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<document id="A9A098088A1120FE6F3881ECB4A130EC" ID-ISSN="1175­5334" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629706805705" checkinUser="felipe" docAuthor="Hara, Marcos Ryotaro &amp; Pinto-Da-Rocha, Ricardo" docDate="2010" docId="039387E5952B461BFF65FE3A88A5F802" docLanguage="en" docName="zt02698p136.pdf" docOrigin="Zootaxa 2698" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Eusarcus uruguayensis Hara &amp; Pinto-Da-Rocha 2010, comb. n." docType="treatment" docVersion="4" lastPageNumber="116" masterDocId="FFAAFF9D955B4668FFF2FFA78A56FF9F" masterDocTitle="Systematic review and cladistic analysis of the genus Eusarcus Perty 1833 (Arachnida, Opiliones, Gonyleptidae) 2698" masterLastPageNumber="136" masterPageNumber="1" pageNumber="113" updateTime="1699098874055" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="23F0B0576F6C408252E0791AD3CAC6AC" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F58EA321DB0F3263F2C5F32D335B04FF">
<mods:title id="4FB3A9A238BFB415C13D73303BD6E9D0">Systematic review and cladistic analysis of the genus Eusarcus Perty 1833 (Arachnida, Opiliones, Gonyleptidae) 2698</mods:title>
</mods:titleInfo>
<mods:name id="C8663EFEE19ADADDC5F4D6C1C1506C82" type="personal">
<mods:role id="8EF85D597CF13C8A507A8A6BC4E57B5D">
<mods:roleTerm id="BB8C8244F982276DF7DC1C0131521432">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="207EBA7E2BF33E78B0B12936532D0819">Hara, Marcos Ryotaro</mods:namePart>
</mods:name>
<mods:name id="539F3351209BD693C6448AC2A27F468C" type="personal">
<mods:role id="9CA7960618004271AEC786B4B0A92040">
<mods:roleTerm id="1555C039F69AD27F05B5C6E5B2DDCE7C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8F0E4207A9BEAEBA72D2CA9C2E9CAFD3">Pinto-Da-Rocha, Ricardo</mods:namePart>
</mods:name>
<mods:typeOfResource id="C1096962A4C1D885E7748CC7C2FD1785">text</mods:typeOfResource>
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<mods:title id="93E01762AB1E7D53CBA9E95EA864B0E4">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="5E2E57D9CB214976C46F6262E722B5A1">
<mods:date id="6288B904A7F596A03E8D42D1E0B8220D">2010</mods:date>
<mods:detail id="944812151C435F8DAF5B758F1D25778E" type="pubDate">
<mods:number id="707AD902167B76FBB892CA54079031A9">2010-12-03</mods:number>
</mods:detail>
<mods:detail id="93B21A256D8871DA09A8B07DE5C8D341" type="volume">
<mods:number id="8A0B378BE9F57360C8AB382C614BBC7D">2698</mods:number>
</mods:detail>
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<mods:start id="0879D1EAA8C958C20FED1C9120B3BA29">1</mods:start>
<mods:end id="9596FBFE14E0C627A008BCFF66477E65">136</mods:end>
</mods:extent>
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<mods:classification id="84EA54ADC5EACF13441F12503689F891">journal article</mods:classification>
<mods:identifier id="C052AEB269985A204C19C3679AF6E994" type="ISSN">1175­5334</mods:identifier>
</mods:mods>
<treatment id="039387E5952B461BFF65FE3A88A5F802" ID-DOI="http://doi.org/10.5281/zenodo.5295627" ID-Zenodo-Dep="5295627" LSID="urn:lsid:plazi:treatment:039387E5952B461BFF65FE3A88A5F802" httpUri="http://treatment.plazi.org/id/039387E5952B461BFF65FE3A88A5F802" lastPageId="115" lastPageNumber="116" pageId="112" pageNumber="113">
<subSubSection id="C3206578952B4618FF65FE3A894DFE28" box="[151,795,413,439]" pageId="112" pageNumber="113" type="nomenclature">
<paragraph id="8B8536F3952B4618FF65FE3A894DFE28" blockId="112.[151,795,413,474]" box="[151,795,413,439]" pageId="112" pageNumber="113">
<heading id="D0CD819F952B4618FF65FE3A894DFE28" bold="true" box="[151,795,413,439]" fontSize="11" level="1" pageId="112" pageNumber="113" reason="1">
<emphasis id="B94EEAE1952B4618FF65FE3A894DFE28" bold="true" box="[151,795,413,439]" pageId="112" pageNumber="113">
<taxonomicName id="4C3A4D70952B4618FF65FE3A8810FE28" ID-CoL="7W5SD" authority="(Ringuelet)" authorityName="Hara &amp; Pinto-Da-Rocha" authorityYear="2010" baseAuthorityName="Ringuelet" box="[151,582,413,439]" class="Arachnida" family="Gonyleptidae" genus="Eusarcus" kingdom="Animalia" order="Opiliones" pageId="112" pageNumber="113" phylum="Arthropoda" rank="species" species="uruguayensis" status="comb. nov.">
<emphasis id="B94EEAE1952B4618FF65FE3A8BE5FE28" bold="true" box="[151,435,413,439]" italics="true" pageId="112" pageNumber="113">Eusarcus uruguayensis</emphasis>
(Ringuelet)
</taxonomicName>
<taxonomicNameLabel id="A27D579A952B4618FDBFFE3A88EFFE28" box="[589,697,413,439]" pageId="112" pageNumber="113" rank="species">comb. n.</taxonomicNameLabel>
, stat. n.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3206578952B4618FF65FE678BC7FE45" box="[151,401,448,474]" pageId="112" pageNumber="113" type="description">
<paragraph id="8B8536F3952B4618FF65FE678BC7FE45" blockId="112.[151,795,413,474]" box="[151,401,448,474]" pageId="112" pageNumber="113">
(
<figureCitation id="13012A76952B4618FF6DFE678B57FE45" box="[159,257,448,474]" captionStart="FIGURE 37" captionStartId="113.[151,255,1887,1911]" captionTargetBox="[213,1375,194,1859]" captionTargetId="figure-89@113.[203,1384,189,1866]" captionTargetPageId="113" captionText="FIGURE 37. Eusarcus uruguayensis (Ringuelet) comb. n., stat. n. Male (holotype): A, habitus, dorsal view; B, dorsal scutum, right lateral view; C, right coxa and trochanter IV, posterior view (in part); D, right femur III, ventral view; E, left tibia III, ventral view; F, right trochanter and femur IV, dorsal view; G, idem, ventral view; H, left trochanter and femur IV, prolateral view; I, right tibia IV, ventral view. Scale bars, 1 mm." pageId="112" pageNumber="113">Figs. 37</figureCitation>
,
<figureCitation id="13012A76952B4618FEFCFE678B0BFE45" box="[270,349,448,474]" captionStart="FIGURE 46" captionStartId="122.[151,255,1386,1410]" captionTargetBox="[155,1431,207,1367]" captionTargetId="figure-237@122.[150,1436,197,1379]" captionTargetPageId="122" captionText="FIGURE 46. Distal part of penis in dorsal and lateral view: A–B, Eusarcus uruguayensis; C–D, E. gemignanii; E–F, E. sulcatus. Scale bars, 0.1 mm." pageId="112" pageNumber="113">46A,B</figureCitation>
,
<figureCitation id="13012A76952B4618FE98FE678BDEFE45" box="[362,392,448,474]" captionStart="FIGURE 51" captionStartId="127.[151,255,1031,1055]" captionTargetBox="[232,1362,180,1010]" captionTargetId="figure-89@127.[228,1362,165,1027]" captionTargetPageId="127" captionText="FIGURE 51. Localities of Eusarcus spp. in eastern Brazil, Uruguay, and northeastern Argentina." pageId="112" pageNumber="113">51</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3206578952B4618FF65FDAD8B69FD1E" pageId="112" pageNumber="113" type="reference_group">
<paragraph id="8B8536F3952B4618FF65FDAD8B69FD1E" blockId="112.[151,1436,522,641]" pageId="112" pageNumber="113">
<treatmentCitationGroup id="AB2A11DD952B4618FF65FDAD8B69FD1E" pageId="112" pageNumber="113">
<treatmentCitation id="0A9B10E2952B4618FF65FDAD8901FDBE" author="Ringuelet, R. A." box="[151,855,522,545]" page="292" pageId="112" pageNumber="113" year="1955">
<taxonomicName id="4C3A4D70952B4618FF65FDAD8901FDBE" authority="Ringuelet 1955 a: 292" authorityName="Ringuelet" authorityPageNumber="292" authorityYear="1955" box="[151,855,522,545]" class="Arachnida" family="Gonyleptidae" genus="Pygophalangodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="112" pageNumber="113" phylum="Arthropoda" rank="subSpecies" species="gemignanii" subSpecies="uruguayensis">
<emphasis id="B94EEAE1952B4618FF65FDAD883CFDBE" box="[151,618,522,545]" italics="true" pageId="112" pageNumber="113">Pygophalangodus gemignanii uruguayensis</emphasis>
<bibRefCitation id="EFAB4B02952B4618FD82FDAD8901FDBE" author="Ringuelet, R. A." box="[624,855,522,545]" pageId="112" pageNumber="113" pagination="279 - 297" refId="ref85995" refString="Ringuelet, R. A. (1955 a) Noticias sobre los opiliones del Uruguay. Notas del Museo de la Plata, 18 (163), 279 - 297." type="journal article" year="1955">Ringuelet 1955a: 292</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs. 2, 3, 295 (diag); 1963: 48 (cat), 49 (key);
<treatmentCitation id="0A9B10E2952B4618FABDFDAD8B25FDDE" author="Capocasale, R." page="632" pageId="112" pageNumber="113" year="1966">
<bibRefCitation id="EFAB4B02952B4618FABDFDAD8B25FDDE" author="Capocasale, R." pageId="112" pageNumber="113" pagination="631 - 644" refId="ref82540" refString="Capocasale, R. (1966) Opiliones del Uruguay. Discocyrtus prospicuus Holmberg, el alotipo hembra de Pygophalangodus gemignanii uruguayensis Ringuelet (Gonyleptidae) y Metalibitia rosascostai sp. nov. (Cosmetidae). Bulletin du Museum National d'Histoire Naturelle, (2) 37 (4), 631 - 644." type="journal article" year="1966">Capocasale 1966: 632</bibRefCitation>
</treatmentCitation>
(cit), 636 (descr fe mistakenly designated as “
<typeStatus id="54818851952B4618FC86FD8D8987FDDE" box="[884,977,554,577]" pageId="112" pageNumber="113" type="allotype">allotype</typeStatus>
”), fig. 2; 1968: 70 (distr);
<treatmentCitation id="0A9B10E2952B4618FB04FD8D8B4EFDFE" author="Maury, E. A. &amp; Pilati, A." page="1" pageId="112" pageNumber="113" year="1996">
<bibRefCitation id="EFAB4B02952B4618FB04FD8D8B4EFDFE" author="Maury, E. A. &amp; Pilati, A." pageId="112" pageNumber="113" pagination="1 - 7" refId="ref84214" refString="Maury, E. A. &amp; Pilati, A. (1996) Comensalismo de Riosegundo birabeni Canals, 1943 (Opiliones, Gonyleptidae) en hormigueros de Acromyrmex lobicornis (Emery, 1877) (Hymenoptera, Formicidae). Homenaje al Dr. Jose Maria Gallardo, Museo Argentino de Ciencias Naturales &quot; Bernardino Rivadavia &quot; e Instituto Nacional de Investigacion de las Ciencias Naturales, Extra, (n. s.) 142, 1 - 7." type="journal article" year="1996">Maury &amp; Pilati 1996: 1</bibRefCitation>
</treatmentCitation>
(cit);
<treatmentCitation id="0A9B10E2952B4618FEAEFDED8844FDFE" author="Kury, A. B." box="[348,530,586,609]" page="189" pageId="112" pageNumber="113" year="2003">
<bibRefCitation id="EFAB4B02952B4618FEAEFDED8844FDFE" author="Kury, A. B." box="[348,530,586,609]" pageId="112" pageNumber="113" pagination="5 - 337" refId="ref83946" refString="Kury, A. B. (2003 a) Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Iberica de Aracnologia, vol. esp. monografico n o 1, 5 - 337." type="journal article" year="2003">Kury 2003a: 189</bibRefCitation>
</treatmentCitation>
(cat); (ma
<typeStatus id="54818851952B4618FD78FDED88B1FDFE" box="[650,743,586,609]" pageId="112" pageNumber="113" type="holotype">holotype</typeStatus>
&amp; 1 ma
<typeStatus id="54818851952B4618FCBBFDED89F1FDFE" box="[841,935,586,609]" pageId="112" pageNumber="113" type="paratype">paratype</typeStatus>
, “
<collectingRegion id="49FEF811952B4618FC4EFDED8E5EFDFE" box="[956,1032,586,609]" country="Uruguay" name="Rivera" pageId="112" pageNumber="113">Rivera</collectingRegion>
,
<collectingRegion id="49FEF811952B4618FBE6FDED8E08FDFE" box="[1044,1118,586,609]" country="Uruguay" name="Rivera" pageId="112" pageNumber="113">Rivera</collectingRegion>
,
<collectingCountry id="F32D7663952B4618FB9BFDED8E92FDFE" box="[1129,1220,586,609]" name="Uruguay" pageId="112" pageNumber="113">Uruguai</collectingCountry>
;
<date id="FF841033952B4618FB3DFDED8F1AFDFE" box="[1231,1356,586,609]" pageId="112" pageNumber="113" value="1951-07-15">15.vii.1951</date>
; FCE”; examined).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3206578952B4618FF65FD088FC7FCC0" pageId="112" pageNumber="113" type="materials_examined">
<paragraph id="8B8536F3952B4618FF65FD088FC7FCC0" blockId="112.[151,1436,687,2020]" pageId="112" pageNumber="113">
<emphasis id="B94EEAE1952B4618FF65FD088BDEFD56" bold="true" box="[151,392,687,713]" pageId="112" pageNumber="113">Material examined</emphasis>
:
<materialsCitation id="3B523CAE952B4618FE6BFD178BB5FD70" collectingDate="1951-07-15" collectionCode="FCE" country="Uruguay" location="Rivera" pageId="112" pageNumber="113" specimenCount="2" stateProvince="Rivera" typeStatus="holotype">
<collectingCountry id="F32D7663952B4618FE6BFD17887CFD55" box="[409,554,688,714]" name="Uruguay" pageId="112" pageNumber="113">URUGUAY</collectingCountry>
.
<collectingRegion id="49FEF811952B4618FDC5FD1788D3FD55" box="[567,645,688,714]" country="Uruguay" name="Rivera" pageId="112" pageNumber="113">Rivera</collectingRegion>
:
<collectingRegion id="49FEF811952B4618FD64FD1788BEFD55" box="[662,744,688,714]" country="Uruguay" name="Rivera" pageId="112" pageNumber="113">Rivera</collectingRegion>
, without name of collector,
<date id="FF841033952B4618FBCBFD178E95FD55" box="[1081,1219,688,714]" pageId="112" pageNumber="113" value="1951-07-15">
<collectingDate id="EFC0E9DB952B4618FBCBFD178E95FD55" box="[1081,1219,688,714]" pageId="112" pageNumber="113" value="1951-07-15">15.vii.1951</collectingDate>
</date>
, ma
<typeStatus id="54818851952B4618FB0FFD178F33FD55" box="[1277,1381,688,714]" pageId="112" pageNumber="113" type="holotype">holotype</typeStatus>
&amp; 1 ma
<typeStatus id="54818851952B4618FF36FD728B7AFD70" box="[196,300,725,751]" pageId="112" pageNumber="113" type="paratype">paratype</typeStatus>
(
<collectionCode id="ED2BAE36952B4618FECCFD728B2AFD70" box="[318,380,725,751]" pageId="112" pageNumber="113">FCE</collectionCode>
). Arapej
</materialsCitation>
:
<materialsCitation id="3B523CAE952B4618FE07FD728E9AFD70" box="[501,1228,725,751]" collectingDate="1972-03-13" country="Uruguay" location="Salto" pageId="112" pageNumber="113" specimenCount="1" stateProvince="Salto" typeStatus="holotype">
<collectingRegion id="49FEF811952B4618FE07FD728860FD70" box="[501,566,725,751]" country="Uruguay" name="Salto" pageId="112" pageNumber="113">Salto</collectingRegion>
, without name of collector,
<date id="FF841033952B4618FC7DFD728E43FD70" box="[911,1045,725,751]" pageId="112" pageNumber="113" value="1972-03-13">
<collectingDate id="EFC0E9DB952B4618FC7DFD728E43FD70" box="[911,1045,725,751]" pageId="112" pageNumber="113" value="1972-03-13">13.iii.1972</collectingDate>
</date>
, 1 fe (URMU)
</materialsCitation>
;
<materialsCitation id="3B523CAE952B4618FB2EFD728FDBFCC0" collectingDate="1954-12-26" collectingDateMax="1969-03-02" collectingDateMin="1954-12-26" collectionCode="FCE" collectorName="Arroyo Cuaro, B. L. &amp; B. L. de Ximenez &amp; Sepulturas, C. S &amp; C. S. Carbonell &amp; P. San Martin" country="Uruguay" location="Picada del Negro Muerto" municipality="Rio Cuarem" pageId="112" pageNumber="113" specimenCode="MZSP 16056" specimenCount="1" stateProvince="Artigas" typeStatus="allotype">
<collectingRegion id="49FEF811952B4618FB2EFD728F62FD70" box="[1244,1332,725,751]" country="Uruguay" name="Artigas" pageId="112" pageNumber="113">Artigas</collectingRegion>
:
<collectorName id="26CF5325952B4618FAB7FD728B7AFC8B" pageId="112" pageNumber="113">Arroyo Cuaró, B.L.</collectorName>
de Ximénez leg.,
<date id="FF841033952B4618FDEAFD5D8825FC8B" box="[536,627,762,788]" pageId="112" pageNumber="113" value="1960-02">
<collectingDate id="EFC0E9DB952B4618FDEAFD5D8825FC8B" box="[536,627,762,788]" pageId="112" pageNumber="113" value="1960-02">ii.1960</collectingDate>
</date>
, 1 ma (
<collectionCode id="ED2BAE36952B4618FD26FD5D8942FC8B" box="[724,788,762,788]" pageId="112" pageNumber="113">FCE</collectionCode>
);
<collectorName id="26CF5325952B4618FCD9FD5D89A2FC8B" box="[811,1012,762,788]" pageId="112" pageNumber="113">Sepulturas, C.S</collectorName>
. Carbonell leg.,
<date id="FF841033952B4618FB3BFD5D8F0AFC8B" box="[1225,1372,762,788]" pageId="112" pageNumber="113" value="1957-12-19">
<collectingDate id="EFC0E9DB952B4618FB3BFD5D8F0AFC8B" box="[1225,1372,762,788]" pageId="112" pageNumber="113" value="1957-12-19">19.xii.1957</collectingDate>
</date>
, 1 fe mistankely designated as “
<typeStatus id="54818851952B4618FE3EFC878866FCA5" box="[460,560,800,826]" pageId="112" pageNumber="113" type="allotype">allotype</typeStatus>
” (
<collectionCode id="ED2BAE36952B4618FDBBFC8788D3FCA5" box="[585,645,800,826]" pageId="112" pageNumber="113">FCE</collectionCode>
); idem (
<collectingMunicipality id="6BE1AC89952B4618FD10FC878923FCA5" box="[738,885,800,826]" pageId="112" pageNumber="113">Rio Cuarem</collectingMunicipality>
,
<location id="8EE56028952B4618FC72FC878EFDFCA5" LSID="urn:lsid:plazi:treatment:039387E5952B461BFF65FE3A88A5F802:8EE56028952B4618FC72FC878EFDFCA5" box="[896,1195,800,826]" country="Uruguay" municipality="Rio Cuarem" name="Picada del Negro Muerto" pageId="112" pageNumber="113" stateProvince="Artigas">Picada del Negro Muerto</location>
),
<collectorName id="26CF5325952B4618FB4CFC878F0AFCA5" box="[1214,1372,800,826]" pageId="112" pageNumber="113">P. San Martin</collectorName>
<emphasis id="B94EEAE1952B4618FA90FC878FC3FCA6" box="[1378,1429,800,825]" italics="true" pageId="112" pageNumber="113">et al</emphasis>
. leg.,
<date id="FF841033952B4618FF23FCE28B15FCC0" box="[209,323,837,863]" pageId="112" pageNumber="113" value="1969-03-02">
<collectingDate id="EFC0E9DB952B4618FF23FCE28B15FCC0" box="[209,323,837,863]" pageId="112" pageNumber="113" value="1969-03-02">2.iii.1969</collectingDate>
</date>
, 1 ma &amp; 1 fe (
<specimenCode id="DB9C9E88952B4618FE19FCE288D8FCC0" box="[491,654,837,863]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="112" pageNumber="113" type="Museum">MZSP 16056</specimenCode>
); Tres Cruces, without name of collector,
<date id="FF841033952B4618FB8AFCE28F56FCC0" box="[1144,1280,837,863]" pageId="112" pageNumber="113" value="1954-12-26">
<collectingDate id="EFC0E9DB952B4618FB8AFCE28F56FCC0" box="[1144,1280,837,863]" pageId="112" pageNumber="113" value="1954-12-26">26.xii.1954</collectingDate>
</date>
, 2 fe (
<collectionCode id="ED2BAE36952B4618FABBFCE28FD3FCC0" box="[1353,1413,837,863]" pageId="112" pageNumber="113">FCE</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3206578952B4618FF34FCCD88A2FBA0" pageId="112" pageNumber="113" type="diagnosis">
<paragraph id="8B8536F3952B4618FF34FCCD88A2FBA0" blockId="112.[151,1436,687,2020]" pageId="112" pageNumber="113">
<emphasis id="B94EEAE1952B4618FF34FCCD8B1EFC1B" bold="true" box="[198,328,874,900]" pageId="112" pageNumber="113">Diagnosis:</emphasis>
<taxonomicName id="4C3A4D70952B4618FEA1FCCD8844FC1C" authorityName="Hara &amp; Pinto-Da-Rocha" authorityYear="2010" baseAuthorityName="Hara &amp; Pinto-Da-Rocha" baseAuthorityYear="2010" box="[339,530,874,900]" class="Arachnida" family="Gonyleptidae" genus="Eusarcus" kingdom="Animalia" order="Opiliones" pageId="112" pageNumber="113" phylum="Arthropoda" rank="species" species="uruguayensis">
<emphasis id="B94EEAE1952B4618FEA1FCCD8B33FC1C" box="[339,357,874,899]" italics="true" pageId="112" pageNumber="113">E</emphasis>
.
<emphasis id="B94EEAE1952B4618FE84FCCD8844FC1C" box="[374,530,874,899]" italics="true" pageId="112" pageNumber="113">uruguayensis</emphasis>
</taxonomicName>
resembles
<taxonomicName id="4C3A4D70952B4618FD6EFCCD890DFC1C" authorityName="Hara &amp; Pinto-Da-Rocha" authorityYear="2010" box="[668,859,874,900]" class="Arachnida" family="Gonyleptidae" genus="Eusarcus" kingdom="Animalia" order="Opiliones" pageId="112" pageNumber="113" phylum="Arthropoda" rank="species" species="caparaoensis">
<emphasis id="B94EEAE1952B4618FD6EFCCD88F8FC1C" box="[668,686,874,899]" italics="true" pageId="112" pageNumber="113">E</emphasis>
.
<emphasis id="B94EEAE1952B4618FD4DFCCD890DFC1C" box="[703,859,874,899]" italics="true" pageId="112" pageNumber="113">caparaoensis</emphasis>
</taxonomicName>
+ and
<taxonomicName id="4C3A4D70952B4618FC59FCCD8E70FC1C" authorityName="Hara &amp; Pinto-Da-Rocha" authorityYear="2010" box="[939,1062,874,900]" class="Arachnida" family="Gonyleptidae" genus="Eusarcus" kingdom="Animalia" order="Opiliones" pageId="112" pageNumber="113" phylum="Arthropoda" rank="species" species="manero">
<emphasis id="B94EEAE1952B4618FC59FCCD89EBFC1C" box="[939,957,874,899]" italics="true" pageId="112" pageNumber="113">E</emphasis>
.
<emphasis id="B94EEAE1952B4618FC3CFCCD8E70FC1C" box="[974,1062,874,899]" italics="true" pageId="112" pageNumber="113">manero</emphasis>
</taxonomicName>
+, which have a conical, short, blunt prolateral basal apophysis on the male trochanter IV, and can be distinguished by: The presence of PAM; ocularium high, with 2 large tubercles; without median spine on scutal area III; femur IV with dorso-median enlarged tubercles; prolateral row of enlarged, slightly high tubercles increasing in size to the middle; retrolateral row with a series of high, pointed tubercles in distal third, another with high, blunt tubercles in remaining two third, besides 2 ventro-apical spines.
</paragraph>
</subSubSection>
<subSubSection id="C3206578952B461BFF37FBED8993F8CD" lastPageId="115" lastPageNumber="116" pageId="112" pageNumber="113" type="description">
<paragraph id="8B8536F3952B4618FF37FBED8B5EFADB" blockId="112.[151,1436,687,2020]" pageId="112" pageNumber="113">
<emphasis id="B94EEAE1952B4618FF37FBED8BD4FBFB" bold="true" box="[197,386,1098,1124]" pageId="112" pageNumber="113">Redescription:</emphasis>
<emphasis id="B94EEAE1952B4618FE79FBEC8B9EFBFB" box="[395,456,1099,1124]" italics="true" pageId="112" pageNumber="113">Male</emphasis>
(
<typeStatus id="54818851952B4618FE28FBED881AFBFB" box="[474,588,1098,1124]" pageId="112" pageNumber="113" type="holotype">holotype</typeStatus>
): Dorsum (
<figureCitation id="13012A76952B4618FD26FBED8934FBFB" box="[724,866,1098,1124]" captionStart="FIGURE 37" captionStartId="113.[151,255,1887,1911]" captionTargetBox="[213,1375,194,1859]" captionTargetId="figure-89@113.[203,1384,189,1866]" captionTargetPageId="113" captionText="FIGURE 37. Eusarcus uruguayensis (Ringuelet) comb. n., stat. n. Male (holotype): A, habitus, dorsal view; B, dorsal scutum, right lateral view; C, right coxa and trochanter IV, posterior view (in part); D, right femur III, ventral view; E, left tibia III, ventral view; F, right trochanter and femur IV, dorsal view; G, idem, ventral view; H, left trochanter and femur IV, prolateral view; I, right tibia IV, ventral view. Scale bars, 1 mm." pageId="112" pageNumber="113">Fig. 37A,B</figureCitation>
): Measurements: SL 3.80; SMW 3.35; femur I 1.70; II 2.70; III 2.10; IV 2.80. Median paracheliceral projection of similar-size as PAM. Anterior margin of prosoma with 1 large tubercle on each corner. Ocularium near to anterior scutal margin; high, with 2 large tubercles and 12 small ones. Carapace with 42 scattered tubercles. Scutal area I with 11–12 scattered tubercles on each side; II with 16; III with 20; median spine absent; IV with 21 tubercles. Posterior margin of dorsal scutum and free tergites I–III with a row of 16, 15, 15, 11 tubercles, respectively. Anal operculum with 27 tubercles.
</paragraph>
<paragraph id="8B8536F3952B4618FF37FAF78805FA10" blockId="112.[151,1436,687,2020]" pageId="112" pageNumber="113">Venter: Coxa I with 17–18 tubercles, median and distal ones largest; II with 26–28 tubercles, distal ones largest; III–IV irregularly tuberculate.</paragraph>
<paragraph id="8B8536F3952B4618FF37FA3D8874FA2B" blockId="112.[151,1436,687,2020]" box="[197,546,1434,1460]" pageId="112" pageNumber="113">Chelicera: Segment I smooth.</paragraph>
<paragraph id="8B8536F3952B4618FF37FA6789DDF9BB" blockId="112.[151,1436,687,2020]" pageId="112" pageNumber="113">Pedipalpus: Trochanter dorsally inflated, with 2–3 tubercles. Femur dorsally with 3–4 aligned tubercles; ventrally with 1 large basal tubercle, 2–3 small pointed aligned ones. Tibial setation: Prolateral IiIi/IIi, retrolateral IiIi/iiIi. Tarsal setation: Prolateral IIi, retrolateral IiIi.</paragraph>
<paragraph id="8B8536F3952B461AFF37F9978E1AF9F4" blockId="112.[151,1436,687,2020]" lastBlockId="114.[151,1436,1577,2019]" lastPageId="114" lastPageNumber="115" pageId="112" pageNumber="113">
Legs (
<figureCitation id="13012A76952B4618FEE0F9978BCBF9D5" box="[274,413,1584,1610]" captionStart="FIGURE 37" captionStartId="113.[151,255,1887,1911]" captionTargetBox="[213,1375,194,1859]" captionTargetId="figure-89@113.[203,1384,189,1866]" captionTargetPageId="113" captionText="FIGURE 37. Eusarcus uruguayensis (Ringuelet) comb. n., stat. n. Male (holotype): A, habitus, dorsal view; B, dorsal scutum, right lateral view; C, right coxa and trochanter IV, posterior view (in part); D, right femur III, ventral view; E, left tibia III, ventral view; F, right trochanter and femur IV, dorsal view; G, idem, ventral view; H, left trochanter and femur IV, prolateral view; I, right tibia IV, ventral view. Scale bars, 1 mm." pageId="112" pageNumber="113">Fig. 37C–I</figureCitation>
): Coxa II with 2–3 tubercles on retrolateral margin; IV densely tuberculate, with 1 oblique prolateral apical apophysis, this conical, long, blunt, curved posteriad and ventrad, ventrally swollen in the middle and subapically (less conspicuous). Trochanters I–IV with scattered tubercles; I with 1 large ventro-basal tubercle; III with 1 large retrolateral apical tubercle; IV prolaterally with 1 basal apophysis, this conical, short, blunt, swollen in the middle; retrolaterally with 1 large apical tubercle. Femora I–II with reduced PDS, RDS; ventrally with 2 rows of slightly enlarged tubercles; III sinuous, with reduced PDS, small RDS; ventrally with a prolateral row of tubercles increasing in size distally, 1 PVS; IV sinuous, with enlarged dorso-median tubercles; prolateral row of enlarged, slightly high tubercles increasing in size to the middle; retrolateral row with a series of high, pointed tubercles in distal third, another with high, blunt tubercles in remaining two thirds; medium PDS, small RDS; ventrally with a prolateral row of enlarged tubercles increasing in size in distal third, becoming high, pointed, 2 apical spines. Tibia I ventrally with 2 rows of slightly enlarged tubercles; III ventrally with 2 rows of high, pointed tubercles increasing in size subapically; IV retrolaterally with a row of enlarged tubercles, with 1 apical spine; ventrally with 2 rows of tubercles increasing in size distally, 2 apical spines. Tarsal segmentation: 4–5(2–3), 6, 6, 6.
</paragraph>
<caption id="DF45667B952A4619FF65F8F889FAF849" pageId="113" pageNumber="114" startId="113.[151,255,1887,1911]" targetBox="[213,1375,194,1859]" targetPageId="113">
<paragraph id="8B8536F3952A4619FF65F8F889FAF849" blockId="113.[151,1436,1887,2006]" pageId="113" pageNumber="114">
<emphasis id="B94EEAE1952A4619FF65F8F88B7EF8E8" bold="true" box="[151,296,1887,1911]" pageId="113" pageNumber="114">FIGURE 37.</emphasis>
<taxonomicName id="4C3A4D70952A4619FEDDF8C788FFF8E9" authority="(Ringuelet)" authorityName="Hara &amp; Pinto-Da-Rocha" authorityYear="2010" baseAuthorityName="Hara &amp; Pinto-Da-Rocha" baseAuthorityYear="2010" box="[303,681,1887,1911]" class="Arachnida" family="Gonyleptidae" genus="Eusarcus" kingdom="Animalia" order="Opiliones" pageId="113" pageNumber="114" phylum="Arthropoda" rank="species" species="uruguayensis" status="comb. nov.">
<emphasis id="B94EEAE1952A4619FEDDF8C7887EF8E8" box="[303,552,1888,1911]" italics="true" pageId="113" pageNumber="114">Eusarcus uruguayensis</emphasis>
(Ringuelet)
</taxonomicName>
<emphasis id="B94EEAE1952A4619FD42F8F88938F8E8" bold="true" box="[688,878,1887,1911]" pageId="113" pageNumber="114">
<taxonomicNameLabel id="A27D579A952A4619FD42F8F88942F8E8" box="[688,788,1887,1911]" pageId="113" pageNumber="114" rank="species">comb. n.</taxonomicNameLabel>
, stat. n.
</emphasis>
Male (holotype): A, habitus, dorsal view; B, dorsal scutum, right lateral view; C, right coxa and trochanter IV, posterior view (in part); D, right femur III, ventral view; E, left tibia III, ventral view; F, right trochanter and femur IV, dorsal view; G, idem, ventral view; H, left trochanter and femur IV, prolateral view; I, right tibia IV, ventral view. Scale bars, 1 mm.
</paragraph>
</caption>
<caption id="DF45667B9529461AFF65FA278A9EFA68" pageId="114" pageNumber="115" startId="114.[151,257,1408,1432]" targetBox="[177,1415,191,1350]" targetPageId="114">
<paragraph id="8B8536F39529461AFF65FA278A9EFA68" blockId="114.[151,1436,1408,1527]" pageId="114" pageNumber="115">
<emphasis id="B94EEAE19529461AFF65FA278B7CFA07" bold="true" box="[151,298,1408,1432]" pageId="114" pageNumber="115">FIGURE 38.</emphasis>
Distal part of penis in dorsal and lateral view: A–B,
<taxonomicName id="4C3A4D709529461AFC89FA268E06FA07" box="[891,1104,1409,1432]" class="Arachnida" family="Gonyleptidae" genus="Eusarcus" kingdom="Animalia" order="Opiliones" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="schubarti">
<emphasis id="B94EEAE19529461AFC89FA268E06FA07" box="[891,1104,1409,1432]" italics="true" pageId="114" pageNumber="115">Eusarcus schubarti</emphasis>
</taxonomicName>
; C–D,
<taxonomicName id="4C3A4D709529461AFB50FA268F6CFA07" box="[1186,1338,1408,1432]" class="Arachnida" family="Gonyleptidae" genus="Eusarcus" kingdom="Animalia" order="Opiliones" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="teresincola">
<emphasis id="B94EEAE19529461AFB50FA268EE4FA07" box="[1186,1202,1409,1432]" italics="true" pageId="114" pageNumber="115">E</emphasis>
.
<emphasis id="B94EEAE19529461AFB33FA268F6CFA07" box="[1217,1338,1409,1432]" italics="true" pageId="114" pageNumber="115">teresincola</emphasis>
</taxonomicName>
; E–F,
<taxonomicName id="4C3A4D709529461AFA76FA268AB8FA27" class="Arachnida" family="Gonyleptidae" genus="Eusarcus" kingdom="Animalia" order="Opiliones" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="armatus">
<emphasis id="B94EEAE19529461AFA76FA268FC2FA07" box="[1412,1428,1409,1432]" italics="true" pageId="114" pageNumber="115">E</emphasis>
.
<emphasis id="B94EEAE19529461AFF65FA068AB8FA27" box="[151,238,1441,1464]" italics="true" pageId="114" pageNumber="115">armatus</emphasis>
</taxonomicName>
. Abbreviations: GL, glans; ST, stylus; TR, trunk; VPL, ventral plate; VPR, ventral process. Arrow in F indicates the angular medio-ventral projection of the stylus. Arrows in B, D indicate the medio-ventral trichomes. Scale bars, 0.1 mm.
</paragraph>
</caption>
<paragraph id="8B8536F39529461AFF37F9DE8EA2F942" blockId="114.[151,1436,1577,2019]" pageId="114" pageNumber="115">
Penis (MZSP 16056;
<figureCitation id="13012A769529461AFE39F9DE880FF90C" box="[459,601,1657,1683]" captionStart="FIGURE 46" captionStartId="122.[151,255,1386,1410]" captionTargetBox="[155,1431,207,1367]" captionTargetId="figure-237@122.[150,1436,197,1379]" captionTargetPageId="122" captionText="FIGURE 46. Distal part of penis in dorsal and lateral view: A–B, Eusarcus uruguayensis; C–D, E. gemignanii; E–F, E. sulcatus. Scale bars, 0.1 mm." pageId="114" pageNumber="115">Fig. 46A,B</figureCitation>
): Stylus with medio-ventral trichomes, apex fairly thin; with angular medio-ventral projection. Ventral process of glans curved; apex blunt. Ventral plate with concave sides; with 3 pairs of curved distal setae; 1 pair of short, straight median setae; 4 pairs of straight basal setae.
</paragraph>
<paragraph id="8B8536F39529461AFF37F94E8E75F8B7" blockId="114.[151,1436,1577,2019]" pageId="114" pageNumber="115">Coloration: Brown; dark brown on apical apophysis of coxa IV, leg IV and most part of metatarsus IV. Light brown in distal third of metatarsus IV, on remaining metatarsi and tarsi.</paragraph>
<paragraph id="8B8536F39529461BFF37F89488C2FEF3" blockId="114.[151,1436,1577,2019]" lastBlockId="115.[151,1436,152,364]" lastPageId="115" lastPageNumber="116" pageId="114" pageNumber="115">
<emphasis id="B94EEAE19529461AFF37F8948B48F8D3" box="[197,286,1843,1868]" italics="true" pageId="114" pageNumber="115">Female</emphasis>
(specimen mistakenly designated as “
<typeStatus id="548188519529461AFD18F8948906F8D2" box="[746,848,1843,1869]" pageId="114" pageNumber="115" type="allotype">allotype</typeStatus>
” of
<taxonomicName id="4C3A4D709529461AFC76F8948E87F8D3" authorityName="Ringuelet" authorityYear="1955" box="[900,1233,1843,1869]" class="Arachnida" family="Gonyleptidae" genus="Pygophalangodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="114" pageNumber="115" phylum="Arthropoda" rank="subSpecies" species="gemignanii" subSpecies="uruguayensis">
<emphasis id="B94EEAE19529461AFC76F89489C5F8D3" box="[900,915,1843,1868]" italics="true" pageId="114" pageNumber="115">P</emphasis>
.
<emphasis id="B94EEAE19529461AFC51F8948E87F8D3" box="[931,1233,1843,1868]" italics="true" pageId="114" pageNumber="115">gemignanii uruguayensis</emphasis>
</taxonomicName>
; FCE): Dorsum: Measurements: SL 3.65; SMW 3.10; femur I 1.60; II 2.60; III 2.00; IV 2.60. Median paracheliceral projection smaller than PAM. Chelicera: Segment I with 3 tubercles. Pedipalpus: Tibial setation: Prolateral and retrolateral IiIi. Legs: Coxa IV with 1 oblique prolateral apical apophysis, this conical, short. Trochanter II with scattered tubercles, 1 large ventro-basal tubercle; IV with scattered tubercles, retrolaterally with 1 large apical tubercle. Femur II with rows of similar-sized tubercles; III ventrally with 2 rows of tubercles increasing in size distally, 1 large prolateral apical tubercle; IV with rows of similar-sized tubercles; ventrally with a prolateral row of tubercles slightly increasing in size distally, 1 large prolateral apical tubercle. Tibiae I, III with rows of similar-sized tubercles; IV with retrolateral row of similar-sized tubercles, ventrally with 2 rows of tubercles slightly increasing in size distally. Tarsal segmentation: 5, 6, 6, 6. Coloration: Brown; dark brown on free tergites and edges of dorsal scutum.
</paragraph>
<caption id="DF45667B9528461BFF65F9E48EB7F9E5" pageId="115" pageNumber="116" startId="115.[151,255,1603,1627]" targetBox="[164,1433,425,1573]" targetPageId="115">
<paragraph id="8B8536F39528461BFF65F9E48EB7F9E5" blockId="115.[151,1436,1603,1659]" pageId="115" pageNumber="116">
<emphasis id="B94EEAE19528461BFF65F9E48B70F9C4" bold="true" box="[151,294,1603,1627]" pageId="115" pageNumber="116">FIGURE 39.</emphasis>
Distal part of penis in dorsal and lateral view: A–B,
<taxonomicName id="4C3A4D709528461BFCAAF9E48E13F9C5" box="[856,1093,1603,1626]" class="Arachnida" family="Gonyleptidae" genus="Eusarcus" kingdom="Animalia" order="Opiliones" pageId="115" pageNumber="116" phylum="Arthropoda" rank="species" species="oxyacanthus">
<emphasis id="B94EEAE19528461BFCAAF9E48E13F9C5" box="[856,1093,1603,1626]" italics="true" pageId="115" pageNumber="116">Eusarcus oxyacanthus</emphasis>
</taxonomicName>
; C–D,
<taxonomicName id="4C3A4D709528461BFB61F9E48F16F9C5" authorityName="Hara &amp; Pinto-Da-Rocha" authorityYear="2010" baseAuthorityName="Mello-Leitao" baseAuthorityYear="1927" box="[1171,1344,1603,1626]" class="Arachnida" family="Gonyleptidae" genus="Eusarcus" kingdom="Animalia" order="Opiliones" pageId="115" pageNumber="116" phylum="Arthropoda" rank="species" species="catharinensis">
<emphasis id="B94EEAE19528461BFB61F9E48EF5F9C5" box="[1171,1187,1603,1626]" italics="true" pageId="115" pageNumber="116">E</emphasis>
.
<emphasis id="B94EEAE19528461BFB42F9E48F16F9C5" box="[1200,1344,1603,1626]" italics="true" pageId="115" pageNumber="116">catharinensis</emphasis>
</taxonomicName>
; E–F,
<taxonomicName id="4C3A4D709528461BFA77F9E48AA1F9E5" authority="Hara &amp; Pinto-Da-Rocha, 2010" authorityName="Hara &amp; Pinto-Da-Rocha" authorityYear="2010" class="Arachnida" family="Gonyleptidae" genus="Eusarcus" kingdom="Animalia" order="Opiliones" pageId="115" pageNumber="116" phylum="Arthropoda" rank="species" species="mirabilis" status="sp. nov.">
<emphasis id="B94EEAE19528461BFA77F9E48FC3F9C5" box="[1413,1429,1603,1626]" italics="true" pageId="115" pageNumber="116">E</emphasis>
.
<emphasis id="B94EEAE19528461BFF65F9C48AA1F9E5" box="[151,247,1635,1658]" italics="true" pageId="115" pageNumber="116">mirabilis</emphasis>
</taxonomicName>
<emphasis id="B94EEAE19528461BFF0CF9C48B6CF9E4" bold="true" box="[254,314,1635,1659]" pageId="115" pageNumber="116">
<taxonomicNameLabel id="A27D579A9528461BFF0CF9C48B6CF9E4" box="[254,314,1635,1659]" pageId="115" pageNumber="116" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
Arrow indicates the angular medio-ventral projection of the stylus. Scale bars, 0.1 mm.
</paragraph>
</caption>
<paragraph id="8B8536F39528461BFF37F9048B2BF97D" blockId="115.[151,1436,1699,1949]" pageId="115" pageNumber="116">
<emphasis id="B94EEAE19528461BFF37F9048BF6F923" box="[197,416,1699,1724]" italics="true" pageId="115" pageNumber="116">Variation in males</emphasis>
(n=3): Measurements: SL 3.80–4.00; SMW 3.35–3.60; femur II 2.60–2.90; III 2.10– 2.15; IV 2.70–2.80.
</paragraph>
<paragraph id="8B8536F39528461BFF37F9498993F8CD" blockId="115.[151,1436,1699,1949]" pageId="115" pageNumber="116">
<emphasis id="B94EEAE19528461BFF37F9498BE3F898" box="[197,437,1774,1799]" italics="true" pageId="115" pageNumber="116">Variation in females</emphasis>
(n=5): Measurements: SL 3.60–4.10; SMW 3.00–3.20; femur I 1.50–1.70; II 2.30– 2.70; III 1.90–2.10; IV 2.40–2.70. Pedipalpus: Tibial setation: Prolateral IiIi/Iiii, retrolateral IiIi/iiIi. Femora I–II with reduced or small PDS; III with reduced or small PDS, RDS.
</paragraph>
</subSubSection>
<subSubSection id="C32065789528461BFF37F8FA88A5F802" pageId="115" pageNumber="116" type="materials_examined">
<paragraph id="8B8536F39528461BFF37F8FA88CDF8E7" blockId="115.[151,1436,1699,1949]" box="[197,667,1885,1912]" pageId="115" pageNumber="116">
<materialsCitation id="3B523CAE9528461BFF37F8FA88CEF8E7" box="[197,664,1885,1912]" country="Uruguay" location="Rivera" pageId="115" pageNumber="116" specimenCount="1" stateProvince="Rivera" typeStatus="holotype">
<emphasis id="B94EEAE19528461BFF37F8FA8B38F8E8" bold="true" box="[197,366,1885,1911]" pageId="115" pageNumber="116">
<typeStatus id="548188519528461BFF37F8FA8B54F8E8" box="[197,258,1885,1911]" pageId="115" pageNumber="116">Type</typeStatus>
locality:
</emphasis>
<collectingCountry id="F32D76639528461BFE84F8F98B8BF8E7" box="[374,477,1886,1912]" name="Uruguay" pageId="115" pageNumber="116">Uruguay</collectingCountry>
,
<collectingRegion id="49FEF8119528461BFE1BF8F9886CF8E7" box="[489,570,1886,1912]" country="Uruguay" name="Rivera" pageId="115" pageNumber="116">Rivera</collectingRegion>
,
<collectingRegion id="49FEF8119528461BFDB4F8F988CEF8E7" box="[582,664,1886,1912]" country="Uruguay" name="Rivera" pageId="115" pageNumber="116">Rivera</collectingRegion>
</materialsCitation>
.
</paragraph>
<paragraph id="8B8536F39528461BFF37F82588A5F802" blockId="115.[151,1436,1699,1949]" box="[197,755,1922,1949]" pageId="115" pageNumber="116">
<emphasis id="B94EEAE19528461BFF37F825885FF803" bold="true" box="[197,521,1922,1948]" pageId="115" pageNumber="116">Geographical distribution</emphasis>
(
<figureCitation id="13012A769528461BFDE8F8248827F802" box="[538,625,1923,1949]" captionStart="FIGURE 51" captionStartId="127.[151,255,1031,1055]" captionTargetBox="[232,1362,180,1010]" captionTargetId="figure-89@127.[228,1362,165,1027]" captionTargetPageId="127" captionText="FIGURE 51. Localities of Eusarcus spp. in eastern Brazil, Uruguay, and northeastern Argentina." pageId="115" pageNumber="116">Fig. 51</figureCitation>
):
<collectingCountry id="F32D76639528461BFD75F82488B9F802" box="[647,751,1923,1949]" name="Uruguay" pageId="115" pageNumber="116">Uruguay</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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