489 lines
83 KiB
XML
489 lines
83 KiB
XML
<document id="37D35F999B1AD69719AEDF9CF1CBDA33" ID-DOI="10.11646/zootaxa.5486.3.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="13210005" ID-ZooBank="C7F7E7D5-6F6E-49C8-AF31-9B769EC8B56C" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722788280951" checkinUser="plazi" docAuthor="Orr, Albert G. W., Dow, Rory A. & Steinhoff, Philip O. M." docDate="2024" docId="0393879EFFDEFFDBDBF74F46FD907954" docLanguage="en" docName="zootaxa.5486.2.3.1.pdf" docOrigin="Zootaxa 5486 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5486.3.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Heliocypha biseriata" docType="treatment" docVersion="2" lastPageNumber="314" masterDocId="FFAAFFE6FFD8FFD5DB60480DFF897815" masterDocTitle="Descriptions of larvae of four mainly DNA barcode-matched species of chlorocyphids from south-east Asia (Odonata: Chlorocyphidae) with notes on the generic and species level larval identification of Oriental region members of the family." masterLastPageNumber="337" masterPageNumber="301" pageNumber="307" updateTime="1722788985873" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="3BBE402D7A3AE49815E5E03516FA691E">Descriptions of larvae of four mainly DNA barcode-matched species of chlorocyphids from south-east Asia (Odonata: Chlorocyphidae) with notes on the generic and species level larval identification of Oriental region members of the family.</mods:title>
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<mods:name id="6CAA5C403B5A895E239787C3484234CF" type="personal">
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<mods:namePart id="ADCB64032663599CFC67847682716218">Orr, Albert G. W.</mods:namePart>
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<mods:affiliation id="B6AB9010F15C53E03C8B11A461E73927">Environmental Futures Research Institute, Griffith University, Nathan, Australia.</mods:affiliation>
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<mods:namePart id="12CAEE35186CC9E29C8A8910FCDC805C">Dow, Rory A.</mods:namePart>
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<mods:affiliation id="A156F06836D17DE97965E9D609B00444">Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, The Netherlands. & Institute of Biodiversity and Environmental Conservation, Universiti Malaysia Sarawak, 94300 Kota Samarahan, Sarawak, Malaysia.</mods:affiliation>
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<mods:namePart id="48A9706A941CCDC41AA7B25980D63B85">Steinhoff, Philip O. M.</mods:namePart>
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<mods:affiliation id="13CFD69B2948EEBDC67A0CF760438BA1">Höxter, Germany.</mods:affiliation>
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<mods:typeOfResource id="5F7A51F1328F9267C50377241BCCF80A">text</mods:typeOfResource>
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<mods:title id="EBE6B469642447699DE4C36C374ACF3C">Zootaxa</mods:title>
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<mods:date id="BB29B8A55C90F6E0C1FC74C05062D833">2024</mods:date>
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<mods:number id="CD4D6D4426E3879D7CE6F182AD19DC69">2024-07-26</mods:number>
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<mods:number id="8984575AF1D65830E66690E6E3885BC7">5486</mods:number>
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<mods:number id="52F5AD39B49A07E570577E0938B15775">3</mods:number>
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<mods:start id="4E01C9F50DFC1953FB231C4D371396F1">301</mods:start>
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<mods:url id="48BEF3F5B6A85441C603C8CD0E4843C7">http://dx.doi.org/10.11646/zootaxa.5486.3.1</mods:url>
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<treatment id="0393879EFFDEFFDBDBF74F46FD907954" LSID="urn:lsid:plazi:treatment:0393879EFFDEFFDBDBF74F46FD907954" httpUri="http://treatment.plazi.org/id/0393879EFFDEFFDBDBF74F46FD907954" lastPageId="14" lastPageNumber="314" pageId="6" pageNumber="307">
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<subSubSection id="C3206503FFDEFFD3DBF74F46FE627F73" box="[151,491,1867,1894]" pageId="6" pageNumber="307" type="nomenclature">
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<paragraph id="8B853688FFDEFFD3DBF74F46FE627F73" blockId="6.[151,974,1867,1931]" box="[151,491,1867,1894]" pageId="6" pageNumber="307">
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<heading id="D0CD81E4FFDEFFD3DBF74F46FE627F73" bold="true" box="[151,491,1867,1894]" fontSize="11" level="1" pageId="6" pageNumber="307" reason="1">
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<taxonomicName id="4C3A4D0BFFDEFFD3DBF74F46FE627F73" ID-CoL="3KB5Q" authority="(Selys)" baseAuthorityName="Selys" box="[151,491,1867,1894]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="biseriata">
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<emphasis id="B94EEA9AFFDEFFD3DBF74F46FE627F73" bold="true" box="[151,491,1867,1894]" pageId="6" pageNumber="307">
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<emphasis id="B94EEA9AFFDEFFD3DBF74F46FE187F73" bold="true" box="[151,401,1867,1894]" italics="true" pageId="6" pageNumber="307">Heliocypha biseriata</emphasis>
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(Selys)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<subSubSection id="C3206503FFDEFFD3DBF74F7DFC477F9F" box="[151,974,1904,1931]" pageId="6" pageNumber="307" type="description">
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<paragraph id="8B853688FFDEFFD3DBF74F7DFC477F9F" blockId="6.[151,974,1867,1931]" box="[151,974,1904,1931]" pageId="6" pageNumber="307">
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(
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<figureCitation id="13012A0DFFDEFFD3DBFF4F7DFF7D7F9F" box="[159,244,1904,1930]" captionStart="FIGURE 4" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[302,1377,197,1855]" captionTargetId="figure-18@7.[172,1416,181,1919]" captionTargetPageId="7" captionText="FIGURE 4. Habitus of (a) Heliocypha biseriata, (b) Aristocypha fenestrella, (c) Sundacypha petiolata, (d) Libellago hyalina. Markings shown only on legs and antennae. Scale bar 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210013" httpUri="https://zenodo.org/record/13210013/files/figure.png" pageId="6" pageNumber="307">Figs 4a</figureCitation>
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,
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<figureCitation id="13012A0DFFDEFFD3DA604F7DFE957F9E" box="[256,284,1904,1931]" captionStart="FIGURE 5" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[206,1300,192,1855]" captionTargetId="figure-22@8.[196,1391,181,1919]" captionTargetPageId="8" captionText="FIGURE 5. Detail of heads, antennae and prothoraces of: (a) Heliocypha biseriata, (b) Aristocypha fenestrella, (c) Sundacypha petiolata, (d) Libellago hyalina. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210015" httpUri="https://zenodo.org/record/13210015/files/figure.png" pageId="6" pageNumber="307">5a</figureCitation>
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,
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<figureCitation id="13012A0DFFDEFFD3DA494F7DFE1E7F9E" box="[297,407,1904,1931]" captionStart="FIGURE 6" captionStartId="9.[151,250,1855,1879]" captionTargetBox="[153,1429,242,1805]" captionTargetId="figure-18@9.[151,1436,181,1831]" captionTargetPageId="9" captionText="FIGURE 6. (a) Frontal view of labrum of Heliocypha biseriata; (b, c) detail of scape showing short recurved and longer, mainly straighter setae on H. biseriata and Aristocypha fenestrella respectively; (d) Fore-tarsus in H. biseriata and (e) detail of pectinate setae on ventral side of same. Scale bars as indicated." figureDoi="http://doi.org/10.5281/zenodo.13210017" httpUri="https://zenodo.org/record/13210017/files/figure.png" pageId="6" pageNumber="307">6a, b, d, e</figureCitation>
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,
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<figureCitation id="13012A0DFFDEFFD3DAC34F7CFE497F9E" box="[419,448,1905,1931]" captionStart="FIGURE 7" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[228,1386,186,1919]" captionTargetId="figure-22@10.[155,1433,181,1919]" captionTargetPageId="10" captionText="FIGURE 7. Left lateral view of head (left) and ventral view of genae showing spines (right): (a) Heliocypha biseriata, (b) Aristocypha fenestrella, (c) Sundacypha petiolata, (d) Libellago hyalina. Scale bars left side, 1 mm; right side 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210019" httpUri="https://zenodo.org/record/13210019/files/figure.png" pageId="6" pageNumber="307">7a</figureCitation>
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,
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<figureCitation id="13012A0DFFDEFFD3DAAC4F7DFE607F9F" box="[460,489,1904,1930]" captionStart="FIGURE 8" captionStartId="11.[151,250,1675,1699]" captionTargetBox="[160,1432,215,1639]" captionTargetId="figure-18@11.[151,1436,181,1652]" captionTargetPageId="11" captionText="FIGURE 8. Ventral view of head with mask at rest and anterior two thoracic segments in ventral view: (a) Heliocypha biseriata, (b) Aristocypha fenestrella, (c) Sundacypha petiolata, (d) Libellago hyalina. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210021" httpUri="https://zenodo.org/record/13210021/files/figure.png" pageId="6" pageNumber="307">8a</figureCitation>
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,
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<figureCitation id="13012A0DFFDEFFD3DA944F7DFD987F9F" box="[500,529,1904,1930]" captionStart="FIGURE 9" captionStartId="17.[151,250,1943,1967]" captionTargetBox="[244,1341,217,1810]" captionTargetId="figure-18@17.[155,1433,181,1919]" captionTargetPageId="17" captionText="FIGURE 9. Prementum in dorsal view: (a) Heliocypha biseriata, (b) Aristocypha fenestrella, (c) Sundacypha petiolata, (d) Libellago hyalina, (e) Libellago lineata. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210023" httpUri="https://zenodo.org/record/13210023/files/figure.png" pageId="6" pageNumber="307">9a</figureCitation>
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,
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<figureCitation id="13012A0DFFDEFFD3D97D4F7DFDF47F9E" box="[541,637,1904,1931]" captionStart="FIGURE 10" captionStartId="18.[151,250,1943,1967]" captionTargetBox="[167,1433,182,1916]" captionTargetId="figure-22@18.[155,1433,181,1919]" captionTargetPageId="18" captionText="FIGURE 10. Details of anterior prementum and labial palp Heliocypha biseriata: (a) anterior prementum and labial palp in dorsal view (base of movable hook dotted to reveal details of palpal lobe below); (b) labial palp and anterior margin of prementum, frontal view; (c) labial palp, interior view. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210025" httpUri="https://zenodo.org/record/13210025/files/figure.png" pageId="6" pageNumber="307">10a, b, c</figureCitation>
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,
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<figureCitation id="13012A0DFFDEFFD3D9EA4F7DFD637F9E" box="[650,746,1904,1931]" captionStart="FIGURE 12" captionStartId="20.[151,250,1909,1933]" captionTargetBox="[205,1382,181,1764]" captionTargetId="figure-22@20.[205,1382,181,1884]" captionTargetPageId="20" captionText="FIGURE 12. Details of maxillae—(a–d) Right maxillae: Heliocypha biseriata, (a) ventral view, (b) oral view; (c) Sundacypha petiolata distal detail, oral view; (d) L. hyalina oral view. (e–h) Left maxillae, oral view, details of terminal strong spine with three shorter basal to median supplementary spines: (e) Heliocypha biseriata, (f) Aristocypha fenestrella, (g) Sundacypha petiolata, (h) Libellago hyalina. Scale bars: above 1 mm; below 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13210029" httpUri="https://zenodo.org/record/13210029/files/figure.png" pageId="6" pageNumber="307">12a, b, e</figureCitation>
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,
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<figureCitation id="13012A0DFFDEFFD3D9974F7DFCB47F9F" box="[759,829,1904,1930]" captionStart="FIGURE 13" captionStartId="21.[151,250,1909,1933]" captionTargetBox="[206,1359,208,1883]" captionTargetId="figure-18@21.[205,1382,181,1884]" captionTargetPageId="21" captionText="FIGURE 13. Mandibles of Heliocypha biseriata: (a) right mandible, (b) left mandible in frontal view; (c) right mandible, (d) left mandible in ventral view; (e) detail of right molar crest in profile; (f) slightly postero-ventro-interior view of left mandible showing marginal teeth and molar crest on edge; (e) detail of denticles of left molar crest. Scale bars a–d, f, 0.5 mm; e, g, 0.1 mm as indicated." figureDoi="http://doi.org/10.5281/zenodo.13210031" httpUri="https://zenodo.org/record/13210031/files/figure.png" pageId="6" pageNumber="307">13a–g</figureCitation>
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,
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<figureCitation id="13012A0DFFDEFFD3D8294F7DFC067F9F" box="[841,911,1904,1930]" captionStart="FIGURE 16" captionStartId="24.[151,250,1964,1988]" captionTargetBox="[158,1439,262,1931]" captionTargetId="figure-44@24.[149,1439,237,1940]" captionTargetPageId="24" captionText="FIGURE 16. Female gonapophyses and terminal abdominal segments in ventral view: (a) H. biseriata, (b) S. petiolata, (c) L. hyalina, and in lateral view: (d) H. biseriata, (e) L. hyalina, (f) S. petiolata. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210037" httpUri="https://zenodo.org/record/13210037/files/figure.png" pageId="6" pageNumber="307">16a, d</figureCitation>
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,
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<figureCitation id="13012A0DFFDEFFD3D8FB4F7DFC4D7F9F" box="[923,964,1904,1930]" captionStart="FIGURE 18" captionStartId="27.[151,250,1943,1967]" captionTargetBox="[191,1381,197,1917]" captionTargetId="figure-18@27.[189,1399,181,1919]" captionTargetPageId="27" captionText="FIGURE 18. Caudal spikes scaled in proportion to overall body length: (a) H. biseriata, (b) A. fenstrella, (c) S. petiolata, (d) L. hyalina. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210041" httpUri="https://zenodo.org/record/13210041/files/figure.png" pageId="6" pageNumber="307">18a</figureCitation>
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)
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</paragraph>
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<paragraph id="8B853688FFDEFFD3DBF74FB5FA0F7FE2" blockId="6.[151,1436,1976,2039]" pageId="6" pageNumber="307">
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<emphasis id="B94EEA9AFFDEFFD3DBF74FB5FEF77FC7" bold="true" box="[151,382,1976,2002]" pageId="6" pageNumber="307">Material examined.</emphasis>
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<materialsCitation id="3B523CD5FFDEFFD3DAE24FB5FA0F7FE2" collectingDate="2013-08-22" collectionCode="RMNH, INS" collectorName="A. Pasang." country="Malaysia" county="Sungai Datai" latitude="1.6493" location="Kapit Division" longLatPrecision="7" longitude="112.2627" municipality="Lanjak Entimau Wildlife Sanctuary" pageId="6" pageNumber="307" specimenCount="2" specimenCount-female="2" stateProvince="Sarawak">
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<collectingCountry id="F32D7618FFDEFFD3DAE24FB5FD9F7FC7" box="[386,534,1976,2002]" name="Malaysia" pageId="6" pageNumber="307">MALAYSIA</collectingCountry>
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,
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<specimenCount id="9D3CFD01FFDEFFD3D97F4FB5FDD27FC6" box="[543,603,1976,2003]" count="2" pageId="6" pageNumber="307" type="female">2 ♀♀</specimenCount>
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F (
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<collectionCode id="ED2BAE4DFFDEFFD3D91C4FB5FD5F7FC6" box="[636,726,1976,2003]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="6" pageNumber="307" type="Museum">RMNH</collectionCode>
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.
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<collectionCode id="ED2BAE4DFFDEFFD3D9B94FB5FC8E7FC6" box="[729,775,1976,2003]" pageId="6" pageNumber="307">INS</collectionCode>
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.557684,
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<collectionCode id="ED2BAE4DFFDEFFD3D80C4FB5FC4F7FC6" box="[876,966,1976,2003]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="6" pageNumber="307" type="Museum">RMNH</collectionCode>
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.
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<collectionCode id="ED2BAE4DFFDEFFD3D8A94FB5FC7E7FC6" box="[969,1015,1976,2003]" pageId="6" pageNumber="307">INS</collectionCode>
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.557685),
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<collectingCounty id="62E44E04FFDEFFD3DF044FB5FB7A7FC7" box="[1124,1267,1976,2002]" pageId="6" pageNumber="307">Sungai Datai</collectingCounty>
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and tributaries,
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<collectingMunicipality id="6BE1ACF2FFDEFFD3DBF74FD1FDAC7FE3" box="[151,549,2012,2038]" pageId="6" pageNumber="307">Lanjak Entimau Wildlife Sanctuary</collectingMunicipality>
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,
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<location id="8EE56053FFDEFFD3D9514FD1FD507FE3" LSID="urn:lsid:plazi:treatment:0393879EFFDEFFDBDBF74F46FD907954:8EE56053FFDEFFD3D9514FD1FD507FE3" box="[561,729,2012,2038]" country="Malaysia" county="Sungai Datai" latitude="1.6493" longLatPrecision="7" longitude="112.2627" municipality="Lanjak Entimau Wildlife Sanctuary" name="Kapit Division" pageId="6" pageNumber="307" stateProvince="Sarawak">Kapit Division</location>
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,
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<collectingRegion id="49FEF86AFFDEFFD3D9844FD1FCCE7FE3" box="[740,839,2012,2038]" country="Malaysia" name="Sarawak" pageId="6" pageNumber="307">Sarawak</collectingRegion>
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,
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,
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|
||
<collectingDate id="EFC0E9A0FFDEFFD3DF294FD1FB5A7FE3" box="[1097,1235,2012,2038]" pageId="6" pageNumber="307" value="2013-08-22">22 viii 2013</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="26CF535EFFDEFFD3DE6D4FD1FA0F7FE2" box="[1293,1414,2012,2039]" pageId="6" pageNumber="307">A. Pasang.</collectorName>
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<caption id="DF456600FFDFFFD2DBF74F9AFC897FC6" ID-DOI="http://doi.org/10.5281/zenodo.13210013" ID-Zenodo-Dep="13210013" httpUri="https://zenodo.org/record/13210013/files/figure.png" pageId="7" pageNumber="308" startId="7.[151,250,1943,1967]" targetBox="[302,1377,197,1855]" targetPageId="7" targetType="figure">
|
||
<paragraph id="8B853688FFDFFFD2DBF74F9AFC897FC6" blockId="7.[151,1436,1943,2004]" pageId="7" pageNumber="308">
|
||
<emphasis id="B94EEA9AFFDFFFD2DBF74F9AFE9D7FBA" bold="true" box="[151,276,1943,1967]" pageId="7" pageNumber="308">FIGURE 4.</emphasis>
|
||
Habitus of (a)
|
||
<taxonomicName id="4C3A4D0BFFDFFFD2DAD04F95FD0C7FA5" box="[432,645,1944,1968]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="biseriata">
|
||
<emphasis id="B94EEA9AFFDFFFD2DAD04F95FD0C7FA5" box="[432,645,1944,1968]" italics="true" pageId="7" pageNumber="308">Heliocypha biseriata</emphasis>
|
||
</taxonomicName>
|
||
, (b)
|
||
<taxonomicName id="4C3A4D0BFFDFFFD2D9D74F95FC2A7FA5" box="[695,931,1944,1968]" class="Insecta" family="Chlorocyphidae" genus="Aristocypha" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="fenestrella">
|
||
<emphasis id="B94EEA9AFFDFFFD2D9D74F95FC2A7FA5" box="[695,931,1944,1968]" italics="true" pageId="7" pageNumber="308">Aristocypha fenestrella</emphasis>
|
||
</taxonomicName>
|
||
, (c)
|
||
<taxonomicName id="4C3A4D0BFFDFFFD2D8B34F95FB397FA5" box="[979,1200,1944,1968]" class="Insecta" family="Chlorocyphidae" genus="Sundacypha" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="petiolata">
|
||
<emphasis id="B94EEA9AFFDFFFD2D8B34F95FB397FA5" box="[979,1200,1944,1968]" italics="true" pageId="7" pageNumber="308">Sundacypha petiolata</emphasis>
|
||
</taxonomicName>
|
||
, (d)
|
||
<taxonomicName id="4C3A4D0BFFDFFFD2DF824F95FA1F7FA5" box="[1250,1430,1944,1968]" class="Insecta" family="Chlorocyphidae" genus="Libellago" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="hyalina">
|
||
<emphasis id="B94EEA9AFFDFFFD2DF824F95FA1F7FA5" box="[1250,1430,1944,1968]" italics="true" pageId="7" pageNumber="308">Libellago hyalina</emphasis>
|
||
</taxonomicName>
|
||
. Markings shown only on legs and antennae. Scale bar 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF456600FFD0FFDDDBF74F9AFD0E7FC6" ID-DOI="http://doi.org/10.5281/zenodo.13210015" ID-Zenodo-Dep="13210015" httpUri="https://zenodo.org/record/13210015/files/figure.png" pageId="8" pageNumber="309" startId="8.[151,250,1943,1967]" targetBox="[206,1300,192,1855]" targetPageId="8" targetType="figure">
|
||
<paragraph id="8B853688FFD0FFDDDBF74F9AFD0E7FC6" blockId="8.[151,1436,1943,2004]" pageId="8" pageNumber="309">
|
||
<emphasis id="B94EEA9AFFD0FFDDDBF74F9AFE9B7FA5" bold="true" box="[151,274,1943,1968]" pageId="8" pageNumber="309">FIGURE 5.</emphasis>
|
||
Detail of heads, antennae and prothoraces of: (a)
|
||
<taxonomicName id="4C3A4D0BFFD0FFDDD8654F95FC507FA5" box="[773,985,1944,1968]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="biseriata">
|
||
<emphasis id="B94EEA9AFFD0FFDDD8654F95FC507FA5" box="[773,985,1944,1968]" italics="true" pageId="8" pageNumber="309">Heliocypha biseriata</emphasis>
|
||
</taxonomicName>
|
||
, (b)
|
||
<taxonomicName id="4C3A4D0BFFD0FFDDDF694F95FB7A7FA5" box="[1033,1267,1944,1968]" class="Insecta" family="Chlorocyphidae" genus="Aristocypha" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="fenestrella">
|
||
<emphasis id="B94EEA9AFFD0FFDDDF694F95FB7A7FA5" box="[1033,1267,1944,1968]" italics="true" pageId="8" pageNumber="309">Aristocypha fenestrella</emphasis>
|
||
</taxonomicName>
|
||
, (c)
|
||
<taxonomicName id="4C3A4D0BFFD0FFDDDE404F95FF787FC1" class="Insecta" family="Chlorocyphidae" genus="Sundacypha" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="petiolata">
|
||
<emphasis id="B94EEA9AFFD0FFDDDE404F95FF787FC1" italics="true" pageId="8" pageNumber="309">Sundacypha petiolata</emphasis>
|
||
</taxonomicName>
|
||
, (d)
|
||
<taxonomicName id="4C3A4D0BFFD0FFDDDA424FB1FE5C7FC1" box="[290,469,1980,2004]" class="Insecta" family="Chlorocyphidae" genus="Libellago" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="hyalina">
|
||
<emphasis id="B94EEA9AFFD0FFDDDA424FB1FE5C7FC1" box="[290,469,1980,2004]" italics="true" pageId="8" pageNumber="309">Libellago hyalina</emphasis>
|
||
</taxonomicName>
|
||
. Scale bar 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF456600FFD1FFDCDBF74F32FD3C7F8A" ID-DOI="http://doi.org/10.5281/zenodo.13210017" ID-Zenodo-Dep="13210017" httpUri="https://zenodo.org/record/13210017/files/figure.png" pageId="9" pageNumber="310" startId="9.[151,250,1855,1879]" targetBox="[153,1429,242,1805]" targetPageId="9" targetType="figure">
|
||
<paragraph id="8B853688FFD1FFDCDBF74F32FD3C7F8A" blockId="9.[151,1436,1855,1951]" pageId="9" pageNumber="310">
|
||
<emphasis id="B94EEA9AFFD1FFDCDBF74F32FE9B7F42" bold="true" box="[151,274,1855,1879]" pageId="9" pageNumber="310">FIGURE 6.</emphasis>
|
||
(a) Frontal view of labrum of
|
||
<taxonomicName id="4C3A4D0BFFD1FFDCD9204F32FC9D7F42" box="[576,788,1855,1879]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="biseriata">
|
||
<emphasis id="B94EEA9AFFD1FFDCD9204F32FC9D7F42" box="[576,788,1855,1879]" italics="true" pageId="9" pageNumber="310">Heliocypha biseriata</emphasis>
|
||
</taxonomicName>
|
||
; (b, c) detail of scape showing short recurved and longer, mainly straighter setae on
|
||
<taxonomicName id="4C3A4D0BFFD1FFDCDA324F69FE437F6E" box="[338,458,1891,1915]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="biseriata">
|
||
<emphasis id="B94EEA9AFFD1FFDCDA324F69FE437F6E" box="[338,458,1891,1915]" italics="true" pageId="9" pageNumber="310">H. biseriata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3A4D0BFFD1FFDCDA994F6EFD6A7F6E" box="[505,739,1891,1915]" class="Insecta" family="Chlorocyphidae" genus="Aristocypha" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="fenestrella">
|
||
<emphasis id="B94EEA9AFFD1FFDCDA994F6EFD6A7F6E" box="[505,739,1891,1915]" italics="true" pageId="9" pageNumber="310">Aristocypha fenestrella</emphasis>
|
||
</taxonomicName>
|
||
respectively; (d) Fore-tarsus in
|
||
<taxonomicName id="4C3A4D0BFFD1FFDCDF414F69FB107F6E" box="[1057,1177,1891,1915]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="biseriata">
|
||
<emphasis id="B94EEA9AFFD1FFDCDF414F69FB107F6E" box="[1057,1177,1891,1915]" italics="true" pageId="9" pageNumber="310">H. biseriata</emphasis>
|
||
</taxonomicName>
|
||
and (e) detail of pectinate setae on ventral side of same. Scale bars as indicated.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF456600FFD2FFDFDBF74F9AFAC27FC6" ID-DOI="http://doi.org/10.5281/zenodo.13210019" ID-Zenodo-Dep="13210019" httpUri="https://zenodo.org/record/13210019/files/figure.png" pageId="10" pageNumber="311" startId="10.[151,250,1943,1967]" targetBox="[228,1386,186,1919]" targetPageId="10" targetType="figure">
|
||
<paragraph id="8B853688FFD2FFDFDBF74F9AFAC27FC6" blockId="10.[151,1436,1943,2004]" pageId="10" pageNumber="311">
|
||
<emphasis id="B94EEA9AFFD2FFDFDBF74F9AFE9C7FA5" bold="true" box="[151,277,1943,1968]" pageId="10" pageNumber="311">FIGURE 7.</emphasis>
|
||
Left lateral view of head (left) and ventral view of genae showing spines (right): (a)
|
||
<taxonomicName id="4C3A4D0BFFD2FFDFDFF84F95FAF97FA5" box="[1176,1392,1944,1968]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="biseriata">
|
||
<emphasis id="B94EEA9AFFD2FFDFDFF84F95FAF97FA5" box="[1176,1392,1944,1968]" italics="true" pageId="10" pageNumber="311">Heliocypha biseriata</emphasis>
|
||
</taxonomicName>
|
||
, (b)
|
||
<taxonomicName id="4C3A4D0BFFD2FFDFDBF74FB1FE0B7FC1" box="[151,386,1980,2004]" class="Insecta" family="Chlorocyphidae" genus="Aristocypha" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="fenestrella">
|
||
<emphasis id="B94EEA9AFFD2FFDFDBF74FB1FE0B7FC1" box="[151,386,1980,2004]" italics="true" pageId="10" pageNumber="311">Aristocypha fenestrella</emphasis>
|
||
</taxonomicName>
|
||
, (c)
|
||
<taxonomicName id="4C3A4D0BFFD2FFDFDAD14FB1FD047FC1" box="[433,653,1980,2004]" class="Insecta" family="Chlorocyphidae" genus="Sundacypha" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="petiolata">
|
||
<emphasis id="B94EEA9AFFD2FFDFDAD14FB1FD047FC1" box="[433,653,1980,2004]" italics="true" pageId="10" pageNumber="311">Sundacypha petiolata</emphasis>
|
||
</taxonomicName>
|
||
, (d)
|
||
<taxonomicName id="4C3A4D0BFFD2FFDFD9DE4FB1FCF87FC1" box="[702,881,1980,2004]" class="Insecta" family="Chlorocyphidae" genus="Libellago" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="hyalina">
|
||
<emphasis id="B94EEA9AFFD2FFDFD9DE4FB1FCF87FC1" box="[702,881,1980,2004]" italics="true" pageId="10" pageNumber="311">Libellago hyalina</emphasis>
|
||
</taxonomicName>
|
||
. Scale bars left side, 1 mm; right side 1.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF456600FFD3FFDEDBF74E86FBCE7ED2" ID-DOI="http://doi.org/10.5281/zenodo.13210021" ID-Zenodo-Dep="13210021" httpUri="https://zenodo.org/record/13210021/files/figure.png" pageId="11" pageNumber="312" startId="11.[151,250,1675,1699]" targetBox="[160,1432,215,1639]" targetPageId="11" targetType="figure">
|
||
<paragraph id="8B853688FFD3FFDEDBF74E86FBCE7ED2" blockId="11.[151,1436,1675,1736]" pageId="11" pageNumber="312">
|
||
<emphasis id="B94EEA9AFFD3FFDEDBF74E86FE9B7EB1" bold="true" box="[151,274,1675,1700]" pageId="11" pageNumber="312">FIGURE 8.</emphasis>
|
||
Ventral view of head with mask at rest and anterior two thoracic segments in ventral view: (a)
|
||
<taxonomicName id="4C3A4D0BFFD3FFDEDFA24E81FA1F7EB1" box="[1218,1430,1676,1700]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="312" phylum="Arthropoda" rank="species" species="biseriata">
|
||
<emphasis id="B94EEA9AFFD3FFDEDFA24E81FA1F7EB1" box="[1218,1430,1676,1700]" italics="true" pageId="11" pageNumber="312">Heliocypha biseriata</emphasis>
|
||
</taxonomicName>
|
||
, (b)
|
||
<taxonomicName id="4C3A4D0BFFD3FFDEDBDB4EBDFE2F7EDD" box="[187,422,1712,1736]" class="Insecta" family="Chlorocyphidae" genus="Aristocypha" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="312" phylum="Arthropoda" rank="species" species="fenestrella">
|
||
<emphasis id="B94EEA9AFFD3FFDEDBDB4EBDFE2F7EDD" box="[187,422,1712,1736]" italics="true" pageId="11" pageNumber="312">Aristocypha fenestrella</emphasis>
|
||
</taxonomicName>
|
||
, (c)
|
||
<taxonomicName id="4C3A4D0BFFD3FFDEDAB54EBDFD387EDD" box="[469,689,1712,1736]" class="Insecta" family="Chlorocyphidae" genus="Sundacypha" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="312" phylum="Arthropoda" rank="species" species="petiolata">
|
||
<emphasis id="B94EEA9AFFD3FFDEDAB54EBDFD387EDD" box="[469,689,1712,1736]" italics="true" pageId="11" pageNumber="312">Sundacypha petiolata</emphasis>
|
||
</taxonomicName>
|
||
, (d)
|
||
<taxonomicName id="4C3A4D0BFFD3FFDED9824EBDFC1C7EDD" box="[738,917,1712,1736]" class="Insecta" family="Chlorocyphidae" genus="Libellago" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="312" phylum="Arthropoda" rank="species" species="hyalina">
|
||
<emphasis id="B94EEA9AFFD3FFDED9824EBDFC1C7EDD" box="[738,917,1712,1736]" italics="true" pageId="11" pageNumber="312">Libellago hyalina</emphasis>
|
||
</taxonomicName>
|
||
. Scale bar 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B853688FFD3FFDEDBA74F17FD017FB5" blockId="11.[151,1437,1818,2025]" pageId="11" pageNumber="312">
|
||
<emphasis id="B94EEA9AFFD3FFDEDBA74F17FEA67F21" bold="true" box="[199,303,1818,1844]" pageId="11" pageNumber="312">Habitus:</emphasis>
|
||
small larva, squat body with long, thin banded legs and mottled pattern on the body and appendages (
|
||
<figureCitation id="13012A0DFFD3FFDEDBFF4F33FF7A7F4D" box="[159,243,1854,1880]" captionStart="FIGURE 4" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[302,1377,197,1855]" captionTargetId="figure-18@7.[172,1416,181,1919]" captionTargetPageId="7" captionText="FIGURE 4. Habitus of (a) Heliocypha biseriata, (b) Aristocypha fenestrella, (c) Sundacypha petiolata, (d) Libellago hyalina. Markings shown only on legs and antennae. Scale bar 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210013" httpUri="https://zenodo.org/record/13210013/files/figure.png" pageId="11" pageNumber="312">Fig. 4a</figureCitation>
|
||
); sagittate head; scape slightly less than half total length of antennae, pedicel with dark tip; lateral caudal gills form two rapier like tapered spikes with numerous short backward pointing spines and setae; central cone-like epiproct represents the vestigial median gill.
|
||
</paragraph>
|
||
<paragraph id="8B853688FFD3FFD9DBA74FA4FBCC7D40" blockId="11.[151,1437,1818,2025]" lastBlockId="12.[151,1437,150,2013]" lastPageId="12" lastPageNumber="313" pageId="11" pageNumber="312">
|
||
<emphasis id="B94EEA9AFFD3FFDEDBA74FA4FE877FD1" bold="true" box="[199,270,1961,1988]" pageId="11" pageNumber="312">
|
||
<emphasis id="B94EEA9AFFD3FFDEDBA74FA4FE8C7FD1" bold="true" box="[199,261,1961,1988]" italics="true" pageId="11" pageNumber="312">Head</emphasis>
|
||
:
|
||
</emphasis>
|
||
in dorsal view rounded pentagonal, ca 5/6
|
||
<superScript id="7C4F9BC0FFD3FFDED9BF4FA7FD657FAD" attach="left" box="[735,748,1962,1976]" fontSize="6" pageId="11" pageNumber="312">th</superScript>
|
||
as long as broad, with occipital hind margin deeply excavated, producing overall sagittate outline (
|
||
<figureCitation id="13012A0DFFD3FFDED9424FC3FDFB7FFC" box="[546,626,1998,2025]" captionStart="FIGURE 5" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[206,1300,192,1855]" captionTargetId="figure-22@8.[196,1391,181,1919]" captionTargetPageId="8" captionText="FIGURE 5. Detail of heads, antennae and prothoraces of: (a) Heliocypha biseriata, (b) Aristocypha fenestrella, (c) Sundacypha petiolata, (d) Libellago hyalina. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210015" httpUri="https://zenodo.org/record/13210015/files/figure.png" pageId="11" pageNumber="312">Fig. 5a</figureCitation>
|
||
), frontoclypeus smooth. Antennae 7-segmented (
|
||
<figureCitation id="13012A0DFFD3FFDEDFF34FC3FB6D7FFC" box="[1171,1252,1998,2025]" captionStart="FIGURE 5" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[206,1300,192,1855]" captionTargetId="figure-22@8.[196,1391,181,1919]" captionTargetPageId="8" captionText="FIGURE 5. Detail of heads, antennae and prothoraces of: (a) Heliocypha biseriata, (b) Aristocypha fenestrella, (c) Sundacypha petiolata, (d) Libellago hyalina. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210015" httpUri="https://zenodo.org/record/13210015/files/figure.png" pageId="11" pageNumber="312">Fig. 5a</figureCitation>
|
||
); scape long and thick, about 2/5
|
||
<superScript id="7C4F9BC0FFD4FFD9DA2A489BFEDE78B1" attach="left" box="[330,343,150,164]" fontSize="6" pageId="12" pageNumber="313">th</superScript>
|
||
length of entire antenna with basal and apical markings, bearing short fine, curved setae, often recurved to touch integument (
|
||
<figureCitation id="13012A0DFFD4FFD9DA9848B6FDC778C0" box="[504,590,187,213]" captionStart="FIGURE 6" captionStartId="9.[151,250,1855,1879]" captionTargetBox="[153,1429,242,1805]" captionTargetId="figure-18@9.[151,1436,181,1831]" captionTargetPageId="9" captionText="FIGURE 6. (a) Frontal view of labrum of Heliocypha biseriata; (b, c) detail of scape showing short recurved and longer, mainly straighter setae on H. biseriata and Aristocypha fenestrella respectively; (d) Fore-tarsus in H. biseriata and (e) detail of pectinate setae on ventral side of same. Scale bars as indicated." figureDoi="http://doi.org/10.5281/zenodo.13210017" httpUri="https://zenodo.org/record/13210017/files/figure.png" pageId="12" pageNumber="313">Fig. 6b</figureCitation>
|
||
), which create a warty appearance when wet (but not submerged) as they trap droplets of fluid held by surface tension; pedicel with distinct dark mark on apex; bearing sparse long setae; remaining segments thin and pale. Labrum (
|
||
<figureCitation id="13012A0DFFD4FFD9D9E5490EFD5C7908" box="[645,725,259,285]" captionStart="FIGURE 6" captionStartId="9.[151,250,1855,1879]" captionTargetBox="[153,1429,242,1805]" captionTargetId="figure-18@9.[151,1436,181,1831]" captionTargetPageId="9" captionText="FIGURE 6. (a) Frontal view of labrum of Heliocypha biseriata; (b, c) detail of scape showing short recurved and longer, mainly straighter setae on H. biseriata and Aristocypha fenestrella respectively; (d) Fore-tarsus in H. biseriata and (e) detail of pectinate setae on ventral side of same. Scale bars as indicated." figureDoi="http://doi.org/10.5281/zenodo.13210017" httpUri="https://zenodo.org/record/13210017/files/figure.png" pageId="12" pageNumber="313">Fig. 6a</figureCitation>
|
||
) broadly curved with distinct dorsal and ventral parts, the latter boomerang shaped, the two parts meeting along thickened anterior ridge bearing small tubercles laterally; ventral part with median row of 12–13 ventrally directed strong setae, forming distinctive ‘moustache’, patches of shorter setae mid-ventrally and tufts of long fine setae towards corners; entire labrum forms a curved cavity into which anterior half of mandibles fit on either side. Vertex with three smooth swellings representing ocelli; eyes irregularly trapezoidal and prominent occupying outer 1/3
|
||
<superScript id="7C4F9BC0FFD4FFD9D9C549BBFD3A79D1" attach="left" box="[677,691,438,452]" fontSize="6" pageId="12" pageNumber="313">rd</superScript>
|
||
of lateral part of head but compound facets restricted to two small globular areas at the outer corners, clearly demarked by an anterior groove (
|
||
<figureCitation id="13012A0DFFD4FFD9DF6B49D6FBEA79E0" box="[1035,1123,474,501]" captionStart="FIGURE 5" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[206,1300,192,1855]" captionTargetId="figure-22@8.[196,1391,181,1919]" captionTargetPageId="8" captionText="FIGURE 5. Detail of heads, antennae and prothoraces of: (a) Heliocypha biseriata, (b) Aristocypha fenestrella, (c) Sundacypha petiolata, (d) Libellago hyalina. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210015" httpUri="https://zenodo.org/record/13210015/files/figure.png" pageId="12" pageNumber="313">Figs 5a</figureCitation>
|
||
,
|
||
<figureCitation id="13012A0DFFD4FFD9DF1249D6FB0679E0" box="[1138,1167,475,501]" captionStart="FIGURE 7" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[228,1386,186,1919]" captionTargetId="figure-22@10.[155,1433,181,1919]" captionTargetPageId="10" captionText="FIGURE 7. Left lateral view of head (left) and ventral view of genae showing spines (right): (a) Heliocypha biseriata, (b) Aristocypha fenestrella, (c) Sundacypha petiolata, (d) Libellago hyalina. Scale bars left side, 1 mm; right side 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210019" httpUri="https://zenodo.org/record/13210019/files/figure.png" pageId="12" pageNumber="313">7a</figureCitation>
|
||
); postocular lobes with prominent dorsolateral tubercle anterior to the posterior corner. Dorsal surface of head, excluding vertex scattered with small sparse low warty structures extending even onto inner marginal areas of eyes; antennal sockets swollen, with sparse long, fine setae laterally. Ventrally, genae bear a cluster of short heavy subocular spines, with one strong row dorsally, the anterior 2–3 acuminate, the remainder rounded, and below this a sparse irregular scattering of shorter rounded tubercles clearly visible in lateral and ventral view (
|
||
<figureCitation id="13012A0DFFD4FFD9D8C74A82FC727ABC" box="[935,1019,655,681]" captionStart="FIGURE 7" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[228,1386,186,1919]" captionTargetId="figure-22@10.[155,1433,181,1919]" captionTargetPageId="10" captionText="FIGURE 7. Left lateral view of head (left) and ventral view of genae showing spines (right): (a) Heliocypha biseriata, (b) Aristocypha fenestrella, (c) Sundacypha petiolata, (d) Libellago hyalina. Scale bars left side, 1 mm; right side 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210019" httpUri="https://zenodo.org/record/13210019/files/figure.png" pageId="12" pageNumber="313">Fig. 7a</figureCitation>
|
||
); postocular lobes with ventrolateral cluster of weakly developed spines and setae; strong ventral spur at margin of gena and occiput. Mask long and thin, articulation resting between anterior part of coxae of mesothorax (
|
||
<figureCitation id="13012A0DFFD4FFD9D8EB4ADAFC577AE4" box="[907,990,727,753]" captionStart="FIGURE 8" captionStartId="11.[151,250,1675,1699]" captionTargetBox="[160,1432,215,1639]" captionTargetId="figure-18@11.[151,1436,181,1652]" captionTargetPageId="11" captionText="FIGURE 8. Ventral view of head with mask at rest and anterior two thoracic segments in ventral view: (a) Heliocypha biseriata, (b) Aristocypha fenestrella, (c) Sundacypha petiolata, (d) Libellago hyalina. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210021" httpUri="https://zenodo.org/record/13210021/files/figure.png" pageId="12" pageNumber="313">Fig. 8a</figureCitation>
|
||
); prementum narrow basally, gradually flared anteriorly with margins of anterior half weakly convex; lateral margins with sparse fine setae on distal 2/3
|
||
<superScript id="7C4F9BC0FFD4FFD9DEEF4AF7FA147B1D" attach="right" box="[1423,1437,762,776]" fontSize="6" pageId="12" pageNumber="313">rd</superScript>
|
||
(
|
||
<figureCitation id="13012A0DFFD4FFD9DBFF4B12FF7B7B2C" box="[159,242,799,825]" captionStart="FIGURE 9" captionStartId="17.[151,250,1943,1967]" captionTargetBox="[244,1341,217,1810]" captionTargetId="figure-18@17.[155,1433,181,1919]" captionTargetPageId="17" captionText="FIGURE 9. Prementum in dorsal view: (a) Heliocypha biseriata, (b) Aristocypha fenestrella, (c) Sundacypha petiolata, (d) Libellago hyalina, (e) Libellago lineata. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210023" httpUri="https://zenodo.org/record/13210023/files/figure.png" pageId="12" pageNumber="313">Fig. 9a</figureCitation>
|
||
) anterior median lobe weakly convex with coarsely serrate margins; median cleft open and moderately deep with small nearly circular aperture at base (
|
||
<figureCitation id="13012A0DFFD4FFD9D9B44B4EFCB17B48" box="[724,824,835,861]" captionStart="FIGURE 10" captionStartId="18.[151,250,1943,1967]" captionTargetBox="[167,1433,182,1916]" captionTargetId="figure-22@18.[155,1433,181,1919]" captionTargetPageId="18" captionText="FIGURE 10. Details of anterior prementum and labial palp Heliocypha biseriata: (a) anterior prementum and labial palp in dorsal view (base of movable hook dotted to reveal details of palpal lobe below); (b) labial palp and anterior margin of prementum, frontal view; (c) labial palp, interior view. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210025" httpUri="https://zenodo.org/record/13210025/files/figure.png" pageId="12" pageNumber="313">Fig. 10a</figureCitation>
|
||
); Lobe of labial palp bifid (
|
||
<figureCitation id="13012A0DFFD4FFD9DFE14B4EFA8D7B48" box="[1153,1284,835,861]" captionStart="FIGURE 10" captionStartId="18.[151,250,1943,1967]" captionTargetBox="[167,1433,182,1916]" captionTargetId="figure-22@18.[155,1433,181,1919]" captionTargetPageId="18" captionText="FIGURE 10. Details of anterior prementum and labial palp Heliocypha biseriata: (a) anterior prementum and labial palp in dorsal view (base of movable hook dotted to reveal details of palpal lobe below); (b) labial palp and anterior margin of prementum, frontal view; (c) labial palp, interior view. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210025" httpUri="https://zenodo.org/record/13210025/files/figure.png" pageId="12" pageNumber="313">Figs 10a–c</figureCitation>
|
||
), with dorsal process a long claw bearing secondary dorsal hook and ventral process a shorter claw with small interior beak like process; inner margin of lobe with fine sparse denticulations evident even on the inner process; movable hook stout, moderately long and curved especially in apical half. Maxillae as shown (
|
||
<figureCitation id="13012A0DFFD4FFD9DF6E4BA2FB4D7BDC" box="[1038,1220,943,969]" captionStart="FIGURE 12" captionStartId="20.[151,250,1909,1933]" captionTargetBox="[205,1382,181,1764]" captionTargetId="figure-22@20.[205,1382,181,1884]" captionTargetPageId="20" captionText="FIGURE 12. Details of maxillae—(a–d) Right maxillae: Heliocypha biseriata, (a) ventral view, (b) oral view; (c) Sundacypha petiolata distal detail, oral view; (d) L. hyalina oral view. (e–h) Left maxillae, oral view, details of terminal strong spine with three shorter basal to median supplementary spines: (e) Heliocypha biseriata, (f) Aristocypha fenestrella, (g) Sundacypha petiolata, (h) Libellago hyalina. Scale bars: above 1 mm; below 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13210029" httpUri="https://zenodo.org/record/13210029/files/figure.png" pageId="12" pageNumber="313">Figs 12a–b, 12e</figureCitation>
|
||
); galeolacinea with four long strong teeth towards apex curved orally; apical tooth with row of three inner broad-based supplementary denticles (
|
||
<figureCitation id="13012A0DFFD4FFD9DA6A4BFAFEE27C04" box="[266,363,1015,1041]" captionStart="FIGURE 12" captionStartId="20.[151,250,1909,1933]" captionTargetBox="[205,1382,181,1764]" captionTargetId="figure-22@20.[205,1382,181,1884]" captionTargetPageId="20" captionText="FIGURE 12. Details of maxillae—(a–d) Right maxillae: Heliocypha biseriata, (a) ventral view, (b) oral view; (c) Sundacypha petiolata distal detail, oral view; (d) L. hyalina oral view. (e–h) Left maxillae, oral view, details of terminal strong spine with three shorter basal to median supplementary spines: (e) Heliocypha biseriata, (f) Aristocypha fenestrella, (g) Sundacypha petiolata, (h) Libellago hyalina. Scale bars: above 1 mm; below 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13210029" httpUri="https://zenodo.org/record/13210029/files/figure.png" pageId="12" pageNumber="313">Fig. 12e</figureCitation>
|
||
), basal pair partially fused at base, longest one apicad; row of ca 5 strong setae along low ridge aligned with apical tooth; long fine setae present on internal margin of galeolacinea; palp tapering, not reaching tip of apical tooth, bearing long strong setae in all outward directions. Mandibles (
|
||
<figureCitation id="13012A0DFFD4FFD9DF544C32FB227C4C" box="[1076,1195,1087,1113]" captionStart="FIGURE 13" captionStartId="21.[151,250,1909,1933]" captionTargetBox="[206,1359,208,1883]" captionTargetId="figure-18@21.[205,1382,181,1884]" captionTargetPageId="21" captionText="FIGURE 13. Mandibles of Heliocypha biseriata: (a) right mandible, (b) left mandible in frontal view; (c) right mandible, (d) left mandible in ventral view; (e) detail of right molar crest in profile; (f) slightly postero-ventro-interior view of left mandible showing marginal teeth and molar crest on edge; (e) detail of denticles of left molar crest. Scale bars a–d, f, 0.5 mm; e, g, 0.1 mm as indicated." figureDoi="http://doi.org/10.5281/zenodo.13210031" httpUri="https://zenodo.org/record/13210031/files/figure.png" pageId="12" pageNumber="313">Figs 13a–f</figureCitation>
|
||
)—both left and right with five distinct teeth on incisor visible in external view (
|
||
<figureCitation id="13012A0DFFD4FFD9D84E4C6EFC397C68" box="[814,944,1123,1149]" captionStart="FIGURE 13" captionStartId="21.[151,250,1909,1933]" captionTargetBox="[206,1359,208,1883]" captionTargetId="figure-18@21.[205,1382,181,1884]" captionTargetPageId="21" captionText="FIGURE 13. Mandibles of Heliocypha biseriata: (a) right mandible, (b) left mandible in frontal view; (c) right mandible, (d) left mandible in ventral view; (e) detail of right molar crest in profile; (f) slightly postero-ventro-interior view of left mandible showing marginal teeth and molar crest on edge; (e) detail of denticles of left molar crest. Scale bars a–d, f, 0.5 mm; e, g, 0.1 mm as indicated." figureDoi="http://doi.org/10.5281/zenodo.13210031" httpUri="https://zenodo.org/record/13210031/files/figure.png" pageId="12" pageNumber="313">Figs 13a, b</figureCitation>
|
||
); in inner view (
|
||
<figureCitation id="13012A0DFFD4FFD9DF0C4C6EFB437C68" box="[1132,1226,1123,1149]" captionStart="FIGURE 13" captionStartId="21.[151,250,1909,1933]" captionTargetBox="[206,1359,208,1883]" captionTargetId="figure-18@21.[205,1382,181,1884]" captionTargetPageId="21" captionText="FIGURE 13. Mandibles of Heliocypha biseriata: (a) right mandible, (b) left mandible in frontal view; (c) right mandible, (d) left mandible in ventral view; (e) detail of right molar crest in profile; (f) slightly postero-ventro-interior view of left mandible showing marginal teeth and molar crest on edge; (e) detail of denticles of left molar crest. Scale bars a–d, f, 0.5 mm; e, g, 0.1 mm as indicated." figureDoi="http://doi.org/10.5281/zenodo.13210031" httpUri="https://zenodo.org/record/13210031/files/figure.png" pageId="12" pageNumber="313">Fig. 13c</figureCitation>
|
||
) right incisor lobe with four long teeth and one shorter one between fourth and fifth one ventrally and small supplementary tooth basal to strong ventral tooth; molar crest a low swelling with two small nipple-like points, lightly sclerotised (
|
||
<figureCitation id="13012A0DFFD4FFD9DE574CA6FF2F7CFC" captionStart="FIGURE 13" captionStartId="21.[151,250,1909,1933]" captionTargetBox="[206,1359,208,1883]" captionTargetId="figure-18@21.[205,1382,181,1884]" captionTargetPageId="21" captionText="FIGURE 13. Mandibles of Heliocypha biseriata: (a) right mandible, (b) left mandible in frontal view; (c) right mandible, (d) left mandible in ventral view; (e) detail of right molar crest in profile; (f) slightly postero-ventro-interior view of left mandible showing marginal teeth and molar crest on edge; (e) detail of denticles of left molar crest. Scale bars a–d, f, 0.5 mm; e, g, 0.1 mm as indicated." figureDoi="http://doi.org/10.5281/zenodo.13210031" httpUri="https://zenodo.org/record/13210031/files/figure.png" pageId="12" pageNumber="313">Fig. 13c, e</figureCitation>
|
||
); left incisor (
|
||
<figureCitation id="13012A0DFFD4FFD9DA274CC2FE487CFC" box="[327,449,1231,1257]" captionStart="FIGURE 13" captionStartId="21.[151,250,1909,1933]" captionTargetBox="[206,1359,208,1883]" captionTargetId="figure-18@21.[205,1382,181,1884]" captionTargetPageId="21" captionText="FIGURE 13. Mandibles of Heliocypha biseriata: (a) right mandible, (b) left mandible in frontal view; (c) right mandible, (d) left mandible in ventral view; (e) detail of right molar crest in profile; (f) slightly postero-ventro-interior view of left mandible showing marginal teeth and molar crest on edge; (e) detail of denticles of left molar crest. Scale bars a–d, f, 0.5 mm; e, g, 0.1 mm as indicated." figureDoi="http://doi.org/10.5281/zenodo.13210031" httpUri="https://zenodo.org/record/13210031/files/figure.png" pageId="12" pageNumber="313">Fig. 13d, f</figureCitation>
|
||
) with six teeth, the third from the dorsal being vestigial and only visible on inner face, pronounced gap between first and second teeth from ventral; molar crest bearing strong rounded cupped blade (
|
||
<figureCitation id="13012A0DFFD4FFD9DE0F4CFEFF4B7D24" captionStart="FIGURE 13" captionStartId="21.[151,250,1909,1933]" captionTargetBox="[206,1359,208,1883]" captionTargetId="figure-18@21.[205,1382,181,1884]" captionTargetPageId="21" captionText="FIGURE 13. Mandibles of Heliocypha biseriata: (a) right mandible, (b) left mandible in frontal view; (c) right mandible, (d) left mandible in ventral view; (e) detail of right molar crest in profile; (f) slightly postero-ventro-interior view of left mandible showing marginal teeth and molar crest on edge; (e) detail of denticles of left molar crest. Scale bars a–d, f, 0.5 mm; e, g, 0.1 mm as indicated." figureDoi="http://doi.org/10.5281/zenodo.13210031" httpUri="https://zenodo.org/record/13210031/files/figure.png" pageId="12" pageNumber="313">Fig. 13d</figureCitation>
|
||
) with nine minute denticles along roughly straight distal margin (
|
||
<figureCitation id="13012A0DFFD4FFD9D8C44D1AFB8F7D24" box="[932,1030,1303,1329]" captionStart="FIGURE 13" captionStartId="21.[151,250,1909,1933]" captionTargetBox="[206,1359,208,1883]" captionTargetId="figure-18@21.[205,1382,181,1884]" captionTargetPageId="21" captionText="FIGURE 13. Mandibles of Heliocypha biseriata: (a) right mandible, (b) left mandible in frontal view; (c) right mandible, (d) left mandible in ventral view; (e) detail of right molar crest in profile; (f) slightly postero-ventro-interior view of left mandible showing marginal teeth and molar crest on edge; (e) detail of denticles of left molar crest. Scale bars a–d, f, 0.5 mm; e, g, 0.1 mm as indicated." figureDoi="http://doi.org/10.5281/zenodo.13210031" httpUri="https://zenodo.org/record/13210031/files/figure.png" pageId="12" pageNumber="313">
|
||
Fig.
|
||
<quantity id="4CC29B6DFFD4FFD9D8B84D1AFB8F7D24" box="[984,1030,1303,1329]" metricMagnitude="-2" metricUnit="kg" metricValue="1.3" pageId="12" pageNumber="313" unit="g" value="13.0">13g</quantity>
|
||
</figureCitation>
|
||
). Single strong seta at exterior basal corner on each mandible (
|
||
<figureCitation id="13012A0DFFD4FFD9DADB4D36FD927D40" box="[443,539,1339,1365]" captionStart="FIGURE 13" captionStartId="21.[151,250,1909,1933]" captionTargetBox="[206,1359,208,1883]" captionTargetId="figure-18@21.[205,1382,181,1884]" captionTargetPageId="21" captionText="FIGURE 13. Mandibles of Heliocypha biseriata: (a) right mandible, (b) left mandible in frontal view; (c) right mandible, (d) left mandible in ventral view; (e) detail of right molar crest in profile; (f) slightly postero-ventro-interior view of left mandible showing marginal teeth and molar crest on edge; (e) detail of denticles of left molar crest. Scale bars a–d, f, 0.5 mm; e, g, 0.1 mm as indicated." figureDoi="http://doi.org/10.5281/zenodo.13210031" httpUri="https://zenodo.org/record/13210031/files/figure.png" pageId="12" pageNumber="313">Fig. 13d</figureCitation>
|
||
), not always visible, depending on angle of view.
|
||
</paragraph>
|
||
<paragraph id="8B853688FFD4FFD9DBA74D53FD1D7F58" blockId="12.[151,1437,150,2013]" pageId="12" pageNumber="313">
|
||
<emphasis id="B94EEA9AFFD4FFD9DBA74D53FEAF7D6C" bold="true" box="[199,294,1374,1401]" pageId="12" pageNumber="313">
|
||
<emphasis id="B94EEA9AFFD4FFD9DBA74D53FE947D6C" bold="true" box="[199,285,1374,1401]" italics="true" pageId="12" pageNumber="313">Thorax</emphasis>
|
||
:
|
||
</emphasis>
|
||
prothorax saddle-shaped with narrow flange anteriorly fitted into occipital margin; dorsum with transverse central groove; two pairs of prominent broad tubercles, a central pair just behind groove and a lateral pair posterior to these and close to ends of posterior flange that curves broadly around posterior margin (
|
||
<figureCitation id="13012A0DFFD4FFD9DE5C4DAAFA067DD4" box="[1340,1423,1446,1473]" captionStart="FIGURE 5" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[206,1300,192,1855]" captionTargetId="figure-22@8.[196,1391,181,1919]" captionTargetPageId="8" captionText="FIGURE 5. Detail of heads, antennae and prothoraces of: (a) Heliocypha biseriata, (b) Aristocypha fenestrella, (c) Sundacypha petiolata, (d) Libellago hyalina. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210015" httpUri="https://zenodo.org/record/13210015/files/figure.png" pageId="12" pageNumber="313">Fig. 5a</figureCitation>
|
||
). Propleura swollen with small anterior ventral truncated spur, well ahead of coxae. Remainer of thorax with shallow grooves and bulges; entire dorsum of thorax with sparse fine warty texture, as on head, and with sparse long, fine setae collapsed in preserved specimens. Marking (not illustrated on body) variable and mottled but always with broad pale band pale between two anterior tubercles on prothorax. Wing sheaths slightly spatulate, not completely obscuring abdomen at sides, forewing reaching to end of S4, hindwing to 3/4 along S5 (
|
||
<figureCitation id="13012A0DFFD4FFD9DF024E56FB3D7E60" box="[1122,1204,1627,1653]" captionStart="FIGURE 4" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[302,1377,197,1855]" captionTargetId="figure-18@7.[172,1416,181,1919]" captionTargetPageId="7" captionText="FIGURE 4. Habitus of (a) Heliocypha biseriata, (b) Aristocypha fenestrella, (c) Sundacypha petiolata, (d) Libellago hyalina. Markings shown only on legs and antennae. Scale bar 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210013" httpUri="https://zenodo.org/record/13210013/files/figure.png" pageId="12" pageNumber="313">Fig. 4a</figureCitation>
|
||
). Legs long and thin, metafemur reaching end of S7; femora all with three distinct evenly spaced dark bands on ochreous ground colour, with tiny dark mark at base sometimes also on trochanter; all femora with anterior ridge bearing very small outward directed spines, some bearing setae, and sparse fine setae towards tibial joint, some quite long; tibiae unmarked or with single basal darker band, with fairly sparse setae, mainly fine and straight but some stronger and curled, longer denser ventral patch of straight setae apically; tarsus (
|
||
<figureCitation id="13012A0DFFD4FFD9D99A4F02FCE57F3C" box="[762,876,1807,1833]" captionStart="FIGURE 6" captionStartId="9.[151,250,1855,1879]" captionTargetBox="[153,1429,242,1805]" captionTargetId="figure-18@9.[151,1436,181,1831]" captionTargetPageId="9" captionText="FIGURE 6. (a) Frontal view of labrum of Heliocypha biseriata; (b, c) detail of scape showing short recurved and longer, mainly straighter setae on H. biseriata and Aristocypha fenestrella respectively; (d) Fore-tarsus in H. biseriata and (e) detail of pectinate setae on ventral side of same. Scale bars as indicated." figureDoi="http://doi.org/10.5281/zenodo.13210017" httpUri="https://zenodo.org/record/13210017/files/figure.png" pageId="12" pageNumber="313">Figs 6d, e</figureCitation>
|
||
) bearing two rows each of ca 25 strong, tapering pectinate setae (
|
||
<figureCitation id="13012A0DFFD4FFD9DA2B4F3EFE157F58" box="[331,412,1843,1869]" captionStart="FIGURE 6" captionStartId="9.[151,250,1855,1879]" captionTargetBox="[153,1429,242,1805]" captionTargetId="figure-18@9.[151,1436,181,1831]" captionTargetPageId="9" captionText="FIGURE 6. (a) Frontal view of labrum of Heliocypha biseriata; (b, c) detail of scape showing short recurved and longer, mainly straighter setae on H. biseriata and Aristocypha fenestrella respectively; (d) Fore-tarsus in H. biseriata and (e) detail of pectinate setae on ventral side of same. Scale bars as indicated." figureDoi="http://doi.org/10.5281/zenodo.13210017" httpUri="https://zenodo.org/record/13210017/files/figure.png" pageId="12" pageNumber="313">Fig. 6e</figureCitation>
|
||
). Tarsal claws simple.
|
||
</paragraph>
|
||
<paragraph id="8B853688FFD4FFD8DBA74F5BFAFF79B8" blockId="12.[151,1437,150,2013]" lastBlockId="13.[151,1437,151,825]" lastPageId="13" lastPageNumber="314" pageId="12" pageNumber="313">
|
||
<emphasis id="B94EEA9AFFD4FFD9DBA74F5BFEB67F64" bold="true" box="[199,319,1878,1905]" pageId="12" pageNumber="313">
|
||
<emphasis id="B94EEA9AFFD4FFD9DBA74F5BFEBC7F64" bold="true" box="[199,309,1878,1905]" italics="true" pageId="12" pageNumber="313">Abdomen</emphasis>
|
||
:
|
||
</emphasis>
|
||
squat, mainly dark with well-defined pale narrow middorsal stripe (not figured). Laterally with patches of small spine like setae and mixture of curved and fine long setae; each segment from S4 with submarginal row of small ventral spines; ventrolateral margin of S9 with 4–5 strong, long spines intermingled with 4–5 smaller spines partly placed a little above margin and four strong spines along ventral margin; S10 hind-margin with 4 strong rearward pointing ventrolateral spines and 3 long spines extending to lateral hind-margins, smaller spines along ventral hind margin and on venter (
|
||
<figureCitation id="13012A0DFFD5FFD8D94048B6FD2978C0" box="[544,672,187,213]" captionStart="FIGURE 16" captionStartId="24.[151,250,1964,1988]" captionTargetBox="[158,1439,262,1931]" captionTargetId="figure-44@24.[149,1439,237,1940]" captionTargetPageId="24" captionText="FIGURE 16. Female gonapophyses and terminal abdominal segments in ventral view: (a) H. biseriata, (b) S. petiolata, (c) L. hyalina, and in lateral view: (d) H. biseriata, (e) L. hyalina, (f) S. petiolata. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210037" httpUri="https://zenodo.org/record/13210037/files/figure.png" pageId="13" pageNumber="314">Figs 16a, d</figureCitation>
|
||
). Outer female gonapophyses arising from base of S9 and extending well beyond hind margin of S10 (
|
||
<figureCitation id="13012A0DFFD5FFD8D97A48D2FDF278EC" box="[538,635,223,249]" captionStart="FIGURE 16" captionStartId="24.[151,250,1964,1988]" captionTargetBox="[158,1439,262,1931]" captionTargetId="figure-44@24.[149,1439,237,1940]" captionTargetPageId="24" captionText="FIGURE 16. Female gonapophyses and terminal abdominal segments in ventral view: (a) H. biseriata, (b) S. petiolata, (c) L. hyalina, and in lateral view: (d) H. biseriata, (e) L. hyalina, (f) S. petiolata. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210037" httpUri="https://zenodo.org/record/13210037/files/figure.png" pageId="13" pageNumber="314">Fig. 16a</figureCitation>
|
||
), ventral margin with series of sparse heavy spines and terminating in a single long heavy spine deflected ventrally in lateral view (
|
||
<figureCitation id="13012A0DFFD5FFD8D829490EFC257908" box="[841,940,259,285]" captionStart="FIGURE 16" captionStartId="24.[151,250,1964,1988]" captionTargetBox="[158,1439,262,1931]" captionTargetId="figure-44@24.[149,1439,237,1940]" captionTargetPageId="24" captionText="FIGURE 16. Female gonapophyses and terminal abdominal segments in ventral view: (a) H. biseriata, (b) S. petiolata, (c) L. hyalina, and in lateral view: (d) H. biseriata, (e) L. hyalina, (f) S. petiolata. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210037" httpUri="https://zenodo.org/record/13210037/files/figure.png" pageId="13" pageNumber="314">Fig. 16d</figureCitation>
|
||
); inner apophyses smooth, curved smoothly upward to a rounded point extending well beyond outer apophyses and epiproct. Male gonapophyses unknown. Epiproct, short, subconical, in shape of elongated onion dome. Caudal spikes long, about 55% body length, broad basally and from just before midpoint tapered gradually and smoothly to a sharp point (
|
||
<figureCitation id="13012A0DFFD5FFD8DF004962FB36799C" box="[1120,1215,367,393]" captionStart="FIGURE 18" captionStartId="27.[151,250,1943,1967]" captionTargetBox="[191,1381,197,1917]" captionTargetId="figure-18@27.[189,1399,181,1919]" captionTargetPageId="27" captionText="FIGURE 18. Caudal spikes scaled in proportion to overall body length: (a) H. biseriata, (b) A. fenstrella, (c) S. petiolata, (d) L. hyalina. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210041" httpUri="https://zenodo.org/record/13210041/files/figure.png" pageId="13" pageNumber="314">Fig. 18a</figureCitation>
|
||
); bearing numerous backwardly curved strong sharp spines, heavy setae, and also long fine setae too entangled to assess accurately.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3206503FFD5FFD8DBA749BAFBAC7B2C" pageId="13" pageNumber="314" type="description">
|
||
<paragraph id="8B853688FFD5FFD8DBA749BAFDC77A28" blockId="13.[151,1437,151,825]" pageId="13" pageNumber="314">
|
||
<emphasis id="B94EEA9AFFD5FFD8DBA749BAFE6079C5" bold="true" box="[199,489,438,465]" pageId="13" pageNumber="314">Measurements (in mm):</emphasis>
|
||
body length excluding antennae, caudal appendages and spikes 11.8; lateral caudal spikes 6.5; head width 2.7; prementum 2.65; palpal lobe (to tip of dorsal branch) 0.85; movable hook 0.72; maxillae base to distal hook of galeolacinea (1.24), maxillary palp (0.54); mandibles long axis (0.85); antenna 1.70, 0.65, 0.70, 0.50, 0.28, 0.19, 0.10 (total 4.11).
|
||
</paragraph>
|
||
<paragraph id="8B853688FFD5FFD8DBA74A4BFBAC7B2C" blockId="13.[151,1437,151,825]" pageId="13" pageNumber="314">
|
||
<emphasis id="B94EEA9AFFD5FFD8DBA74A4BFEC87A75" bold="true" box="[199,321,582,608]" pageId="13" pageNumber="314">Variation:</emphasis>
|
||
outwardly no noticeable variation, other than minor differences in the mottling on the head, in the numbers of large setae in the ‘moustache’ of the labrum, and in the number of spines present on S9 and S10, was evident in the
|
||
<specimenCount id="9D3CFD01FFD5FFD8DA574A82FE3C7ABC" box="[311,437,655,681]" count="2" pageId="13" pageNumber="314" type="female">two female</specimenCount>
|
||
specimens examined. The unusual dentition of the mandibles differed slightly in the
|
||
<specimenCount id="9D3CFD01FFD5FFD8DE124A82FE857AD8" count="2" pageId="13" pageNumber="314" type="generic">two specimens</specimenCount>
|
||
examined in that in the second specimen (not illustrated) the vestigial inner tooth at the base of the 2
|
||
<superScript id="7C4F9BC0FFD5FFD8DEEC4ABFFA157AD5" attach="none" box="[1420,1436,690,704]" fontSize="6" pageId="13" pageNumber="314">nd</superScript>
|
||
and 3
|
||
<superScript id="7C4F9BC0FFD5FFD8DBB44ADBFF6B7AF1" attach="left" box="[212,226,726,740]" fontSize="6" pageId="13" pageNumber="314">rd</superScript>
|
||
incisor teeth from the dorsal is minute, no more than 15 μm long. No significant differences were evident in the male specimen photographed in some detail by POMS; no image was made of the gonapophyses hence these cannot be described, but they are unlikely to present useful diagnostic characters.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3206503FFD5FFDBDBF74B86FD907954" lastPageId="14" lastPageNumber="315" pageId="13" pageNumber="314" type="discussion">
|
||
<paragraph id="8B853688FFD5FFD8DBF74B86FE807BB0" blockId="13.[151,265,907,933]" box="[151,265,907,933]" pageId="13" pageNumber="314">
|
||
<emphasis id="B94EEA9AFFD5FFD8DBF74B86FE807BB0" bold="true" box="[151,265,907,933]" pageId="13" pageNumber="314">Remarks</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B853688FFD5FFD8DBF74BDEFE5F7C68" blockId="13.[151,1437,979,2013]" pageId="13" pageNumber="314">
|
||
While unusual characteristics of the mouthparts, especially the dentition of the mandibles appear to be consistent it is possible that the inner vestigial left incisor tooth is entirely lacking in some specimens. The labrum, while similar in other species examined (see below), is unique among odonates and is probably an autapomorphy for the family. Pectinate setae confined to tarsi, previously recorded by
|
||
<bibRefCitation id="EFAB4B79FFD5FFD8D87F4C32FC497C4C" author="Kumar, A." box="[799,960,1087,1113]" pageId="13" pageNumber="314" pagination="83 - 118" refId="ref15386" refString="Kumar, A. (1973) Descriptions of the last instar larvae of Odonata from the Dehra Dun Valley (India), with notes on Biology I. (suborder Zygoptera). Oriental Insects, 7, 83 - 118. https: // doi. org / 10.1080 / 00305316.1973.10434207" type="journal article" year="1973">Kumar (1973)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFAB4B79FFD5FFD8D89A4C32FA997C4C" author="Kumar, A. & Prasad, M." box="[1018,1296,1087,1113]" pageId="13" pageNumber="314" pagination="547 - 554" refId="ref15440" refString="Kumar, A. & Prasad, M. (1977) On the larvae of Rhinocypha (Odonata: Chlorocyphidae) from Garhwal hills. Oriental Insects, 11, 547 - 554. https: // doi. org / 10.1080 / 00305316.1977.11090927" type="journal article" year="1977">Kumar & Prasad (1977)</bibRefCitation>
|
||
, are another probable family apomorphy.
|
||
</paragraph>
|
||
<paragraph id="8B853688FFD5FFDBDBA74C8AFD907954" blockId="13.[151,1437,979,2013]" lastBlockId="14.[151,1437,151,321]" lastPageId="14" lastPageNumber="315" pageId="13" pageNumber="314">
|
||
Two other species of
|
||
<taxonomicName id="4C3A4D0BFFD5FFD8DAD34C8AFDBD7CB4" box="[435,564,1159,1185]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="314" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94EEA9AFFD5FFD8DAD34C8AFDBD7CB4" box="[435,564,1159,1185]" italics="true" pageId="13" pageNumber="314">Heliocypha</emphasis>
|
||
</taxonomicName>
|
||
larvae have been described in some detail:
|
||
<taxonomicName id="4C3A4D0BFFD5FFD8DF754C8AFB187CB4" box="[1045,1169,1159,1185]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="biforata">
|
||
<emphasis id="B94EEA9AFFD5FFD8DF754C8AFB187CB4" box="[1045,1169,1159,1185]" italics="true" pageId="13" pageNumber="314">H. biforata</emphasis>
|
||
</taxonomicName>
|
||
(as
|
||
<taxonomicName id="4C3A4D0BFFD5FFD8DFDE4C8AFA1F7CB4" box="[1214,1430,1159,1185]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="314" phylum="Arthropoda" rank="subSpecies" species="biforata" subSpecies="beesoni">
|
||
<emphasis id="B94EEA9AFFD5FFD8DFDE4C8AFA1F7CB4" box="[1214,1430,1159,1185]" italics="true" pageId="13" pageNumber="314">H. biforata beesoni</emphasis>
|
||
</taxonomicName>
|
||
, now considered a synonym of
|
||
<taxonomicName id="4C3A4D0BFFD5FFD8DA8F4CA6FD447CD0" box="[495,717,1195,1221]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="314" phylum="Arthropoda" rank="subSpecies" species="biforata" subSpecies="biforata">
|
||
<emphasis id="B94EEA9AFFD5FFD8DA8F4CA6FD447CD0" box="[495,717,1195,1221]" italics="true" pageId="13" pageNumber="314">H. biforata biforata</emphasis>
|
||
</taxonomicName>
|
||
), by
|
||
<bibRefCitation id="EFAB4B79FFD5FFD8D8684CA6FB9E7CD0" author="Kumar, A. & Prasad, M." box="[776,1047,1195,1221]" pageId="13" pageNumber="314" pagination="547 - 554" refId="ref15440" refString="Kumar, A. & Prasad, M. (1977) On the larvae of Rhinocypha (Odonata: Chlorocyphidae) from Garhwal hills. Oriental Insects, 11, 547 - 554. https: // doi. org / 10.1080 / 00305316.1977.11090927" type="journal article" year="1977">Kumar & Prasad (1977)</bibRefCitation>
|
||
and
|
||
<emphasis id="B94EEA9AFFD5FFD8DF2E4CA6FB7E7CD0" box="[1102,1271,1195,1221]" italics="true" pageId="13" pageNumber="314">
|
||
H. p.
|
||
<taxonomicName id="4C3A4D0BFFD5FFD8DFED4CA6FB7E7CD0" box="[1165,1271,1195,1221]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="perforata">perforata</taxonomicName>
|
||
</emphasis>
|
||
by
|
||
<bibRefCitation id="EFAB4B79FFD5FFD8DE414CA6FA117CD0" author="Xu, Q. - H." box="[1313,1432,1194,1221]" pageId="13" pageNumber="314" pagination="137 - 141" refId="ref15969" refString="Xu, Q. - H. (2015) Description of the final stadium larva of Heliocypha perforata perforata (Percheron), with discussion of the taxonomic characters of the larvae of the genus Heliocypha Fraser (Odonata: Zygoptera: Chlorocyphidae). Zootaxa, 3926 (1), 137 - 141. https: // doi. org / 10.11646 / zootaxa. 3926.1.8" type="journal article" year="2015">Xu (2015)</bibRefCitation>
|
||
. The drawing of the antenna in the former paper depicts the scape as 0.33 times the overall length of the antenna, but by the measurements given in the text and the depiction of the antennae on the habitus drawing this proportion should be 0.42, which is very close to the other species considered, hence the detailed drawing itself must be regarded as a lapsus, perhaps because it was inadvertently observed at an angle. The authors do not comment on an obviously relatively short scape. They also illustrate the scape with a rough warty texture, but in
|
||
<taxonomicName id="4C3A4D0BFFD5FFD8DF9F4D52FA0C7D6C" box="[1279,1413,1375,1401]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="biseriata">
|
||
<emphasis id="B94EEA9AFFD5FFD8DF9F4D52FA0C7D6C" box="[1279,1413,1375,1401]" italics="true" pageId="13" pageNumber="314">H. biseriata</emphasis>
|
||
</taxonomicName>
|
||
it was found that these apparent structures were an illusion (seemingly present in photographs taken independently by AGWO and POMS) created by short recurved setae trapping fluid in wet specimens and giving the appearance of small tubercles where none were present. The shape of the prementum in
|
||
<taxonomicName id="4C3A4D0BFFD5FFD8D88B4DC6FBEE7DF0" box="[1003,1127,1483,1509]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="biforata">
|
||
<emphasis id="B94EEA9AFFD5FFD8D88B4DC6FBEE7DF0" box="[1003,1127,1483,1509]" italics="true" pageId="13" pageNumber="314">H. biforata</emphasis>
|
||
</taxonomicName>
|
||
, although basically similar, differs from that of
|
||
<taxonomicName id="4C3A4D0BFFD5FFD8DA1B4DE2FD8A7E1C" box="[379,515,1519,1545]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="biseriata">
|
||
<emphasis id="B94EEA9AFFD5FFD8DA1B4DE2FD8A7E1C" box="[379,515,1519,1545]" italics="true" pageId="13" pageNumber="314">H. biseriata</emphasis>
|
||
</taxonomicName>
|
||
in being relatively shorter, and the anterior median lobe is strongly convex with only a small notch representing the median cleft. The subocular spines on the gena appear very similar. Similarly, in
|
||
<emphasis id="B94EEA9AFFD5FFD8DBD54E3AFED67E44" box="[181,351,1591,1617]" italics="true" pageId="13" pageNumber="314">
|
||
H. p.
|
||
<taxonomicName id="4C3A4D0BFFD5FFD8DB954E3AFED67E44" box="[245,351,1591,1617]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="perforata">perforata</taxonomicName>
|
||
</emphasis>
|
||
the prementum is relatively shorter than in
|
||
<taxonomicName id="4C3A4D0BFFD5FFD8D8324E3AFC507E44" box="[850,985,1591,1617]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="biseriata">
|
||
<emphasis id="B94EEA9AFFD5FFD8D8324E3AFC507E44" box="[850,985,1591,1617]" italics="true" pageId="13" pageNumber="314">H. biseriata</emphasis>
|
||
</taxonomicName>
|
||
but is otherwise very similar. There are also strong similarities in the female genital apophyses. The depiction of the labial palp by
|
||
<bibRefCitation id="EFAB4B79FFD5FFD8DFF24E56FA8F7E60" author="Xu, Q. - H." box="[1170,1286,1626,1653]" pageId="13" pageNumber="314" pagination="137 - 141" refId="ref15969" refString="Xu, Q. - H. (2015) Description of the final stadium larva of Heliocypha perforata perforata (Percheron), with discussion of the taxonomic characters of the larvae of the genus Heliocypha Fraser (Odonata: Zygoptera: Chlorocyphidae). Zootaxa, 3926 (1), 137 - 141. https: // doi. org / 10.11646 / zootaxa. 3926.1.8" type="journal article" year="2015">Xu (2015)</bibRefCitation>
|
||
is erroneous, as it omits the secondary upper branch of the palpal lobe, a structure clearly evident in photographs of this species (
|
||
<bibRefCitation id="EFAB4B79FFD5FFD8DBFF4EAEFE817EA8" author="Ng, C. Y." box="[159,264,1699,1725]" pageId="13" pageNumber="314" refId="ref15661" refString="Ng, C. Y. (2024) Freshwater Macroinvertebrates in Hong Kong: Odonata: Heliocypha perforata. Available from: https: // www. biosch. hku. hk / ecology / staffhp / dd / macroinvertebrates / Odonata / Chlorocyphidae / Rhinocypha _ perforata. html (accessed 1 July 2024)" type="url" year="2024">Ng 2024</bibRefCitation>
|
||
). However both
|
||
<bibRefCitation id="EFAB4B79FFD5FFD8DAA64EAEFD507EA8" author="Kumar, A. & Prasad, M." box="[454,729,1699,1725]" pageId="13" pageNumber="314" pagination="547 - 554" refId="ref15440" refString="Kumar, A. & Prasad, M. (1977) On the larvae of Rhinocypha (Odonata: Chlorocyphidae) from Garhwal hills. Oriental Insects, 11, 547 - 554. https: // doi. org / 10.1080 / 00305316.1977.11090927" type="journal article" year="1977">Kumar & Prasad (1977)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFAB4B79FFD5FFD8D8734EAEFC017EA8" author="Xu, Q. - H." box="[787,904,1698,1725]" pageId="13" pageNumber="314" pagination="137 - 141" refId="ref15969" refString="Xu, Q. - H. (2015) Description of the final stadium larva of Heliocypha perforata perforata (Percheron), with discussion of the taxonomic characters of the larvae of the genus Heliocypha Fraser (Odonata: Zygoptera: Chlorocyphidae). Zootaxa, 3926 (1), 137 - 141. https: // doi. org / 10.11646 / zootaxa. 3926.1.8" type="journal article" year="2015">Xu (2015)</bibRefCitation>
|
||
depict the caudal spikes of
|
||
<taxonomicName id="4C3A4D0BFFD5FFD8DFA94EAEFACE7EA8" box="[1225,1351,1699,1725]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="biforata">
|
||
<emphasis id="B94EEA9AFFD5FFD8DFA94EAEFACE7EA8" box="[1225,1351,1699,1725]" italics="true" pageId="13" pageNumber="314">H. biforata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3A4D0BFFD5FFD8DEE14EAEFE887EF4" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="perforata">
|
||
<emphasis id="B94EEA9AFFD5FFD8DEE14EAEFE887EF4" italics="true" pageId="13" pageNumber="314">H. perforata</emphasis>
|
||
</taxonomicName>
|
||
respectively and they are very similar in form to those of
|
||
<taxonomicName id="4C3A4D0BFFD5FFD8D8FD4ECAFBAC7EF4" box="[925,1061,1735,1761]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="biseriata">
|
||
<emphasis id="B94EEA9AFFD5FFD8D8FD4ECAFBAC7EF4" box="[925,1061,1735,1761]" italics="true" pageId="13" pageNumber="314">H. biseriata</emphasis>
|
||
</taxonomicName>
|
||
, being rather heavy and tapering gradually to a point from around the midpoint (in this respect the drawing by
|
||
<bibRefCitation id="EFAB4B79FFD5FFD8DF4C4EE6FB187F10" author="Xu, Q. - H." box="[1068,1169,1770,1797]" pageId="13" pageNumber="314" pagination="137 - 141" refId="ref15969" refString="Xu, Q. - H. (2015) Description of the final stadium larva of Heliocypha perforata perforata (Percheron), with discussion of the taxonomic characters of the larvae of the genus Heliocypha Fraser (Odonata: Zygoptera: Chlorocyphidae). Zootaxa, 3926 (1), 137 - 141. https: // doi. org / 10.11646 / zootaxa. 3926.1.8" type="journal article" year="2015">Xu 2015</bibRefCitation>
|
||
is misleading, but this character can clearly be seen in the habitus photograph provided). The spikes are relatively long in
|
||
<taxonomicName id="4C3A4D0BFFD5FFD8DF884F02FAFD7F3C" box="[1256,1396,1807,1833]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="perforata">
|
||
<emphasis id="B94EEA9AFFD5FFD8DF884F02FAFD7F3C" box="[1256,1396,1807,1833]" italics="true" pageId="13" pageNumber="314">H. perforata</emphasis>
|
||
</taxonomicName>
|
||
(ca 55% body length) but are figured as shorter in
|
||
<taxonomicName id="4C3A4D0BFFD5FFD8D9D14F3EFCB47F58" box="[689,829,1843,1869]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="bisignata">
|
||
<emphasis id="B94EEA9AFFD5FFD8D9D14F3EFCB47F58" box="[689,829,1843,1869]" italics="true" pageId="13" pageNumber="314">H. bisignata</emphasis>
|
||
</taxonomicName>
|
||
(ca 46% body length) depicted by
|
||
<bibRefCitation id="EFAB4B79FFD5FFD8DFB24F3EFAE27F58" author="Fraser, F. C." box="[1234,1387,1843,1869]" pageId="13" pageNumber="314" pagination="683 - 691" refId="ref15149" refString="Fraser, F. C. (1928) Indian dragonflies, Part XXX. Journal of the Bombay Natural History Society, 32, 683 - 691." type="journal article" year="1928">Fraser (1928)</bibRefCitation>
|
||
and later briefly described by the same author (
|
||
<bibRefCitation id="EFAB4B79FFD5FFD8D90D4F5AFD7E7F64" author="Fraser, F. C." box="[621,759,1879,1905]" pageId="13" pageNumber="314" refId="ref15178" refString="Fraser, F. C. (1934) The fauna of British India, including Ceylon and Burma. Odonata. Vol. II. Taylor & Francis Ltd., London, xxiii + 398 pp." type="book" year="1934">Fraser 1934</bibRefCitation>
|
||
).Although lacking subtlety, the 1928 illustration agrees quite well with other
|
||
<taxonomicName id="4C3A4D0BFFD5FFD8DA2B4F76FDA17F80" box="[331,552,1915,1941]" pageId="13" pageNumber="314">
|
||
<emphasis id="B94EEA9AFFD5FFD8DA2B4F76FE457F80" box="[331,460,1915,1941]" italics="true" pageId="13" pageNumber="314">Heliocypha</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
, especially in the form of the caudal spikes. Neither
|
||
<bibRefCitation id="EFAB4B79FFD5FFD8DF1D4F76FB7B7F80" author="Xu, Q. - H." box="[1149,1266,1914,1941]" pageId="13" pageNumber="314" pagination="137 - 141" refId="ref15969" refString="Xu, Q. - H. (2015) Description of the final stadium larva of Heliocypha perforata perforata (Percheron), with discussion of the taxonomic characters of the larvae of the genus Heliocypha Fraser (Odonata: Zygoptera: Chlorocyphidae). Zootaxa, 3926 (1), 137 - 141. https: // doi. org / 10.11646 / zootaxa. 3926.1.8" type="journal article" year="2015">Xu (2015)</bibRefCitation>
|
||
nor
|
||
<bibRefCitation id="EFAB4B79FFD5FFD8DE474F76FA157F80" author="Ng, C. Y." box="[1319,1436,1915,1941]" pageId="13" pageNumber="314" refId="ref15661" refString="Ng, C. Y. (2024) Freshwater Macroinvertebrates in Hong Kong: Odonata: Heliocypha perforata. Available from: https: // www. biosch. hku. hk / ecology / staffhp / dd / macroinvertebrates / Odonata / Chlorocyphidae / Rhinocypha _ perforata. html (accessed 1 July 2024)" type="url" year="2024">Ng (2024)</bibRefCitation>
|
||
examine the subocular spines on the gena, of
|
||
<taxonomicName id="4C3A4D0BFFD5FFD8D9EE4F92FC907FAC" box="[654,793,1951,1977]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="perforata">
|
||
<emphasis id="B94EEA9AFFD5FFD8D9EE4F92FC907FAC" box="[654,793,1951,1977]" italics="true" pageId="13" pageNumber="314">H. perforata</emphasis>
|
||
</taxonomicName>
|
||
but it seems these, and the form of the caudal spikes, may be characteristic of the genus. Both
|
||
<taxonomicName id="4C3A4D0BFFD5FFD8D94B4FCEFD217FC8" box="[555,680,1987,2013]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="biforata">
|
||
<emphasis id="B94EEA9AFFD5FFD8D94B4FCEFD217FC8" box="[555,680,1987,2013]" italics="true" pageId="13" pageNumber="314">H. biforata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3A4D0BFFD5FFD8D9BF4FCEFCE57FC8" box="[735,876,1987,2013]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="perforata">
|
||
<emphasis id="B94EEA9AFFD5FFD8D9BF4FCEFCE57FC8" box="[735,876,1987,2013]" italics="true" pageId="13" pageNumber="314">H. perforata</emphasis>
|
||
</taxonomicName>
|
||
also have long legs, with the metafemur reaching the hind margin of S
|
||
<quantity id="4CC29B6DFFD6FFDBDAE5489AFE3A78A4" box="[389,435,151,177]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="14" pageNumber="315" unit="in" value="7.0">7 in</quantity>
|
||
F stadium larvae. The depiction of the prementum and labial palps of
|
||
<taxonomicName id="4C3A4D0BFFD6FFDBDF83489AFAFE78A4" box="[1251,1399,151,177]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="fenestrata">
|
||
<emphasis id="B94EEA9AFFD6FFDBDF83489AFAFE78A4" box="[1251,1399,151,177]" italics="true" pageId="14" pageNumber="315">H. fenestrata</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EFAB4B79FFD6FFDBDBF748B6FEDC78C0" author="Lieftinck, M. A." box="[151,341,187,213]" pageId="14" pageNumber="315" pagination="1 - 95" refId="ref15617" refString="Lieftinck, M. A. (1962) Odonata. Insects of Micronesia, 5 (1), 1 - 95. https: // doi. org / 10.1007 / 978 - 94 - 015 - 1100 - 1 _ 1" type="journal article" year="1962">Lieftinck (1962)</bibRefCitation>
|
||
, is very reminiscent of
|
||
<taxonomicName id="4C3A4D0BFFD6FFDBD90C48B6FD7F78C0" box="[620,758,187,213]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="biseriata">
|
||
<emphasis id="B94EEA9AFFD6FFDBD90C48B6FD7F78C0" box="[620,758,187,213]" italics="true" pageId="14" pageNumber="315">H. biseriata</emphasis>
|
||
</taxonomicName>
|
||
but the habitus drawing does not depict the caudal spikes convincingly. This, and the style, strongly suggest the drawing was executed by a Bogor Museum staff member such as G. Abdulkadir rather than Lieftinck himself (see
|
||
<bibRefCitation id="EFAB4B79FFD6FFDBD983490EFCF57908" author="van Tol, J." box="[739,892,259,285]" pageId="14" pageNumber="315" pagination="1 - 263" refId="ref15935" refString="van Tol, J. (1992) An annotated index to names of Odonata used in publications by M. A. Lieftinck. Zoologiische Verhandelingen, 279, 1 - 263." type="journal article" year="1992">van Tol 1992</bibRefCitation>
|
||
), and it is well known that many such drawings are more decorative than accurate.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |