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<document id="983EE93696389F24D5191AE4CA5A8854" ID-CLB-Dataset="8333" ID-DOI="10.11646/zootaxa.4920.3.4" ID-GBIF-Dataset="37eefe05-4ff9-4762-a724-07deef527831" ID-ISSN="1175-5326" ID-Zenodo-Dep="4478107" ID-ZooBank="845CF8B2-07CD-45DC-8128-2EA56C825356" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1611900587808" checkinUser="plazi" docAuthor="Sommer, David, Hillert, Oliver, Hrůzová, Lucie &amp; Král, David" docDate="2021" docId="039387DBFF9EAD27D8DA80ECFBB6FED7" docLanguage="en" docName="zootaxa.4920.3.4.pdf" docOrigin="Zootaxa 4920 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Bolbelasmus (Bolbelasmus) makrisi Miessen 2011" docType="treatment" docVersion="9" lastPageNumber="385" masterDocId="FFAAFFA3FF9BAD22D84D8542FFA6F90D" masterDocTitle="Bolbelasmus (Bolbelasmus) zagrosensis (Coleoptera: Scarabaeoidea: Bolboceratidae), a new species from Iran, along with an updated key to the western Palaearctic species of the subgenus" masterLastPageNumber="394" masterPageNumber="380" pageNumber="385" updateTime="1698925811253" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="0F3A2F6B4FEE5E31C060B3E86CF5410F">Bolbelasmus (Bolbelasmus) zagrosensis (Coleoptera: Scarabaeoidea: Bolboceratidae), a new species from Iran, along with an updated key to the western Palaearctic species of the subgenus</mods:title>
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<bibRefCitation id="EFAB4B3CFF9EAD27DA1F80EDFD58FCC4" author="Miessen, G." box="[594,766,1455,1481]" pageId="5" pageNumber="385" pagination="109 - 119" refId="ref8514" refString="Miessen, G. (2011) Quelques commentaires sur le genre Bolbelasmus Boucomont, 1911 et description d'une nouvelle espece de Chypre. Lambillionea, 111, 109 - 119." type="journal article" year="2011">Miessen, 2011</bibRefCitation>
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(
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,
<figureCitation id="13012A48FF9EAD27D9408091FE97FCE0" box="[269,305,1491,1517]" captionStart="FIGURE 2" captionStartId="3.[151,250,1151,1175]" captionTargetBox="[157,1426,190,1120]" captionTargetId="figure-20@3.[151,1436,181,1127]" captionTargetPageId="3" captionText="FIGURE 2AD. Pronotum of males in frontal view. A, holotype of Bolbelasmus zagrosensis Sommer, Hillert, Hrůzová &amp; Král, new species; B, B. makrisi Miessen, 2011; C, B. nireus (Reitter, 1895) from Greece (Rhodes); D, B. nireus from Turkey. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4478113" httpUri="https://zenodo.org/record/4478113/files/figure.png" pageId="5" pageNumber="385">2B</figureCitation>
,
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,
<figureCitation id="13012A48FF9EAD27D9268091FE28FCE0" box="[363,398,1491,1517]" captionStart="FIGURE 4" captionStartId="5.[151,250,1277,1301]" captionTargetBox="[158,1426,188,1244]" captionTargetId="figure-20@5.[151,1436,181,1253]" captionTargetPageId="5" captionText="FIGURE 4AD. Pronotum of males in lateral view. A, Bolbelasmus zagrosensis Sommer, Hillert, Hrůzová &amp; Král, new species, from Fars province, Iran; B, B. zagrosensis Sommer, Hillert, Hrůzová &amp; Král, new species, from Lorestan province, Iran; C, B. makrisi Miessen, 2011; D, B. nireus (Reitter, 1895) from Greece, Rhodes. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4478117" httpUri="https://zenodo.org/record/4478117/files/figure.png" pageId="5" pageNumber="385">4C</figureCitation>
,
<figureCitation id="13012A48FF9EAD27D9D48091FE1AFCE3" box="[409,444,1491,1518]" captionStart="FIGURE 5" captionStartId="6.[151,250,986,1010]" captionTargetBox="[155,1428,185,958]" captionTargetId="figure-17@6.[151,1436,181,962]" captionTargetPageId="6" captionText="FIGURE 5AC. Parameres in dorsal view. A, Bolbelasmus zagrosensis Sommer, Hillert, Hrůzová &amp; Král, new species; B, B. makrisi Miessen, 2011; C, B. nireus (Reitter, 1895). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4478119" httpUri="https://zenodo.org/record/4478119/files/figure.png" pageId="5" pageNumber="385">5B</figureCitation>
,
<figureCitation id="13012A48FF9EAD27D9858091FE71FCE0" box="[456,471,1491,1517]" captionStart="FIGURE 9" captionStartId="10.[151,250,940,964]" captionTargetBox="[170,1418,187,912]" captionTargetId="figure-17@10.[164,1424,181,916]" captionTargetPageId="10" captionText="FIGURE 9. Sketch map with marked distribution of Bolbelasmus makrisi Miessen, 2011 (green), B. nireus (Reitter, 1895) (yellow) and B. zagrosensis Sommer, Hillert, Hrůzová &amp; Král, new species (red). Squares represents the type locality of each species." figureDoi="http://doi.org/10.5281/zenodo.4478127" httpUri="https://zenodo.org/record/4478127/files/figure.png" pageId="5" pageNumber="385">9</figureCitation>
)
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, figs. on pp. 116, 118119 [not numbered] (
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, 1 ♁
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(
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), “Rizoelia nat. for. Park | Larnaka—CYPRUS || leg.:
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| collection:
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<taxonomicName id="4C3A4D4EFF9EAD27DAC383E2FCDDFFB7" authority="Sommer &amp; Hillert &amp; Hrůzová &amp; Král, 2021" authorityName="Sommer &amp; Hillert &amp; Hrůzová &amp; Král" authorityYear="2021" box="[654,891,1696,1722]" class="Insecta" family="Bolboceratidae" genus="Bolbelasmus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="385" phylum="Arthropoda" rank="species" species="makrisi" status="sp. nov.">Bolbelasmus makrisi</taxonomicName>
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det. 2011 |
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[printed, red label]”
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.
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<emphasis id="B94EEADFFF9EAD27D88A83AAFD93FE0F" bold="true" box="[199,565,1768,1794]" pageId="5" pageNumber="385">Additional material examined.</emphasis>
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<emphasis id="B94EEADFFF9EAD27DA7083AAFD39FE0F" bold="true" box="[573,671,1768,1794]" pageId="5" pageNumber="385">
<collectingCountry id="F32D765DFF9EAD27DA7083AAFD3CFE0F" box="[573,666,1768,1794]" name="Cyprus" pageId="5" pageNumber="385">Cyprus</collectingCountry>
:
</emphasis>
1 ♁
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(
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),
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,
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<quantity id="4CC29B28FF9EAD27DC4683AAFBE2FE0F" box="[1035,1092,1768,1795]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="385" unit="km" value="5.0">5 km</quantity>
N of Lara
</locationDeviation>
,
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<collectingDate id="EFC0E9E5FF9EAD27DC8583AAFAD3FE0E" box="[1224,1397,1768,1795]" pageId="5" pageNumber="385" value="2015-03-13" valueMax="2015-03-20" valueMin="2015-03-13">13.20.iii.2015</collectingDate>
</date>
,
<collectorName id="26CF531BFF9EAD27DDCC83AAFF46FE2B" pageId="5" pageNumber="385">O. Hillert</collectorName>
lgt.;
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1 ♁ (
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),
<collectorName id="26CF531BFF9EAD27D989824EFDEBFE2A" box="[452,589,1804,1831]" pageId="5" pageNumber="385">Akamas NP</collectorName>
,
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,
<date id="FF84100DFF9EAD27DAD0824EFCECFE2A" box="[669,842,1804,1831]" pageId="5" pageNumber="385" value="2015-03-13" valueMax="2015-03-20" valueMin="2015-03-13">
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</date>
,
<collectorName id="26CF531BFF9EAD27DB1A824EFC63FE2B" box="[855,965,1804,1830]" pageId="5" pageNumber="385">O. Hillert</collectorName>
lgt.;
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1 ♁ (
<collectionCode id="ED2BAE08FF9EAD27DC0C824EFB32FE2B" box="[1089,1172,1804,1830]" pageId="5" pageNumber="385">OHCB</collectionCode>
),
<collectorName id="26CF531BFF9EAD27DCE7824EFB5FFE2B" box="[1194,1273,1804,1830]" pageId="5" pageNumber="385">Miliou</collectorName>
, N of
<collectingRegion id="49FEF82FFF9EAD27DD09824EFA3EFE2B" box="[1348,1432,1804,1830]" country="Cyprus" name="Pafos" pageId="5" pageNumber="385">Paphos</collectingRegion>
,
<date id="FF84100DFF9EAD27D8DA8272FEE2FE46" box="[151,324,1840,1867]" pageId="5" pageNumber="385" value="2015-03-13" valueMax="2015-03-20" valueMin="2015-03-13">
<collectingDate id="EFC0E9E5FF9EAD27D8DA8272FEE2FE46" box="[151,324,1840,1867]" pageId="5" pageNumber="385" value="2015-03-13" valueMax="2015-03-20" valueMin="2015-03-13">13.20.iii.2015</collectingDate>
</date>
,
<collectorName id="26CF531BFF9EAD27D91F8272FE19FE47" box="[338,447,1840,1866]" pageId="5" pageNumber="385">O. Hillert</collectorName>
lgt.;
</materialsCitation>
<materialsCitation id="3B523C90FF9EAD27D9B38272FDE7FE63" ID-GBIF-Occurrence="3033773304" collectingDate="2014-06-07" collectingDateMax="2015-03-20" collectingDateMin="2014-06-07" collectionCode="OHCB, UV" collectorName="G. Miessen" country="Cyprus" latitude="34.93935" location="Lara" longLatPrecision="1" longitude="32.45989" pageId="5" pageNumber="385" specimenCount="1" specimenCount-female="1" stateProvince="Pafos">
1 ♁ (
<collectionCode id="ED2BAE08FF9EAD27DA718272FD29FE47" box="[572,655,1840,1866]" pageId="5" pageNumber="385">OHCB</collectionCode>
),
<collectorName id="26CF531BFF9EAD27DAE88272FD52FE47" box="[677,756,1840,1866]" pageId="5" pageNumber="385">Miliou</collectorName>
,
<geoCoordinate id="EE0E500AFF9EAD27DB4C8272FC16FE46" box="[769,944,1840,1867]" degrees="34" direction="north" minutes="56" orientation="latitude" pageId="5" pageNumber="385" precision="1" seconds="22.23" value="34.93951">34°5622.23”N</geoCoordinate>
,
<geoCoordinate id="EE0E500AFF9EAD27DBF38272FBCCFE46" box="[958,1130,1840,1867]" degrees="32" direction="east" minutes="27" orientation="longitude" pageId="5" pageNumber="385" precision="1" seconds="35.90" value="32.459972">32°2735.90”E</geoCoordinate>
,
<date id="FF84100DFF9EAD27DC3A8272FB42FE47" box="[1143,1252,1840,1866]" pageId="5" pageNumber="385" value="2014-06-07">
<collectingDate id="EFC0E9E5FF9EAD27DC3A8272FB42FE47" box="[1143,1252,1840,1866]" pageId="5" pageNumber="385" value="2014-06-07">7.vi.2014</collectingDate>
</date>
, vivant dans piscine [alive in pool],
<collectorName id="26CF531BFF9EAD27D9C68216FDADFE63" box="[395,523,1876,1902]" pageId="5" pageNumber="385">G. Miessen</collectorName>
lgt.;
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<materialsCitation id="3B523C90FF9EAD27DA048216FD68FE9F" ID-GBIF-Occurrence="3033773308" collectingDate="2014-06-07" collectingDateMax="2015-03-20" collectingDateMin="2014-06-07" collectionCode="OHCB, UV" collectorName="G. Miessen" country="Cyprus" latitude="34.93935" location="Lara" longLatPrecision="1" longitude="32.45989" pageId="5" pageNumber="385" specimenCount="1" specimenCount-female="1" stateProvince="Pafos">
1 ♁ (
<collectionCode id="ED2BAE08FF9EAD27DAC88216FD7EFE63" box="[645,728,1876,1902]" pageId="5" pageNumber="385">OHCB</collectionCode>
),
<collectorName id="26CF531BFF9EAD27DAA08216FC9AFE63" box="[749,828,1876,1902]" pageId="5" pageNumber="385">Miliou</collectorName>
,
<geoCoordinate id="EE0E500AFF9EAD27DB048216FC51FE62" box="[841,1015,1876,1903]" degrees="34" direction="north" minutes="56" orientation="latitude" pageId="5" pageNumber="385" precision="1" seconds="21.66" value="34.93935">34°5621.66”N</geoCoordinate>
,
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,
<date id="FF84100DFF9EAD27DCF08216FA8CFE63" box="[1213,1322,1876,1902]" pageId="5" pageNumber="385" value="2014-06-07">
<collectingDate id="EFC0E9E5FF9EAD27DCF08216FA8CFE63" box="[1213,1322,1876,1902]" pageId="5" pageNumber="385" value="2014-06-07">7.vi.2014</collectingDate>
</date>
, attire par lampe UV [attracted by
<collectionCode id="ED2BAE08FF9EAD27D9EE823BFE6DFE9E" box="[419,459,1913,1939]" country="Colombia" name="Departamento de Biologia de la Universidad del Valle" pageId="5" pageNumber="385">UV</collectionCode>
light],
<collectorName id="26CF531BFF9EAD27DA51823AFD3DFE9F" box="[540,667,1912,1938]" pageId="5" pageNumber="385">G. Miessen</collectorName>
lgt.;
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<materialsCitation id="3B523C90FF9EAD27DA99823AFC6FFEBB" ID-GBIF-Occurrence="3033773305" collectingDate="2014-06-07" collectingDateMax="2015-03-20" collectingDateMin="2014-06-07" collectionCode="OHCB, UV" collectorName="O. Hillert " country="Cyprus" latitude="34.93935" location="Lara" longLatPrecision="1" longitude="32.45989" pageId="5" pageNumber="385" specimenCount="1" specimenCount-female="1" stateProvince="Pafos">
16 ♁♁
<specimenCount id="9D3CFD44FF9EAD27DB6A823AFCD2FE9E" box="[807,884,1912,1939]" pageId="5" pageNumber="385" type="female">15 ♀♀</specimenCount>
(
<collectionCode id="ED2BAE08FF9EAD27DBCF823AFC75FE9F" box="[898,979,1912,1938]" pageId="5" pageNumber="385">OHCB</collectionCode>
) (
<figureCitation id="13012A48FF9EAD27DBA7823AFBEDFE9F" box="[1002,1099,1912,1938]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[166,1419,193,1903]" captionTargetId="figure-17@2.[153,1435,179,1918]" captionTargetPageId="2" captionText="FIGURE 1AD. Habitus of males in dorsal view. A, holotype of Bolbelasmus zagrosensis Sommer, Hillert, Hrůzová &amp; Král, new species; B, B. makrisi Miessen, 2011; C, B. nireus (Reitter, 1895) from Greece (Rhodes); D, B. nireus from Turkey. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4478111" httpUri="https://zenodo.org/record/4478111/files/figure.png" pageId="5" pageNumber="385">Figs. 1B</figureCitation>
,
<figureCitation id="13012A48FF9EAD27DC1A823AFBDCFE9F" box="[1111,1146,1912,1938]" captionStart="FIGURE 2" captionStartId="3.[151,250,1151,1175]" captionTargetBox="[157,1426,190,1120]" captionTargetId="figure-20@3.[151,1436,181,1127]" captionTargetPageId="3" captionText="FIGURE 2AD. Pronotum of males in frontal view. A, holotype of Bolbelasmus zagrosensis Sommer, Hillert, Hrůzová &amp; Král, new species; B, B. makrisi Miessen, 2011; C, B. nireus (Reitter, 1895) from Greece (Rhodes); D, B. nireus from Turkey. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4478113" httpUri="https://zenodo.org/record/4478113/files/figure.png" pageId="5" pageNumber="385">2B</figureCitation>
,
<figureCitation id="13012A48FF9EAD27DCC8823AFB0EFE9F" box="[1157,1192,1912,1938]" captionStart="FIGURE 3" captionStartId="4.[151,250,1926,1950]" captionTargetBox="[151,1425,535,1892]" captionTargetId="figure-159@4.[151,1436,525,1902]" captionTargetPageId="4" captionText="FIGURE 3AD. Habitus of males in lateral view. A, holotype of Bolbelasmus zagrosensis Sommer, Hillert, Hrůzová &amp; Král, new species; B, B. makrisi Miessen, 2011; C, B. nireus (Reitter, 1895) from Greece (Rhodes); D, B. nireus from Turkey. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4478115" httpUri="https://zenodo.org/record/4478115/files/figure.png" pageId="5" pageNumber="385">3B</figureCitation>
,
<figureCitation id="13012A48FF9EAD27DCFE823AFB70FE9F" box="[1203,1238,1912,1938]" captionStart="FIGURE 4" captionStartId="5.[151,250,1277,1301]" captionTargetBox="[158,1426,188,1244]" captionTargetId="figure-20@5.[151,1436,181,1253]" captionTargetPageId="5" captionText="FIGURE 4AD. Pronotum of males in lateral view. A, Bolbelasmus zagrosensis Sommer, Hillert, Hrůzová &amp; Král, new species, from Fars province, Iran; B, B. zagrosensis Sommer, Hillert, Hrůzová &amp; Král, new species, from Lorestan province, Iran; C, B. makrisi Miessen, 2011; D, B. nireus (Reitter, 1895) from Greece, Rhodes. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4478117" httpUri="https://zenodo.org/record/4478117/files/figure.png" pageId="5" pageNumber="385">4C</figureCitation>
,
<figureCitation id="13012A48FF9EAD27DCAC823AFAA3FE9E" box="[1249,1285,1912,1939]" captionStart="FIGURE 5" captionStartId="6.[151,250,986,1010]" captionTargetBox="[155,1428,185,958]" captionTargetId="figure-17@6.[151,1436,181,962]" captionTargetPageId="6" captionText="FIGURE 5AC. Parameres in dorsal view. A, Bolbelasmus zagrosensis Sommer, Hillert, Hrůzová &amp; Král, new species; B, B. makrisi Miessen, 2011; C, B. nireus (Reitter, 1895). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4478119" httpUri="https://zenodo.org/record/4478119/files/figure.png" pageId="5" pageNumber="385">5B</figureCitation>
), 2 ♁♁
<specimenCount id="9D3CFD44FF9EAD27DD13823AFA3AFE9E" box="[1374,1436,1912,1939]" pageId="5" pageNumber="385" type="female">2 ♀♀</specimenCount>
(
<collectionCode id="ED2BAE08FF9EAD27D8D282DEFF52FEBB" box="[159,244,1948,1974]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="5" pageNumber="385">NMPC</collectionCode>
),
<quantity id="4CC29B28FF9EAD27D94A82DEFEE6FEBB" box="[263,320,1948,1975]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="385" unit="km" value="5.0">5 km</quantity>
NE of
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,
<collectingRegion id="49FEF82FFF9EAD27D9A782DEFD9DFEBB" box="[490,571,1948,1974]" country="Cyprus" name="Pafos" pageId="5" pageNumber="385">Paphos</collectingRegion>
env.,
<date id="FF84100DFF9EAD27DA3382DEFC8CFEBA" box="[638,810,1948,1975]" pageId="5" pageNumber="385" value="2015-03-13" valueMax="2015-03-20" valueMin="2015-03-13">
<collectingDate id="EFC0E9E5FF9EAD27DA3382DEFC8CFEBA" box="[638,810,1948,1975]" pageId="5" pageNumber="385" value="2015-03-13" valueMax="2015-03-20" valueMin="2015-03-13">13.20.iii.2015</collectingDate>
</date>
,
<collectorName id="26CF531BFF9EAD27DB7B82DEFC07FEBB" box="[822,929,1948,1974]" pageId="5" pageNumber="385">O. Hillert</collectorName>
lgt
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3206546FF9EAD27D88A8282FBB6FED7" box="[199,1040,1984,2011]" pageId="5" pageNumber="385" type="distribution">
<paragraph id="8B8536CDFF9EAD27D88A8282FBB6FED7" blockId="5.[151,1436,1660,2011]" box="[199,1040,1984,2011]" pageId="5" pageNumber="385">
<emphasis id="B94EEADFFF9EAD27D88A8282FEC7FED7" bold="true" box="[199,353,1984,2010]" pageId="5" pageNumber="385">Distribution.</emphasis>
Endemic to
<collectingCountry id="F32D765DFF9EAD27D9A28282FDE6FED7" box="[495,576,1984,2010]" name="Cyprus" pageId="5" pageNumber="385">Cyprus</collectingCountry>
(
<bibRefCitation id="EFAB4B3CFF9EAD27DA038282FC98FED6" author="Nikolajev, G. V. &amp; Kral, D. &amp; Bezdek, A." box="[590,830,1984,2011]" pageId="5" pageNumber="385" pagination="1802" refId="ref8582" refString="Nikolajev, G. V., Kral, D. &amp; Bezdek, A. (2016) Family Geotrupidae Latreille, 1802. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 3. Scarabaeoidea-Scirtoidea-Dasciloidea-Buprestoidea-Byrrhoidea. Revised and Updated Edition. E. J. Brill, Leiden, pp. 33 - 52. https: // doi. org / 10.1163 / 9789004309142 _ 003" type="book chapter" year="2016">
Nikolajev
<emphasis id="B94EEADFFF9EAD27DA888283FD58FED7" box="[709,766,1984,2010]" italics="true" pageId="5" pageNumber="385">et al.</emphasis>
2016
</bibRefCitation>
) (see map,
<figureCitation id="13012A48FF9EAD27DBF28282FBA4FED7" box="[959,1026,1984,2010]" captionStart="FIGURE 9" captionStartId="10.[151,250,940,964]" captionTargetBox="[170,1418,187,912]" captionTargetId="figure-17@10.[164,1424,181,916]" captionTargetPageId="10" captionText="FIGURE 9. Sketch map with marked distribution of Bolbelasmus makrisi Miessen, 2011 (green), B. nireus (Reitter, 1895) (yellow) and B. zagrosensis Sommer, Hillert, Hrůzová &amp; Král, new species (red). Squares represents the type locality of each species." figureDoi="http://doi.org/10.5281/zenodo.4478127" httpUri="https://zenodo.org/record/4478127/files/figure.png" pageId="5" pageNumber="385">Fig. 9</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>