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<document id="06D09CBC4768987E471EEF84722AF249" ID-DOI="10.5281/zenodo.176487" ID-GBIF-Dataset="815f47b1-6767-416f-a0bf-f925b54845fa" ID-ISSN="1175-5326" ID-Zenodo-Dep="176487" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459834238221" checkinUser="plazi" docAuthor="Shear, William A." docDate="2007" docId="0393879DFFF49623FF7C5423FA50C0C1" docLanguage="en" docName="zt01463p012.pdf" docOrigin="Zootaxa 1463" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Idagona lehmanensis Shear, 2007, n. sp." docType="treatment" docVersion="9" lastPageNumber="11" masterDocId="FFAAFFE5FFFC9629FFEB514CFFCCC62B" masterDocTitle="Cave millipeds of the United States. V. The genus Idagona Buckett &amp; Gardner (Chordeumatida, Conotylidae, Idagoninae)" masterLastPageNumber="12" masterPageNumber="1" pageNumber="9" updateTime="1698223281701" updateUser="plazi">
<mods:mods id="C7A82E6853EDA0A88B2AB2B17DB37279" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="AF5ED5B357BA3D21797D6E598E915AF4">Cave millipeds of the United States. V. The genus Idagona Buckett &amp; Gardner (Chordeumatida, Conotylidae, Idagoninae)</mods:title>
</mods:titleInfo>
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<mods:namePart id="EE6755B6FFCD60A1E6402D89E6550731">Shear, William A.</mods:namePart>
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<mods:typeOfResource id="635C473F348E1BFE021665504D8083E1">text</mods:typeOfResource>
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<mods:date id="59DD7202548504023E131BEF7414A1C6">2007</mods:date>
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<mods:number id="A0684AA980ED32856E5EE32CA2CD28E4">1463</mods:number>
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<mods:classification id="D8E209BDE317927A00DC196186918FAD">journal article</mods:classification>
<mods:identifier id="FC529DBAD9380D873E486C041A674A06" type="DOI">10.5281/zenodo.176487</mods:identifier>
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<mods:identifier id="D2D3701B82A55188404B952C32F5AAD9" type="Zenodo-Dep">176487</mods:identifier>
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<treatment id="0393879DFFF49623FF7C5423FA50C0C1" ID-DOI="http://doi.org/10.5281/zenodo.6236713" ID-GBIF-Taxon="119358478" ID-Zenodo-Dep="6236713" LSID="urn:lsid:plazi:treatment:0393879DFFF49623FF7C5423FA50C0C1" httpUri="http://treatment.plazi.org/id/0393879DFFF49623FF7C5423FA50C0C1" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9">
<subSubSection id="C3206500FFF49621FF7C5423FE19C387" pageId="8" pageNumber="9" type="nomenclature">
<paragraph id="8B85368BFFF49621FF7C5423FE20C3A2" blockId="8.[151,492,1391,1452]" box="[151,492,1391,1417]" pageId="8" pageNumber="9">
<heading id="D0CD81E7FFF49621FF7C5423FE20C3A2" bold="true" box="[151,492,1391,1417]" fontSize="11" level="1" pageId="8" pageNumber="9" reason="1">
<emphasis id="B94EEA99FFF49621FF7C5423FE20C3A2" bold="true" box="[151,492,1391,1417]" pageId="8" pageNumber="9">
<taxonomicName id="4C3A4D08FFF49621FF7C5423FE56C3A2" ID-CoL="3PC3Q" box="[151,410,1391,1417]" class="Diplopoda" family="Conotylidae" genus="Idagona" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="lehmanensis" status="sp. nov.">
<emphasis id="B94EEA99FFF49621FF7C5423FE56C3A2" bold="true" box="[151,410,1391,1417]" italics="true" pageId="8" pageNumber="9">Idagona lehmanensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A27D57E2FFF49621FE425423FE20C3A2" box="[425,492,1391,1417]" pageId="8" pageNumber="9" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B85368BFFF49621FF7C54DEFE19C387" blockId="8.[151,492,1391,1452]" box="[151,469,1426,1452]" pageId="8" pageNumber="9">
<figureCitation id="13012A0EFFF49621FF7C54DEFF25C387" box="[151,233,1426,1452]" captionStart="FIGURES 1 – 4" captionStartId="3.[151,269,1923,1947]" captionTargetBox="[151,1435,729,1899]" captionTargetId="figure@3.[151,1436,729,1900]" captionTargetPageId="3" captionText="FIGURES 1 – 4. Gonopods of Idagona species. 1, Idagona westcotti, Giant Arch Cave, posteriorventral view. 2. I. westcotti, Ice Capades Cave, posterioventral view. 3. I. jasperi, n. sp. posterioventral view. 4. I. lehmanenesis, n. sp., posterior view." httpUri="https://zenodo.org/record/176488/files/figure.png" pageId="8" pageNumber="9">Figs. 4</figureCitation>
, 8, 12-14, 18, 21-23
</paragraph>
</subSubSection>
<subSubSection id="C3206500FFF49621FF7C54AEFB57C13F" pageId="8" pageNumber="9" type="description">
<paragraph id="8B85368BFFF49621FF7C54AEFC41C067" blockId="8.[151,1437,1506,2012]" pageId="8" pageNumber="9">
<emphasis id="B94EEA99FFF49621FF7C54AEFF2DC3D0" box="[151,225,1506,1531]" italics="true" pageId="8" pageNumber="9">
<typeStatus id="54818829FFF49621FF7C54AEFF1BC3D0" box="[151,215,1506,1531]" pageId="8" pageNumber="9">Types</typeStatus>
:
</emphasis>
Male
<typeStatus id="54818829FFF49621FED154AEFE68C3D7" box="[314,420,1506,1532]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
, two male
<typeStatus id="54818829FFF49621FDDD54AEFD6AC3D7" box="[566,678,1506,1532]" pageId="8" pageNumber="9" type="paratype">paratypes</typeStatus>
and female
<typeStatus id="54818829FFF49621FCA154AEFC63C3D7" box="[842,943,1506,1532]" pageId="8" pageNumber="9" type="paratype">paratype</typeStatus>
from Water Trough Cave, Great Basin National Park, Nevada, collected
<date id="FF84104BFFF49621FDF55746FD75C00F" box="[542,697,1546,1572]" pageId="8" pageNumber="9" value="2006-05-24">24 May 2006</date>
by S. J. Taylor, J. K. Krejca, M. G. Slay and G.
<collectingCountry id="F32D761BFFF49621FB0B5746FAEAC00F" box="[1248,1318,1546,1572]" name="United States Pacific Island Wildlife Refuges" pageId="8" pageNumber="9">Baker</collectingCountry>
(FMNH). Additional immature specimens were collected at the same time.
</paragraph>
<paragraph id="8B85368BFFF49621FF2D5716FB57C13F" blockId="8.[151,1437,1506,2012]" pageId="8" pageNumber="9">
Diagnosis;
<taxonomicName id="4C3A4D08FFF49621FEA55716FD80C058" box="[334,588,1626,1651]" class="Diplopoda" family="Conotylidae" genus="Idagona" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="lehmanensis">
<emphasis id="B94EEA99FFF49621FEA55716FD80C058" box="[334,588,1626,1651]" italics="true" pageId="8" pageNumber="9">Idagona lehmanensis</emphasis>
</taxonomicName>
is easily distinguished from all other known
<taxonomicName id="4C3A4D08FFF49621FB6F5716FB2AC058" box="[1156,1254,1626,1651]" class="Diplopoda" family="Conotylidae" genus="Idagona" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA99FFF49621FB6F5716FB2AC058" box="[1156,1254,1626,1651]" italics="true" pageId="8" pageNumber="9">Idagona</emphasis>
</taxonomicName>
species by the complicated subterminal branch of the anterior gonopod (
<figureCitation id="13012A0EFFF49621FCA057CEFC11C0B7" box="[843,989,1666,1692]" captionStart="FIGURES 21, 22" captionStartId="9.[151,269,1434,1458]" captionTargetBox="[151,1435,557,1407]" captionTargetId="figure@9.[151,1436,557,1408]" captionTargetPageId="9" captionText="FIGURES 21, 22. Idagona lehmanensis n. sp. 21. Tip of anterior gonopod, ventral view. 22. Tip of anterior gonopod, lateral view." httpUri="https://zenodo.org/record/176492/files/figure.png" pageId="8" pageNumber="9">Figs. 21, 22</figureCitation>
) and the presence of modified, porebearing femora on the fourth, as well as the third, legpair (
<figureCitation id="13012A0EFFF49621FCAE57E6FC50C0EF" box="[837,924,1706,1732]" captionStart="FIGURES 9 – 14" captionStartId="6.[151,269,1722,1746]" captionTargetBox="[151,1436,357,1706]" captionTargetId="figure@6.[151,1436,357,1707]" captionTargetPageId="6" captionText="FIGURES 9 – 14. Pregonopodal leg modifications of Idagona species. 9 – 12, third legs, anterior views. 9. Idagona westcotti, Giant Arch Cave. 10.. I. westcotti, Ice Capades Cave. 11. I. jasperi n. sp. 12. I. lehmanensis n. sp. 13. Leg 4 of I. lehmanensis n. sp., anterior view. 14. Denticles near adenostyle of leg 3 femur of I. lehmanensis n. sp." httpUri="https://zenodo.org/record/176489/files/figure.png" pageId="8" pageNumber="9">Fig. 13</figureCitation>
). The tenth coxae of
<taxonomicName id="4C3A4D08FFF49621FB7757E6FAE1C0E8" box="[1180,1325,1706,1731]" class="Diplopoda" family="Conotylidae" genus="Idagona" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="lehmanensis">
<emphasis id="B94EEA99FFF49621FB7757E6FAE1C0E8" box="[1180,1325,1706,1731]" italics="true" pageId="8" pageNumber="9">lehmanensis</emphasis>
</taxonomicName>
(
<figureCitation id="13012A0EFFF49621FAD557E6FA5FC0EF" box="[1342,1427,1706,1732]" captionStart="FIGURES 15 – 18" captionStartId="7.[151,269,1493,1517]" captionTargetBox="[151,1435,357,1466]" captionTargetId="figure@7.[151,1436,357,1467]" captionTargetPageId="7" captionText="FIGURES 15 – 18. Leg 10 coxae of Idagona species, posterior views. 15.. Idagona westcotti, Giant Arch Cave. 16. I. westcotti, Ice Capades Cave. 17. I. jasperi n. sp. 18. I. lehmanensis n. sp., anterior view." httpUri="https://zenodo.org/record/176490/files/figure.png" pageId="8" pageNumber="9">Fig. 18</figureCitation>
) have much less well-developed apical knobs and are extended beyond the trochanteral joint; the anterior surfaces are poorly sclerotized and the coxal gland openings are frontal rather than ventral.
</paragraph>
</subSubSection>
<subSubSection id="C3206500FFF49620FF2D566EFD3AC009" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="etymology">
<paragraph id="8B85368BFFF49621FF2D566EFB27C14F" blockId="8.[151,1437,1506,2012]" pageId="8" pageNumber="9">
<emphasis id="B94EEA99FFF49621FF2D566EFE8FC110" box="[198,323,1826,1851]" italics="true" pageId="8" pageNumber="9">Etymology</emphasis>
: The species epithet,
<emphasis id="B94EEA99FFF49621FDBA566EFD25C110" box="[593,745,1826,1851]" italics="true" pageId="8" pageNumber="9">
<taxonomicName id="4C3A4D08FFF49621FDBA566EFD29C110" box="[593,741,1826,1851]" class="Diplopoda" family="Conotylidae" genus="Idagona" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="lehmanensis">lehmanensis</taxonomicName>
,
</emphasis>
was suggested by S. J. Taylor and makes reference to the former Lehman Caves National Monument, which became Great Basin National Park in 1986.
</paragraph>
<paragraph id="8B85368BFFF49621FF2D563EFA50C1F7" blockId="8.[151,1437,1506,2012]" pageId="8" pageNumber="9">
Description: Male (
<figureCitation id="13012A0EFFF49621FE42563EFDCCC1A7" box="[425,512,1906,1932]" captionStart="FIGURE 23" captionStartId="10.[151,255,828,852]" captionTargetBox="[594,1002,197,809]" captionTargetId="figure@10.[594,1003,197,809]" captionTargetPageId="10" captionText="FIGURE 23. Idagona lehmanenesis, n. sp. Anterior end of male paratype. Note the swollen femora on both legpairs 3 and 4 and the absence of conotyliform telopodites on the posterior gonopods. Photo by S. J. Taylor." httpUri="https://zenodo.org/record/176493/files/figure.png" pageId="8" pageNumber="9">Fig. 23</figureCitation>
): Length, 13.0 mm, width 1.0 mm. Ocelli 23-27, round, black, in compact oval patch, dorsal row of ocelli somewhat indistinct. Third legpair (
<figureCitation id="13012A0EFFF49621FC6D56D6FC17C19F" box="[902,987,1946,1972]" captionStart="FIGURES 9 – 14" captionStartId="6.[151,269,1722,1746]" captionTargetBox="[151,1436,357,1706]" captionTargetId="figure@6.[151,1436,357,1707]" captionTargetPageId="6" captionText="FIGURES 9 – 14. Pregonopodal leg modifications of Idagona species. 9 – 12, third legs, anterior views. 9. Idagona westcotti, Giant Arch Cave. 10.. I. westcotti, Ice Capades Cave. 11. I. jasperi n. sp. 12. I. lehmanensis n. sp. 13. Leg 4 of I. lehmanensis n. sp., anterior view. 14. Denticles near adenostyle of leg 3 femur of I. lehmanensis n. sp." httpUri="https://zenodo.org/record/176489/files/figure.png" pageId="8" pageNumber="9">Fig. 12</figureCitation>
) with expanded femur bearing adenostyle as low mound; adenostyle subtended on anterior side by cluster of small, cuticular granules (
<figureCitation id="13012A0EFFF49621FADF568EFA42C1F7" box="[1332,1422,1986,2012]" captionStart="FIGURES 9 – 14" captionStartId="6.[151,269,1722,1746]" captionTargetBox="[151,1436,357,1706]" captionTargetId="figure@6.[151,1436,357,1707]" captionTargetPageId="6" captionText="FIGURES 9 – 14. Pregonopodal leg modifications of Idagona species. 9 – 12, third legs, anterior views. 9. Idagona westcotti, Giant Arch Cave. 10.. I. westcotti, Ice Capades Cave. 11. I. jasperi n. sp. 12. I. lehmanensis n. sp. 13. Leg 4 of I. lehmanensis n. sp., anterior view. 14. Denticles near adenostyle of leg 3 femur of I. lehmanensis n. sp." httpUri="https://zenodo.org/record/176489/files/figure.png" pageId="8" pageNumber="9">Fig. 14</figureCitation>
).
</paragraph>
<paragraph id="8B85368BFFF59620FF7C51D4FEC9C7D9" blockId="9.[151,1437,152,498]" pageId="9" pageNumber="10">
Fourth legpair similar (
<figureCitation id="13012A0EFFF59620FE4551D4FDCBC699" box="[430,519,152,178]" captionStart="FIGURES 9 – 14" captionStartId="6.[151,269,1722,1746]" captionTargetBox="[151,1436,357,1706]" captionTargetId="figure@6.[151,1436,357,1707]" captionTargetPageId="6" captionText="FIGURES 9 – 14. Pregonopodal leg modifications of Idagona species. 9 – 12, third legs, anterior views. 9. Idagona westcotti, Giant Arch Cave. 10.. I. westcotti, Ice Capades Cave. 11. I. jasperi n. sp. 12. I. lehmanensis n. sp. 13. Leg 4 of I. lehmanensis n. sp., anterior view. 14. Denticles near adenostyle of leg 3 femur of I. lehmanensis n. sp." httpUri="https://zenodo.org/record/176489/files/figure.png" pageId="9" pageNumber="10">Fig. 13</figureCitation>
). Gonopods (
<figureCitation id="13012A0EFFF59620FD5551D4FCD8C699" box="[702,788,152,178]" captionStart="FIGURES 1 – 4" captionStartId="3.[151,269,1923,1947]" captionTargetBox="[151,1435,729,1899]" captionTargetId="figure@3.[151,1436,729,1900]" captionTargetPageId="3" captionText="FIGURES 1 – 4. Gonopods of Idagona species. 1, Idagona westcotti, Giant Arch Cave, posteriorventral view. 2. I. westcotti, Ice Capades Cave, posterioventral view. 3. I. jasperi, n. sp. posterioventral view. 4. I. lehmanenesis, n. sp., posterior view." httpUri="https://zenodo.org/record/176488/files/figure.png" pageId="9" pageNumber="10">Figs. 4</figureCitation>
, 8, 21, 22): anterior gonopods large, upright, distally expanded, with complex posteriorly located subterminal branch (
<figureCitation id="13012A0EFFF59620FC47518CFB8DC6F1" box="[940,1089,192,218]" captionStart="FIGURES 21, 22" captionStartId="9.[151,269,1434,1458]" captionTargetBox="[151,1435,557,1407]" captionTargetId="figure@9.[151,1436,557,1408]" captionTargetPageId="9" captionText="FIGURES 21, 22. Idagona lehmanensis n. sp. 21. Tip of anterior gonopod, ventral view. 22. Tip of anterior gonopod, lateral view." httpUri="https://zenodo.org/record/176492/files/figure.png" pageId="9" pageNumber="10">Figs. 21. 22</figureCitation>
) bearing numerous cuticular fimbriae, narrow bifid subbranch, and distally expanded subbranch. Apex of gonopod deeply divided, medial division broad, trullate; lateral division narrow, evenly curved. Posterior gonopod coxites with poorly sclerotized sternum, basal gonopodal elements much reduced, coxites relatively small, not scooplike, with small posterior nodule, clusters of microteeth near apex. Tenth coxae (
<figureCitation id="13012A0EFFF59620FC60502CFC13C751" box="[907,991,352,378]" captionStart="FIGURES 15 – 18" captionStartId="7.[151,269,1493,1517]" captionTargetBox="[151,1435,357,1466]" captionTargetId="figure@7.[151,1436,357,1467]" captionTargetPageId="7" captionText="FIGURES 15 – 18. Leg 10 coxae of Idagona species, posterior views. 15.. Idagona westcotti, Giant Arch Cave. 16. I. westcotti, Ice Capades Cave. 17. I. jasperi n. sp. 18. I. lehmanensis n. sp., anterior view." httpUri="https://zenodo.org/record/176490/files/figure.png" pageId="9" pageNumber="10">Fig. 18</figureCitation>
) extended beyond trochanteral articulation, posterior knob low, anterior surface poorly sclerotized, membranous, gland opening on anterior surface. Coloration white to pale tan, anterior segments and head lightly mottled purplish-brown, legs white (
<figureCitation id="13012A0EFFF59620FF745094FF3BC7D9" box="[159,247,472,498]" captionStart="FIGURE 23" captionStartId="10.[151,255,828,852]" captionTargetBox="[594,1002,197,809]" captionTargetId="figure@10.[594,1003,197,809]" captionTargetPageId="10" captionText="FIGURE 23. Idagona lehmanenesis, n. sp. Anterior end of male paratype. Note the swollen femora on both legpairs 3 and 4 and the absence of conotyliform telopodites on the posterior gonopods. Photo by S. J. Taylor." httpUri="https://zenodo.org/record/176493/files/figure.png" pageId="9" pageNumber="10">Fig. 23</figureCitation>
).
</paragraph>
<caption id="DF456603FFF59620FF7C54D6FED6C3FF" httpUri="https://zenodo.org/record/176492/files/figure.png" pageId="9" pageNumber="10" targetBox="[151,1435,557,1407]" targetPageId="9">
<paragraph id="8B85368BFFF59620FF7C54D6FED6C3FF" blockId="9.[151,1436,1434,1492]" pageId="9" pageNumber="10">
<emphasis id="B94EEA99FFF59620FF7C54D6FE91C399" bold="true" box="[151,349,1434,1458]" pageId="9" pageNumber="10">FIGURES 21, 22.</emphasis>
<taxonomicName id="4C3A4D08FFF59620FE8054D6FD82C39A" box="[363,590,1434,1457]" class="Diplopoda" family="Conotylidae" genus="Idagona" kingdom="Animalia" order="Chordeumatida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="lehmanensis" status="sp. nov.">
<emphasis id="B94EEA99FFF59620FE8054D6FD82C39A" box="[363,590,1434,1457]" italics="true" pageId="9" pageNumber="10">Idagona lehmanensis</emphasis>
</taxonomicName>
<emphasis id="B94EEA99FFF59620FDBF54D6FD5CC399" bold="true" box="[596,656,1434,1458]" pageId="9" pageNumber="10">
<taxonomicNameLabel id="A27D57E2FFF59620FDBF54D6FD5CC399" box="[596,656,1434,1458]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
21. Tip of anterior gonopod, ventral view. 22. Tip of anterior gonopod, lateral view.
</paragraph>
</caption>
<paragraph id="8B85368BFFF59620FF2D5744FD3AC009" blockId="9.[151,1437,1544,2010]" box="[198,758,1544,1570]" pageId="9" pageNumber="10">Female similar to male in nonsexual characters.</paragraph>
</subSubSection>
<subSubSection id="C3206500FFF59623FF2D577CFA50C0C1" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10" type="distribution">
<paragraph id="8B85368BFFF59620FF2D577CFDB2C059" blockId="9.[151,1437,1544,2010]" pageId="9" pageNumber="10">
<emphasis id="B94EEA99FFF59620FF2D577CFEACC062" box="[198,352,1584,1609]" italics="true" pageId="9" pageNumber="10">Distribution:</emphasis>
In addition to Water Trough Cave, juvenile idagonines undoubtedly this species were collected in the nearly adjacent Model Cave.
</paragraph>
<paragraph id="8B85368BFFF59620FF2D57CCFD1FC1F1" blockId="9.[151,1437,1544,2010]" pageId="9" pageNumber="10">
<emphasis id="B94EEA99FFF59620FF2D57CCFDDAC0B2" box="[198,534,1664,1689]" italics="true" pageId="9" pageNumber="10">
Notes:
<taxonomicName id="4C3A4D08FFF59620FEF057CCFDDAC0B2" box="[283,534,1664,1689]" class="Diplopoda" family="Conotylidae" genus="Idagona" kingdom="Animalia" order="Chordeumatida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="lehmanensis">Idagona lehmanensis</taxonomicName>
</emphasis>
differs strongly from the other two species and might have been considered the
<typeStatus id="54818829FFF59620FF2957E4FF38C0E9" box="[194,244,1704,1730]" pageId="9" pageNumber="10">type</typeStatus>
of a new genus. However, such designation, if justified, can wait for the discovery of more species of idagonines and a fuller understanding of their relationships. The more complex gonopods of
<taxonomicName id="4C3A4D08FFF59620FB01579CFA59C0C2" box="[1258,1429,1744,1769]" class="Diplopoda" family="Conotylidae" genus="Idagona" kingdom="Animalia" order="Chordeumatida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="lehmanensis">
<emphasis id="B94EEA99FFF59620FB01579CFA59C0C2" box="[1258,1429,1744,1769]" italics="true" pageId="9" pageNumber="10">I. lehmanensis</emphasis>
</taxonomicName>
, when compared to the other two, mirrors a similar situation in the eastern North American genus
<taxonomicName id="4C3A4D08FFF59620FADB57B4FA50C13A" box="[1328,1436,1784,1809]" class="Diplopoda" family="Conotylidae" genus="Conotyla" kingdom="Animalia" order="Chordeumatida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA99FFF59620FADB57B4FA50C13A" box="[1328,1436,1784,1809]" italics="true" pageId="9" pageNumber="10">Conotyla</emphasis>
</taxonomicName>
(
<taxonomicName id="4C3A4D08FFF59620FF74566CFEFFC111" box="[159,307,1824,1850]" class="Diplopoda" family="Conotylidae" kingdom="Animalia" order="Chordeumatida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="family">Conotylidae</taxonomicName>
,
<taxonomicName id="4C3A4D08FFF59620FEA9566CFE1AC111" box="[322,470,1824,1850]" class="Diplopoda" family="Conotylidae" kingdom="Animalia" order="Chordeumatida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="subFamily" subFamily="Conotylinae">Conotylinae</taxonomicName>
), in which species with simpler gonopods are found in the northern part of the generic range, and moving south, one finds gonopods become progressively more complex. The more northerly occurring species of
<taxonomicName id="4C3A4D08FFF59620FE3F563CFD8CC1A2" box="[468,576,1904,1929]" class="Diplopoda" family="Conotylidae" genus="Conotyla" kingdom="Animalia" order="Chordeumatida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA99FFF59620FE3F563CFD8CC1A2" box="[468,576,1904,1929]" italics="true" pageId="9" pageNumber="10">Conotyla</emphasis>
</taxonomicName>
are also more widely distributed; southerly species may have ranges restricted to single ridgetops or summits (
<bibRefCitation id="EFAB4B7AFFF59620FD9556D4FCC5C199" author="Shear" box="[638,777,1944,1970]" pageId="9" pageNumber="12" refString="Shear, W. A. (1971) The milliped family Conotylidae in North America, with a description of the new family Adritylidae (Diplopoda: Chordeumida). Bulletin of the Museum of Comparative Zoology, 141, 55 - 98." type="journal article" year="1971">Shear 1971</bibRefCitation>
). At this point we do not know enough about the diversity or distribution of
<taxonomicName id="4C3A4D08FFF59620FE7D568CFE34C1F2" box="[406,504,1984,2009]" class="Diplopoda" family="Conotylidae" genus="Idagona" kingdom="Animalia" order="Chordeumatida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA99FFF59620FE7D568CFE34C1F2" box="[406,504,1984,2009]" italics="true" pageId="9" pageNumber="10">Idagona</emphasis>
</taxonomicName>
to draw a parallel.
</paragraph>
<caption id="DF456603FFF69623FF7C5270FB76C552" httpUri="https://zenodo.org/record/176493/files/figure.png" pageId="10" pageNumber="11" targetBox="[594,1002,197,809]" targetPageId="10">
<paragraph id="8B85368BFFF69623FF7C5270FB76C552" blockId="10.[151,1435,828,889]" pageId="10" pageNumber="11">
<emphasis id="B94EEA99FFF69623FF7C5270FEEBC57F" bold="true" box="[151,295,828,852]" pageId="10" pageNumber="11">FIGURE 23.</emphasis>
<emphasis id="B94EEA99FFF69623FEC55271FDE8C57F" box="[302,548,829,852]" italics="true" pageId="10" pageNumber="11">
<taxonomicName id="4C3A4D08FFF69623FEC55271FDD3C57F" box="[302,543,829,852]" class="Diplopoda" family="Conotylidae" genus="Idagona" kingdom="Animalia" order="Chordeumatida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="lehmanenesis" status="sp. nov.">Idagona lehmanenesis</taxonomicName>
,
</emphasis>
<emphasis id="B94EEA99FFF69623FDC05270FDABC57F" bold="true" box="[555,615,828,852]" pageId="10" pageNumber="11">
<taxonomicNameLabel id="A27D57E2FFF69623FDC05270FDABC57F" box="[555,615,828,852]" pageId="10" pageNumber="11" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Anterior end of male paratype. Note the swollen femora on both legpairs 3 and 4 and the absence of conotyliform telopodites on the posterior gonopods. Photo by S. J. Taylor.
</paragraph>
</caption>
<paragraph id="8B85368BFFF69623FF2D5294FF1DC371" blockId="10.[151,1436,984,1770]" pageId="10" pageNumber="11">
The caves of this immediate region are closely clustered and many are isolated from one another only by relatively recently formed erosional features.
<taxonomicName id="4C3A4D08FFF69623FD5E554CFC63C232" box="[693,943,1024,1049]" class="Diplopoda" family="Conotylidae" genus="Idagona" kingdom="Animalia" order="Chordeumatida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="lehmanensis">
<emphasis id="B94EEA99FFF69623FD5E554CFC63C232" box="[693,943,1024,1049]" italics="true" pageId="10" pageNumber="11">Idagona lehmanensis</emphasis>
</taxonomicName>
very likely will be found in most or all of these caves. As with
<emphasis id="B94EEA99FFF69623FE7A5564FDD5C26A" box="[401,537,1064,1089]" italics="true" pageId="10" pageNumber="11">
<taxonomicName id="4C3A4D08FFF69623FE7A5564FDD8C26A" box="[401,532,1064,1089]" class="Diplopoda" family="Conotylidae" genus="Idagona" kingdom="Animalia" order="Chordeumatida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="westcotti">I. westcotti</taxonomicName>
,
</emphasis>
the presence of this species, not highly adapted for life underground, in these caves is probably attributable to the latest Pleistocene climatic events: as the Great Basin climate became drier, forested habitats retreated to higher altitudes on the many inselberg ranges (
<bibRefCitation id="EFAB4B7AFFF69623FBBA5534FF1AC291" author="Vandevender" pageId="10" pageNumber="12" refString="Vandevender, R. R. &amp; Spaulding, W. G. (1979) Development of vegetation and climate in the southwestern United States. Science, 204, 701 - 710." type="journal article" year="1979">Vandevender and Spaulding 1979</bibRefCitation>
), but because of the significantly more southerly location of the caves, this may have happened much earlier than in northern Utah and southern Idaho. As with the Snake River lava beds, the present surrounding ecosystem is sagebrush desert and dry juniper woodland—inhospitable for chordeumatid millipeds. Besides
<emphasis id="B94EEA99FFF69623FF7C5454FE85C31A" box="[151,329,1304,1329]" italics="true" pageId="10" pageNumber="11">
<taxonomicName id="4C3A4D08FFF69623FF7C5454FE89C31A" box="[151,325,1304,1329]" class="Diplopoda" family="Conotylidae" genus="Idagona" kingdom="Animalia" order="Chordeumatida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="lehmanensis">I. lehmanensis</taxonomicName>
,
</emphasis>
the two caves also host a new genus and species of polydesmidan milliped, to be described later.
</paragraph>
<paragraph id="8B85368BFFF69623FF2D5424FA50C0C1" blockId="10.[151,1436,984,1770]" pageId="10" pageNumber="11">
Despite its likely isolation in caves, as with its congeners, little special adaptation can be discerned in this species; indeed the number of ocelli present is considerably greater than that in the two northern species. The Great Basin is a region of North
<collectingCountry id="F32D761BFFF69623FDBE54F4FD76C3F9" box="[597,698,1464,1490]" name="United States of America" pageId="10" pageNumber="11">America</collectingCountry>
where crustal extension has resulted in the tipping and downdropping of blocks of strata, producing the characteristic basin-and-range topography. Isolated mountain ranges, many high enough to have forested habitats in the otherwise desertlike or arid grassland terrain, are separated by basins that often have only internal drainage and may harbor short-lived, saline bodies of water. Particularly in eastern Nevada, many of these inselbergs contain karst (Map 1). At higher elevations in Great Basin inselberg ranges, where moist forests occur, relict populations of this or other
<taxonomicName id="4C3A4D08FFF69623FB7E57CCFB3BC0B2" box="[1173,1271,1664,1689]" class="Diplopoda" family="Conotylidae" genus="Idagona" kingdom="Animalia" order="Chordeumatida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA99FFF69623FB7E57CCFB3BC0B2" box="[1173,1271,1664,1689]" italics="true" pageId="10" pageNumber="11">Idagona</emphasis>
</taxonomicName>
species may exist, just as they may in areas of the Rocky Mountains, to the northeast. These habitats in eastern Nevada remain almost entirely unexplored for invertebrate biodiversity, as do the many other karst regions of the state.
</paragraph>
</subSubSection>
</treatment>
</document>
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