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<document id="5116BAEB2B32CB1DA8D2B08BFB413FE0" ID-DOI="10.5281/zenodo.3463445" ID-GBIF-Dataset="d20f0d8c-0592-40d4-94dd-6d9ec60dfe49" ID-ISSN="0035-418" ID-Zenodo-Dep="3463445" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1583846354969" checkinUser="carolina" docAuthor="Huang, Meng-Chi &amp; Yin, Zi-Wei" docDate="2019" docId="0393879EFF9E4B6666CFFE8019B99638" docLanguage="en" docName="RevSuisseZool.126.2.165-196.pdf.imf" docOrigin="Revue suisse de Zoologie 126 (2)" docStyle="DocumentStyle:225056F0A23C51DDC06AC27EAEA15D5B.2:RevSuisseZool.2015-2019.journal_article" docStyleId="225056F0A23C51DDC06AC27EAEA15D5B" docStyleName="RevSuisseZool.2015-2019.journal_article" docStyleVersion="2" docTitle="Pselaphodes procerus Huang &amp; Yin, 2019, sp. nov." docType="treatment" docVersion="5" lastPageNumber="181" masterDocId="FFAAFFE6FF934B766507FF831B3E9704" masterDocTitle="The Pselaphodes (Coleoptera: Staphylinidae: Pselaphinae) of Nepal" masterLastPageNumber="196" masterPageNumber="165" pageNumber="178" updateTime="1698726426458" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="2E3A3A4A39EA56CDA2290C347C3B9285">The Pselaphodes (Coleoptera: Staphylinidae: Pselaphinae) of Nepal</mods:title>
</mods:titleInfo>
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<mods:namePart id="8B1D88607EA0FB41D79C2EA1261EE5CB">Huang, Meng-Chi</mods:namePart>
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<mods:namePart id="4E41817414313BB770ADF296FA94994D">Yin, Zi-Wei</mods:namePart>
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<mods:title id="9DC2BBF502E7CDD31A4B130482159549">Revue suisse de Zoologie</mods:title>
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<mods:date id="F3DDB99E42026FB0020AF6813B0BFE60">2019</mods:date>
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<mods:number id="F1617B2EBEA2427CFC235A1973B7F7B0">2019-09-30</mods:number>
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<treatment id="0393879EFF9E4B6666CFFE8019B99638" ID-DOI="http://doi.org/10.5281/zenodo.3706091" ID-GBIF-Taxon="162481179" ID-Zenodo-Dep="3706091" LSID="urn:lsid:plazi:treatment:0393879EFF9E4B6666CFFE8019B99638" httpUri="http://treatment.plazi.org/id/0393879EFF9E4B6666CFFE8019B99638" lastPageId="16" lastPageNumber="181" pageId="13" pageNumber="178">
<subSubSection id="C3206503FF9E4B7B66CFFE801E2A961F" box="[968,1300,259,283]" pageId="13" pageNumber="178" type="nomenclature">
<paragraph id="8B853688FF9E4B7B66CFFE801E2A961F" blockId="13.[968,1300,259,315]" box="[968,1300,259,283]" pageId="13" pageNumber="178">
<heading id="D0CD81E4FF9E4B7B66CFFE801E2A961F" bold="true" box="[968,1300,259,283]" centered="true" level="2" pageId="13" pageNumber="178" reason="2">
<emphasis id="B94EEA9AFF9E4B7B66CFFE801E2A961F" bold="true" box="[968,1300,259,283]" pageId="13" pageNumber="178">
<taxonomicName id="4C3A4D0BFF9E4B7B66CFFE801F89961F" ID-CoL="8Y8Q9" box="[968,1207,259,283]" class="Insecta" family="Staphylinidae" genus="Pselaphodes" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="178" phylum="Arthropoda" rank="species" species="procerus" status="sp. nov.">
<emphasis id="B94EEA9AFF9E4B7B66CFFE801F89961F" bold="true" box="[968,1207,259,283]" italics="true" pageId="13" pageNumber="178">Pselaphodes procerus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27D57E1FF9E4B7B61B9FE801E2A961F" box="[1214,1300,259,283]" pageId="13" pageNumber="178" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3206503FF9E4B7B6113FEA01FF4963F" box="[1044,1226,291,315]" pageId="13" pageNumber="178" type="description">
<paragraph id="8B853688FF9E4B7B6113FEA01FF4963F" blockId="13.[968,1300,259,315]" box="[1044,1226,291,315]" pageId="13" pageNumber="178">
<figureCitation id="13012A0DFF9E4B7B6113FEA01F49963F" box="[1044,1143,291,315]" captionStart="Fig 10" captionStartId="12.[151,183,2056,2078]" captionTargetBox="[155,1411,892,2022]" captionTargetId="figure@12.[151,1411,890,2028]" captionTargetPageId="12" captionText="Fig 10. Dorsal habitus of Pselaphodes males. (A) P. loebli sp. nov. (B) P. procerus sp. nov. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3702671" httpUri="https://zenodo.org/record/3702671/files/figure.png" pageId="13" pageNumber="178">Figs 10B</figureCitation>
,
<figureCitation id="13012A0DFF9E4B7B6185FEA01FA5963F" box="[1154,1179,291,315]" captionStart="Fig 12" captionStartId="15.[151,183,1799,1821]" captionTargetBox="[157,1411,261,1744]" captionTargetId="figure@15.[151,1411,257,1750]" captionTargetPageId="15" captionText="Fig 12. Male diagnostic features of Pselaphodes procerus sp. nov. (A) Antennal club. (B) Pronotum. (C) Metaventral process, in lateral view. (D) Protrochanter and profemur. (E) Apex of protibia. (F) Mesotrochanter and mesofemur. (G) Apex of mesotibia. (H) Metatrochanter and metafemur. (I) Sternite IX. (J-L) Aedeagus, in dorsal (J), lateral (K), and ventral (L) view. Scale bars: 0.3 mm in A-B, D, F, H; 0.2 mm in C, J-L; 0.1 mm in E, G, I." figureDoi="http://doi.org/10.5281/zenodo.3702675" httpUri="https://zenodo.org/record/3702675/files/figure.png" pageId="13" pageNumber="178">12</figureCitation>
A-L
</paragraph>
</subSubSection>
<subSubSection id="C3206503FF9E4B7B6647FED11894950E" pageId="13" pageNumber="178" type="materials_examined">
<paragraph id="8B853688FF9E4B7B6647FED11894950E" blockId="13.[832,1437,338,522]" pageId="13" pageNumber="178">
<emphasis id="B94EEA9AFF9E4B7B6647FED118D1966E" bold="true" box="[832,1007,338,362]" pageId="13" pageNumber="178">Type material</emphasis>
(
<materialsCitation id="3B523CD5FF9E4B7B6117FED11894950E" ID-GBIF-Occurrence="2575116020" collectingDate="2000-09-14" collectionCode="MHNG" collectorName="NE Besisahar &amp; A. Hetzel" country="Nepal" elevation="3950" latitude="28.35" location="Manaslu" longLatPrecision="1232" longitude="84.55" municipality="Barapokhari Lekh" pageId="13" pageNumber="178" specimenCount="1" typeStatus="holotype">
<specimenCount id="9D3CFD01FF9E4B7B6117FED11FA7966E" box="[1040,1177,338,362]" pageId="13" pageNumber="178" type="generic">1 specimen</specimenCount>
):
<typeStatus id="5481882AFF9E4B7B61B8FED11E16966E" box="[1215,1320,338,362]" pageId="13" pageNumber="178" type="holotype">Holotype</typeStatus>
; MHNG- ENTO-44008; ♂;
<collectingCountry id="F32D7618FF9E4B7B610BFEF11F5D968E" box="[1036,1123,370,394]" name="Nepal" pageId="13" pageNumber="178">NEPAL</collectingCountry>
, C-Nepal:
<location id="8EE56053FF9E4B7B61EFFEF11E7A968E" LSID="urn:lsid:plazi:treatment:0393879EFF9E4B6666CFFE8019B99638:8EE56053FF9E4B7B61EFFEF11E7A968E" box="[1256,1348,370,394]" country="Nepal" latitude="28.35" longLatPrecision="1232" longitude="84.55" municipality="Barapokhari Lekh" name="Manaslu" pageId="13" pageNumber="178">Manaslu</location>
massif,
<collectingMunicipality id="6BE1ACF2FF9E4B7B6647FE111F3996AE" box="[832,1031,402,426]" pageId="13" pageNumber="178">Barapokhari Lekh</collectingMunicipality>
,
<quantity id="4CC29B6DFF9E4B7B6113FE111F6396AE" box="[1044,1117,402,426]" metricMagnitude="4" metricUnit="m" metricValue="2.3" pageId="13" pageNumber="178" unit="km" value="23.0">23 km</quantity>
,
<collectorName id="26CF535EFF9E4B7B616DFE101FC196AE" box="[1130,1279,402,426]" pageId="13" pageNumber="178">NE Besisahar</collectorName>
vill.
<geoCoordinate id="EE0E504FFF9E4B7B603BFE121EA696AF" box="[1340,1432,401,427]" degrees="28" direction="north" minutes="21" orientation="latitude" pageId="13" pageNumber="178" precision="925" value="28.35">28°21N</geoCoordinate>
,
<geoCoordinate id="EE0E504FFF9E4B7B6647FE3218A796CF" box="[832,921,433,459]" degrees="84" direction="east" minutes="33" orientation="longitude" pageId="13" pageNumber="178" precision="925" value="84.55">84°33E</geoCoordinate>
,
<date id="FF841048FF9E4B7B66AAFE311F1296CE" box="[941,1068,434,458]" pageId="13" pageNumber="178" value="2000-09-14">
<collectingDate id="EFC0E9A0FF9E4B7B66AAFE311F1296CE" box="[941,1068,434,458]" pageId="13" pageNumber="178" value="2000-09-14">14.IX.2000</collectingDate>
</date>
, leg.
<collectorName id="26CF535EFF9E4B7B6172FE311FD696CE" box="[1141,1256,434,458]" pageId="13" pageNumber="178">A. Hetzel</collectorName>
,
<quantity id="4CC29B6DFF9E4B7B61FBFE311EA696CE" box="[1276,1432,434,458]" metricMagnitude="3" metricUnit="m" metricValue="3.95" metricValueMax="4.1" metricValueMin="3.8" pageId="13" pageNumber="178" unit="m" value="3950.0" valueMax="4100.0" valueMin="3800.0">
<elevation id="0017D1BBFF9E4B7B61FBFE311EA696CE" box="[1276,1432,434,458]" metricMagnitude="3" metricUnit="m" metricValue="3.95" metricValueMax="4.1" metricValueMin="3.8" pageId="13" pageNumber="178" unit="m" value="3950.0" valueMax="4100.0" valueMin="3800.0">3800-4100 m</elevation>
</quantity>
, sieved from moss and
<taxonomicName id="4C3A4D0BFF9E4B7B616AFE511E3096EE" box="[1133,1294,466,490]" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" order="Ericales" pageId="13" pageNumber="178" phylum="Tracheophyta" rank="genus">
<emphasis id="B94EEA9AFF9E4B7B616AFE511E3096EE" box="[1133,1294,466,490]" italics="true" pageId="13" pageNumber="178">Rhododendron</emphasis>
</taxonomicName>
leaf litter (
<collectionCode id="ED2BAE4DFF9E4B7B6640FE7118A1950E" box="[839,927,498,522]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="13" pageNumber="178" type="Museum">MHNG</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3206503FF9E4B7B6647FDA11E3B955D" pageId="13" pageNumber="178" type="distribution">
<paragraph id="8B853688FF9E4B7B6647FDA11E3B955D" blockId="13.[832,1436,545,601]" pageId="13" pageNumber="178">
<emphasis id="B94EEA9AFF9E4B7B6647FDA118D5953D" bold="true" box="[832,1003,545,570]" pageId="13" pageNumber="178">
<typeStatus id="5481882AFF9E4B7B6647FDA11847953E" box="[832,889,546,570]" pageId="13" pageNumber="178">Type</typeStatus>
locality:
</emphasis>
<collectingCountry id="F32D7618FF9E4B7B66F8FDA21F7D953D" box="[1023,1091,545,569]" name="Nepal" pageId="13" pageNumber="178">Nepal</collectingCountry>
,
<location id="8EE56053FF9E4B7B615CFDA21E19953D" LSID="urn:lsid:plazi:treatment:0393879EFF9E4B6666CFFE8019B99638:8EE56053FF9E4B7B615CFDA21E19953D" box="[1115,1319,545,569]" country="Nepal" county="Bara Pokhari" municipality="Manaslu Mountain" name="Lamjung District" pageId="13" pageNumber="178">Lamjung District</location>
,
<collectingMunicipality id="6BE1ACF2FF9E4B7B6038FDA21894955D" pageId="13" pageNumber="178">Manaslu Mountain</collectingMunicipality>
,
<collectingCounty id="62E44E04FF9E4B7B66B2FDC11F7A955D" box="[949,1092,577,601]" pageId="13" pageNumber="178">Bara Pokhari</collectingCounty>
,
<quantity id="4CC29B6DFF9E4B7B6149FDC21FBA955D" box="[1102,1156,577,601]" metricMagnitude="3" metricUnit="m" metricValue="3.95" metricValueMax="4.1" metricValueMin="3.8" pageId="13" pageNumber="178" unit="m" value="3950.0" valueMax="4100.0" valueMin="3800.0">3800</quantity>
-4100 m alt.
</paragraph>
</subSubSection>
<subSubSection id="C3206503FF9E4B7B6647FDF31EAC94AC" pageId="13" pageNumber="178" type="diagnosis">
<paragraph id="8B853688FF9E4B7B6647FDF31EAC94AC" blockId="13.[832,1437,624,936]" pageId="13" pageNumber="178">
<emphasis id="B94EEA9AFF9E4B7B6647FDF31F1C958D" bold="true" box="[832,1058,624,649]" pageId="13" pageNumber="178">Diagnosis of males:</emphasis>
Length
<quantity id="4CC29B6DFF9E4B7B6185FDF31FD6958C" box="[1154,1256,624,648]" metricMagnitude="-3" metricUnit="m" metricValue="3.15" pageId="13" pageNumber="178" unit="mm" value="3.15">3.15 mm</quantity>
. Antennomere 9 with projection at apex, antennomeres 10 with distinct projection at base. Metaventral processes broad, and apically forked. Protrochanters with acute ventral spine, profemora with blunt ventral spine, protibiae with distinct projection at apex; mesotrochanters with long acute ventral spine, mesotibiae with two distinct projections at apex; metacoxae with large blunt ventral projection. Median lobe of aedeagus broad apically; parameres moderately elongate and apically broadened.
</paragraph>
</subSubSection>
<subSubSection id="C3206503FF9E4B7B6647FC3C1F329013" pageId="13" pageNumber="178" type="description">
<paragraph id="8B853688FF9E4B7B6647FC3C18A091F3" blockId="13.[831,1437,959,1815]" pageId="13" pageNumber="178">
<emphasis id="B94EEA9AFF9E4B7B6647FC3C18F094D3" bold="true" box="[832,974,959,983]" pageId="13" pageNumber="178">Description:</emphasis>
Male (
<figureCitation id="13012A0DFF9E4B7B612FFC3C1FAC94DC" box="[1064,1170,959,984]" captionStart="Fig 10" captionStartId="12.[151,183,2056,2078]" captionTargetBox="[155,1411,892,2022]" captionTargetId="figure@12.[151,1411,890,2028]" captionTargetPageId="12" captionText="Fig 10. Dorsal habitus of Pselaphodes males. (A) P. loebli sp. nov. (B) P. procerus sp. nov. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3702671" httpUri="https://zenodo.org/record/3702671/files/figure.png" pageId="13" pageNumber="178">Fig. 10B</figureCitation>
). Body reddish brown, BL
<quantity id="4CC29B6DFF9E4B7B6669FC5C18E894F3" box="[878,982,991,1015]" metricMagnitude="-3" metricUnit="m" metricValue="3.15" pageId="13" pageNumber="178" unit="mm" value="3.15">3.15 mm</quantity>
. Head longer than wide, HL
<quantity id="4CC29B6DFF9E4B7B6037FC5C1EA694F3" box="[1328,1432,991,1015]" metricMagnitude="-4" metricUnit="m" metricValue="6.7" pageId="13" pageNumber="178" unit="mm" value="0.67">0.67 mm</quantity>
, HW
<quantity id="4CC29B6DFF9E4B7B6671FC7C18E29313" box="[886,988,1023,1047]" metricMagnitude="-4" metricUnit="m" metricValue="6.2" pageId="13" pageNumber="178" unit="mm" value="0.62">0.62 mm</quantity>
; each eye composed of about 45 facets; with well-developed ocular canthus. Antennomeres (
<figureCitation id="13012A0DFF9E4B7B664FFBBC188E935C" box="[840,944,1087,1112]" captionStart="Fig 12" captionStartId="15.[151,183,1799,1821]" captionTargetBox="[157,1411,261,1744]" captionTargetId="figure@15.[151,1411,257,1750]" captionTargetPageId="15" captionText="Fig 12. Male diagnostic features of Pselaphodes procerus sp. nov. (A) Antennal club. (B) Pronotum. (C) Metaventral process, in lateral view. (D) Protrochanter and profemur. (E) Apex of protibia. (F) Mesotrochanter and mesofemur. (G) Apex of mesotibia. (H) Metatrochanter and metafemur. (I) Sternite IX. (J-L) Aedeagus, in dorsal (J), lateral (K), and ventral (L) view. Scale bars: 0.3 mm in A-B, D, F, H; 0.2 mm in C, J-L; 0.1 mm in E, G, I." figureDoi="http://doi.org/10.5281/zenodo.3702675" httpUri="https://zenodo.org/record/3702675/files/figure.png" pageId="13" pageNumber="178">Fig. 12A</figureCitation>
) 9-11 forming distinct club, antennomeres 9 with projection at apex, antennomeres 10 with distinct projection at base. Pronotum (
<figureCitation id="13012A0DFF9E4B7B6001FBFC1E50939C" box="[1286,1390,1151,1176]" captionStart="Fig 12" captionStartId="15.[151,183,1799,1821]" captionTargetBox="[157,1411,261,1744]" captionTargetId="figure@15.[151,1411,257,1750]" captionTargetPageId="15" captionText="Fig 12. Male diagnostic features of Pselaphodes procerus sp. nov. (A) Antennal club. (B) Pronotum. (C) Metaventral process, in lateral view. (D) Protrochanter and profemur. (E) Apex of protibia. (F) Mesotrochanter and mesofemur. (G) Apex of mesotibia. (H) Metatrochanter and metafemur. (I) Sternite IX. (J-L) Aedeagus, in dorsal (J), lateral (K), and ventral (L) view. Scale bars: 0.3 mm in A-B, D, F, H; 0.2 mm in C, J-L; 0.1 mm in E, G, I." figureDoi="http://doi.org/10.5281/zenodo.3702675" httpUri="https://zenodo.org/record/3702675/files/figure.png" pageId="13" pageNumber="178">Fig. 12B</figureCitation>
) as long as wide, PL
<quantity id="4CC29B6DFF9E4B7B6116FB1C1F4793B3" box="[1041,1145,1183,1207]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="13" pageNumber="178" unit="mm" value="0.65">0.65 mm</quantity>
, PW
<quantity id="4CC29B6DFF9E4B7B61B9FB1C1E1B93B3" box="[1214,1317,1183,1207]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="13" pageNumber="178" unit="mm" value="0.65">0.65 mm</quantity>
, angularly expanded at anterolateral margins, strongly constricted at apical third. Elytra wider than long, EL
<quantity id="4CC29B6DFF9E4B7B6036FB5C1EA693F3" box="[1329,1432,1247,1271]" metricMagnitude="-4" metricUnit="m" metricValue="9.6" pageId="13" pageNumber="178" unit="mm" value="0.96">0.96 mm</quantity>
, EW
<quantity id="4CC29B6DFF9E4B7B6679FA8318CE9213" box="[894,1008,1280,1304]" metricMagnitude="-3" metricUnit="m" metricValue="1.34" pageId="13" pageNumber="178" unit="mm" value="1.34">1.34 mm</quantity>
. Metaventral processes (
<figureCitation id="13012A0DFF9E4B7B6022FB7C1EAA921C" box="[1317,1428,1279,1304]" captionStart="Fig 12" captionStartId="15.[151,183,1799,1821]" captionTargetBox="[157,1411,261,1744]" captionTargetId="figure@15.[151,1411,257,1750]" captionTargetPageId="15" captionText="Fig 12. Male diagnostic features of Pselaphodes procerus sp. nov. (A) Antennal club. (B) Pronotum. (C) Metaventral process, in lateral view. (D) Protrochanter and profemur. (E) Apex of protibia. (F) Mesotrochanter and mesofemur. (G) Apex of mesotibia. (H) Metatrochanter and metafemur. (I) Sternite IX. (J-L) Aedeagus, in dorsal (J), lateral (K), and ventral (L) view. Scale bars: 0.3 mm in A-B, D, F, H; 0.2 mm in C, J-L; 0.1 mm in E, G, I." figureDoi="http://doi.org/10.5281/zenodo.3702675" httpUri="https://zenodo.org/record/3702675/files/figure.png" pageId="13" pageNumber="178">Fig. 12C</figureCitation>
) broad, apically forked, anterior branch smaller than posterior branch. Protrochanters with acute ventral spine, profemora with large, blunt ventral spine (
<figureCitation id="13012A0DFF9E4B7B6640FAFC1892929C" box="[839,940,1407,1432]" captionStart="Fig 12" captionStartId="15.[151,183,1799,1821]" captionTargetBox="[157,1411,261,1744]" captionTargetId="figure@15.[151,1411,257,1750]" captionTargetPageId="15" captionText="Fig 12. Male diagnostic features of Pselaphodes procerus sp. nov. (A) Antennal club. (B) Pronotum. (C) Metaventral process, in lateral view. (D) Protrochanter and profemur. (E) Apex of protibia. (F) Mesotrochanter and mesofemur. (G) Apex of mesotibia. (H) Metatrochanter and metafemur. (I) Sternite IX. (J-L) Aedeagus, in dorsal (J), lateral (K), and ventral (L) view. Scale bars: 0.3 mm in A-B, D, F, H; 0.2 mm in C, J-L; 0.1 mm in E, G, I." figureDoi="http://doi.org/10.5281/zenodo.3702675" httpUri="https://zenodo.org/record/3702675/files/figure.png" pageId="13" pageNumber="178">Fig. 12D</figureCitation>
), protibiae (
<figureCitation id="13012A0DFF9E4B7B6136FAFC1FAF929C" box="[1073,1169,1407,1432]" captionStart="Fig 12" captionStartId="15.[151,183,1799,1821]" captionTargetBox="[157,1411,261,1744]" captionTargetId="figure@15.[151,1411,257,1750]" captionTargetPageId="15" captionText="Fig 12. Male diagnostic features of Pselaphodes procerus sp. nov. (A) Antennal club. (B) Pronotum. (C) Metaventral process, in lateral view. (D) Protrochanter and profemur. (E) Apex of protibia. (F) Mesotrochanter and mesofemur. (G) Apex of mesotibia. (H) Metatrochanter and metafemur. (I) Sternite IX. (J-L) Aedeagus, in dorsal (J), lateral (K), and ventral (L) view. Scale bars: 0.3 mm in A-B, D, F, H; 0.2 mm in C, J-L; 0.1 mm in E, G, I." figureDoi="http://doi.org/10.5281/zenodo.3702675" httpUri="https://zenodo.org/record/3702675/files/figure.png" pageId="13" pageNumber="178">Fig. 12E</figureCitation>
) with distinct projection at apex; mesotrochanters (
<figureCitation id="13012A0DFF9E4B7B616CFA1C1FF092BC" box="[1131,1230,1439,1464]" captionStart="Fig 12" captionStartId="15.[151,183,1799,1821]" captionTargetBox="[157,1411,261,1744]" captionTargetId="figure@15.[151,1411,257,1750]" captionTargetPageId="15" captionText="Fig 12. Male diagnostic features of Pselaphodes procerus sp. nov. (A) Antennal club. (B) Pronotum. (C) Metaventral process, in lateral view. (D) Protrochanter and profemur. (E) Apex of protibia. (F) Mesotrochanter and mesofemur. (G) Apex of mesotibia. (H) Metatrochanter and metafemur. (I) Sternite IX. (J-L) Aedeagus, in dorsal (J), lateral (K), and ventral (L) view. Scale bars: 0.3 mm in A-B, D, F, H; 0.2 mm in C, J-L; 0.1 mm in E, G, I." figureDoi="http://doi.org/10.5281/zenodo.3702675" httpUri="https://zenodo.org/record/3702675/files/figure.png" pageId="13" pageNumber="178">Fig. 12F</figureCitation>
) with long, sharp ventral spine; mesotibiae (
<figureCitation id="13012A0DFF9E4B7B6160FA3C1FF392DC" box="[1127,1229,1471,1496]" captionStart="Fig 12" captionStartId="15.[151,183,1799,1821]" captionTargetBox="[157,1411,261,1744]" captionTargetId="figure@15.[151,1411,257,1750]" captionTargetPageId="15" captionText="Fig 12. Male diagnostic features of Pselaphodes procerus sp. nov. (A) Antennal club. (B) Pronotum. (C) Metaventral process, in lateral view. (D) Protrochanter and profemur. (E) Apex of protibia. (F) Mesotrochanter and mesofemur. (G) Apex of mesotibia. (H) Metatrochanter and metafemur. (I) Sternite IX. (J-L) Aedeagus, in dorsal (J), lateral (K), and ventral (L) view. Scale bars: 0.3 mm in A-B, D, F, H; 0.2 mm in C, J-L; 0.1 mm in E, G, I." figureDoi="http://doi.org/10.5281/zenodo.3702675" httpUri="https://zenodo.org/record/3702675/files/figure.png" pageId="13" pageNumber="178">Fig. 12G</figureCitation>
) with two distinct projections at apex; metacoxae (
<figureCitation id="13012A0DFF9E4B7B61B7FA5C1E2992FC" box="[1200,1303,1503,1528]" captionStart="Fig 12" captionStartId="15.[151,183,1799,1821]" captionTargetBox="[157,1411,261,1744]" captionTargetId="figure@15.[151,1411,257,1750]" captionTargetPageId="15" captionText="Fig 12. Male diagnostic features of Pselaphodes procerus sp. nov. (A) Antennal club. (B) Pronotum. (C) Metaventral process, in lateral view. (D) Protrochanter and profemur. (E) Apex of protibia. (F) Mesotrochanter and mesofemur. (G) Apex of mesotibia. (H) Metatrochanter and metafemur. (I) Sternite IX. (J-L) Aedeagus, in dorsal (J), lateral (K), and ventral (L) view. Scale bars: 0.3 mm in A-B, D, F, H; 0.2 mm in C, J-L; 0.1 mm in E, G, I." figureDoi="http://doi.org/10.5281/zenodo.3702675" httpUri="https://zenodo.org/record/3702675/files/figure.png" pageId="13" pageNumber="178">Fig. 12H</figureCitation>
) with large blunt ventral projection. Abdomen broad at base and narrowing apically, AL
<quantity id="4CC29B6DFF9E4B7B6139F9A31F9E9133" box="[1086,1184,1568,1592]" metricMagnitude="-4" metricUnit="m" metricValue="8.7" pageId="13" pageNumber="178" unit="mm" value="0.87">0.87 mm</quantity>
, AW
<quantity id="4CC29B6DFF9E4B7B61DCF9A31E019133" box="[1243,1343,1568,1592]" metricMagnitude="-3" metricUnit="m" metricValue="1.31" pageId="13" pageNumber="178" unit="mm" value="1.31">1.31 mm</quantity>
. Sternite IX semi-membranous, shape as in
<figureCitation id="13012A0DFF9E4B7B61C1F9BC1E1F915C" box="[1222,1313,1599,1624]" captionStart="Fig 12" captionStartId="15.[151,183,1799,1821]" captionTargetBox="[157,1411,261,1744]" captionTargetId="figure@15.[151,1411,257,1750]" captionTargetPageId="15" captionText="Fig 12. Male diagnostic features of Pselaphodes procerus sp. nov. (A) Antennal club. (B) Pronotum. (C) Metaventral process, in lateral view. (D) Protrochanter and profemur. (E) Apex of protibia. (F) Mesotrochanter and mesofemur. (G) Apex of mesotibia. (H) Metatrochanter and metafemur. (I) Sternite IX. (J-L) Aedeagus, in dorsal (J), lateral (K), and ventral (L) view. Scale bars: 0.3 mm in A-B, D, F, H; 0.2 mm in C, J-L; 0.1 mm in E, G, I." figureDoi="http://doi.org/10.5281/zenodo.3702675" httpUri="https://zenodo.org/record/3702675/files/figure.png" pageId="13" pageNumber="178">Fig. 12I</figureCitation>
. Length of aedeagus (
<figureCitation id="13012A0DFF9E4B7B66B0F9DC1F31917C" box="[951,1039,1631,1656]" captionStart="Fig 12" captionStartId="15.[151,183,1799,1821]" captionTargetBox="[157,1411,261,1744]" captionTargetId="figure@15.[151,1411,257,1750]" captionTargetPageId="15" captionText="Fig 12. Male diagnostic features of Pselaphodes procerus sp. nov. (A) Antennal club. (B) Pronotum. (C) Metaventral process, in lateral view. (D) Protrochanter and profemur. (E) Apex of protibia. (F) Mesotrochanter and mesofemur. (G) Apex of mesotibia. (H) Metatrochanter and metafemur. (I) Sternite IX. (J-L) Aedeagus, in dorsal (J), lateral (K), and ventral (L) view. Scale bars: 0.3 mm in A-B, D, F, H; 0.2 mm in C, J-L; 0.1 mm in E, G, I." figureDoi="http://doi.org/10.5281/zenodo.3702675" httpUri="https://zenodo.org/record/3702675/files/figure.png" pageId="13" pageNumber="178">Figs 12</figureCitation>
J-L)
<quantity id="4CC29B6DFF9E4B7B614DF9DC1F8A9173" box="[1098,1204,1631,1655]" metricMagnitude="-4" metricUnit="m" metricValue="6.8999999999999995" pageId="13" pageNumber="178" unit="mm" value="0.69">0.69 mm</quantity>
; median lobe broad and asymmetric (basal capsule collapsed in
<typeStatus id="5481882AFF9E4B7B6029F9FC1EAF9193" box="[1326,1425,1663,1687]" pageId="13" pageNumber="178" type="holotype">holotype</typeStatus>
); parameres narrowed at base and broadened apically; endophallus composed of one long and one short sclerites.
</paragraph>
<paragraph id="8B853688FF9E4B7B6638F97C1F329013" blockId="13.[831,1437,959,1815]" box="[831,1036,1791,1815]" pageId="13" pageNumber="178">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C3206503FF9E4B7B6638F8AD1FB79043" box="[831,1161,1838,1863]" pageId="13" pageNumber="178" type="distribution">
<paragraph id="8B853688FF9E4B7B6638F8AD1FB79043" blockId="13.[831,1161,1838,1863]" box="[831,1161,1838,1863]" pageId="13" pageNumber="178">
<emphasis id="B94EEA9AFF9E4B7B6638F8AD18EA9042" bold="true" box="[831,980,1838,1862]" pageId="13" pageNumber="178">Distribution:</emphasis>
<collectingCountry id="F32D7618FF9E4B7B66DDF8AC1F229043" box="[986,1052,1839,1863]" name="Nepal" pageId="13" pageNumber="178">Nepal</collectingCountry>
,
<collectingRegion id="49FEF86AFF9E4B7B6120F8AC1FBB9043" box="[1063,1157,1839,1863]" country="Nepal" name="Gandaki" pageId="13" pageNumber="178">Gandaki</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C3206503FF9E4B7B6638F8DD18EE9F12" pageId="13" pageNumber="178" type="discussion">
<paragraph id="8B853688FF9E4B7B6638F8DD18EE9F12" blockId="13.[831,1436,1886,2070]" pageId="13" pageNumber="178">
<emphasis id="B94EEA9AFF9E4B7B6638F8DD1F1B9072" bold="true" box="[831,1061,1886,1910]" pageId="13" pageNumber="178">Comparative notes:</emphasis>
The new species is a member of the
<taxonomicName id="4C3A4D0BFF9E4B7B6668F8FC1F389092" box="[879,1030,1918,1942]" class="Insecta" family="Staphylinidae" genus="Pselaphodes" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="178" phylum="Arthropoda" rank="species" species="bagmatius">
<emphasis id="B94EEA9AFF9E4B7B6668F8FC18419092" box="[879,895,1919,1942]" italics="true" pageId="13" pageNumber="178">P</emphasis>
.
<emphasis id="B94EEA9AFF9E4B7B6692F8FD1F389092" box="[917,1030,1918,1942]" italics="true" pageId="13" pageNumber="178">bagmatius</emphasis>
</taxonomicName>
species-group, and can be readily recognized based on the unique forms of the male antennal clubs, the distinct large ventral projection of the metacoxae, and the broad apex of the median lobe of the aedeagus.
</paragraph>
</subSubSection>
<caption id="DF456600FF9D4B786590F92A18079011" ID-DOI="http://doi.org/10.5281/zenodo.3702673" ID-Zenodo-Dep="3702673" httpUri="https://zenodo.org/record/3702673/files/figure.png" pageId="14" pageNumber="179" startId="14.[151,183,1705,1727]" targetBox="[155,1411,259,1650]" targetPageId="14">
<paragraph id="8B853688FF9D4B786590F92A18079011" blockId="14.[151,1435,1705,1813]" pageId="14" pageNumber="179">
Fig 11. Male diagnostic features of
<taxonomicName id="4C3A4D0BFF9D4B7864FFF92A199191BA" box="[504,687,1705,1726]" class="Insecta" family="Staphylinidae" genus="Pselaphodes" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="179" phylum="Arthropoda" rank="species" species="loebli" status="sp. nov.">
<emphasis id="B94EEA9AFF9D4B7864FFF92A199191BA" box="[504,687,1705,1726]" italics="true" pageId="14" pageNumber="179">Pselaphodes loebli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27D57E1FF9D4B7867B2F92A183E91BA" box="[693,768,1705,1726]" pageId="14" pageNumber="179" rank="species">sp. nov.</taxonomicNameLabel>
(A) Antennal club. (B) Pronotum. (C) Metaventral process, in lateral view. (D) Protrochanter and profemur. (E) Apex of protibia. (F) Mesotrochanter and mesofemur. (G) Apex of mesotibia. (H) Metatrochanter and metafemur. (I) Sternite IX. (J-L) Aedeagus, in dorsal (J), lateral (K), and ventral (L) view. Scale bars: 0.3 mm in A-B, D, F, H; 0.2 mm in C, J-L; 0.1 mm in E, G, I.
</paragraph>
</caption>
<caption id="DF456600FF9C4B796590F88418079077" ID-DOI="http://doi.org/10.5281/zenodo.3702675" ID-Zenodo-Dep="3702675" httpUri="https://zenodo.org/record/3702675/files/figure.png" pageId="15" pageNumber="180" startId="15.[151,183,1799,1821]" targetBox="[157,1411,261,1744]" targetPageId="15">
<paragraph id="8B853688FF9C4B796590F88418079077" blockId="15.[151,1437,1799,1908]" pageId="15" pageNumber="180">
Fig 12. Male diagnostic features of
<taxonomicName id="4C3A4D0BFF9C4B796705F88419E69019" box="[514,728,1799,1821]" class="Insecta" family="Staphylinidae" genus="Pselaphodes" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="180" phylum="Arthropoda" rank="species" species="procerus" status="sp. nov.">
<emphasis id="B94EEA9AFF9C4B796705F88419E69019" box="[514,728,1799,1821]" italics="true" pageId="15" pageNumber="180">Pselaphodes procerus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27D57E1FF9C4B7967E6F88B18109019" box="[737,814,1800,1821]" pageId="15" pageNumber="180" rank="species">sp. nov.</taxonomicNameLabel>
(A) Antennal club. (B) Pronotum. (C) Metaventral process, in lateral view. (D) Protrochanter and profemur. (E) Apex of protibia. (F) Mesotrochanter and mesofemur. (G) Apex of mesotibia. (H) Metatrochanter and metafemur. (I) Sternite IX. (J-L) Aedeagus, in dorsal (J), lateral (K), and ventral (L) view. Scale bars: 0.3 mm in A-B, D, F, H; 0.2 mm in C, J-L; 0.1 mm in E, G, I.
</paragraph>
</caption>
<subSubSection id="C3206503FF834B666590FE8719B99638" pageId="16" pageNumber="181" type="etymology">
<paragraph id="8B853688FF834B666590FE8719B99638" blockId="16.[151,756,259,316]" pageId="16" pageNumber="181">
<emphasis id="B94EEA9AFF834B666590FE871A259618" bold="true" box="[151,283,260,284]" pageId="16" pageNumber="181">Etymology:</emphasis>
The specific epithet refers to the large blunt projection of the metacoxa of the new species.
</paragraph>
</subSubSection>
</treatment>
</document>