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<document id="ED8757FE8330EB32E92F0D6984D06738" ID-DOI="10.5281/zenodo.206618" ID-GBIF-Dataset="f5b1d7ac-ca42-4e4c-b2d0-b96771bc085c" ID-ISSN="1175-5326" ID-Zenodo-Dep="206618" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460257628293" checkinUser="plazi" docAuthor="Li, Chun-Lin, Wang, Chuan-Chan &amp; Chen, Hou-Jay" docDate="2011" docId="039387C6803DFFFC61D1D9495CB9FEA2" docLanguage="en" docName="zt02790p034.pdf" docOrigin="Zootaxa 2790" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Amphicoma lalashanensis Li, new species" docType="treatment" docVersion="9" lastPageNumber="28" masterDocId="FFAAFFBE803FFFFA6146DF04580DFFBE" masterDocTitle="Synopsis of the genus Amphicoma Latreille (Coleoptera: Glaphyridae) of Taiwan with special reference to the male genitalia" masterLastPageNumber="34" masterPageNumber="23" pageNumber="25" updateTime="1698243705731" updateUser="plazi">
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<mods:title id="97B5A659D15062E71736F484B4BC03D0">Synopsis of the genus Amphicoma Latreille (Coleoptera: Glaphyridae) of Taiwan with special reference to the male genitalia</mods:title>
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<mods:namePart id="6AE4CF2C6C1A0128815229E8EC3BEF21">Li, Chun-Lin</mods:namePart>
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<mods:namePart id="11FE3739521B2C3C47E6CB1D2FF823CE">Wang, Chuan-Chan</mods:namePart>
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<mods:namePart id="F15F24F1F2E8194B0D5888AE35E6220E">Chen, Hou-Jay</mods:namePart>
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<mods:date id="5E35B93731D668A03F48E4F57B39D407">2011</mods:date>
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<mods:identifier id="521470E8A606569627625DEC3A7E1641" type="DOI">10.5281/zenodo.206618</mods:identifier>
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<treatment id="039387C6803DFFFC61D1D9495CB9FEA2" ID-DOI="http://doi.org/10.5281/zenodo.6193208" ID-GBIF-Taxon="119392670" ID-Zenodo-Dep="6193208" LSID="urn:lsid:plazi:treatment:039387C6803DFFFC61D1D9495CB9FEA2" httpUri="http://treatment.plazi.org/id/039387C6803DFFFC61D1D9495CB9FEA2" lastPageId="6" lastPageNumber="28" pageId="2" pageNumber="25">
<subSubSection id="C320655B803DFFF861D1D94959F7F939" pageId="2" pageNumber="25" type="nomenclature">
<paragraph id="8B8536D0803DFFF861D1D9495A9AF9D9" blockId="2.[151,663,1613,1671]" box="[151,663,1613,1639]" pageId="2" pageNumber="25">
<heading id="D0CD81BC803DFFF861D1D9495A9AF9D9" bold="true" box="[151,663,1613,1639]" fontSize="11" level="1" pageId="2" pageNumber="25" reason="1">
<emphasis id="B94EEAC2803DFFF861D1D9495A9AF9D9" bold="true" box="[151,663,1613,1639]" pageId="2" pageNumber="25">
<taxonomicName id="4C3A4D53803DFFF861D1D94959F4F9D9" ID-CoL="CZXL" authority="Li" authorityName="Li" box="[151,505,1613,1639]" class="Insecta" family="Glaphyridae" genus="Amphicoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="25" phylum="Arthropoda" rank="species" species="lalashanensis" status="sp. nov.">
<emphasis id="B94EEAC2803DFFF861D1D94959D8F9D9" bold="true" box="[151,469,1613,1639]" italics="true" pageId="2" pageNumber="25">Amphicoma lalashanensis</emphasis>
Li
</taxonomicName>
,
<taxonomicNameLabel id="A27D57B9803DFFF86341D9495A9AF9D9" box="[519,663,1613,1639]" pageId="2" pageNumber="25" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8536D0803DFFF861D1D96B59F7F939" blockId="2.[151,663,1613,1671]" box="[151,506,1647,1671]" pageId="2" pageNumber="25">
<figureCitation id="13012A55803DFFF861D1D96B58EEF939" box="[151,227,1647,1671]" captionStart="FIGURE 1 7" captionStartId="3.[151,250,1593,1616]" captionTargetBox="[191,1402,196,1557]" captionTargetId="figure@3.[173,1413,171,1572]" captionTargetPageId="3" captionText="FIGURE 1 7. Dorsal habitus of Amphicoma species. 1. Amphicoma lalashanensis, holotype; 2. Amphicoma aurata, male; 3. Amphicoma aurata, male; 4. Amphicoma aurata, female; 5. Amphicoma formosana, male; 6. Amphicoma formosana, male; 7. Amphicoma formosana, female." httpUri="https://zenodo.org/record/206620/files/figure.png" pageId="2" pageNumber="25">Figs. 1</figureCitation>
,
<figureCitation id="13012A55803DFFF861B7D96B590CF939" box="[241,257,1647,1671]" captionStart="FIGURE 8 10" captionStartId="4.[151,250,638,661]" captionTargetBox="[158,1432,199,609]" captionTargetId="figure@4.[151,1436,193,617]" captionTargetPageId="4" captionText="FIGURE 8 10. Left lateral view of prosternal process of Amphicoma species. 8. Amphicoma lalashanensis; 9. Amphicoma aurata; 10. Amphicoma formosana." httpUri="https://zenodo.org/record/206621/files/figure.png" pageId="2" pageNumber="25">8</figureCitation>
,
<figureCitation id="13012A55803DFFF8604BD96B5927F939" box="[269,298,1647,1671]" captionStart="FIGURES 11 13" captionStartId="4.[151,264,1881,1904]" captionTargetBox="[159,1421,798,1845]" captionTargetId="figure@4.[151,1436,771,1860]" captionTargetPageId="4" captionText="FIGURES 11 13. Dorsal view of male genitalia of Amphicoma species. 11. Amphicoma lalashanensis; 12. Amphicoma aurata; 13. Amphicoma formosana. Abbreviations: BP-ext = basal piece extension; FMaB = flagellum major body. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/206622/files/figure.png" pageId="2" pageNumber="25">11</figureCitation>
,
<figureCitation id="13012A55803DFFF86070D96B5959F939" box="[310,340,1647,1671]" captionStart="FIGURES 14 19" captionStartId="5.[151,264,1350,1373]" captionTargetBox="[160,1426,598,1322]" captionTargetId="figure@5.[151,1436,589,1329]" captionTargetPageId="5" captionText="FIGURES 14 19. Parameres of Amphicoma male genitalia in lateral view. 14 15. Amphicoma lalashanensis; 16 17. Amphicoma auratas; 18 19. Amphicoma formosana. Abbreviations: FMaB = flagellum major body. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/206623/files/figure.png" pageId="2" pageNumber="25">14</figureCitation>
, 15,
<figureCitation id="13012A55803DFFF860CCD96B59A5F939" box="[394,424,1647,1671]" captionStart="FIGURES 20 22. 20" captionStartId="6.[151,264,1150,1173]" captionTargetBox="[170,1430,323,1122]" captionTargetId="figure@6.[151,1436,309,1129]" captionTargetPageId="6" captionText="FIGURES 20 22. 20. Amphicoma lalashanensis genital capsule; 21. Amphicoma aurata ventral plate of genital capsule; 22. Amphicoma formosana ventral plate of genital capsule. Abbreviations: LLP = left lateral plate; MS = median sclerite; RLP = right lateral plate; VP = ventral plate. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/206624/files/figure.png" pageId="2" pageNumber="25">20</figureCitation>
,
<figureCitation id="13012A55803DFFF860F2D96B59DFF939" box="[436,466,1647,1671]" captionStart="FIGURES 23" captionStartId="6.[151,264,1979,2002]" captionTargetBox="[566,1018,1285,1965]" captionTargetId="figure@6.[566,1020,1285,1965]" captionTargetPageId="6" captionText="FIGURES 23. Male genitalia of Amphicoma lalashanensis in ventral view. Abbreviations: BP = basal piece; BP-ext = basal piece extension; FPM = fusion of paramere and median lobe; ML = median lobe; PM = paramere." httpUri="https://zenodo.org/record/206625/files/figure.png" pageId="2" pageNumber="25">23</figureCitation>
,
<figureCitation id="13012A55803DFFF86098D96B59F7F939" box="[478,506,1647,1671]" captionStart="FIGURES 24 26" captionStartId="7.[151,264,1383,1406]" captionTargetBox="[193,1416,238,1334]" captionTargetId="figure@7.[171,1416,193,1362]" captionTargetPageId="7" captionText="FIGURES 24 26. Temones and flagellum of Amphicoma species. 24. Amphicoma lalashanensis; 25. Amphicoma aurata; 26. Amphicoma formosana. Abbreviations: FMaB = flagellum major body; MiS = minor sclerite; TE = temones. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/206626/files/figure.png" pageId="2" pageNumber="25">24</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C320655B803DFFFF61D1D9BC5CB8FDA9" lastPageId="5" lastPageNumber="28" pageId="2" pageNumber="25" type="materials_examined">
<paragraph id="8B8536D0803DFFF861D1D9BC5D9EF84E" blockId="2.[151,1436,1719,2033]" pageId="2" pageNumber="25">
<emphasis id="B94EEAC2803DFFF861D1D9BC5948F96E" bold="true" box="[151,325,1720,1744]" pageId="2" pageNumber="25">
<typeStatus id="54818872803DFFF861D1D9BC5909F96E" box="[151,260,1720,1744]" pageId="2" pageNumber="25" type="holotype">Holotype</typeStatus>
male
</emphasis>
(
<figureCitation id="13012A55803DFFF86012D9B3599BF971" box="[340,406,1719,1743]" captionStart="FIGURE 1 7" captionStartId="3.[151,250,1593,1616]" captionTargetBox="[191,1402,196,1557]" captionTargetId="figure@3.[173,1413,171,1572]" captionTargetPageId="3" captionText="FIGURE 1 7. Dorsal habitus of Amphicoma species. 1. Amphicoma lalashanensis, holotype; 2. Amphicoma aurata, male; 3. Amphicoma aurata, male; 4. Amphicoma aurata, female; 5. Amphicoma formosana, male; 6. Amphicoma formosana, male; 7. Amphicoma formosana, female." httpUri="https://zenodo.org/record/206620/files/figure.png" pageId="2" pageNumber="25">Fig. 1</figureCitation>
). Body oval-elongate, length 12.5 mm; width across humeri 4.1 mm. Body with dorsal surface and venter of thorax metallic green with some goldish green undertones on head, pronotum and basal half of elytra. Pygidium yellowish brown with no metallic reflection. Legs black. Antennae dark brown. Dorsum setose with golden brown setae moderately long, intermixed with longer stout black setae on canthus and along lateral sides of pronotum and apical half surface of elytra; dark brown setae on head and anterior half of pronotum shorter and softer.
<emphasis id="B94EEAC2803DFFF86054D8685955F83A" box="[274,344,1900,1924]" italics="true" pageId="2" pageNumber="25">Head:</emphasis>
Surface densely rugopunctate, puncture large, each with an erect seta. Anterior margin of labrum straight with lateral angles rounded. Clypeus rectangular; surface flat, slightly widened basally; anterior angle rounded; margins of clypeus with complete bead. Clypeofrontal suture weakly defined, almost invisible. Frons narrowing basally. Antenna with 10 antennomeres, three-antennomere club approximately 2.4 times as long as stem.
</paragraph>
<caption id="DF456658803CFFF961D1D93D59D6F933" httpUri="https://zenodo.org/record/206620/files/figure.png" pageId="3" pageNumber="26" targetBox="[191,1402,196,1557]" targetPageId="3">
<paragraph id="8B8536D0803CFFF961D1D93D59D6F933" blockId="3.[151,1436,1593,1677]" pageId="3" pageNumber="26">
<emphasis id="B94EEAC2803CFFF961D1D93D5920F9F1" bold="true" box="[151,301,1593,1616]" pageId="3" pageNumber="26">FIGURE 17.</emphasis>
Dorsal habitus of
<taxonomicName id="4C3A4D53803CFFF960AAD93D5A68F9F1" box="[492,613,1593,1615]" class="Insecta" family="Glaphyridae" genus="Amphicoma" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEAC2803CFFF960AAD93D5A68F9F1" box="[492,613,1593,1615]" italics="true" pageId="3" pageNumber="26">Amphicoma</emphasis>
</taxonomicName>
species. 1.
<taxonomicName id="4C3A4D53803CFFF96398D93D5BE4F9F1" box="[734,1001,1593,1615]" class="Insecta" family="Glaphyridae" genus="Amphicoma" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="lalashanensis">
<emphasis id="B94EEAC2803CFFF96398D93D5BE4F9F1" box="[734,1001,1593,1615]" italics="true" pageId="3" pageNumber="26">Amphicoma lalashanensis</emphasis>
</taxonomicName>
, holotype; 2.
<taxonomicName id="4C3A4D53803CFFF96531D93D5D31F9EE" box="[1143,1340,1593,1616]" class="Insecta" family="Glaphyridae" genus="Amphicoma" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="aurata">
<emphasis id="B94EEAC2803CFFF96531D93D5D31F9EE" box="[1143,1340,1593,1616]" italics="true" pageId="3" pageNumber="26">Amphicoma aurata</emphasis>
</taxonomicName>
, male; 3.
<taxonomicName id="4C3A4D53803CFFF961D1D95C5957F9D0" box="[151,346,1624,1646]" class="Insecta" family="Glaphyridae" genus="Amphicoma" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="aurata">
<emphasis id="B94EEAC2803CFFF961D1D95C5957F9D0" box="[151,346,1624,1646]" italics="true" pageId="3" pageNumber="26">Amphicoma aurata</emphasis>
</taxonomicName>
, male; 4.
<taxonomicName id="4C3A4D53803CFFF96086D95C5A89F9D0" box="[448,644,1624,1646]" class="Insecta" family="Glaphyridae" genus="Amphicoma" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="aurata">
<emphasis id="B94EEAC2803CFFF96086D95C5A89F9D0" box="[448,644,1624,1646]" italics="true" pageId="3" pageNumber="26">Amphicoma aurata</emphasis>
</taxonomicName>
, female; 5.
<taxonomicName id="4C3A4D53803CFFF963B8D95C5BE7F9D0" box="[766,1002,1624,1646]" class="Insecta" family="Glaphyridae" genus="Amphicoma" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="formosana">
<emphasis id="B94EEAC2803CFFF963B8D95C5BE7F9D0" box="[766,1002,1624,1646]" italics="true" pageId="3" pageNumber="26">Amphicoma formosana</emphasis>
</taxonomicName>
, male; 6.
<taxonomicName id="4C3A4D53803CFFF96516D95C5D31F9D0" box="[1104,1340,1624,1646]" class="Insecta" family="Glaphyridae" genus="Amphicoma" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="formosana">
<emphasis id="B94EEAC2803CFFF96516D95C5D31F9D0" box="[1104,1340,1624,1646]" italics="true" pageId="3" pageNumber="26">Amphicoma formosana</emphasis>
</taxonomicName>
, male; 7.
<taxonomicName id="4C3A4D53803CFFF961D1D973598FF933" box="[151,386,1655,1677]" class="Insecta" family="Glaphyridae" genus="Amphicoma" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="formosana">
<emphasis id="B94EEAC2803CFFF961D1D973598FF933" box="[151,386,1655,1677]" italics="true" pageId="3" pageNumber="26">Amphicoma formosana</emphasis>
</taxonomicName>
, female.
</paragraph>
</caption>
<paragraph id="8B8536D0803CFFFF61D1D9BE5CB8FDA9" blockId="3.[151,1436,1721,2034]" lastBlockId="5.[151,1437,151,535]" lastPageId="5" lastPageNumber="28" pageId="3" pageNumber="26">
<emphasis id="B94EEAC2803CFFF961D1D9BE58FCF96C" box="[151,241,1722,1746]" italics="true" pageId="3" pageNumber="26">Thorax:</emphasis>
Pronotum widest at anterior one-third, round in shape, anterior angles weakly developed with basolateral angles rounded, lateral margin explanate on basal two-thirds, middle of basal margin slightly emarginate; setiferous punctures densely distributed on surface; punctures smaller, more well developed than those on head. Scutellum elongate, triangular, flat; apex rounded; punctures dense, similar to punctures on pronotum. Prosternal process (
<figureCitation id="13012A55803CFFF961D9D84D58EEF8DF" box="[159,227,1865,1889]" captionStart="FIGURE 8 10" captionStartId="4.[151,250,638,661]" captionTargetBox="[158,1432,199,609]" captionTargetId="figure@4.[151,1436,193,617]" captionTargetPageId="4" captionText="FIGURE 8 10. Left lateral view of prosternal process of Amphicoma species. 8. Amphicoma lalashanensis; 9. Amphicoma aurata; 10. Amphicoma formosana." httpUri="https://zenodo.org/record/206621/files/figure.png" pageId="3" pageNumber="26">Fig. 8</figureCitation>
) protruding with quadrate apex glabrous, inner margin straight; metasternum with midline impressed, well defined, setose laterally; setae as long as setae on prosternum, mesepisternum, metepisternum, mesepimeron, metepimeron; setae on discal center of metasternum thin, shorter than setae on sides.
<emphasis id="B94EEAC2803CFFF9652ED8965CB5F814" box="[1128,1208,1938,1962]" italics="true" pageId="3" pageNumber="26">Elytra:</emphasis>
Disc surface nearly flat with sutural margins gradually raised apically, discal surface densely rugopunctate; ridged-like epipleura widely expanded, gradually narrowing apically.
<emphasis id="B94EEAC2803CFFF963F4D8DE5B28F84C" box="[690,805,2010,2034]" italics="true" pageId="3" pageNumber="26">Abdomen:</emphasis>
Pygidial width 2.3 times length medially; disc covered with long setae, slightly convex. Sternites 16 with surface setose. Legs: Protibia tridentate. Protarsi pectinate with tarsomeres 14 strongly condensed in length. Surface of profemora and mesofemora moderately covered with long setae, metafemora with short, sparse setae intermixed with longer setae. Mesotibiae and metatibiae inner surface with soft, dense setae and outer surface with more robust, sparse setae. Mesotibia with one inner apical spur, spur subequal in length to half of basal mesotarsomeres; basal mesotarsomere 1.7 times as long as second mesotarsomere. Basal metatarsomere 1.4 times as long as second metatarsomere. Male genitalia: Parameres (
<figureCitation id="13012A55803AFFFF65A4DE485D32FEDA" box="[1250,1343,332,356]" captionStart="FIGURES 11 13" captionStartId="4.[151,264,1881,1904]" captionTargetBox="[159,1421,798,1845]" captionTargetId="figure@4.[151,1436,771,1860]" captionTargetPageId="4" captionText="FIGURES 11 13. Dorsal view of male genitalia of Amphicoma species. 11. Amphicoma lalashanensis; 12. Amphicoma aurata; 13. Amphicoma formosana. Abbreviations: BP-ext = basal piece extension; FMaB = flagellum major body. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/206622/files/figure.png" pageId="5" pageNumber="28">Figs. 11</figureCitation>
,
<figureCitation id="13012A55803AFFFF640ADE485D99FEDA" box="[1356,1428,332,356]" captionStart="FIGURES 14 19" captionStartId="5.[151,264,1350,1373]" captionTargetBox="[160,1426,598,1322]" captionTargetId="figure@5.[151,1436,589,1329]" captionTargetPageId="5" captionText="FIGURES 14 19. Parameres of Amphicoma male genitalia in lateral view. 14 15. Amphicoma lalashanensis; 16 17. Amphicoma auratas; 18 19. Amphicoma formosana. Abbreviations: FMaB = flagellum major body. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/206623/files/figure.png" pageId="5" pageNumber="28">14, 15</figureCitation>
) asymmetrical, tip of left paramere upwardly curved with basal margin concave at basal one-third (
<figureCitation id="13012A55803AFFFF65B2DE6B5D45FE39" box="[1268,1352,367,391]" captionStart="FIGURES 14 19" captionStartId="5.[151,264,1350,1373]" captionTargetBox="[160,1426,598,1322]" captionTargetId="figure@5.[151,1436,589,1329]" captionTargetPageId="5" captionText="FIGURES 14 19. Parameres of Amphicoma male genitalia in lateral view. 14 15. Amphicoma lalashanensis; 16 17. Amphicoma auratas; 18 19. Amphicoma formosana. Abbreviations: FMaB = flagellum major body. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/206623/files/figure.png" pageId="5" pageNumber="28">Fig. 14</figureCitation>
). Basal piece extension reaching apical one-fourth of paramere (
<figureCitation id="13012A55803AFFFF626EDE905B70FE12" box="[808,893,404,428]" captionStart="FIGURES 11 13" captionStartId="4.[151,264,1881,1904]" captionTargetBox="[159,1421,798,1845]" captionTargetId="figure@4.[151,1436,771,1860]" captionTargetPageId="4" captionText="FIGURES 11 13. Dorsal view of male genitalia of Amphicoma species. 11. Amphicoma lalashanensis; 12. Amphicoma aurata; 13. Amphicoma formosana. Abbreviations: BP-ext = basal piece extension; FMaB = flagellum major body. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/206622/files/figure.png" pageId="5" pageNumber="28">Fig. 11</figureCitation>
,
<figureCitation id="13012A55803AFFFF62CADE905BA1FE12" box="[908,940,404,428]" captionStart="FIGURES 23" captionStartId="6.[151,264,1979,2002]" captionTargetBox="[566,1018,1285,1965]" captionTargetId="figure@6.[566,1020,1285,1965]" captionTargetPageId="6" captionText="FIGURES 23. Male genitalia of Amphicoma lalashanensis in ventral view. Abbreviations: BP = basal piece; BP-ext = basal piece extension; FPM = fusion of paramere and median lobe; ML = median lobe; PM = paramere." httpUri="https://zenodo.org/record/206625/files/figure.png" pageId="5" pageNumber="28">23</figureCitation>
). Ventral plate (
<figureCitation id="13012A55803AFFFF652EDE905CB0FE12" box="[1128,1213,404,428]" captionStart="FIGURES 20 22. 20" captionStartId="6.[151,264,1150,1173]" captionTargetBox="[170,1430,323,1122]" captionTargetId="figure@6.[151,1436,309,1129]" captionTargetPageId="6" captionText="FIGURES 20 22. 20. Amphicoma lalashanensis genital capsule; 21. Amphicoma aurata ventral plate of genital capsule; 22. Amphicoma formosana ventral plate of genital capsule. Abbreviations: LLP = left lateral plate; MS = median sclerite; RLP = right lateral plate; VP = ventral plate. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/206624/files/figure.png" pageId="5" pageNumber="28">Fig. 20</figureCitation>
) of genital capsule broadly widened, margins rounded, smooth; median stria well sclerotized, reaching apical one-third. Temones strongly twisted twice and fused apically, minor sclerite longitudinally lying within base of flagellum, flagellum separated basally, longer than paramere and subequal in length to half of basal piece (
<figureCitation id="13012A55803AFFFF6512DEFB5CA4FDA9" box="[1108,1193,511,535]" captionStart="FIGURES 24 26" captionStartId="7.[151,264,1383,1406]" captionTargetBox="[193,1416,238,1334]" captionTargetId="figure@7.[171,1416,193,1362]" captionTargetPageId="7" captionText="FIGURES 24 26. Temones and flagellum of Amphicoma species. 24. Amphicoma lalashanensis; 25. Amphicoma aurata; 26. Amphicoma formosana. Abbreviations: FMaB = flagellum major body; MiS = minor sclerite; TE = temones. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/206626/files/figure.png" pageId="5" pageNumber="28">Fig. 24</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF456658803BFFFE61D1DD7A59F2FD0C" httpUri="https://zenodo.org/record/206621/files/figure.png" pageId="4" pageNumber="27" targetBox="[158,1432,199,609]" targetPageId="4">
<paragraph id="8B8536D0803BFFFE61D1DD7A59F2FD0C" blockId="4.[151,1436,638,691]" pageId="4" pageNumber="27">
<emphasis id="B94EEAC2803BFFFE61D1DD7A5936FD2D" bold="true" box="[151,315,638,661]" pageId="4" pageNumber="27">FIGURE 810.</emphasis>
Left lateral view of prosternal process of
<taxonomicName id="4C3A4D53803BFFFE63B0DD7A5B62FD2A" box="[758,879,638,660]" class="Insecta" family="Glaphyridae" genus="Amphicoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEAC2803BFFFE63B0DD7A5B62FD2A" box="[758,879,638,660]" italics="true" pageId="4" pageNumber="27">Amphicoma</emphasis>
</taxonomicName>
species. 8.
<taxonomicName id="4C3A4D53803BFFFE62AADD7A5CF5FD2A" box="[1004,1272,638,660]" class="Insecta" family="Glaphyridae" genus="Amphicoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="lalashanensis">
<emphasis id="B94EEAC2803BFFFE62AADD7A5CF5FD2A" box="[1004,1272,638,660]" italics="true" pageId="4" pageNumber="27">Amphicoma lalashanensis</emphasis>
</taxonomicName>
; 9.
<taxonomicName id="4C3A4D53803BFFFE6465DD7A58D6FD0D" class="Insecta" family="Glaphyridae" genus="Amphicoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="aurata">
<emphasis id="B94EEAC2803BFFFE6465DD7A58D6FD0D" italics="true" pageId="4" pageNumber="27">Amphicoma aurata</emphasis>
</taxonomicName>
; 10.
<taxonomicName id="4C3A4D53803BFFFE604BDD9959F5FD0D" box="[269,504,669,691]" class="Insecta" family="Glaphyridae" genus="Amphicoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="formosana">
<emphasis id="B94EEAC2803BFFFE604BDD9959F5FD0D" box="[269,504,669,691]" italics="true" pageId="4" pageNumber="27">Amphicoma formosana</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<caption id="DF456658803BFFFE61D1D85D58E1F813" httpUri="https://zenodo.org/record/206622/files/figure.png" pageId="4" pageNumber="27" targetBox="[159,1421,798,1845]" targetPageId="4">
<paragraph id="8B8536D0803BFFFE61D1D85D58E1F813" blockId="4.[151,1436,1881,1965]" pageId="4" pageNumber="27">
<emphasis id="B94EEAC2803BFFFE61D1D85D595AF8D1" bold="true" box="[151,343,1881,1904]" pageId="4" pageNumber="27">FIGURES 1113.</emphasis>
Dorsal view of male genitalia of
<taxonomicName id="4C3A4D53803BFFFE6388D85D5B4AF8D1" box="[718,839,1881,1903]" class="Insecta" family="Glaphyridae" genus="Amphicoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEAC2803BFFFE6388D85D5B4AF8D1" box="[718,839,1881,1903]" italics="true" pageId="4" pageNumber="27">Amphicoma</emphasis>
</taxonomicName>
species. 11.
<taxonomicName id="4C3A4D53803BFFFE6291D85D5CEBF8D1" box="[983,1254,1881,1903]" class="Insecta" family="Glaphyridae" genus="Amphicoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="lalashanensis">
<emphasis id="B94EEAC2803BFFFE6291D85D5CEBF8D1" box="[983,1254,1881,1903]" italics="true" pageId="4" pageNumber="27">Amphicoma lalashanensis</emphasis>
</taxonomicName>
; 12.
<taxonomicName id="4C3A4D53803BFFFE6465D85D58D6F830" class="Insecta" family="Glaphyridae" genus="Amphicoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="aurata">
<emphasis id="B94EEAC2803BFFFE6465D85D58D6F830" italics="true" pageId="4" pageNumber="27">Amphicoma aurata</emphasis>
</taxonomicName>
; 13.
<taxonomicName id="4C3A4D53803BFFFE604BD87C59F4F830" box="[269,505,1912,1934]" class="Insecta" family="Glaphyridae" genus="Amphicoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="formosana">
<emphasis id="B94EEAC2803BFFFE604BD87C59F4F830" box="[269,505,1912,1934]" italics="true" pageId="4" pageNumber="27">Amphicoma formosana</emphasis>
</taxonomicName>
. Abbreviations: BP-ext = basal piece extension; FMaB = flagellum major body. Scale bar = 1.0 mm.
</paragraph>
</caption>
<caption id="DF456658803AFFFF61D1DA425D12FAC5" httpUri="https://zenodo.org/record/206623/files/figure.png" pageId="5" pageNumber="28" targetBox="[160,1426,598,1322]" targetPageId="5">
<paragraph id="8B8536D0803AFFFF61D1DA425D12FAC5" blockId="5.[151,1436,1350,1403]" pageId="5" pageNumber="28">
<emphasis id="B94EEAC2803AFFFF61D1DA425959FAE5" bold="true" box="[151,340,1350,1373]" pageId="5" pageNumber="28">FIGURES 1419.</emphasis>
Parameres of
<taxonomicName id="4C3A4D53803AFFFF60AFDA425A6FFAE2" box="[489,610,1350,1372]" class="Insecta" family="Glaphyridae" genus="Amphicoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEAC2803AFFFF60AFDA425A6FFAE2" box="[489,610,1350,1372]" italics="true" pageId="5" pageNumber="28">Amphicoma</emphasis>
</taxonomicName>
male genitalia in lateral view. 1415.
<taxonomicName id="4C3A4D53803AFFFF62AADA425CFBFAE2" box="[1004,1270,1350,1372]" class="Insecta" family="Glaphyridae" genus="Amphicoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="28" phylum="Arthropoda" rank="species" species="lalashanensis">
<emphasis id="B94EEAC2803AFFFF62AADA425CFBFAE2" box="[1004,1270,1350,1372]" italics="true" pageId="5" pageNumber="28">Amphicoma lalashanensis</emphasis>
</taxonomicName>
; 1617.
<taxonomicName id="4C3A4D53803AFFFF6417DA42592CFAC5" class="Insecta" family="Glaphyridae" genus="Amphicoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="28" phylum="Arthropoda" rank="species" species="auratas">
<emphasis id="B94EEAC2803AFFFF6417DA42592CFAC5" italics="true" pageId="5" pageNumber="28">Amphicoma auratas</emphasis>
</taxonomicName>
; 1819.
<taxonomicName id="4C3A4D53803AFFFF603DDA615A6BFAC5" box="[379,614,1381,1403]" class="Insecta" family="Glaphyridae" genus="Amphicoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="28" phylum="Arthropoda" rank="species" species="formosana">
<emphasis id="B94EEAC2803AFFFF603DDA615A6BFAC5" box="[379,614,1381,1403]" italics="true" pageId="5" pageNumber="28">Amphicoma formosana</emphasis>
</taxonomicName>
. Abbreviations: FMaB = flagellum major body. Scale bar = 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C320655B803AFFFF6183DAAC59ADFA01" box="[197,416,1447,1472]" pageId="5" pageNumber="28" type="description">
<paragraph id="8B8536D0803AFFFF6183DAAC59ADFA01" blockId="5.[151,1437,1447,2012]" box="[197,416,1447,1472]" pageId="5" pageNumber="28">
<emphasis id="B94EEAC2803AFFFF6183DAAC592EFA7E" bold="true" box="[197,291,1448,1472]" pageId="5" pageNumber="28">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C320655B803AFFFF6183DACF5A79F905" pageId="5" pageNumber="28" type="diagnosis">
<paragraph id="8B8536D0803AFFFF6183DACF5A79F905" blockId="5.[151,1437,1447,2012]" pageId="5" pageNumber="28">
<emphasis id="B94EEAC2803AFFFF6183DACF5932FA5A" bold="true" box="[197,319,1483,1508]" pageId="5" pageNumber="28">Diagnosis.</emphasis>
<taxonomicName id="4C3A4D53803AFFFF6000DAC85A50FA5A" box="[326,605,1484,1508]" class="Insecta" family="Glaphyridae" genus="Amphicoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="28" phylum="Arthropoda" rank="species" species="lalashaensis">
<emphasis id="B94EEAC2803AFFFF6000DAC85A50FA5A" box="[326,605,1484,1508]" italics="true" pageId="5" pageNumber="28">Amphicoma lalashaensis</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="4C3A4D53803AFFFF63B7DAC85C32FA5D" authority="Nikodym, 2005" authorityName="Nikodym" authorityYear="2005" box="[753,1087,1483,1508]" class="Insecta" family="Glaphyridae" genus="Amphicoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="28" phylum="Arthropoda" rank="species" species="schneideri">
<emphasis id="B94EEAC2803AFFFF63B7DAC85B88FA5A" box="[753,901,1484,1508]" italics="true" pageId="5" pageNumber="28">A. schneideri</emphasis>
Nikodým, 2005
</taxonomicName>
, found in the highland of Sichuan, southwestern
<collectingCountry id="F32D7640803AFFFF6021DAEB59A0F9B9" box="[359,429,1519,1543]" name="China" pageId="5" pageNumber="28">China</collectingCountry>
, based on the general metallic green coloration, setal distribution, and strongly curved and elongated antennal club. However, it differs from
<taxonomicName id="4C3A4D53803AFFFF638DD9105B6DF992" box="[715,864,1556,1580]" class="Insecta" family="Glaphyridae" genus="Amphicoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="28" phylum="Arthropoda" rank="species" species="schneideri">
<emphasis id="B94EEAC2803AFFFF638DD9105B6DF992" box="[715,864,1556,1580]" italics="true" pageId="5" pageNumber="28">A. schneideri</emphasis>
</taxonomicName>
in the dorsal surface partly goldish green on head, pronotum and basal half of elytra (overall coppery green in
<taxonomicName id="4C3A4D53803AFFFF6274D93C5BC8F9EE" box="[818,965,1592,1616]" class="Insecta" family="Glaphyridae" genus="Amphicoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="28" phylum="Arthropoda" rank="species" species="schneideri">
<emphasis id="B94EEAC2803AFFFF6274D93C5BC8F9EE" box="[818,965,1592,1616]" italics="true" pageId="5" pageNumber="28">A. schneideri</emphasis>
</taxonomicName>
); legs black (rather than reddish brown as in
<taxonomicName id="4C3A4D53803AFFFF61F2D9585945F9CA" box="[180,328,1628,1652]" class="Insecta" family="Glaphyridae" genus="Amphicoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="28" phylum="Arthropoda" rank="species" species="schneideri">
<emphasis id="B94EEAC2803AFFFF61F2D9585945F9CA" box="[180,328,1628,1652]" italics="true" pageId="5" pageNumber="28">A. schneideri</emphasis>
</taxonomicName>
), basal metatarsomere 1.4 times as long as second metatarsomere (1.8 times in
<taxonomicName id="4C3A4D53803AFFFF6581D9585D56F9CA" box="[1223,1371,1628,1652]" class="Insecta" family="Glaphyridae" genus="Amphicoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="28" phylum="Arthropoda" rank="species" species="schneideri">
<emphasis id="B94EEAC2803AFFFF6581D9585D56F9CA" box="[1223,1371,1628,1652]" italics="true" pageId="5" pageNumber="28">A. schneideri</emphasis>
</taxonomicName>
); and in the tip and basal margin of left paramere in lateral view (
<figureCitation id="13012A55803AFFFF626ED97B5B74F929" box="[808,889,1663,1687]" captionStart="FIGURES 14 19" captionStartId="5.[151,264,1350,1373]" captionTargetBox="[160,1426,598,1322]" captionTargetId="figure@5.[151,1436,589,1329]" captionTargetPageId="5" captionText="FIGURES 14 19. Parameres of Amphicoma male genitalia in lateral view. 14 15. Amphicoma lalashanensis; 16 17. Amphicoma auratas; 18 19. Amphicoma formosana. Abbreviations: FMaB = flagellum major body. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/206623/files/figure.png" pageId="5" pageNumber="28">Fig. 14</figureCitation>
) (tip slightly upwardly curved with more smooth basal margin of paramere in
<taxonomicName id="4C3A4D53803AFFFF6091D9A05A66F902" box="[471,619,1700,1724]" class="Insecta" family="Glaphyridae" genus="Amphicoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="28" phylum="Arthropoda" rank="species" species="schneideri">
<emphasis id="B94EEAC2803AFFFF6091D9A05A66F902" box="[471,619,1700,1724]" italics="true" pageId="5" pageNumber="28">A. schneideri</emphasis>
</taxonomicName>
)
</paragraph>
</subSubSection>
<subSubSection id="C320655B803AFFFF6183D9CC59DDF899" pageId="5" pageNumber="28" type="materials_examined">
<paragraph id="8B8536D0803AFFFF6183D9CC59DDF899" blockId="5.[151,1437,1447,2012]" pageId="5" pageNumber="28">
<emphasis id="B94EEAC2803AFFFF6183D9CC5974F95E" bold="true" box="[197,377,1735,1760]" pageId="5" pageNumber="28">
<typeStatus id="54818872803AFFFF6183D9CC590DF95E" box="[197,256,1736,1760]" pageId="5" pageNumber="28">Type</typeStatus>
material.
</emphasis>
The male
<typeStatus id="54818872803AFFFF6345D9C35A68F961" box="[515,613,1735,1759]" pageId="5" pageNumber="28" type="holotype">holotype</typeStatus>
is pinned and with the following information on the label:
<collectingCountry id="F32D7640803AFFFF6404D9C35D95F961" box="[1346,1432,1735,1759]" name="Taiwan" pageId="5" pageNumber="28">Taiwan</collectingCountry>
: Taoyuan, Lalashan,
<date id="FF841010803AFFFF603CD9EF5A1AF8BD" box="[378,535,1771,1795]" pageId="5" pageNumber="28" value="2009-07-15">15. VII. 2009</date>
, leg. H.-J. Chen. The
<typeStatus id="54818872803AFFFF6254D9EF5B79F8BD" box="[786,884,1771,1795]" pageId="5" pageNumber="28" type="holotype">holotype</typeStatus>
is deposited at the
<collectingCountry id="F32D7640803AFFFF6514D9EF5CA9F8BD" box="[1106,1188,1771,1795]" name="Taiwan" pageId="5" pageNumber="28">Taiwan</collectingCountry>
Agriculture Research Institute, Taichung,
<collectingCountry id="F32D7640803AFFFF6030D80B59C1F899" box="[374,460,1807,1831]" name="Taiwan" pageId="5" pageNumber="28">Taiwan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C320655B803AFFFF6183D8375A26F8F5" box="[197,555,1843,1868]" pageId="5" pageNumber="28" type="distribution">
<paragraph id="8B8536D0803AFFFF6183D8375A26F8F5" blockId="5.[151,1437,1447,2012]" box="[197,555,1843,1868]" pageId="5" pageNumber="28">
<emphasis id="B94EEAC2803AFFFF6183D8375953F8F2" bold="true" box="[197,350,1843,1868]" pageId="5" pageNumber="28">Distribution.</emphasis>
Northern
<collectingCountry id="F32D7640803AFFFF6097D8375A2AF8F5" box="[465,551,1843,1867]" name="Taiwan" pageId="5" pageNumber="28">Taiwan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C320655B803AFFFF6183D85C5A78F82D" pageId="5" pageNumber="28" type="etymology">
<paragraph id="8B8536D0803AFFFF6183D85C5A78F82D" blockId="5.[151,1437,1447,2012]" pageId="5" pageNumber="28">
<emphasis id="B94EEAC2803AFFFF6183D85C5941F8CE" bold="true" box="[197,332,1880,1904]" pageId="5" pageNumber="28">Etymology.</emphasis>
The specific name refers to the famous historical collecting site, Lalashan (presently the Dakwanshan Natural Reserve) of northern
<collectingCountry id="F32D7640803AFFFF635DD87F5A7CF82D" box="[539,625,1915,1939]" name="Taiwan" pageId="5" pageNumber="28">Taiwan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C320655B803AFFFC6183D8A45CB9FEA2" lastPageId="6" lastPageNumber="29" pageId="5" pageNumber="28" type="discussion">
<paragraph id="8B8536D0803AFFFC6183D8A45CB9FEA2" blockId="5.[151,1437,1447,2012]" lastBlockId="6.[151,1436,151,285]" lastPageId="6" lastPageNumber="29" pageId="5" pageNumber="28">
<emphasis id="B94EEAC2803AFFFF6183D8A45934F806" bold="true" box="[197,313,1952,1976]" pageId="5" pageNumber="28">Remarks.</emphasis>
This new species seems to be quite rare because the
<typeStatus id="54818872803AFFFF62C1D8A55BBAF807" box="[903,951,1953,1977]" pageId="5" pageNumber="28">type</typeStatus>
locality has been well collected by naturalists over a long period of time. This region is of interest as a representative of primary mid-elevated cloud forests in northern
<collectingCountry id="F32D76408039FFFC61B8DF935959FF11" box="[254,340,151,175]" name="Taiwan" pageId="6" pageNumber="29">Taiwan</collectingCountry>
. There were no known collecting records of this new species until the
<typeStatus id="548188728039FFFC6538DF935CEDFF11" box="[1150,1248,151,175]" pageId="6" pageNumber="29" type="holotype">holotype</typeStatus>
was captured in 2009. The
<typeStatus id="548188728039FFFC6049DFB8597CFF6A" box="[271,369,188,212]" pageId="6" pageNumber="29" type="holotype">holotype</typeStatus>
male was found resting on shrub leaves by a trail within the forest. According to Marco Uliana (Museo di Storia Naturale di Venezia, Venezia,
<collectingCountry id="F32D76408039FFFC63F5DFDB5AEAFF49" box="[691,743,223,247]" name="Italy" pageId="6" pageNumber="29">Italy</collectingCountry>
, personal communication), some glaphyrid adults that do not visit flowers and rarely encountered because they appear as adults for only very short periods.
</paragraph>
</subSubSection>
</treatment>
</document>