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<document id="A18A34513B669DFE7C72B3817030C506" ID-DOI="10.1080/00222930500145123" ID-ISSN="1464-5262" ID-Zenodo-Dep="4581703" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614874722613" checkinUser="carolina" docAuthor="Huber, Bernhard A. &amp; Benjamin, Suresh P." docDate="2005" docId="039387D9FFCCFFF4C2BC1C59FDFE6E68" docLanguage="en" docName="JNATHIST.39.37-3305-3319.pdf" docOrigin="Journal of Natural History 39 (37)" docSource="http://dx.doi.org/10.1080/00222930500145123" docStyle="DocumentStyle:7E99CF6C63B49250F2A87C3AEDC82D8C.2:JNatHist.2005-2007.journal_article.2cover" docStyleId="7E99CF6C63B49250F2A87C3AEDC82D8C" docStyleName="JNatHist.2005-2007.journal_article.2cover" docStyleVersion="2" docTitle="Wanniyala hakgala Huber &amp; Benjamin 2005, new species" docType="treatment" docVersion="3" lastPageNumber="3318" masterDocId="FFAAFFA1FFC2FFFBC3731D01FFC96C16" masterDocTitle="The pholcid spiders from Sri Lanka: redescription of Pholcus ceylonicus and description of a new genus (Araneae: Pholcidae)" masterLastPageNumber="3319" masterPageNumber="3305" pageNumber="3317" updateTime="1698880134788" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="6EDB931EAF6B5195F5B948D6E88B101C">The pholcid spiders from Sri Lanka: redescription of Pholcus ceylonicus and description of a new genus (Araneae: Pholcidae)</mods:title>
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<mods:namePart id="DD9C2BF70DAE4B300C5EFDF2CCEFAA74">Huber, Bernhard A.</mods:namePart>
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<mods:namePart id="5B79A02FB0B4DA3DE0255F938B2BB916">Benjamin, Suresh P.</mods:namePart>
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<mods:typeOfResource id="75403106E63B0BC69BB67FC8E9A6530E">text</mods:typeOfResource>
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<mods:title id="52C6A1359CB34D0600715725D11BADC3">Journal of Natural History</mods:title>
</mods:titleInfo>
<mods:part id="61DC59E7735343522C9ACD3848129915">
<mods:date id="15CDE72D697A2C5D22C227A794FCAEE1">2005</mods:date>
<mods:detail id="A2159A8ED4109E611F07DE4AB212D2D3" type="pubDate">
<mods:number id="BEDE5C6F120FF434F31B07A1A58B7418">2005-11-25</mods:number>
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<mods:number id="13476EFB88F94814BD1634C3FA848E5C">39</mods:number>
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<mods:number id="946A5EBD37B387249764272A1E358900">37</mods:number>
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<mods:start id="6BEF296914E40D3BB943E6C87C772138">3305</mods:start>
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<mods:identifier id="AD5AC0A939E0570C44238A0DDCFE0D7F" type="DOI">10.1080/00222930500145123</mods:identifier>
<mods:identifier id="99DE7D7EA143201D2D3B549C0496DC6E" type="ISSN">1464-5262</mods:identifier>
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<treatment id="039387D9FFCCFFF4C2BC1C59FDFE6E68" LSID="urn:lsid:plazi:treatment:039387D9FFCCFFF4C2BC1C59FDFE6E68" httpUri="http://treatment.plazi.org/id/039387D9FFCCFFF4C2BC1C59FDFE6E68" lastPageId="15" lastPageNumber="3318" pageId="14" pageNumber="3317">
<subSubSection id="C3206544FFCCFFF5C2BC1C59FC956D66" box="[463,860,344,368]" pageId="14" pageNumber="3317" type="nomenclature">
<paragraph id="8B8536CFFFCCFFF5C2BC1C59FC956D66" blockId="14.[463,860,344,400]" box="[463,860,344,368]" pageId="14" pageNumber="3317">
<heading id="D0CD81A3FFCCFFF5C2BC1C59FC956D66" box="[463,860,344,368]" centered="true" fontSize="10" level="2" pageId="14" pageNumber="3317" reason="2">
<taxonomicName id="4C3A4D4CFFCCFFF5C2BC1C59FD026D66" ID-CoL="7FTT8" authority="Huber &amp; Benjamin, 2005" authorityName="Huber &amp; Benjamin" authorityYear="2005" box="[463,715,344,368]" class="Arachnida" family="Pholcidae" genus="Wanniyala" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="3317" phylum="Arthropoda" rank="species" species="hakgala" status="sp. nov.">
<emphasis id="B94EEADDFFCCFFF5C2BC1C59FD026D66" bold="true" box="[463,715,344,368]" italics="true" pageId="14" pageNumber="3317">Wanniyala hakgala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27D57A6FFCCFFF5C1A71C59FC956D66" box="[724,860,344,368]" pageId="14" pageNumber="3317" rank="species">new species</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3206544FFCCFFF4C1671C79FDFE6E68" lastPageId="15" lastPageNumber="3318" pageId="14" pageNumber="3317" type="description">
<paragraph id="8B8536CFFFCCFFF5C1671C79FCDE6D86" blockId="14.[463,860,344,400]" box="[532,791,376,400]" pageId="14" pageNumber="3317">
(
<figureCitation id="13012A4AFFCCFFF5C16F1C79FD4F6D86" box="[540,646,376,400]" captionStart="Figure 5" captionStartId="9.[125,185,1636,1655]" captionTargetBox="[125,1173,566,1600]" captionTargetId="figure-82@9.[125,1173,565,1601]" captionTargetPageId="9" captionText="Figure 5. (ae) Wanniyala agrabopath; (fi) W. hakgala. (a, b, f, g) Male habitus, dorsal and lateral views; (c, h) female abdomen, ventral views; (e) female, ventral view; (d, i) female habitus, lateral views." figureDoi="http://doi.org/10.5281/zenodo.4581719" httpUri="https://zenodo.org/record/4581719/files/figure.png" pageId="14" pageNumber="3317">Figures 5</figureCitation>
fi, 7eh, 8)
</paragraph>
<paragraph id="8B8536CFFFCCFFF5C3F91CC8FF096DF6" blockId="14.[138,192,457,480]" box="[138,192,457,480]" pageId="14" pageNumber="3317">
<emphasis id="B94EEADDFFCCFFF5C3F91CC8FF096DF6" box="[138,192,457,480]" italics="true" pageId="14" pageNumber="3317">Type</emphasis>
</paragraph>
<paragraph id="8B8536CFFFCCFFF5C3F91CF8FF286E46" blockId="14.[138,1185,504,592]" pageId="14" pageNumber="3317">
<materialsCitation id="3B523C92FFCCFFF5C3F91CF8FF146E46" collectingDate="1972-11-18" collectorName="P. Lehtinen" country="Sri Lanka" location="Hakgala" municipality="Nuwara Eliya district" pageId="14" pageNumber="3317" specimenCount="1" typeStatus="holotype">
Male
<typeStatus id="5481886DFFCCFFF5C3A01CF8FEFE6E07" box="[211,311,505,529]" pageId="14" pageNumber="3317" type="holotype">holotype</typeStatus>
from
<location id="8EE56014FFCCFFF5C2FA1CF8FE216E07" LSID="urn:lsid:plazi:treatment:039387D9FFCCFFF4C2BC1C59FDFE6E68:8EE56014FFCCFFF5C2FA1CF8FE216E07" box="[393,488,505,529]" country="Sri Lanka" municipality="Nuwara Eliya district" name="Hakgala" pageId="14" pageNumber="3317">Hakgala</location>
(6
<emphasis id="B94EEADDFFCCFFF5C17D1CF8FDD16E07" bold="true" box="[526,536,505,529]" italics="true" pageId="14" pageNumber="3317">°</emphasis>
549N, 80
<emphasis id="B94EEADDFFCCFFF5C1F41CF8FD586E07" bold="true" box="[647,657,505,529]" italics="true" pageId="14" pageNumber="3317">°</emphasis>
489E),
<collectingMunicipality id="6BE1ACB5FFCCFFF5C19A1CF8FC276E07" box="[745,1006,505,529]" pageId="14" pageNumber="3317">Nuwara Eliya district</collectingMunicipality>
,
<collectingCountry id="F32D765FFFCCFFF5C08C1CF8FBB06E07" box="[1023,1145,505,529]" name="Sri Lanka" pageId="14" pageNumber="3317">Sri Lanka</collectingCountry>
; in grass and litter along a mountain brook,
<date id="FF84100FFFCCFFF5C1FA1F18FCBE6E27" box="[649,887,537,561]" pageId="14" pageNumber="3317" value="1972-11-18">
<collectingDate id="EFC0E9E7FFCCFFF5C1FA1F18FCBE6E27" box="[649,887,537,561]" pageId="14" pageNumber="3317" value="1972-11-18">18 November 1972</collectingDate>
</date>
(
<collectorName id="26CF5319FFCCFFF5C0FD1F18FBEB6E27" box="[910,1058,537,561]" pageId="14" pageNumber="3317">P. Lehtinen</collectorName>
), in MZT (3631)
</materialsCitation>
.
</paragraph>
<paragraph id="8B8536CFFFCCFFF5C3F91F88FF326EB7" blockId="14.[138,251,649,673]" box="[138,251,649,673]" pageId="14" pageNumber="3317">
<emphasis id="B94EEADDFFCCFFF5C3F91F88FF326EB7" box="[138,251,649,673]" italics="true" pageId="14" pageNumber="3317">Etymology</emphasis>
</paragraph>
<paragraph id="8B8536CFFFCCFFF5C3F91FB8FC1B6EC7" blockId="14.[138,978,697,721]" box="[138,978,697,721]" pageId="14" pageNumber="3317">
The specific name is a noun in apposition, taken from the
<typeStatus id="5481886DFFCCFFF5C04E1FB8FCA76EC7" box="[829,878,697,721]" pageId="14" pageNumber="3317">type</typeStatus>
locality.
</paragraph>
<paragraph id="8B8536CFFFCCFFF5C3F91E08FF3A6F37" blockId="14.[138,243,777,801]" box="[138,243,777,801]" pageId="14" pageNumber="3317">
<emphasis id="B94EEADDFFCCFFF5C3F91E08FF3A6F37" box="[138,243,777,801]" italics="true" pageId="14" pageNumber="3317">Diagnosis</emphasis>
</paragraph>
<paragraph id="8B8536CFFFCCFFF5C3F91E38FEE26FA7" blockId="14.[138,1185,825,945]" pageId="14" pageNumber="3317">
Closely related to
<taxonomicName id="4C3A4D4CFFCCFFF5C2001E3BFDD66F47" authorityName="Huber &amp; Benjamin" authorityYear="2005" box="[371,543,825,849]" class="Arachnida" family="Pholcidae" genus="Wanniyala" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="3317" phylum="Arthropoda" rank="species" species="agrabopath">
<emphasis id="B94EEADDFFCCFFF5C2001E3BFDD66F47" box="[371,543,825,849]" italics="true" pageId="14" pageNumber="3317">W. agrabopath</emphasis>
</taxonomicName>
; easily distinguished by shape of dorsal sclerite on procursus (ds in
<figureCitation id="13012A4AFFCCFFF5C2231E58FE756F67" box="[336,444,857,881]" captionStart="Figure 7" captionStartId="11.[125,185,1253,1272]" captionTargetBox="[133,1168,138,1203]" captionTargetId="figure-96@11.[125,1173,133,1217]" captionTargetPageId="11" captionText="Figure 7. (ad) Wanniyala agrabopath; (eh) W. hakgala. (a, b, e, f) Left procursus, prolateral and retrolateral views; (c, g) cleared epigyna, ventral views; (d, h) cleared epigyna, dorsal views. ds, dorsal sclerite; pp, proximal projection; t13, transparent projections. Scale bars: 0.3 mm (a, b, e, f: all to same scale)." figureDoi="http://doi.org/10.5281/zenodo.4581725" httpUri="https://zenodo.org/record/4581725/files/figure.png" pageId="14" pageNumber="3317">Figure 7f</figureCitation>
), by epigynum with pointed projection (
<figureCitation id="13012A4AFFCCFFF5C0E01E58FBC06F67" box="[915,1033,857,881]" captionStart="Figure 5" captionStartId="9.[125,185,1636,1655]" captionTargetBox="[125,1173,566,1600]" captionTargetId="figure-82@9.[125,1173,565,1601]" captionTargetPageId="9" captionText="Figure 5. (ae) Wanniyala agrabopath; (fi) W. hakgala. (a, b, f, g) Male habitus, dorsal and lateral views; (c, h) female abdomen, ventral views; (e) female, ventral view; (d, i) female habitus, lateral views." figureDoi="http://doi.org/10.5281/zenodo.4581719" httpUri="https://zenodo.org/record/4581719/files/figure.png" pageId="14" pageNumber="3317">Figures 5i</figureCitation>
,
<figureCitation id="13012A4AFFCCFFF5C7641E58FBF16F67" box="[1047,1080,857,881]" captionStart="Figure 8" captionStartId="12.[138,198,1090,1109]" captionTargetBox="[138,1186,133,1054]" captionTargetId="figure-159@12.[138,1186,133,1054]" captionTargetPageId="12" captionText="Figure 8. Wanniyala hakgala. (a) Male gonopore with four epiandrous spigots; (b, c) female and male anterior lateral spinnerets with widened and pointed spigots; (d) female abdomen, ventral view. Scale bars: 20 Mm (ac); 0.5 mm (d)." figureDoi="http://doi.org/10.5281/zenodo.4581727" httpUri="https://zenodo.org/record/4581727/files/figure.png" pageId="14" pageNumber="3317">8d</figureCitation>
); also by longer clypeus projections, longer cheliceral apophyses, and apparently by colour pattern (
<figureCitation id="13012A4AFFCCFFF5C3E11E98FED46FA7" box="[146,285,921,945]" captionStart="Figure 5" captionStartId="9.[125,185,1636,1655]" captionTargetBox="[125,1173,566,1600]" captionTargetId="figure-82@9.[125,1173,565,1601]" captionTargetPageId="9" captionText="Figure 5. (ae) Wanniyala agrabopath; (fi) W. hakgala. (a, b, f, g) Male habitus, dorsal and lateral views; (c, h) female abdomen, ventral views; (e) female, ventral view; (d, i) female habitus, lateral views." figureDoi="http://doi.org/10.5281/zenodo.4581719" httpUri="https://zenodo.org/record/4581719/files/figure.png" pageId="14" pageNumber="3317">Figure 5f, g</figureCitation>
).
</paragraph>
<paragraph id="8B8536CFFFCCFFF5C3F91ED8FEDB6A19" blockId="14.[138,1189,983,1551]" pageId="14" pageNumber="3317">
<emphasis id="B94EEADDFFCCFFF5C3F91ED8FE9C6FE7" box="[138,341,985,1009]" italics="true" pageId="14" pageNumber="3317">
Male (
<typeStatus id="5481886DFFCCFFF5C3961ED8FE806FE7" box="[229,329,985,1009]" pageId="14" pageNumber="3317" type="holotype">holotype</typeStatus>
).
</emphasis>
Total length 2.2, carapace width 0.97. Leg 1: 17.1 (4.25+ 0.4+4.5+6.15+1.85), tibia 2: 3.0, tibia 3: 2.1, tibia 4: 2.85; tibia 1 L/d: 51. Habitus as in
<figureCitation id="13012A4AFFCCFFF5C3F91918FEDC6827" box="[138,277,1049,1073]" captionStart="Figure 5" captionStartId="9.[125,185,1636,1655]" captionTargetBox="[125,1173,566,1600]" captionTargetId="figure-82@9.[125,1173,565,1601]" captionTargetPageId="9" captionText="Figure 5. (ae) Wanniyala agrabopath; (fi) W. hakgala. (a, b, f, g) Male habitus, dorsal and lateral views; (c, h) female abdomen, ventral views; (e) female, ventral view; (d, i) female habitus, lateral views." figureDoi="http://doi.org/10.5281/zenodo.4581719" httpUri="https://zenodo.org/record/4581719/files/figure.png" pageId="14" pageNumber="3317">Figure 5f, g</figureCitation>
. Carapace ochre-yellow with darker median and lateral bands, clypeus also darkened; sternum light brown. Legs pale ochre-yellow, slightly darker rings on femora (subdistally), patellae, and tibiae (subdistally). Opisthosoma grey with black pattern, ventrally also with dark pattern. Ocular area slightly elevated but distinct, thoracic furrow very shallow; distance PMEPME 140 Mm; diameter PME 105 Mm; distance PMEALE 25 Mm; AME absent. Clypeus with pair of short projections 90 Mm long and distinct median hump. Sternum wider than long (0.75/0.60). Chelicerae very similar to
<taxonomicName id="4C3A4D4CFFCCFFF5C0BD19D8FBBB68E6" authorityName="Huber &amp; Benjamin" authorityYear="2005" box="[974,1138,1240,1264]" class="Arachnida" family="Pholcidae" genus="Wanniyala" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="3317" phylum="Arthropoda" rank="species" species="agrabopath">
<emphasis id="B94EEADDFFCCFFF5C0BD19D8FBBB68E6" box="[974,1138,1240,1264]" italics="true" pageId="14" pageNumber="3317">W. agrabopath</emphasis>
</taxonomicName>
(cf.
<figureCitation id="13012A4AFFCCFFF5C3F919F9FEDA6906" box="[138,275,1272,1296]" captionStart="Figure 6" captionStartId="10.[138,198,1377,1396]" captionTargetBox="[148,1175,133,1327]" captionTargetId="figure-15@10.[138,1186,133,1341]" captionTargetPageId="10" captionText="Figure 6. Wanniyala agrabopath. (a, b) Left male palp, prolateral and retrolateral views; (c) male prosoma and chelicerae, dorsal view; (d) tip of male cheliceral apophysis; (e) male chelicerae, frontal view. b, genital bulb; ba, bulbal apophysis; e, embolus. Scale bars: 0.5 mm (ac); 50 Mm (d); 0.2 mm (e)." figureDoi="http://doi.org/10.5281/zenodo.4581723" httpUri="https://zenodo.org/record/4581723/files/figure.png" pageId="14" pageNumber="3317">Figure 6c, e</figureCitation>
) with pair of long apophyses provided with two modified hairs on each tip (cf.
<figureCitation id="13012A4AFFCCFFF5C3F91819FF326926" box="[138,251,1304,1328]" captionStart="Figure 6" captionStartId="10.[138,198,1377,1396]" captionTargetBox="[148,1175,133,1327]" captionTargetId="figure-15@10.[138,1186,133,1341]" captionTargetPageId="10" captionText="Figure 6. Wanniyala agrabopath. (a, b) Left male palp, prolateral and retrolateral views; (c) male prosoma and chelicerae, dorsal view; (d) tip of male cheliceral apophysis; (e) male chelicerae, frontal view. b, genital bulb; ba, bulbal apophysis; e, embolus. Scale bars: 0.5 mm (ac); 50 Mm (d); 0.2 mm (e)." figureDoi="http://doi.org/10.5281/zenodo.4581723" httpUri="https://zenodo.org/record/4581723/files/figure.png" pageId="14" pageNumber="3317">Figure 6d</figureCitation>
), tips 0.88 apart. Palps in general very similar to
<taxonomicName id="4C3A4D4CFFCCFFF5C0481818FC166926" authorityName="Huber &amp; Benjamin" authorityYear="2005" box="[827,991,1304,1328]" class="Arachnida" family="Pholcidae" genus="Wanniyala" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="3317" phylum="Arthropoda" rank="species" species="agrabopath">
<emphasis id="B94EEADDFFCCFFF5C0481818FC166926" box="[827,991,1304,1328]" italics="true" pageId="14" pageNumber="3317">W. agrabopath</emphasis>
</taxonomicName>
but significantly larger; trochanter and femur projections identical to
<taxonomicName id="4C3A4D4CFFCCFFF5C19D1838FC5B6946" authorityName="Huber &amp; Benjamin" authorityYear="2005" box="[750,914,1336,1360]" class="Arachnida" family="Pholcidae" genus="Wanniyala" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="3317" phylum="Arthropoda" rank="species" species="agrabopath">
<emphasis id="B94EEADDFFCCFFF5C19D1838FC5B6946" box="[750,914,1336,1360]" italics="true" pageId="14" pageNumber="3317">W. agrabopath</emphasis>
</taxonomicName>
, procursus very similar to
<taxonomicName id="4C3A4D4CFFCCFFF5C3C21859FE956966" authorityName="Huber &amp; Benjamin" authorityYear="2005" box="[177,348,1368,1392]" class="Arachnida" family="Pholcidae" genus="Wanniyala" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="3317" phylum="Arthropoda" rank="species" species="agrabopath">
<emphasis id="B94EEADDFFCCFFF5C3C21859FE956966" box="[177,348,1368,1392]" italics="true" pageId="14" pageNumber="3317">W. agrabopath</emphasis>
</taxonomicName>
, but with very different dorsal sclerite (
<figureCitation id="13012A4AFFCCFFF5C02A1859FC0E6966" box="[857,967,1368,1392]" captionStart="Figure 7" captionStartId="11.[125,185,1253,1272]" captionTargetBox="[133,1168,138,1203]" captionTargetId="figure-96@11.[125,1173,133,1217]" captionTargetPageId="11" captionText="Figure 7. (ad) Wanniyala agrabopath; (eh) W. hakgala. (a, b, e, f) Left procursus, prolateral and retrolateral views; (c, g) cleared epigyna, ventral views; (d, h) cleared epigyna, dorsal views. ds, dorsal sclerite; pp, proximal projection; t13, transparent projections. Scale bars: 0.3 mm (a, b, e, f: all to same scale)." figureDoi="http://doi.org/10.5281/zenodo.4581725" httpUri="https://zenodo.org/record/4581725/files/figure.png" pageId="14" pageNumber="3317">Figure 7f</figureCitation>
); bulb apparently identical to
<taxonomicName id="4C3A4D4CFFCCFFF5C26B1879FE766986" authorityName="Huber &amp; Benjamin" authorityYear="2005" box="[280,447,1400,1424]" class="Arachnida" family="Pholcidae" genus="Wanniyala" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="3317" phylum="Arthropoda" rank="species" species="agrabopath">
<emphasis id="B94EEADDFFCCFFF5C26B1879FE766986" box="[280,447,1400,1424]" italics="true" pageId="14" pageNumber="3317">W. agrabopath</emphasis>
</taxonomicName>
(cf.
<figureCitation id="13012A4AFFCCFFF5C28A1879FDA06986" box="[505,617,1400,1424]" captionStart="Figure 6" captionStartId="10.[138,198,1377,1396]" captionTargetBox="[148,1175,133,1327]" captionTargetId="figure-15@10.[138,1186,133,1341]" captionTargetPageId="10" captionText="Figure 6. Wanniyala agrabopath. (a, b) Left male palp, prolateral and retrolateral views; (c) male prosoma and chelicerae, dorsal view; (d) tip of male cheliceral apophysis; (e) male chelicerae, frontal view. b, genital bulb; ba, bulbal apophysis; e, embolus. Scale bars: 0.5 mm (ac); 50 Mm (d); 0.2 mm (e)." figureDoi="http://doi.org/10.5281/zenodo.4581723" httpUri="https://zenodo.org/record/4581723/files/figure.png" pageId="14" pageNumber="3317">Figure 6a</figureCitation>
). Retrolateral trichobothrium of tibia 1 at 13%; legs without spines, without curved hairs, with several rows of vertical hairs on all metatarsi; tarsus 1 with.20 pseudosegments, only distally quite distinct. Anterior lateral spinnerets with only two spigots each (
<figureCitation id="13012A4AFFCCFFF5C29D18D6FD9669F9" box="[494,607,1495,1519]" captionStart="Figure 8" captionStartId="12.[138,198,1090,1109]" captionTargetBox="[138,1186,133,1054]" captionTargetId="figure-159@12.[138,1186,133,1054]" captionTargetPageId="12" captionText="Figure 8. Wanniyala hakgala. (a) Male gonopore with four epiandrous spigots; (b, c) female and male anterior lateral spinnerets with widened and pointed spigots; (d) female abdomen, ventral view. Scale bars: 20 Mm (ac); 0.5 mm (d)." figureDoi="http://doi.org/10.5281/zenodo.4581727" httpUri="https://zenodo.org/record/4581727/files/figure.png" pageId="14" pageNumber="3317">Figure 8c</figureCitation>
). Male gonopore with four epiandrous spigots (
<figureCitation id="13012A4AFFCCFFF5C3E118F6FECD6A19" box="[146,260,1527,1551]" captionStart="Figure 8" captionStartId="12.[138,198,1090,1109]" captionTargetBox="[138,1186,133,1054]" captionTargetId="figure-159@12.[138,1186,133,1054]" captionTargetPageId="12" captionText="Figure 8. Wanniyala hakgala. (a) Male gonopore with four epiandrous spigots; (b, c) female and male anterior lateral spinnerets with widened and pointed spigots; (d) female abdomen, ventral view. Scale bars: 20 Mm (ac); 0.5 mm (d)." figureDoi="http://doi.org/10.5281/zenodo.4581727" httpUri="https://zenodo.org/record/4581727/files/figure.png" pageId="14" pageNumber="3317">Figure 8a</figureCitation>
).
</paragraph>
<paragraph id="8B8536CFFFCCFFF5C3F91B2BFD996A54" blockId="14.[138,592,1578,1602]" box="[138,592,1578,1602]" pageId="14" pageNumber="3317">
<emphasis id="B94EEADDFFCCFFF5C3F91B2BFF326A54" box="[138,251,1578,1602]" italics="true" pageId="14" pageNumber="3317">Variation.</emphasis>
Tibia
<quantity id="4CC29B2AFFCCFFF5C23C1B2BFEB46A54" box="[335,381,1578,1602]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="14" pageNumber="3317" unit="in" value="1.0">1 in</quantity>
other male: 4.15.
</paragraph>
<paragraph id="8B8536CFFFCCFFF4C3F91B5CFEAE6CF6" blockId="14.[138,1185,1629,1685]" lastBlockId="15.[125,1172,136,224]" lastPageId="15" lastPageNumber="3318" pageId="14" pageNumber="3317">
<emphasis id="B94EEADDFFCCFFF5C3F91B5CFF286A63" box="[138,225,1629,1653]" italics="true" pageId="14" pageNumber="3317">Female.</emphasis>
In general similar to male, but clypeus unmodified and metatarsi without rows of vertical hairs. Tibia
<quantity id="4CC29B2AFFCCFFF5C2041B7CFE6C6A83" box="[375,421,1661,1685]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="14" pageNumber="3317" unit="in" value="1.0">1 in</quantity>
female from Hakgala: 3.13.45; female from Gumbukena: 3.05; females from Kandy district: 2.73.0 (
<emphasis id="B94EEADDFFCDFFF4C1491D88FD816CB6" box="[570,584,137,160]" italics="true" pageId="15" pageNumber="3318">n</emphasis>
54). Epigynum very large, with distinctive median conical elevation (
<figureCitation id="13012A4AFFCDFFF4C23C1DA9FE0A6CD6" box="[335,451,168,192]" captionStart="Figure 8" captionStartId="12.[138,198,1090,1109]" captionTargetBox="[138,1186,133,1054]" captionTargetId="figure-159@12.[138,1186,133,1054]" captionTargetPageId="12" captionText="Figure 8. Wanniyala hakgala. (a) Male gonopore with four epiandrous spigots; (b, c) female and male anterior lateral spinnerets with widened and pointed spigots; (d) female abdomen, ventral view. Scale bars: 20 Mm (ac); 0.5 mm (d)." figureDoi="http://doi.org/10.5281/zenodo.4581727" httpUri="https://zenodo.org/record/4581727/files/figure.png" pageId="15" pageNumber="3318">Figure 8d</figureCitation>
), pockets not seen. Dorsal view as in
<figureCitation id="13012A4AFFCDFFF4C00F1DA9FC276CD6" box="[892,1006,168,192]" captionStart="Figure 7" captionStartId="11.[125,185,1253,1272]" captionTargetBox="[133,1168,138,1203]" captionTargetId="figure-96@11.[125,1173,133,1217]" captionTargetPageId="11" captionText="Figure 7. (ad) Wanniyala agrabopath; (eh) W. hakgala. (a, b, e, f) Left procursus, prolateral and retrolateral views; (c, g) cleared epigyna, ventral views; (d, h) cleared epigyna, dorsal views. ds, dorsal sclerite; pp, proximal projection; t13, transparent projections. Scale bars: 0.3 mm (a, b, e, f: all to same scale)." figureDoi="http://doi.org/10.5281/zenodo.4581725" httpUri="https://zenodo.org/record/4581725/files/figure.png" pageId="15" pageNumber="3318">Figure 7h</figureCitation>
. Spinnerets as in male (
<figureCitation id="13012A4AFFCDFFF4C3961DC9FE906CF6" box="[229,345,200,224]" captionStart="Figure 8" captionStartId="12.[138,198,1090,1109]" captionTargetBox="[138,1186,133,1054]" captionTargetId="figure-159@12.[138,1186,133,1054]" captionTargetPageId="12" captionText="Figure 8. Wanniyala hakgala. (a) Male gonopore with four epiandrous spigots; (b, c) female and male anterior lateral spinnerets with widened and pointed spigots; (d) female abdomen, ventral view. Scale bars: 20 Mm (ac); 0.5 mm (d)." figureDoi="http://doi.org/10.5281/zenodo.4581727" httpUri="https://zenodo.org/record/4581727/files/figure.png" pageId="15" pageNumber="3318">Figure 8b</figureCitation>
).
</paragraph>
<paragraph id="8B8536CFFFCDFFF4C30E1C16FF346D39" blockId="15.[125,253,279,303]" box="[125,253,279,303]" pageId="15" pageNumber="3318">
<emphasis id="B94EEADDFFCDFFF4C30E1C16FF346D39" box="[125,253,279,303]" italics="true" pageId="15" pageNumber="3318">Distribution</emphasis>
</paragraph>
<paragraph id="8B8536CFFFCDFFF4C30E1C46FD2A6D49" blockId="15.[125,739,327,351]" box="[125,739,327,351]" pageId="15" pageNumber="3318">
Known from three localities in
<collectingCountry id="F32D765FFFCDFFF4C29E1C46FD966D49" box="[493,607,327,351]" name="Sri Lanka" pageId="15" pageNumber="3318">Sri Lanka</collectingCountry>
(
<figureCitation id="13012A4AFFCDFFF4C1031C46FD1D6D49" box="[624,724,327,351]" captionStart="Figure 9" captionStartId="13.[125,185,941,960]" captionTargetBox="[365,932,133,906]" captionTargetId="figure-152@13.[365,932,133,906]" captionTargetPageId="13" captionText="Figure 9. Known geographic distributions of Pholcus ceylonicus, Wanniyala agrabopath, W. hakgala, and an undescribed species of Wanniyala." figureDoi="http://doi.org/10.5281/zenodo.4581729" httpUri="https://zenodo.org/record/4581729/files/figure.png" pageId="15" pageNumber="3318">Figure 9</figureCitation>
).
</paragraph>
<paragraph id="8B8536CFFFCDFFF4C30E1C96FB5D6DC9" blockId="15.[125,332,407,430]" lastBlockId="15.[125,1172,455,638]" pageId="15" pageNumber="3318">
<emphasis id="B94EEADDFFCDFFF4C30E1C96FE856DB9" box="[125,332,407,431]" italics="true" pageId="15" pageNumber="3318">Material examined</emphasis>
<materialsCitation id="3B523C92FFCDFFF4C30E1CC6FB5D6DC9" box="[125,1172,454,479]" country="Sri Lanka" location="Nuwara Eliya district" municipality="Hakgala" pageId="15" pageNumber="3318" specimenCount="5" specimenCount-female="5">
<collectingCountry id="F32D765FFFCDFFF4C30E1CC6FF306DC9" box="[125,249,455,479]" name="Sri Lanka" pageId="15" pageNumber="3318">Sri Lanka</collectingCountry>
:
<location id="8EE56014FFCDFFF4C2781CC6FDDB6DC9" LSID="urn:lsid:plazi:treatment:039387D9FFCCFFF4C2BC1C59FDFE6E68:8EE56014FFCDFFF4C2781CC6FDDB6DC9" box="[267,530,455,479]" country="Sri Lanka" municipality="Hakgala" name="Nuwara Eliya district" pageId="15" pageNumber="3318">Nuwara Eliya district</location>
:
<collectingMunicipality id="6BE1ACB5FFCDFFF4C1501CC6FD4F6DC9" box="[547,646,455,479]" pageId="15" pageNumber="3318">Hakgala</collectingMunicipality>
:
<typeStatus id="5481886DFFCDFFF4C1EB1CC6FD006DC9" box="[664,713,455,479]" pageId="15" pageNumber="3318">type</typeStatus>
above, together with 1
<emphasis id="B94EEADDFFCDFFF4C09E1CC6FC366DC8" box="[1005,1023,455,478]" italics="true" pageId="15" pageNumber="3318"></emphasis>
,
<specimenCount id="9D3CFD46FFCDFFF4C7661CC6FBF86DC9" box="[1045,1073,454,479]" pageId="15" pageNumber="3318" type="female">5♀</specimenCount>
; Kandy
</materialsCitation>
</paragraph>
<paragraph id="8B8536CFFFCDFFF4C30E1CE6FDFE6E68" blockId="15.[125,1172,455,638]" pageId="15" pageNumber="3318">
<materialsCitation id="3B523C92FFCDFFF4C30E1CE6FC166E09" collectingDate="1981-08-07" collectingDateMax="1981-08-11" collectingDateMin="1981-08-07" collectionCode="RMNH" elevation="1600" location="Kandy" pageId="15" pageNumber="3318" specimenCount="4" specimenCount-female="4">
district:
<location id="8EE56014FFCDFFF4C3AD1CE6FEE26DE9" LSID="urn:lsid:plazi:treatment:039387D9FFCCFFF4C2BC1C59FDFE6E68:8EE56014FFCDFFF4C3AD1CE6FEE26DE9" box="[222,299,487,511]" name="Kandy" pageId="15" pageNumber="3318">Kandy</location>
(7
<emphasis id="B94EEADDFFCDFFF4C23D1CE6FE916DE9" bold="true" box="[334,344,487,511]" italics="true" pageId="15" pageNumber="3318">°</emphasis>
189N, 80
<emphasis id="B94EEADDFFCDFFF4C2B11CE6FE056DE9" bold="true" box="[450,460,487,511]" italics="true" pageId="15" pageNumber="3318">°</emphasis>
389E),
<elevation id="0017D1FCFFCDFFF4C1531CE6FD7F6DE9" box="[544,694,487,511]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="15" pageNumber="3318" unit="m" value="1600.0">
<quantity id="4CC29B2AFFCDFFF4C1531CE6FDBF6DE9" box="[544,630,487,511]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="15" pageNumber="3318" unit="m" value="1600.0">1600 m</quantity>
a.s.l.
</elevation>
, forest and waterworks, in umbrella web under large tangle,
<date id="FF84100FFFCDFFF4C2171F06FDF46E09" box="[356,573,519,543]" pageId="15" pageNumber="3318" value="1981-08-07" valueMax="1981-08-11" valueMin="1981-08-07">
<collectingDate id="EFC0E9E7FFCDFFF4C2171F06FDF46E09" box="[356,573,519,543]" pageId="15" pageNumber="3318" value="1981-08-07" valueMax="1981-08-11" valueMin="1981-08-07">711 August 1981</collectingDate>
</date>
(collector not given),
<specimenCount id="9D3CFD46FFCDFFF4C0341F06FCAD6E09" box="[839,868,518,543]" pageId="15" pageNumber="3318" type="female">4♀</specimenCount>
(
<collectionCode id="ED2BAE0AFFCDFFF4C0061F06FC1E6E09" box="[885,983,519,543]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="15" pageNumber="3318" type="Museum">RMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B523C92FFCDFFF4C09D1F06FC336E28" collectingDate="1972-11-16" collectionCode="MZT" collectorName="P. Lehtinen" location="Nonawatte'" pageId="15" pageNumber="3318" specimenCode="MZT 3630" specimenCount="1" specimenCount-female="1">
<location id="8EE56014FFCDFFF4C08A1F06FB426E09" LSID="urn:lsid:plazi:treatment:039387D9FFCCFFF4C2BC1C59FDFE6E68:8EE56014FFCDFFF4C08A1F06FB426E09" box="[1017,1163,519,543]" name="Nonawatte'" pageId="15" pageNumber="3318">Nonawatte</location>
, wet slope at waterfall,
<date id="FF84100FFFCDFFF4C2FD1F27FDBE6E28" box="[398,631,550,574]" pageId="15" pageNumber="3318" value="1972-11-16">
<collectingDate id="EFC0E9E7FFCDFFF4C2FD1F27FDBE6E28" box="[398,631,550,574]" pageId="15" pageNumber="3318" value="1972-11-16">16 November 1972</collectingDate>
</date>
(
<collectorName id="26CF5319FFCDFFF4C1F81F27FCD46E28" box="[651,797,550,574]" pageId="15" pageNumber="3318">P. Lehtinen</collectorName>
),
<specimenCount id="9D3CFD46FFCDFFF4C0461F27FC986E29" box="[821,849,550,575]" pageId="15" pageNumber="3318" type="female">1♀</specimenCount>
(
<specimenCode id="DB9C9EB4FFCDFFF4C0161F27FC3A6E28" box="[869,1011,550,574]" collectionCode="MZT" pageId="15" pageNumber="3318">MZT 3630</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B523C92FFCDFFF4C7791F27FDFA6E68" collectingDate="1972-11-19" collectionCode="MZT" collectorName="P. Lehtinen" county="Kumbukana" location="Moneragala district" municipality="Gumbukena'" pageId="15" pageNumber="3318" specimenCode="MZT 3632" specimenCount="1" specimenCount-female="1">
<location id="8EE56014FFCDFFF4C7791F27FF046E48" LSID="urn:lsid:plazi:treatment:039387D9FFCCFFF4C2BC1C59FDFE6E68:8EE56014FFCDFFF4C7791F27FF046E48" county="Kumbukana" municipality="Gumbukena'" name="Moneragala district" pageId="15" pageNumber="3318">Moneragala district</location>
(6
<emphasis id="B94EEADDFFCDFFF4C39D1F47FF316E48" bold="true" box="[238,248,582,606]" italics="true" pageId="15" pageNumber="3318">°</emphasis>
409N,, 81
<emphasis id="B94EEADDFFCDFFF4C2041F47FE486E48" bold="true" box="[375,385,582,606]" italics="true" pageId="15" pageNumber="3318">°</emphasis>
209E):
<collectingMunicipality id="6BE1ACB5FFCDFFF4C2AE1F47FDB46E48" box="[477,637,582,606]" pageId="15" pageNumber="3318">Gumbukena</collectingMunicipality>
(5
<collectingCounty id="62E44E43FFCDFFF4C1DA1F47FCF76E48" box="[681,830,582,606]" pageId="15" pageNumber="3318">Kumbukana</collectingCounty>
?), in teak litter,
<date id="FF84100FFFCDFFF4C0861F47FF716E68" pageId="15" pageNumber="3318" value="1972-11-19">
<collectingDate id="EFC0E9E7FFCDFFF4C0861F47FF716E68" pageId="15" pageNumber="3318" value="1972-11-19">19 November 1972</collectingDate>
</date>
(
<collectorName id="26CF5319FFCDFFF4C3B81F67FE926E68" box="[203,347,614,638]" pageId="15" pageNumber="3318">P. Lehtinen</collectorName>
),
<specimenCount id="9D3CFD46FFCDFFF4C2031F67FE446E68" box="[368,397,613,638]" pageId="15" pageNumber="3318" type="female">1♀</specimenCount>
(
<specimenCode id="DB9C9EB4FFCDFFF4C2ED1F67FDE36E68" box="[414,554,614,638]" collectionCode="MZT" pageId="15" pageNumber="3318">MZT 3632</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>