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<document id="74BBE9BEC32FAE34B7FD0B271B32003F" ID-DOI="10.5281/zenodo.178835" ID-GBIF-Dataset="726a1c3a-48e9-43d2-bde1-5f5182054db4" ID-ISSN="1175-5326" ID-Zenodo-Dep="178835" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459925681839" checkinUser="plazi" docAuthor="Hernández-Ávila, Iván &amp; Campos, Ernesto" docDate="2007" docId="0393879CFFA9FFDDFF10FA8FFBEAFEF6" docLanguage="en" docName="zt01607p062.pdf" docOrigin="Zootaxa 1607" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Parapinnixa bolagnosi Hernández-Ávila &amp; Campos, 2007, new species" docType="treatment" docVersion="9" lastPageNumber="62" masterDocId="FFAAFFE4FFA8FFD8FF87FF8AFFC4FFF4" masterDocTitle="Parapinnixa bolagnosi, a new species of pinnotherid crab from Cubagua Island and Los Frailes Archipelago, Venezuela (Crustacea: Brachyura: Pinnotheridae)" masterLastPageNumber="62" masterPageNumber="57" pageNumber="58" updateTime="1698229487936" updateUser="plazi">
<mods:mods id="76320DAC070416679867CA5035248800" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="218893F431DA1C80A951877E907C202E">
<mods:title id="6A15BA2699ECF0CD6B86F85395BCA66B">Parapinnixa bolagnosi, a new species of pinnotherid crab from Cubagua Island and Los Frailes Archipelago, Venezuela (Crustacea: Brachyura: Pinnotheridae)</mods:title>
</mods:titleInfo>
<mods:name id="EDF7F0EE72B76E110E5A080A4CE748CA" type="personal">
<mods:role id="2D1B235B8B6F2803AD23C4D16381648F">
<mods:roleTerm id="5D3794307329CA54DA5BB3B94D42C559">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="96B18E125B8228BDE344212E82C8F25E">Hernández-Ávila, Iván</mods:namePart>
</mods:name>
<mods:name id="6CCE9301C49FE1BA69EB4D646DD0A37B" type="personal">
<mods:role id="34C02F95B0BA47CCF8D7108239D42E19">
<mods:roleTerm id="5875D6E7F8A71F77A0D996640EFAF82D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="133334215DE9A24386B31E3734B8C5D4">Campos, Ernesto</mods:namePart>
</mods:name>
<mods:typeOfResource id="DF6A2F1D2264217A83963E5A38AB0444">text</mods:typeOfResource>
<mods:relatedItem id="AE4C799340A9957DA0AD09357D680B26" type="host">
<mods:titleInfo id="BB0DDABCDB7360C8433293FD845CAB87">
<mods:title id="6CD7975E1DDD901553C4CC91DCFB49F5">Zootaxa</mods:title>
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<mods:part id="43DE44BA9149EB3E88F53FE7E2D02F75">
<mods:date id="3F230CC4E1283CB4425B6EBB5B2CB36C">2007</mods:date>
<mods:detail id="81C0EA9F1FFD4FBA0E81926A928E5EA8" type="volume">
<mods:number id="3FDCAE50E45647C50B8F4038D1032E16">1607</mods:number>
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<mods:classification id="04E1C64F84785A5B3B513277FCD4687C">journal article</mods:classification>
<mods:identifier id="F71C208A15B182B0431B634648530FEB" type="DOI">10.5281/zenodo.178835</mods:identifier>
<mods:identifier id="9763936B91DA037DD0A9A83894636CEC" type="GBIF-Dataset">726a1c3a-48e9-43d2-bde1-5f5182054db4</mods:identifier>
<mods:identifier id="C4DAAB62057F1A9121394E81FEDA174F" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="9339DFD5567BA5FA4A629154628D8020" type="Zenodo-Dep">178835</mods:identifier>
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<treatment id="0393879CFFA9FFDDFF10FA8FFBEAFEF6" ID-DOI="http://doi.org/10.5281/zenodo.6247443" ID-GBIF-Taxon="119374268" ID-Zenodo-Dep="6247443" LSID="urn:lsid:plazi:treatment:0393879CFFA9FFDDFF10FA8FFBEAFEF6" httpUri="http://treatment.plazi.org/id/0393879CFFA9FFDDFF10FA8FFBEAFEF6" lastPageId="5" lastPageNumber="62" pageId="1" pageNumber="58">
<subSubSection id="C3206501FFA9FFD9FF10FA8FFE04FAB6" pageId="1" pageNumber="58" type="nomenclature">
<paragraph id="8B85368AFFA9FFD9FF10FA8FFD86FAEB" blockId="1.[151,578,1285,1346]" box="[151,578,1285,1311]" pageId="1" pageNumber="58">
<heading id="D0CD81E6FFA9FFD9FF10FA8FFD86FAEB" bold="true" box="[151,578,1285,1311]" fontSize="11" level="1" pageId="1" pageNumber="58" reason="1">
<emphasis id="B94EEA98FFA9FFD9FF10FA8FFD86FAEB" bold="true" box="[151,578,1285,1311]" pageId="1" pageNumber="58">
<taxonomicName id="4C3A4D09FFA9FFD9FF10FA8FFE6EFAEB" ID-CoL="75QYB" box="[151,426,1285,1311]" class="Malacostraca" family="Pinnotheridae" genus="Parapinnixa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="58" phylum="Arthropoda" rank="species" species="bolagnosi" status="sp. nov.">
<emphasis id="B94EEA98FFA9FFD9FF10FA8FFE6EFAEB" bold="true" box="[151,426,1285,1311]" italics="true" pageId="1" pageNumber="58">Parapinnixa bolagnosi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27D57E3FFA9FFD9FE35FA8FFD86FAEB" box="[434,578,1285,1311]" pageId="1" pageNumber="58" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B85368AFFA9FFD9FED4FAA2FE04FAB6" blockId="1.[151,578,1285,1346]" box="[339,448,1320,1346]" pageId="1" pageNumber="58">
<figureCitation id="13012A0FFFA9FFD9FED4FAA2FE04FAB6" box="[339,448,1320,1346]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[151,255,1919,1943]" captionStartId-1="3.[151,255,1427,1451]" captionTargetBox-0="[190,1411,323,1861]" captionTargetBox-1="[213,1305,277,1400]" captionTargetId-0="figure@2.[151,1436,317,1892]" captionTargetId-1="figure@3.[213,1374,194,1404]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1. Parapinnixa bolagnosi new species. a, Holotype, dorsal view; b g, Paratype; b, cheliped, outer surface; c, cheliped, inner surface; d g, walking legs 1 4; b c setae removed." captionText-1="FIGURE 2. Parapinnixa bolagnosi new species. a, Holotype; frontal view; b-d, Paratype, b, antenae, c, abdomen; d, third maxilliped." httpUri-0="https://zenodo.org/record/178836/files/figure.png" httpUri-1="https://zenodo.org/record/178837/files/figure.png" pageId="1" pageNumber="58">Figs. 12</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3206501FFA9FFD9FF10FAFDFE10F886" pageId="1" pageNumber="58" type="materials_examined">
<paragraph id="8B85368AFFA9FFD9FF10FAFDFE10F886" blockId="1.[151,1436,1399,2026]" pageId="1" pageNumber="58">
<emphasis id="B94EEA98FFA9FFD9FF10FAFDFE41FA65" bold="true" box="[151,389,1399,1425]" pageId="1" pageNumber="58">Material examined</emphasis>
. Cubagua Is.,
<collectingCountry id="F32D761AFFA9FFD9FDB1FAF2FD76FA66" box="[566,690,1400,1426]" name="Venezuela" pageId="1" pageNumber="58">Venezuela</collectingCountry>
.—
<typeStatus id="54818828FFA9FFD9FD5BFAF2FC8AFA66" box="[732,846,1400,1426]" pageId="1" pageNumber="58" type="holotype">Holotype</typeStatus>
: MMM.cr.03252, female, (CL= 1.88 CW= 3.68), Punta Yirú,
<geoCoordinate id="EE0E504DFFA9FFD9FEA2FA2AFE05FA4E" box="[293,449,1440,1466]" direction="north" orientation="latitude" pageId="1" pageNumber="58" precision="1" value="10.806917">10º48'24.9''N</geoCoordinate>
<geoCoordinate id="EE0E504DFFA9FFD9FE4EFA2AFD99FA4E" box="[457,605,1440,1466]" direction="west" orientation="longitude" pageId="1" pageNumber="58" precision="1" value="-64.1512">64º9'4.32''W</geoCoordinate>
, on rocks on
<taxonomicName id="4C3A4D09FFA9FFD9FD7BFA2AFCAAFA4D" box="[764,878,1440,1465]" class="Liliopsida" family="Hydrocharitaceae" genus="Thalassia" kingdom="Animalia" order="Alismatales" pageId="1" pageNumber="58" phylum="Tracheophyta" rank="genus">
<emphasis id="B94EEA98FFA9FFD9FD7BFA2AFCAAFA4D" box="[764,878,1440,1465]" italics="true" pageId="1" pageNumber="58">Thalassia</emphasis>
</taxonomicName>
bed.,
<quantity id="4CC29B6FFFA9FFD9FC30FA2AFC3FFA4E" box="[951,1019,1440,1466]" metricMagnitude="0" metricUnit="m" metricValue="1.7" pageId="1" pageNumber="58" unit="m" value="1.7">1.7 m</quantity>
depth,
<quantity id="4CC29B6FFFA9FFD9FBD4FA2AFB4BFA4E" box="[1107,1167,1440,1466]" metricMagnitude="1" metricUnit="m" metricValue="7.5" pageId="1" pageNumber="58" unit="m" value="75.0">75 m</quantity>
from the coast, coll. I. Hernández-Ávila &amp; A. Gómez,
<date id="FF84104AFFA9FFD9FD8AFA42FD3CFA16" box="[525,760,1480,1506]" pageId="1" pageNumber="58" value="2003-11-11">November 11, 2003</date>
.
<typeStatus id="54818828FFA9FFD9FC83FA42FCBEFA16" box="[772,890,1480,1506]" pageId="1" pageNumber="58" type="paratype">Paratypes</typeStatus>
: Cubagua Is.,
<collectingCountry id="F32D761AFFA9FFD9FB98FA42FB5EFA16" box="[1055,1178,1480,1506]" name="Venezuela" pageId="1" pageNumber="58">Venezuela</collectingCountry>
.— MMM.cr.03251,
<specimenCount id="9D3CFD03FFA9FFD9FA0AFA42FF2EF9FE" pageId="1" pageNumber="58" type="female">1 female</specimenCount>
, (CL= 1.40 CW= 2.84), Playa El Medio,
<geoCoordinate id="EE0E504DFFA9FFD9FD48FA7AFCAFF9FE" box="[719,875,1520,1546]" direction="north" orientation="latitude" pageId="1" pageNumber="58" precision="1" value="10.837417">10º50'14.7''N</geoCoordinate>
<geoCoordinate id="EE0E504DFFA9FFD9FCF4FA7AFBD1F9FE" box="[883,1045,1520,1546]" direction="west" orientation="longitude" pageId="1" pageNumber="58" precision="1" value="-64.15414">64º9'14.88''W</geoCoordinate>
, associated with
<taxonomicName id="4C3A4D09FFA9FFD9FB5BFA7AFF72F9C5" class="Bryopsidophyceae" family="Halimedaceae" genus="Halimeda" higherTaxonomySource="GBIF" kingdom="Plantae" order="Bryopsidales" pageId="1" pageNumber="58" phylum="Chlorophyta" rank="species" species="opuntia">
<emphasis id="B94EEA98FFA9FFD9FB5BFA7AFF72F9C5" italics="true" pageId="1" pageNumber="58">Halimeda opuntia</emphasis>
</taxonomicName>
,
<quantity id="4CC29B6FFFA9FFD9FF41F992FF30F9C6" box="[198,244,1560,1586]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="58" unit="m" value="3.0">3 m</quantity>
depth,
<quantity id="4CC29B6FFFA9FFD9FEC9F992FE5EF9C6" box="[334,410,1560,1586]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="58" unit="m" value="100.0">100 m</quantity>
from the coast, coll. I. Hernández-Ávila,
<date id="FF84104AFFA9FFD9FC0BF992FB91F9C6" box="[908,1109,1560,1586]" pageId="1" pageNumber="58" value="2003-08-27">August 27, 2003</date>
. MMM.cr.03253,
<specimenCount id="9D3CFD03FFA9FFD9FAAAF992FA5CF9C6" box="[1325,1432,1560,1586]" pageId="1" pageNumber="58" type="female">1 female</specimenCount>
, (CL= 1.78 CW= 3.40), Playa El Guichere,
<geoCoordinate id="EE0E504DFFA9FFD9FD12F9CAFC84F9AE" box="[661,832,1600,1626]" direction="north" orientation="latitude" pageId="1" pageNumber="58" precision="1" value="10.804184">10º48'15.06''N</geoCoordinate>
<geoCoordinate id="EE0E504DFFA9FFD9FCCFF9CAFC2FF9AE" box="[840,1003,1600,1626]" direction="west" orientation="longitude" pageId="1" pageNumber="58" precision="1" value="-64.16037">64º9'37.32''W</geoCoordinate>
, on rocks,
<quantity id="4CC29B6FFFA9FFD9FBEBF9CAFB5EF9AE" box="[1132,1178,1600,1626]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="58" unit="m" value="2.0">2 m</quantity>
depth,
<quantity id="4CC29B6FFFA9FFD9FB75F9CAFAEBF9AE" box="[1266,1327,1600,1626]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="58" unit="m" value="50.0">50 m</quantity>
from the coast, coll. I. Hernández-Ávila &amp; A. Gómez,
<date id="FF84104AFFA9FFD9FD34F9E2FC54F976" box="[691,912,1640,1666]" pageId="1" pageNumber="58" value="2003-12-08">December 8, 2003</date>
. MMM.cr. 0 3254,
<specimenCount id="9D3CFD03FFA9FFD9FBF7F9E2FB1EF976" box="[1136,1242,1640,1666]" pageId="1" pageNumber="58" type="female">1 female</specimenCount>
, (CL= 1.0 CW= 2.14), Barranca Blanca,
<geoCoordinate id="EE0E504DFFA9FFD9FE45F91AFD99F95E" box="[450,605,1680,1706]" direction="north" orientation="latitude" pageId="1" pageNumber="58" precision="1" value="10.801384">10º48'4.98''N</geoCoordinate>
<geoCoordinate id="EE0E504DFFA9FFD9FDEDF91AFCD8F95E" box="[618,796,1680,1706]" direction="west" orientation="longitude" pageId="1" pageNumber="58" precision="1" value="-64.1738">64º10'25.68''W</geoCoordinate>
, on rocks,
<quantity id="4CC29B6FFFA9FFD9FC2FF91AFC1EF95E" box="[936,986,1680,1706]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="58" unit="m" value="2.0">2 m</quantity>
depth,
<quantity id="4CC29B6FFFA9FFD9FBBDF91AFBBFF95E" box="[1082,1147,1680,1706]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="58" unit="m" value="50.0">50 m</quantity>
from the coast, coll. I. Hernández-Ávila &amp; A. Gómez,
<date id="FF84104AFFA9FFD9FD8AF932FD1DF926" box="[525,729,1720,1746]" pageId="1" pageNumber="58" value="2004-01-22">January 22, 2004</date>
. MMM.cr.03255,
<specimenCount id="9D3CFD03FFA9FFD9FC28F932FBDCF926" box="[943,1048,1720,1746]" pageId="1" pageNumber="58" type="female">1 female</specimenCount>
, (CL= 1.72 CW= 3.44), Punta El Olivito,
<geoCoordinate id="EE0E504DFFA9FFD9FF7DF96AFE51F90E" box="[250,405,1760,1786]" direction="north" orientation="latitude" pageId="1" pageNumber="58" precision="1" value="10.80095">10º48'3.42''N</geoCoordinate>
<geoCoordinate id="EE0E504DFFA9FFD9FE19F96AFD84F90E" box="[414,576,1760,1786]" direction="west" orientation="longitude" pageId="1" pageNumber="58" precision="1" value="-64.18792">64º11'16.5''W</geoCoordinate>
, on rocks,
<quantity id="4CC29B6FFFA9FFD9FD45F96AFCC2F90E" box="[706,774,1760,1786]" metricMagnitude="0" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="58" unit="m" value="2.5">2.5 m</quantity>
depth,
<quantity id="4CC29B6FFFA9FFD9FCD8F96AFC58F90E" box="[863,924,1760,1786]" metricMagnitude="1" metricUnit="m" metricValue="7.5" pageId="1" pageNumber="58" unit="m" value="75.0">75 m</quantity>
from the coast, coll. I. Hernández-Ávila &amp; A. Gómez,
<date id="FF84104AFFA9FFD9FEAEF882FDD4F8D6" box="[297,528,1800,1826]" pageId="1" pageNumber="58" value="2004-02-19">February 19, 2004</date>
. Los Frailes Archipelago,
<collectingCountry id="F32D761AFFA9FFD9FCDCF882FC13F8D6" box="[859,983,1800,1826]" name="Venezuela" pageId="1" pageNumber="58">Venezuela</collectingCountry>
.— MMM.cr. 0 3662,
<specimenCount id="9D3CFD03FFA9FFD9FB66F882FA89F8D6" box="[1249,1357,1800,1826]" pageId="1" pageNumber="58" type="female">1 female</specimenCount>
(fragmented), (CL= 2.36 CW= 4.32), La Pecha,
<geoCoordinate id="EE0E504DFFA9FFD9FD25F8BAFD3BF8BE" box="[674,767,1840,1866]" direction="north" orientation="latitude" pageId="1" pageNumber="58" precision="925" value="11.1">11º06'N</geoCoordinate>
<geoCoordinate id="EE0E504DFFA9FFD9FC8FF8BAFCA9F8BE" box="[776,877,1840,1866]" direction="west" orientation="longitude" pageId="1" pageNumber="58" precision="925" value="-63.8">63º48'W</geoCoordinate>
(approx.),
<quantity id="4CC29B6FFFA9FFD9FC74F8BAFBE5F8BE" box="[1011,1057,1840,1866]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="58" unit="m" value="8.0">8 m</quantity>
depth between oysters, coll. A. Tagliafico, February, 2004.
</paragraph>
</subSubSection>
<subSubSection id="C3206501FFA9FFDCFF42F8F5FEC9FD4E" lastPageId="4" lastPageNumber="61" pageId="1" pageNumber="58" type="description">
<paragraph id="8B85368AFFA9FFDAFF42F8F5FEE1FEF6" blockId="1.[151,1436,1399,2026]" lastBlockId="2.[151,1436,152,258]" lastPageId="2" lastPageNumber="59" pageId="1" pageNumber="58">
<emphasis id="B94EEA98FFA9FFD9FF42F8F5FE92F86D" bold="true" box="[197,342,1919,1945]" pageId="1" pageNumber="58">Description</emphasis>
. Carapace (
<figureCitation id="13012A0FFFA9FFD9FE63F80AFDEDF86E" box="[484,553,1920,1946]" captionStart="FIGURE 1" captionStartId="2.[151,255,1919,1943]" captionTargetBox="[190,1411,323,1861]" captionTargetId="figure@2.[151,1436,317,1892]" captionTargetPageId="2" captionText="FIGURE 1. Parapinnixa bolagnosi new species. a, Holotype, dorsal view; b g, Paratype; b, cheliped, outer surface; c, cheliped, inner surface; d g, walking legs 1 4; b c setae removed." httpUri="https://zenodo.org/record/178836/files/figure.png" pageId="1" pageNumber="58">Fig. 1</figureCitation>
a), very small, ovate, smooth, width 1.832.02 times length, dorsally convex; anterior margin somewhat straight, laterally arcuated, unarmed; front-orbital width 1/3 carapace width, front triangular, deflexed, tip hidden dorsally. Orbits rounded, eyes small, dorsally visible. Antennules folded in oblique fossettes (
<figureCitation id="13012A0FFFAAFFDAFE0CFF12FE17FF46" box="[395,467,152,178]" captionStart="FIGURE 2" captionStartId="3.[151,255,1427,1451]" captionTargetBox="[213,1305,277,1400]" captionTargetId="figure@3.[213,1374,194,1404]" captionTargetPageId="3" captionText="FIGURE 2. Parapinnixa bolagnosi new species. a, Holotype; frontal view; b-d, Paratype, b, antenae, c, abdomen; d, third maxilliped." httpUri="https://zenodo.org/record/178837/files/figure.png" pageId="2" pageNumber="59">Fig. 2</figureCitation>
a). Epistome (
<figureCitation id="13012A0FFFAAFFDAFDF0FF12FD7AFF46" box="[631,702,152,178]" captionStart="FIGURE 2" captionStartId="3.[151,255,1427,1451]" captionTargetBox="[213,1305,277,1400]" captionTargetId="figure@3.[213,1374,194,1404]" captionTargetPageId="3" captionText="FIGURE 2. Parapinnixa bolagnosi new species. a, Holotype; frontal view; b-d, Paratype, b, antenae, c, abdomen; d, third maxilliped." httpUri="https://zenodo.org/record/178837/files/figure.png" pageId="2" pageNumber="59">Fig. 2</figureCitation>
a) with biconcave dorsal margin, medially pointed, wider than ventral concave margin, lateral margins obliquely curved. Buccal area subtriangular, basal breadth 1/4 carapace length.
</paragraph>
<caption id="DF456602FFAAFFDAFF10F8F5FC9AF84D" httpUri="https://zenodo.org/record/178836/files/figure.png" pageId="2" pageNumber="59" targetBox="[190,1411,323,1861]" targetPageId="2">
<paragraph id="8B85368AFFAAFFDAFF10F8F5FC9AF84D" blockId="2.[151,1436,1919,1977]" pageId="2" pageNumber="59">
<emphasis id="B94EEA98FFAAFFDAFF10F8F5FEDEF863" bold="true" box="[151,282,1919,1943]" pageId="2" pageNumber="59">FIGURE 1.</emphasis>
<taxonomicName id="4C3A4D09FFAAFFDAFEA5F8F5FDDCF862" box="[290,536,1919,1942]" class="Malacostraca" family="Pinnotheridae" genus="Parapinnixa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="bolagnosi" status="sp. nov.">
<emphasis id="B94EEA98FFAAFFDAFEA5F8F5FDDCF862" box="[290,536,1919,1942]" italics="true" pageId="2" pageNumber="59">Parapinnixa bolagnosi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27D57E3FFAAFFDAFDA7F8F5FD60F862" box="[544,676,1919,1942]" pageId="2" pageNumber="59" rank="species">new species</taxonomicNameLabel>
. a, Holotype, dorsal view; bg, Paratype; b, cheliped, outer surface; c, cheliped, inner surface; dg, walking legs 14; bc setae removed.
</paragraph>
</caption>
<caption id="DF456602FFABFFDBFF10FA19FE8DFA39" httpUri="https://zenodo.org/record/178837/files/figure.png" pageId="3" pageNumber="60" targetBox="[213,1305,277,1400]" targetPageId="3">
<paragraph id="8B85368AFFABFFDBFF10FA19FE8DFA39" blockId="3.[151,1436,1427,1485]" pageId="3" pageNumber="60">
<emphasis id="B94EEA98FFABFFDBFF10FA19FEDFFA5F" bold="true" box="[151,283,1427,1451]" pageId="3" pageNumber="60">FIGURE 2.</emphasis>
<taxonomicName id="4C3A4D09FFABFFDBFEA3FA1EFDD8FA5F" box="[292,540,1428,1451]" class="Malacostraca" family="Pinnotheridae" genus="Parapinnixa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="60" phylum="Arthropoda" rank="species" species="bolagnosi" status="sp. nov.">
<emphasis id="B94EEA98FFABFFDBFEA3FA1EFDD8FA5F" box="[292,540,1428,1451]" italics="true" pageId="3" pageNumber="60">Parapinnixa bolagnosi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27D57E3FFABFFDBFDA2FA1EFD6FFA5F" box="[549,683,1428,1451]" pageId="3" pageNumber="60" rank="species">new species</taxonomicNameLabel>
. a, Holotype; frontal view; b-d, Paratype, b, antenae, c, abdomen; d, third maxilliped.
</paragraph>
</caption>
<paragraph id="8B85368AFFABFFDBFF42FA76FB6EF97A" blockId="3.[151,1437,1532,2038]" pageId="3" pageNumber="60">
MXP3 (
<figureCitation id="13012A0FFFABFFDBFEA3FA76FEAFF9E2" box="[292,363,1532,1558]" captionStart="FIGURE 2" captionStartId="3.[151,255,1427,1451]" captionTargetBox="[213,1305,277,1400]" captionTargetId="figure@3.[213,1374,194,1404]" captionTargetPageId="3" captionText="FIGURE 2. Parapinnixa bolagnosi new species. a, Holotype; frontal view; b-d, Paratype, b, antenae, c, abdomen; d, third maxilliped." httpUri="https://zenodo.org/record/178837/files/figure.png" pageId="3" pageNumber="60">Fig. 2</figureCitation>
d) subtrapezoidal, merus broadly subtrapezoidal, fused with ischium, distal margin slightly sinuous, dorsal margin slightly concave, with large plumose setae, ventral margin broadly convex, outer surface with short setae. Palp with two articles not extending beyond distal margin of merus, dactylus absent, carpus subtriangular, with large plumose setae; propodus minute, digitiform, exopod absent.
</paragraph>
<paragraph id="8B85368AFFABFFDBFF42F916FE89F852" blockId="3.[151,1437,1532,2038]" pageId="3" pageNumber="60">
Chelipeds (
<figureCitation id="13012A0FFFABFFDBFEC9F916FE57F942" box="[334,403,1692,1718]" captionStart="FIGURE 1" captionStartId="2.[151,255,1919,1943]" captionTargetBox="[190,1411,323,1861]" captionTargetId="figure@2.[151,1436,317,1892]" captionTargetPageId="2" captionText="FIGURE 1. Parapinnixa bolagnosi new species. a, Holotype, dorsal view; b g, Paratype; b, cheliped, outer surface; c, cheliped, inner surface; d g, walking legs 1 4; b c setae removed." httpUri="https://zenodo.org/record/178836/files/figure.png" pageId="3" pageNumber="60">Fig. 1</figureCitation>
b) symmetrical, stout, contour rounded, nearly as long as WL 1, merus short, lateroexternal surface subtrapezoidal, flattened; carpus rounded, merus and carpus with short setae on inner margin. Chelae (
<figureCitation id="13012A0FFFABFFDBFF7CF966FE84F8F2" box="[251,320,1772,1798]" captionStart="FIGURE 1" captionStartId="2.[151,255,1919,1943]" captionTargetBox="[190,1411,323,1861]" captionTargetId="figure@2.[151,1436,317,1892]" captionTargetPageId="2" captionText="FIGURE 1. Parapinnixa bolagnosi new species. a, Holotype, dorsal view; b g, Paratype; b, cheliped, outer surface; c, cheliped, inner surface; d g, walking legs 1 4; b c setae removed." httpUri="https://zenodo.org/record/178836/files/figure.png" pageId="3" pageNumber="60">Fig. 1</figureCitation>
c) tomentose, length of palm subequal to height, longer than fingers, margins little concave, with minute granules on outer surface, mesiodorsal row of short setae on inner surface. Fingers subtriangular, curving at tip where they cross, base of the pollex wider than dactylus, cutting surface of pollex with small, medial serrate crest that ends in subtruncated tooth; cutting surface of dactylus smooth, gently curved, dorsal margin convex.
</paragraph>
<paragraph id="8B85368AFFABFFDCFF42F83EFDCDFEF6" blockId="3.[151,1437,1532,2038]" lastBlockId="4.[151,1436,152,698]" lastPageId="4" lastPageNumber="61" pageId="3" pageNumber="60">
WL (
<figureCitation id="13012A0FFFABFFDBFE84F83EFE90F83A" box="[259,340,1972,1998]" captionStart="FIGURE 1" captionStartId="2.[151,255,1919,1943]" captionTargetBox="[190,1411,323,1861]" captionTargetId="figure@2.[151,1436,317,1892]" captionTargetPageId="2" captionText="FIGURE 1. Parapinnixa bolagnosi new species. a, Holotype, dorsal view; b g, Paratype; b, cheliped, outer surface; c, cheliped, inner surface; d g, walking legs 1 4; b c setae removed." httpUri="https://zenodo.org/record/178836/files/figure.png" pageId="3" pageNumber="60">Figs. 1</figureCitation>
dg) decreasing in order from the largest WL1 to the smallest WL4; merus of WL1 (
<figureCitation id="13012A0FFFABFFDBFAB9F83EFA44F83A" box="[1342,1408,1972,1998]" captionStart="FIGURE 1" captionStartId="2.[151,255,1919,1943]" captionTargetBox="[190,1411,323,1861]" captionTargetId="figure@2.[151,1436,317,1892]" captionTargetPageId="2" captionText="FIGURE 1. Parapinnixa bolagnosi new species. a, Holotype, dorsal view; b g, Paratype; b, cheliped, outer surface; c, cheliped, inner surface; d g, walking legs 1 4; b c setae removed." httpUri="https://zenodo.org/record/178836/files/figure.png" pageId="3" pageNumber="60">Fig. 1</figureCitation>
d) widening distally, inner surface proximally concave to accommodate the ischium-merus of cheliped, dorsal ridge setosed, distal width around 0.6 times length; carpus short, stout, dorsally subelliptical and convex, propodus tapering distally, subelliptical, ventral margin with short simple setae, dactylus small, subconical, somewhat falcate, tip corneous.
</paragraph>
<paragraph id="8B85368AFFACFFDCFF42FE9AFA83FE8E" blockId="4.[151,1436,152,698]" pageId="4" pageNumber="61">
WL2 (
<figureCitation id="13012A0FFFACFFDCFE95FE9AFE91FEDE" box="[274,341,272,298]" captionStart="FIGURE 1" captionStartId="2.[151,255,1919,1943]" captionTargetBox="[190,1411,323,1861]" captionTargetId="figure@2.[151,1436,317,1892]" captionTargetPageId="2" captionText="FIGURE 1. Parapinnixa bolagnosi new species. a, Holotype, dorsal view; b g, Paratype; b, cheliped, outer surface; c, cheliped, inner surface; d g, walking legs 1 4; b c setae removed." httpUri="https://zenodo.org/record/178836/files/figure.png" pageId="4" pageNumber="61">Fig. 1</figureCitation>
e) compressed, merus little shorter but much slender than merus of WL1, greatest width 0.4 times its length; carpus longer than wide, subtriangular, propodus subrectangular, dorsal margin gently curved, ventral margin concave, ending in, a distal lobe, dactylus thin, somewhat falcate, tip corneous.
</paragraph>
<paragraph id="8B85368AFFACFFDCFF42FE02FB04FE3E" blockId="4.[151,1436,152,698]" pageId="4" pageNumber="61">
WL3 (
<figureCitation id="13012A0FFFACFFDCFE92FE02FE9EFE56" box="[277,346,392,418]" captionStart="FIGURE 1" captionStartId="2.[151,255,1919,1943]" captionTargetBox="[190,1411,323,1861]" captionTargetId="figure@2.[151,1436,317,1892]" captionTargetPageId="2" captionText="FIGURE 1. Parapinnixa bolagnosi new species. a, Holotype, dorsal view; b g, Paratype; b, cheliped, outer surface; c, cheliped, inner surface; d g, walking legs 1 4; b c setae removed." httpUri="https://zenodo.org/record/178836/files/figure.png" pageId="4" pageNumber="61">Fig. 1</figureCitation>
f) smaller that WL2, merus width almost 0.5 times its length; carpus subtriangular, much shorter than subrectangular propodus, dactylus subconical, somewhat falcate, tip corneous.
</paragraph>
<paragraph id="8B85368AFFACFFDCFF42FE52FBB5FDEE" blockId="4.[151,1436,152,698]" pageId="4" pageNumber="61">
WL4 (
<figureCitation id="13012A0FFFACFFDCFE96FE52FE96FE06" box="[273,338,472,498]" captionStart="FIGURE 1" captionStartId="2.[151,255,1919,1943]" captionTargetBox="[190,1411,323,1861]" captionTargetId="figure@2.[151,1436,317,1892]" captionTargetPageId="2" captionText="FIGURE 1. Parapinnixa bolagnosi new species. a, Holotype, dorsal view; b g, Paratype; b, cheliped, outer surface; c, cheliped, inner surface; d g, walking legs 1 4; b c setae removed." httpUri="https://zenodo.org/record/178836/files/figure.png" pageId="4" pageNumber="61">
Fig.
<quantity id="4CC29B6FFFACFFDCFEC0FE52FE96FE06" box="[327,338,472,498]" metricMagnitude="-3" metricUnit="kg" metricValue="1.0" pageId="4" pageNumber="61" unit="g" value="1.0">1</quantity>
</figureCitation>
g) smallest, reaching distal margin of merus of WL3; merus width around 0.5 times its length; carpus, propodus and dactylus similar in shape that WL3 but proportionally smaller.
</paragraph>
<paragraph id="8B85368AFFACFFDCFF42FDA2FEC9FD4E" blockId="4.[151,1436,152,698]" pageId="4" pageNumber="61">
Abdomen (
<figureCitation id="13012A0FFFACFFDCFED0FDA2FE5BFDB6" box="[343,415,552,578]" captionStart="FIGURE 2" captionStartId="3.[151,255,1427,1451]" captionTargetBox="[213,1305,277,1400]" captionTargetId="figure@3.[213,1374,194,1404]" captionTargetPageId="3" captionText="FIGURE 2. Parapinnixa bolagnosi new species. a, Holotype; frontal view; b-d, Paratype, b, antenae, c, abdomen; d, third maxilliped." httpUri="https://zenodo.org/record/178837/files/figure.png" pageId="4" pageNumber="61">Fig. 2</figureCitation>
c) with six free segments and telson; third somite the widest; lateral margin convex; somites 46 tapering distally, lateral margins slightly convex; lateral margins of somites 26 with plumose setae; telson slightly more that twice as wide as long; subtriangular, with broad rounded corners, margin with long setae
</paragraph>
</subSubSection>
<caption id="DF456602FFACFFDCFF10FA53FE80F9E7" httpUri="https://zenodo.org/record/178838/files/figure.png" pageId="4" pageNumber="61" targetBox="[390,1197,761,1496]" targetPageId="4">
<paragraph id="8B85368AFFACFFDCFF10FA53FE80F9E7" blockId="4.[151,1436,1497,1555]" pageId="4" pageNumber="61">
<emphasis id="B94EEA98FFACFFDCFF10FA53FEDFFA05" bold="true" box="[151,283,1497,1521]" pageId="4" pageNumber="61">FIGURE 3.</emphasis>
Third maxilliped of
<taxonomicName id="4C3A4D09FFACFFDCFD84FA50FB68FA04" authority="Werding and Muller 1990" authorityName="Werding and Muller" authorityYear="1990" box="[515,1196,1497,1521]" class="Malacostraca" family="Pinnotheridae" genus="Parapinnixa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="beaufortensis">
<emphasis id="B94EEA98FFACFFDCFD84FA50FD4CFA05" box="[515,648,1498,1521]" italics="true" pageId="4" pageNumber="61">Parapinnixa</emphasis>
cf.
<emphasis id="B94EEA98FFACFFDCFD32FA50FC81FA05" box="[693,837,1498,1521]" italics="true" pageId="4" pageNumber="61">beaufortensis</emphasis>
sensu Werding and Müller 1990
</taxonomicName>
(not,
<taxonomicName id="4C3A4D09FFACFFDCFB77FA50FEF3F9E7" authority="Rathbun, 1918" authorityName="Rathbun" authorityYear="1918" class="Malacostraca" family="Pinnotheridae" genus="Parapinnixa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="beaufortensis">
<emphasis id="B94EEA98FFACFFDCFB77FA50FA58FA05" box="[1264,1436,1498,1521]" italics="true" pageId="4" pageNumber="61">P. beaufortensis</emphasis>
Rathbun, 1918
</taxonomicName>
).
</paragraph>
</caption>
<subSubSection id="C3206501FFACFFDCFF42F9E4FB0FF92D" pageId="4" pageNumber="61" type="etymology">
<paragraph id="8B85368AFFACFFDCFF42F9E4FB0FF92D" blockId="4.[151,1437,1646,2033]" pageId="4" pageNumber="61">
<emphasis id="B94EEA98FFACFFDCFF42F9E4FE8EF97C" bold="true" box="[197,330,1646,1672]" pageId="4" pageNumber="61">Etymology</emphasis>
. This species is named in honor to Juan Antonio Bolaños Curvelo, Professor of Carcinology of the Escuela de Ciencias Aplicadas del Mar, Universidad de Oriente,
<collectingCountry id="F32D761AFFACFFDCFC5DF91DFB91F945" box="[986,1109,1687,1713]" name="Venezuela" pageId="4" pageNumber="61">Venezuela</collectingCountry>
, for his contributions to our knowledge of the larval development and taxonomy of decapods in northeastern
<collectingCountry id="F32D761AFFACFFDCFBCCF935FB03F92D" box="[1099,1223,1727,1753]" name="Venezuela" pageId="4" pageNumber="61">Venezuela</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3206501FFACFFDDFF42F96CFBEAFEF6" lastPageId="5" lastPageNumber="62" pageId="4" pageNumber="61" type="discussion">
<paragraph id="8B85368AFFACFFDCFF42F96CFCE8F805" blockId="4.[151,1437,1646,2033]" pageId="4" pageNumber="61">
<emphasis id="B94EEA98FFACFFDCFF42F96CFEF3F8F4" bold="true" box="[197,311,1766,1792]" pageId="4" pageNumber="61">Remarks</emphasis>
.
<bibRefCitation id="EFAB4B7BFFACFFDCFECDF96DFD41F8F5" author="Werding" box="[330,645,1767,1793]" pageId="4" pageNumber="62" refString="Werding, B., &amp; H. G. Muller (1990) Die Gattung Parapinnixa Holmes, 1894 and der Nordkuste Kolumbiens, mit beschreibung von Parapinnixa magadalenensis n. sp. (Crustacea: Decapoda: Pinnotheridae). Senckenbergiana Biologica, 70, 221 - 227." type="journal article" year="1990">Werding &amp; Müller (1990)</bibRefCitation>
reported, with reservations, a male specimen as
<taxonomicName id="4C3A4D09FFACFFDCFB67F96DFE34F8DD" authority="Rathbun, 1918" authorityName="Rathbun" authorityYear="1918" class="Malacostraca" family="Pinnotheridae" genus="Parapinnixa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="beaufortensis">
<emphasis id="B94EEA98FFACFFDCFB67F96DFAB7F8F4" box="[1248,1395,1767,1792]" italics="true" pageId="4" pageNumber="61">Parapinnixa</emphasis>
cf.
<emphasis id="B94EEA98FFACFFDCFF10F885FEF1F8DC" box="[151,309,1807,1832]" italics="true" pageId="4" pageNumber="61">beaufortensis</emphasis>
<bibRefCitation id="EFAB4B7BFFACFFDCFEB9F885FE34F8DD" author="Rathbun" box="[318,496,1807,1833]" pageId="4" pageNumber="62" refString="Rathbun, M. J. (1918) The grapsoid crabs of America. United States National Museum Bulletin, 97, 1 - 461." type="journal article" year="1918">Rathbun, 1918</bibRefCitation>
</taxonomicName>
, which was collected in Bahía Chengue, near Santa
<collectingCountry id="F32D761AFFACFFDCFBECF885FB70F8DD" box="[1131,1204,1807,1833]" name="Malta" pageId="4" pageNumber="61">Marta</collectingCountry>
,
<collectingCountry id="F32D761AFFACFFDCFB45F885FAFDF8DD" box="[1218,1337,1807,1833]" name="Colombia" pageId="4" pageNumber="61">Colombia</collectingCountry>
. After a careful examination of Werding &amp; Müllers figure of the MXP3 (
<figureCitation id="13012A0FFFACFFDCFC0BF8BDFC16F8A5" box="[908,978,1847,1873]" captionStart="FIGURE 3" captionStartId="4.[151,255,1497,1521]" captionTargetBox="[390,1197,761,1496]" captionTargetId="figure@4.[378,1209,754,1555]" captionTargetPageId="4" captionText="FIGURE 3. Third maxilliped of Parapinnixa cf. beaufortensis sensu Werding and Müller 1990 (not, P. beaufortensis Rathbun, 1918)." httpUri="https://zenodo.org/record/178838/files/figure.png" pageId="4" pageNumber="61">Fig. 3</figureCitation>
), their specimen appears to have closer affinities with
<taxonomicName id="4C3A4D09FFACFFDCFEC7F8D5FE0BF88C" box="[320,463,1887,1912]" class="Malacostraca" family="Pinnotheridae" genus="Parapinnixa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="bolagnosi">
<emphasis id="B94EEA98FFACFFDCFEC7F8D5FE0BF88C" box="[320,463,1887,1912]" italics="true" pageId="4" pageNumber="61">P. bolagnosi</emphasis>
</taxonomicName>
since both share a biarticulated palp and lack of exopod. The palp of the third maxilliped of
<taxonomicName id="4C3A4D09FFACFFDCFE8EF80DFD2CF855" authority="Rathbun, 1918" authorityName="Rathbun" authorityYear="1918" box="[265,744,1927,1953]" class="Malacostraca" family="Pinnotheridae" genus="Parapinixa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="beaufortensis">
<emphasis id="B94EEA98FFACFFDCFE8EF80DFDF7F854" box="[265,563,1927,1952]" italics="true" pageId="4" pageNumber="61">Parapinixa beaufortensis</emphasis>
<bibRefCitation id="EFAB4B7BFFACFFDCFDBCF80DFD2CF855" author="Rathbun" box="[571,744,1927,1953]" pageId="4" pageNumber="62" refString="Rathbun, M. J. (1918) The grapsoid crabs of America. United States National Museum Bulletin, 97, 1 - 461." type="journal article" year="1918">Rathbun, 1918</bibRefCitation>
</taxonomicName>
has three articles. Since no males of
<taxonomicName id="4C3A4D09FFACFFDCFB1AF80DFAEBF854" box="[1181,1327,1927,1952]" class="Malacostraca" family="Pinnotheridae" genus="Parapinnixa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="bolagnosi">
<emphasis id="B94EEA98FFACFFDCFB1AF80DFAEBF854" box="[1181,1327,1927,1952]" italics="true" pageId="4" pageNumber="61">P. bolagnosi</emphasis>
</taxonomicName>
are presently available for study it is not possible to determine whether or not Werding &amp; Müllers Colombian specimen is conspecific or if it represents a new species.
</paragraph>
<paragraph id="8B85368AFFADFFDDFF42FF1DFBEAFEF6" blockId="5.[151,1436,151,258]" pageId="5" pageNumber="62">
<emphasis id="B94EEA98FFADFFDDFF42FF1DFE73FF45" bold="true" box="[197,439,151,177]" pageId="5" pageNumber="62">Ecological remarks</emphasis>
.
<taxonomicName id="4C3A4D09FFADFFDDFE40FF12FD12FF45" box="[455,726,152,177]" class="Malacostraca" family="Pinnotheridae" genus="Parapinnixa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="62" phylum="Arthropoda" rank="species" species="bolagnosi">
<emphasis id="B94EEA98FFADFFDDFE40FF12FD12FF45" box="[455,726,152,177]" italics="true" pageId="5" pageNumber="62">Parapinnixa bolagnosi</emphasis>
</taxonomicName>
has been collected associated with rocks in
<taxonomicName id="4C3A4D09FFADFFDDFB63FF12FA92FF45" box="[1252,1366,152,177]" class="Liliopsida" family="Hydrocharitaceae" genus="Thalassia" kingdom="Animalia" order="Alismatales" pageId="5" pageNumber="62" phylum="Tracheophyta" rank="genus">
<emphasis id="B94EEA98FFADFFDDFB63FF12FA92FF45" box="[1252,1366,152,177]" italics="true" pageId="5" pageNumber="62">Thalassia</emphasis>
</taxonomicName>
beds, on aggregations of the calcareous algae
<taxonomicName id="4C3A4D09FFADFFDDFD0CFF4AFCA0FF2D" box="[651,868,192,217]" class="Bryopsidophyceae" family="Halimedaceae" genus="Halimeda" higherTaxonomySource="GBIF" kingdom="Plantae" order="Bryopsidales" pageId="5" pageNumber="62" phylum="Chlorophyta" rank="species" species="opuntia">
<emphasis id="B94EEA98FFADFFDDFD0CFF4AFCA0FF2D" box="[651,868,192,217]" italics="true" pageId="5" pageNumber="62">Halimeda opuntia</emphasis>
</taxonomicName>
, between oysters, in Cubagua Island and Los Frailes Archipelago. One lost specimen was collected in the sponge
<taxonomicName id="4C3A4D09FFADFFDDFC30FF62FBC1FEF5" box="[951,1029,232,257]" class="Demospongiae" family="Irciniidae" genus="Ircinia" kingdom="Animalia" order="Dictyoceratida" pageId="5" pageNumber="62" phylum="Porifera" rank="genus">
<emphasis id="B94EEA98FFADFFDDFC30FF62FBC1FEF5" box="[951,1029,232,257]" italics="true" pageId="5" pageNumber="62">Ircinia</emphasis>
</taxonomicName>
sp.
</paragraph>
</subSubSection>
</treatment>
</document>