountry>
, coll.
<collectorName id="26CF5330C66CFF99ACE5F8CA0FFDF8F7" box="[396,507,1838,1862]" pageId="10" pageNumber="574">J Grinang</collectorName>
,
<date id="FF841026C66CFF99AF63F8CA0CC1F8F7" box="[522,711,1838,1862]" pageId="10" pageNumber="574" value="2014-01-14">
<collectingDate id="EFC0E9CEC66CFF99AF63F8CA0CC1F8F7" box="[522,711,1838,1862]" pageId="10" pageNumber="574" value="2014-01-14">14 January 2014</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B523CBBC66CFF99AFBCF8CA0EACF817" collectingDate="2013-11-05" collectionCode="ZRC" collectorName="HH Tan &amp; J Grinang" country="Malaysia" location="Kampung Sentah" municipality="Padawan" pageId="10" pageNumber="574" specimenCode="ZRC 2015.0609" specimenCount="15" specimenCount-female="3" specimenCount-male="12" typeStatus="paratype">
<specimenCount id="9D3CFD6FC66CFF99AFBCF8CA0EB1F8D7" pageId="10" pageNumber="574" type="male">12 males</specimenCount>
,
<specimenCount id="9D3CFD6FC66CFF99ADAFF8AA0F31F8D7" box="[198,311,1870,1894]" pageId="10" pageNumber="574" type="female">3 females</specimenCount>
(
<specimenCode id="DB9C9E9DC66CFF99AC23F8AA0C0DF8D4" box="[330,523,1869,1894]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="10" pageNumber="574">ZRC 2015.0609</specimenCode>
),
<location id="8EE5603DC66CFF99AF4BF8AA0CEDF8D7" LSID="urn:lsid:plazi:treatment:039387F0C66CFF9FADBCFA690AF5F942:8EE5603DC66CFF99AF4BF8AA0CEDF8D7" box="[546,747,1870,1894]" country="Malaysia" municipality="Padawan" name="Kampung Sentah" pageId="10" pageNumber="574">Kampung Sentah</location>
,
<collectingMunicipality id="6BE1AC9CC66CFF99AD1BF88A0ED3F837" box="[114,213,1902,1926]" pageId="10" pageNumber="574">Padawan</collectingMunicipality>
,
<collectingCountry id="F32D7676C66CFF99AD89F88A0F39F837" box="[224,319,1902,1926]" name="Malaysia" pageId="10" pageNumber="574">Sarawak</collectingCountry>
, coll.
<collectorName id="26CF5330C66CFF99AC17F88A0FD2F834" box="[382,468,1902,1925]" pageId="10" pageNumber="574">HH Tan</collectorName>
&amp;
<collectorName id="26CF5330C66CFF99AC9EF88A0C64F837" box="[503,610,1902,1926]" pageId="10" pageNumber="574">J Grinang</collectorName>
,
<date id="FF841026C66CFF99AF05F8890EACF817" pageId="10" pageNumber="574" value="2013-11-05">
<collectingDate id="EFC0E9CEC66CFF99AF05F8890EACF817" pageId="10" pageNumber="574" value="2013-11-05">5 November 2013</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B523CBBC66CFF99ADDFF86A0C91F877" collectingDate="2013-12-04" collectionCode="ZRC" collectorName="Tan" country="Malaysia" location="Sarawak" pageId="10" pageNumber="574" specimenCode="ZRC 2015.0610" specimenCount="6" specimenCount-female="3" specimenCount-male="3" typeStatus="paratype">
<specimenCount id="9D3CFD6FC66CFF99ADDFF86A0F0DF817" box="[182,267,1934,1958]" pageId="10" pageNumber="574" type="male">3 males</specimenCount>
,
<specimenCount id="9D3CFD6FC66CFF99AC7EF86A0F78F817" box="[279,382,1934,1958]" pageId="10" pageNumber="574" type="female">3 females</specimenCount>
(
<specimenCode id="DB9C9E9DC66CFF99ACE4F86A0C46F814" box="[397,576,1933,1958]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="10" pageNumber="574">ZRC 2015.0610</specimenCode>
),
<collectingCountry id="F32D7676C66CFF99AF3BF86A0CB7F817" box="[594,689,1934,1958]" name="Malaysia" pageId="10" pageNumber="574">Sarawak</collectingCountry>
, from aquarium trade, obtained HH
<collectorName id="26CF5330C66CFF99ACCFF84A0FD4F874" box="[422,466,1966,1989]" pageId="10" pageNumber="574">Tan</collectorName>
,
<date id="FF841026C66CFF99ACB2F84A0C91F877" box="[475,663,1966,1990]" pageId="10" pageNumber="574" value="2013-12-04">
<collectingDate id="EFC0E9CEC66CFF99ACB2F84A0C91F877" box="[475,663,1966,1990]" pageId="10" pageNumber="574" value="2013-12-04">4 December 2013</collectingDate>
</date>
</materialsCitation>
;
<specimenCount id="9D3CFD6FC66CFF99AFC9F8490CF7F877" box="[672,753,1965,1990]" pageId="10" pageNumber="574" type="male">5 males</specimenCount>
(largest 15.8 ×
<quantity id="4CC29B03C66CFF99AC7CF82A0F7EF857" box="[277,376,1998,2022]" metricMagnitude="-2" metricUnit="m" metricValue="1.3199999999999998" pageId="10" pageNumber="574" unit="mm" value="13.2">13.2 mm</quantity>
),
<materialsCitation id="3B523CBBC66CFF99ACE2F82A0F8EF794" collectingDate="2015-03-08" collectorName="Bung Brungu &amp; Padawan &amp; J Grinang" country="Malaysia" location="Sarawak" pageId="10" pageNumber="574" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<specimenCount id="9D3CFD6FC66CFF99ACE2F82A0FF5F857" box="[395,499,1998,2022]" pageId="10" pageNumber="574" type="female">2 females</specimenCount>
(larger 10.8 ×
<quantity id="4CC29B03C66CFF99AFFCF82A0CEFF857" box="[661,745,1998,2022]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" pageId="10" pageNumber="574" unit="mm" value="9.1">9.1 mm</quantity>
) (SBC.C.00440–46),
<collectorName id="26CF5330C66CFF99AC24F80A0FE6F7B7" box="[333,480,2030,2054]" pageId="10" pageNumber="574">Bung Brungu</collectorName>
,
<collectorName id="26CF5330C66CFF99AC82F80A0C48F7B7" box="[491,590,2030,2054]" pageId="10" pageNumber="574">Padawan</collectorName>
,
<collectingCountry id="F32D7676C66CFF99AF31F80A0CB1F7B7" box="[600,695,2030,2054]" name="Malaysia" pageId="10" pageNumber="574">Sarawak</collectingCountry>
, coll.
<collectorName id="26CF5330C66CFF99AD18F7EA0ED8F797" box="[113,222,2062,2086]" pageId="10" pageNumber="574">J Grinang</collectorName>
,
<date id="FF841026C66CFF99AD82F7EA0F8EF794" box="[235,392,2061,2086]" pageId="10" pageNumber="574" value="2015-03-08">
<collectingDate id="EFC0E9CEC66CFF99AD82F7EA0F8EF794" box="[235,392,2061,2086]" pageId="10" pageNumber="574" value="2015-03-08">8 March 2015</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C320656DC66CFF9FAE44FA6A0F52F942" lastPageId="12" lastPageNumber="576" pageId="10" pageNumber="574" type="diagnosis">
<paragraph id="8B8536E6C66CFF99AE44FA6A0A40F797" blockId="10.[812,1452,1422,2086]" pageId="10" pageNumber="574">
<emphasis id="B94EEAF4C66CFF99AE44FA6A0DA7FA17" bold="true" box="[813,929,1422,1446]" pageId="10" pageNumber="574">Diagnosis.</emphasis>
Carapace relatively quadrate, surface relatively flat, smooth, anterolateral margins serrated; antero- and posterolateral regions slightly rugose; epibranchial tooth undiscernible; external orbital tooth with outer margin slightly concave, smooth; serrated projection on frontal region distinct; epigastric and postorbital cristae distinct, margins serrated; cervical and H-grooves relatively deep, not confluent (
<figureCitation id="13012A63C66CFF99AEA4F98A0A6CF937" box="[973,1130,1646,1670]" captionStart="Fig" captionStartId="11.[136,171,2006,2028]" captionTargetBox="[218,1391,189,1991]" captionTargetId="figure-88@11.[218,1393,189,1991]" captionTargetPageId="11" captionText="Fig. 9. Lepidothelphusa padawan, new species, holotype male (13.6 × 12.5 mm) (ZRC 2015.0608), Kampung Sentah, Padawan, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, left cheliped showing serrated inner margin of merus; H, outer view of left chela; I, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.5386743" httpUri="https://zenodo.org/record/5386743/files/figure.png" pageId="10" pageNumber="574">Fig. 9A, B, D</figureCitation>
); ischium of third maxilliped rhomboidal, slightly longer than broad, sulcus undiscernible (
<figureCitation id="13012A63C66CFF99AE5DF94A0D8CF977" box="[820,906,1710,1734]" captionStart="Fig" captionStartId="11.[136,171,2006,2028]" captionTargetBox="[218,1391,189,1991]" captionTargetId="figure-88@11.[218,1393,189,1991]" captionTargetPageId="11" captionText="Fig. 9. Lepidothelphusa padawan, new species, holotype male (13.6 × 12.5 mm) (ZRC 2015.0608), Kampung Sentah, Padawan, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, left cheliped showing serrated inner margin of merus; H, outer view of left chela; I, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.5386743" httpUri="https://zenodo.org/record/5386743/files/figure.png" pageId="10" pageNumber="574">Fig. 9F</figureCitation>
); inner margin of merus of chelipeds with large, prominently expanded serrated projection (
<figureCitation id="13012A63C66CFF99A992F92A0B54F957" box="[1275,1362,1742,1766]" captionStart="Fig" captionStartId="11.[136,171,2006,2028]" captionTargetBox="[218,1391,189,1991]" captionTargetId="figure-88@11.[218,1393,189,1991]" captionTargetPageId="11" captionText="Fig. 9. Lepidothelphusa padawan, new species, holotype male (13.6 × 12.5 mm) (ZRC 2015.0608), Kampung Sentah, Padawan, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, left cheliped showing serrated inner margin of merus; H, outer view of left chela; I, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.5386743" httpUri="https://zenodo.org/record/5386743/files/figure.png" pageId="10" pageNumber="574">Fig. 9G</figureCitation>
); carpus of chelipeds smooth, inner angle with broadly triangular tooth (
<figureCitation id="13012A63C66CFF99AE1DF8EA0DCAF897" box="[884,972,1806,1830]" captionStart="Fig" captionStartId="11.[136,171,2006,2028]" captionTargetBox="[218,1391,189,1991]" captionTargetId="figure-88@11.[218,1393,189,1991]" captionTargetPageId="11" captionText="Fig. 9. Lepidothelphusa padawan, new species, holotype male (13.6 × 12.5 mm) (ZRC 2015.0608), Kampung Sentah, Padawan, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, left cheliped showing serrated inner margin of merus; H, outer view of left chela; I, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.5386743" httpUri="https://zenodo.org/record/5386743/files/figure.png" pageId="10" pageNumber="574">Fig. 9A</figureCitation>
); fingers of adult major male chela strongly gaping (
<figureCitation id="13012A63C66CFF99AEECF8CA0DF5F8F7" box="[901,1011,1838,1862]" captionStart="Fig" captionStartId="11.[136,171,2006,2028]" captionTargetBox="[218,1391,189,1991]" captionTargetId="figure-88@11.[218,1393,189,1991]" captionTargetPageId="11" captionText="Fig. 9. Lepidothelphusa padawan, new species, holotype male (13.6 × 12.5 mm) (ZRC 2015.0608), Kampung Sentah, Padawan, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, left cheliped showing serrated inner margin of merus; H, outer view of left chela; I, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.5386743" httpUri="https://zenodo.org/record/5386743/files/figure.png" pageId="10" pageNumber="574">Fig. 9H, I</figureCitation>
); ambulatory legs relatively long, merus of fourth ambulatory legs subequal to length of carapace (
<figureCitation id="13012A63C66CFF99AE5CF88A0DB7F837" box="[821,945,1902,1926]" captionStart="Fig" captionStartId="11.[136,171,2006,2028]" captionTargetBox="[218,1391,189,1991]" captionTargetId="figure-88@11.[218,1393,189,1991]" captionTargetPageId="11" captionText="Fig. 9. Lepidothelphusa padawan, new species, holotype male (13.6 × 12.5 mm) (ZRC 2015.0608), Kampung Sentah, Padawan, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, left cheliped showing serrated inner margin of merus; H, outer view of left chela; I, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.5386743" httpUri="https://zenodo.org/record/5386743/files/figure.png" pageId="10" pageNumber="574">Fig. 9A, E</figureCitation>
); male abdomen triangular, somite 6 slightly longer than wide, subequal to length of telson (
<figureCitation id="13012A63C66CFF99A82CF86A0B98F817" box="[1349,1438,1934,1958]" captionStart="Fig" captionStartId="11.[136,171,2006,2028]" captionTargetBox="[218,1391,189,1991]" captionTargetId="figure-88@11.[218,1393,189,1991]" captionTargetPageId="11" captionText="Fig. 9. Lepidothelphusa padawan, new species, holotype male (13.6 × 12.5 mm) (ZRC 2015.0608), Kampung Sentah, Padawan, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, left cheliped showing serrated inner margin of merus; H, outer view of left chela; I, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.5386743" httpUri="https://zenodo.org/record/5386743/files/figure.png" pageId="10" pageNumber="574">Fig. 9C</figureCitation>
); G1 cylindrical, sinuous, gently curving outwards; terminal segment distinctly curving outwards (
<figureCitation id="13012A63C66CFF99A9A6F82A0B9BF857" box="[1231,1437,1998,2022]" captionStart="Fig" captionStartId="12.[113,148,1333,1355]" captionTargetBox="[111,1451,190,1316]" captionTargetId="figure-12@12.[95,1467,176,1328]" captionTargetPageId="12" captionText="Fig. 10. Lepidothelphusa padawan, new species, holotype male (13.6 × 12.5 mm) (ZRC 2015.0608), Kampung Sentah, Padawan, Sarawak. A, dorsal view of left G1; B, dorsal view of distal part of left G1; C, dorsal view of left G2; D, ventral view of left G1; E, ventral view of distal part of left G1. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5386745" httpUri="https://zenodo.org/record/5386745/files/figure.png" pageId="10" pageNumber="574">Fig. 10A, B, D, E</figureCitation>
); G2 with relatively long distal segment, about half length of basal segment (
<figureCitation id="13012A63C66CFF99AEBFF7EA0A3CF797" box="[982,1082,2062,2086]" captionStart="Fig" captionStartId="12.[113,148,1333,1355]" captionTargetBox="[111,1451,190,1316]" captionTargetId="figure-12@12.[95,1467,176,1328]" captionTargetPageId="12" captionText="Fig. 10. Lepidothelphusa padawan, new species, holotype male (13.6 × 12.5 mm) (ZRC 2015.0608), Kampung Sentah, Padawan, Sarawak. A, dorsal view of left G1; B, dorsal view of distal part of left G1; C, dorsal view of left G2; D, ventral view of left G1; E, ventral view of distal part of left G1. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5386745" httpUri="https://zenodo.org/record/5386745/files/figure.png" pageId="10" pageNumber="574">Fig. 10C</figureCitation>
).
</paragraph>
<caption id="DF45666EC66DFF98ADE1F8320DEFF7F2" ID-DOI="http://doi.org/10.5281/zenodo.5386743" ID-Zenodo-Dep="5386743" httpUri="https://zenodo.org/record/5386743/files/figure.png" pageId="11" pageNumber="575" startId="11.[136,171,2006,2028]" targetBox="[218,1391,189,1991]" targetPageId="11">
<paragraph id="8B8536E6C66DFF98ADE1F8320DEFF7F2" blockId="11.[136,1475,2006,2115]" pageId="11" pageNumber="575">
Fig. 9.
<taxonomicName id="4C3A4D65C66DFF98ADA4F8320FCAF85A" authority="Grinang &amp; Ng, 2015" authorityName="Grinang &amp; Ng" authorityYear="2015" box="[205,460,2006,2027]" class="Malacostraca" family="Gecarcinucidae" genus="Lepidothelphusa" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="575" phylum="Arthropoda" rank="species" species="padawan" status="sp. nov.">
<emphasis id="B94EEAF4C66DFF98ADA4F8320FCAF85A" box="[205,460,2006,2027]" italics="true" pageId="11" pageNumber="575">Lepidothelphusa padawan</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A27D578FC66DFF98ACB0F8330C49F85D" box="[473,591,2006,2028]" pageId="11" pageNumber="575" rank="species">new species</taxonomicNameLabel>
, holotype male (13.6 × 12.5 mm) (ZRC 2015.0608), Kampung Sentah, Padawan, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, left cheliped showing serrated inner margin of merus; H, outer view of left chela; I, outer view of right chela.
</paragraph>
</caption>
<caption id="DF45666EC66AFF9FAD18FAD10C31FA35" ID-DOI="http://doi.org/10.5281/zenodo.5386745" ID-Zenodo-Dep="5386745" httpUri="https://zenodo.org/record/5386745/files/figure.png" pageId="12" pageNumber="576" startId="12.[113,148,1333,1355]" targetBox="[111,1451,190,1316]" targetPageId="12">
<paragraph id="8B8536E6C66AFF9FAD18FAD10C31FA35" blockId="12.[113,1452,1332,1412]" pageId="12" pageNumber="576">
Fig. 10.
<taxonomicName id="4C3A4D65C66AFF9FADA9FAD10FB8FAFB" authority="Grinang &amp; Ng, 2015" authorityName="Grinang &amp; Ng" authorityYear="2015" box="[192,446,1333,1354]" class="Malacostraca" family="Gecarcinucidae" genus="Lepidothelphusa" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="576" phylum="Arthropoda" rank="species" species="padawan" status="sp. nov.">
<emphasis id="B94EEAF4C66AFF9FADA9FAD10FB8FAFB" box="[192,446,1333,1354]" italics="true" pageId="12" pageNumber="576">Lepidothelphusa padawan</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A27D578FC66AFF9FACA3FAD10C46FAFA" box="[458,576,1333,1355]" pageId="12" pageNumber="576" rank="species">new species</taxonomicNameLabel>
, holotype male (13.6 × 12.5 mm) (ZRC 2015.0608), Kampung Sentah, Padawan, Sarawak. A, dorsal view of left G1; B, dorsal view of distal part of left G1; C, dorsal view of left G2; D, ventral view of left G1; E, ventral view of distal part of left G1. Scale bars = 0.5 mm.
</paragraph>
</caption>
<paragraph id="8B8536E6C66AFF9FAD18FA5F0F52F942" blockId="12.[113,753,1467,1779]" pageId="12" pageNumber="576">
<emphasis id="B94EEAF4C66AFF9FAD18FA5F0ECEFA62" bold="true" box="[113,200,1467,1491]" pageId="12" pageNumber="576">Colour.</emphasis>
In life, the males have dark purple (almost black) ambulatory legs, white chelipeds, and at least three distinctive white-dark purple/black patterns on the carapace: i) carapace white or cream-white (anterior) and dark purple to black (posterior) equally or about equally horizontally (
<figureCitation id="13012A63C66AFF9FAFD1F9DF0EC0F9C2" captionStart="Fig" captionStartId="16.[113,148,1308,1330]" captionTargetBox="[113,1452,189,1293]" captionTargetId="figure-491@16.[113,1452,188,1294]" captionTargetPageId="16" captionText="Fig. 14. Live colours of three Lepidothelphusa species from Kuching Division, southwestern Sarawak. A, B, Lepidothelphusa loi, holotype male (10.3 × 9.6 mm) (ZRC 2015.0607), Gunung Ampungan, Sadong River Basin, Serian, Sarawak; C, D, Lepidothelphusa padawan, new species, paratype male (15.8 × 13.2 mm) (SBC.C.00440–4), Bung Brungu, Padawan, Sarawak; E, F, Lepidothelphusa sangon, new species, paratype male (8.0 × 7.0 mm) (SBC.C.00476), Nawang Waterfall, Bung Bratak, Bau, Sarawak." figureDoi="http://doi.org/10.5281/zenodo.5386759" httpUri="https://zenodo.org/record/5386759/files/figure.png" pageId="12" pageNumber="576">Figs. 14C, D</figureCitation>
,
<figureCitation id="13012A63C66AFF9FADBDF9BF0F21F9C2" box="[212,295,1627,1651]" captionStart="Fig" captionStartId="17.[135,170,1446,1468]" captionTargetBox="[135,1473,188,1430]" captionTargetId="figure-69@17.[134,1475,188,1430]" captionTargetPageId="17" captionText="Fig. 15. Live colours of Lepidothelphusa padawan, new species, paratypes (ZRC 2015.0609), showing variation. A, male (ca. 11.0 mm carapace width); B, male (ca. 11.0 mm carapace width); C, male (ca. 12.0 mm carapace width); D, male (ca. 12.0 mm carapace width); E, male (ca. 12.0 mm carapace width); F, female (ca. 9.0 mm carapace width). Kampung Sentah, Padawan, Sarawak" figureDoi="http://doi.org/10.5281/zenodo.5386761" httpUri="https://zenodo.org/record/5386761/files/figure.png" pageId="12" pageNumber="576">15A, B</figureCitation>
); ii) carapace white-dark purple to black broad “V” shaped (
<figureCitation id="13012A63C66AFF9FAC56F99F0FC7F922" box="[319,449,1659,1683]" captionStart="Fig" captionStartId="17.[135,170,1446,1468]" captionTargetBox="[135,1473,188,1430]" captionTargetId="figure-69@17.[134,1475,188,1430]" captionTargetPageId="17" captionText="Fig. 15. Live colours of Lepidothelphusa padawan, new species, paratypes (ZRC 2015.0609), showing variation. A, male (ca. 11.0 mm carapace width); B, male (ca. 11.0 mm carapace width); C, male (ca. 12.0 mm carapace width); D, male (ca. 12.0 mm carapace width); E, male (ca. 12.0 mm carapace width); F, female (ca. 9.0 mm carapace width). Kampung Sentah, Padawan, Sarawak" figureDoi="http://doi.org/10.5281/zenodo.5386761" httpUri="https://zenodo.org/record/5386761/files/figure.png" pageId="12" pageNumber="576">Fig. 15C, D</figureCitation>
); and iii) carapace almost or entirely white (
<figureCitation id="13012A63C66AFF9FAC79F97F0F69F902" box="[272,367,1691,1715]" captionStart="Fig" captionStartId="17.[135,170,1446,1468]" captionTargetBox="[135,1473,188,1430]" captionTargetId="figure-69@17.[134,1475,188,1430]" captionTargetPageId="17" captionText="Fig. 15. Live colours of Lepidothelphusa padawan, new species, paratypes (ZRC 2015.0609), showing variation. A, male (ca. 11.0 mm carapace width); B, male (ca. 11.0 mm carapace width); C, male (ca. 12.0 mm carapace width); D, male (ca. 12.0 mm carapace width); E, male (ca. 12.0 mm carapace width); F, female (ca. 9.0 mm carapace width). Kampung Sentah, Padawan, Sarawak" figureDoi="http://doi.org/10.5281/zenodo.5386761" httpUri="https://zenodo.org/record/5386761/files/figure.png" pageId="12" pageNumber="576">Fig. 15E</figureCitation>
). Female crabs are usually a uniform dark brown to purplish-brown to black, with cream-white chelipeds (
<figureCitation id="13012A63C66AFF9FAD8EF93F0F4EF942" box="[231,328,1755,1779]" captionStart="Fig" captionStartId="17.[135,170,1446,1468]" captionTargetBox="[135,1473,188,1430]" captionTargetId="figure-69@17.[134,1475,188,1430]" captionTargetPageId="17" captionText="Fig. 15. Live colours of Lepidothelphusa padawan, new species, paratypes (ZRC 2015.0609), showing variation. A, male (ca. 11.0 mm carapace width); B, male (ca. 11.0 mm carapace width); C, male (ca. 12.0 mm carapace width); D, male (ca. 12.0 mm carapace width); E, male (ca. 12.0 mm carapace width); F, female (ca. 9.0 mm carapace width). Kampung Sentah, Padawan, Sarawak" figureDoi="http://doi.org/10.5281/zenodo.5386761" httpUri="https://zenodo.org/record/5386761/files/figure.png" pageId="12" pageNumber="576">Fig. 15F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C320656DC66AFF9FAD18F8FF0EEFF8C2" pageId="12" pageNumber="576" type="etymology">
<paragraph id="8B8536E6C66AFF9FAD18F8FF0EEFF8C2" blockId="12.[113,752,1819,1907]" pageId="12" pageNumber="576">
<emphasis id="B94EEAF4C66AFF9FAD18F8FF0EF5F882" bold="true" box="[113,243,1819,1843]" pageId="12" pageNumber="576">Etymology.</emphasis>
The species is named after Padawan, the area where it was collected. The name is used as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C320656DC66AFF9FAD18F87F0AF5F942" pageId="12" pageNumber="576" type="discussion">
<paragraph id="8B8536E6C66AFF9FAD18F87F0B24F9E2" blockId="12.[113,755,1947,2099]" lastBlockId="12.[812,1452,1467,1619]" pageId="12" pageNumber="576">
<emphasis id="B94EEAF4C66AFF9FAD18F87F0EE6F802" bold="true" box="[113,224,1947,1971]" pageId="12" pageNumber="576">Remarks.</emphasis>
<taxonomicName id="4C3A4D65C66AFF9FAD80F87F0C00F802" authority="Grinang &amp; Ng, 2015" authorityName="Grinang &amp; Ng" authorityYear="2015" box="[233,518,1947,1971]" class="Malacostraca" family="Gecarcinucidae" genus="Lepidothelphusa" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="576" phylum="Arthropoda" rank="species" species="padawan" status="sp. nov.">
<emphasis id="B94EEAF4C66AFF9FAD80F87F0C00F802" box="[233,518,1947,1971]" italics="true" pageId="12" pageNumber="576">Lepidothelphusa padawan</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A27D578FC66AFF9FAF7FF8780C9AF802" box="[534,668,1947,1971]" pageId="12" pageNumber="576" rank="species">new species</taxonomicNameLabel>
, can be easily distinguished from
<taxonomicName id="4C3A4D65C66AFF9FACE7F8580C0CF862" authorityName="Nobili" authorityYear="1903" box="[398,522,1980,2003]" class="Malacostraca" family="Potamidae" genus="Potamon" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="576" phylum="Arthropoda" rank="species" sensu="stricto" species="cognettii">
<emphasis id="B94EEAF4C66AFF9FACE7F8580C0CF862" box="[398,522,1980,2003]" italics="true" pageId="12" pageNumber="576">L. cognettii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27D578FC66AFF9FAF78F8580C4AF862" box="[529,588,1980,2003]" pageId="12" pageNumber="576" sensu="stricto">s. str.</taxonomicNameLabel>
and congeners in having a distinctive white-dark purple to black body in life. This contrasting colour pattern is unique among known
<taxonomicName id="4C3A4D65C66AFF9FAD18F7FF0F87F782" box="[113,385,2075,2099]" class="Malacostraca" family="Gecarcinucidae" genus="Lepidothelphusa" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="576" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B94EEAF4C66AFF9FAD18F7FF0F25F782" box="[113,291,2075,2099]" italics="true" pageId="12" pageNumber="576">Lepidothelphusa</emphasis>
species.
</taxonomicName>
It also differs from other species in having proportionately the widest gape of fingers of the adult major male chela (
<figureCitation id="13012A63C66AFF9FA941FA3F0A7EFA42" box="[1064,1144,1499,1523]" captionStart="Fig" captionStartId="11.[136,171,2006,2028]" captionTargetBox="[218,1391,189,1991]" captionTargetId="figure-88@11.[218,1393,189,1991]" captionTargetPageId="11" captionText="Fig. 9. Lepidothelphusa padawan, new species, holotype male (13.6 × 12.5 mm) (ZRC 2015.0608), Kampung Sentah, Padawan, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, left cheliped showing serrated inner margin of merus; H, outer view of left chela; I, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.5386743" httpUri="https://zenodo.org/record/5386743/files/figure.png" pageId="12" pageNumber="576">Fig. 9H</figureCitation>
versus
<figureCitation id="13012A63C66AFF9FA9AFFA3F0B24FA42" box="[1222,1314,1499,1523]" captionStart="Fig" captionStartId="2.[113,149,2006,2028]" captionTargetBox="[190,1374,189,1991]" captionTargetId="figure-93@2.[190,1374,189,1991]" captionTargetPageId="2" captionText="Fig. 1. Lepidothelphusa cognettii (Nobili, 1903), male (12.2 × 10.2 mm) (ZRC 2015.0601), Batu Panggah Trail, Gunung Penrissen, Kuching, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, left cheliped showing serrated inner margin of merus; H, outer view of left chela; I, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.5386715" httpUri="https://zenodo.org/record/5386715/files/figure.png" pageId="12" pageNumber="576">Figs. 1H</figureCitation>
,
<figureCitation id="13012A63C66AFF9FA845FA3F0B49FA42" box="[1324,1359,1499,1523]" captionStart="Fig" captionStartId="5.[136,171,2006,2028]" captionTargetBox="[256,1353,189,1991]" captionTargetId="figure-90@5.[256,1354,189,1991]" captionTargetPageId="5" captionText="Fig. 3. Lepidothelphusa flavochela, new species, holotype male (8.1 × 7.2 mm) (ZRC 2015.0604), Pagar Besi, Kampung Gumbang, Bau, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, outer view of right chela; H, outer view of left chela; I, left cheliped showing serrated inner margin of merus." figureDoi="http://doi.org/10.5281/zenodo.5386723" httpUri="https://zenodo.org/record/5386723/files/figure.png" pageId="12" pageNumber="576">3H</figureCitation>
,
<figureCitation id="13012A63C66AFF9FA831FA3F0B7DFA42" box="[1368,1403,1499,1523]" captionStart="Fig" captionStartId="7.[136,171,2006,2028]" captionTargetBox="[177,1434,189,1991]" captionTargetId="figure-90@7.[177,1434,189,1991]" captionTargetPageId="7" captionText="Fig. 5. Lepidothelphusa limau, new species, holotype male (12.1 × 10.8 mm) (ZRC 2015.0605), Kampung Peros, Krokong, Bau, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, left cheliped showing serrated inner margin of merus; H, outer view of left chela; I, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.5386729" httpUri="https://zenodo.org/record/5386729/files/figure.png" pageId="12" pageNumber="576">5H</figureCitation>
,
<figureCitation id="13012A63C66AFF9FA8ECFA380BA1FA42" box="[1413,1447,1500,1523]" captionStart="Fig" captionStartId="9.[136,171,2006,2028]" captionTargetBox="[235,1376,189,1991]" captionTargetId="figure-91@9.[235,1376,189,1991]" captionTargetPageId="9" captionText="Fig. 7. Lepidothelphusa loi, new species, holotype male (10.3 × 9.6 mm) (ZRC 2015.0607), Gunung Ampungan, Sadong River Basin, Serian, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, left cheliped showing serrated inner margin of merus; H, outer view of left chela; I, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.5386737" httpUri="https://zenodo.org/record/5386737/files/figure.png" pageId="12" pageNumber="576">7H</figureCitation>
,
<figureCitation id="13012A63C66AFF9FAE44FA1F0D58F9A2" box="[813,862,1531,1555]" captionStart="Fig" captionStartId="13.[136,171,2006,2028]" captionTargetBox="[176,1431,189,1991]" captionTargetId="figure-90@13.[176,1431,189,1991]" captionTargetPageId="13" captionText="Fig. 11. Lepidothelphusa sangon, new species, holotype male (8.4 × 7.6 mm) (ZRC 2015.0611), Nawang Waterfall, Bung Bratak, Bau, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, left cheliped showing serrated inner margin of merus; H, outer view of left chela; I, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.5386749" httpUri="https://zenodo.org/record/5386749/files/figure.png" pageId="12" pageNumber="576">11H</figureCitation>
). The G1 of
<taxonomicName id="4C3A4D65C66AFF9FAE88FA180A59F9A2" authorityName="Grinang &amp; Ng" authorityYear="2015" box="[993,1119,1531,1555]" class="Malacostraca" family="Gecarcinucidae" genus="Lepidothelphusa" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="576" phylum="Arthropoda" rank="species" species="padawan">
<emphasis id="B94EEAF4C66AFF9FAE88FA180A59F9A2" box="[993,1119,1531,1555]" italics="true" pageId="12" pageNumber="576">L. padawan</emphasis>
</taxonomicName>
is diagnostic among congeners in being the most slender and having the terminal segment the most strongly curved (
<figureCitation id="13012A63C66AFF9FA924F9DF0B10F9E2" box="[1101,1302,1595,1619]" captionStart="Fig" captionStartId="12.[113,148,1333,1355]" captionTargetBox="[111,1451,190,1316]" captionTargetId="figure-12@12.[95,1467,176,1328]" captionTargetPageId="12" captionText="Fig. 10. Lepidothelphusa padawan, new species, holotype male (13.6 × 12.5 mm) (ZRC 2015.0608), Kampung Sentah, Padawan, Sarawak. A, dorsal view of left G1; B, dorsal view of distal part of left G1; C, dorsal view of left G2; D, ventral view of left G1; E, ventral view of distal part of left G1. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5386745" httpUri="https://zenodo.org/record/5386745/files/figure.png" pageId="12" pageNumber="576">Fig. 10A, B, D, E</figureCitation>
).
</paragraph>
<paragraph id="8B8536E6C66AFF9FAE44F99F0AF5F942" blockId="12.[813,1452,1659,1779]" pageId="12" pageNumber="576">
<emphasis id="B94EEAF4C66AFF9FAE44F99F0D8DF922" bold="true" box="[813,907,1659,1683]" pageId="12" pageNumber="576">Habitat.</emphasis>
The habitat is moist rocks and leaf litter, in partially open or shaded forest springs. The area is old primary dipterocarp forest with a mudstone substrate, and elevation is between
<quantity id="4CC29B03C66AFF9FAEC0F93F0A3AF942" box="[937,1084,1755,1779]" metricMagnitude="2" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="12" pageNumber="576" unit="m" value="200.0" valueMax="300.0" valueMin="100.0">100 to 300 m</quantity>
above sea level.
</paragraph>
</subSubSection>
</treatment>
</document>
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