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<mods:title id="B41680EDE5B0F601B0877E294FDA2016">Prochelator tupuhi sp. nov., the first record of Desmosomatidae Sars, 1897 (Crustacea: Isopoda) from New Zealand waters *</mods:title>
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<mods:namePart id="E6C24D2B43F16C0F67617264BAD5035C">Brix, Saskia</mods:namePart>
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<mods:namePart id="7FCC8E2C2B17581D7CC5BB9E8C184FF1">Bruce, Niel L.</mods:namePart>
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<mods:date id="AFFEC2CD12A50734674DDA050C785A66">2008</mods:date>
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<subSubSection id="C3206505FFD3FFC0FF43F924762D4F4A" pageId="1" pageNumber="483" type="nomenclature">
<paragraph id="8B85368EFFD3FFC0FF43F92475944EDC" blockId="1.[151,464,1690,1716]" box="[151,464,1690,1716]" pageId="1" pageNumber="483">
<heading id="D0CD81E2FFD3FFC0FF43F92475944EDC" bold="true" box="[151,464,1690,1716]" fontSize="11" level="1" pageId="1" pageNumber="483" reason="1">
<taxonomicName id="4C3A4D0DFFD3FFC0FF43F92475944EDC" ID-CoL="7PVCW" authority="Hessler, 1970" authorityName="Hessler" authorityYear="1970" box="[151,464,1690,1716]" class="Malacostraca" family="Desmosomatidae" genus="Prochelator" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="483" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA9CFFD3FFC0FF43F92475944EDC" bold="true" box="[151,464,1690,1716]" pageId="1" pageNumber="483">
<emphasis id="B94EEA9CFFD3FFC0FF43F92475624EDC" bold="true" box="[151,294,1690,1716]" italics="true" pageId="1" pageNumber="483">Prochelator</emphasis>
<bibRefCitation id="EFAB4B7FFFD3FFC0FEF9F92475944EDC" author="Hessler" box="[301,464,1690,1716]" pageId="1" pageNumber="492" refString="Hessler, R. R. (1970) The Desmosomatidae (Isopoda, Asellota) of the Gay Head-Bermuda Transect. Bulletin of the Scripps Institution of Oceanography, 15, 1 - 185." type="journal article" year="1970">Hessler, 1970</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B85368EFFD3FFC0FF43F95E762D4F4A" blockId="1.[151,1438,1760,2026]" pageId="1" pageNumber="483">
<emphasis id="B94EEA9CFFD3FFC0FF43F95E75534E91" box="[151,279,1760,1785]" italics="true" pageId="1" pageNumber="483">Synonymy:</emphasis>
<taxonomicName id="4C3A4D0DFFD3FFC0FECAF95E763B4E92" authority="Hessler, 1970: 27" authorityName="Hessler" authorityPageNumber="27" authorityYear="1970" box="[286,639,1760,1786]" class="Malacostraca" family="Desmosomatidae" genus="Prochelator" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="483" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA9CFFD3FFC0FECAF95E75EF4E91" box="[286,427,1760,1785]" italics="true" pageId="1" pageNumber="483">Prochelator</emphasis>
<bibRefCitation id="EFAB4B7FFFD3FFC0FE66F95E76124E92" author="Hessler" box="[434,598,1760,1786]" pageId="1" pageNumber="492" refString="Hessler, R. R. (1970) The Desmosomatidae (Isopoda, Asellota) of the Gay Head-Bermuda Transect. Bulletin of the Scripps Institution of Oceanography, 15, 1 - 185." type="journal article" year="1970">Hessler, 1970</bibRefCitation>
: 27
</taxonomicName>
28;
<bibRefCitation id="EFAB4B7FFFD3FFC0FD6EF95E77224E92" author="Mezhov" box="[698,870,1760,1786]" pageId="1" pageNumber="492" refString="Mezhov, B. V. (1986) Bathyal and abyssal Nannoniscidae and Desmosomatidae (Isopoda, Asellota) from Alaska Bay. Archives of Zoological Museum Moscow State University, 24, 126 - 167." type="journal article" year="1986">Mezhov, 1986</bibRefCitation>
: 139;
<bibRefCitation id="EFAB4B7FFFD3FFC0FC78F95E70224E92" author="Kussakin" box="[940,1126,1760,1786]" pageId="1" pageNumber="492" refString="Kussakin, O. G. (1999) Marine and salt-water Asellota (Isopoda) of the cold and temperate Waters of the northern hemisphere (in Russian). Vol. III. Suborder Asellota. Part 2. Nauka, Leningrad - AH SSSR, 383 pp." type="book" year="1999">Kussakin, 1999</bibRefCitation>
: 278;
<bibRefCitation id="EFAB4B7FFFD3FFC0FB79F95E71144E92" author="George" box="[1197,1360,1760,1786]" pageId="1" pageNumber="492" refString="George, R. Y. (2001) Desmosomatidae and Nannoniscidae (Crustacea, Isopoda, Asellota) from bathyal and abyssal depth off north Carolina and their evolution. Journal of Natural History, 35 No. 12, 1831 - 1859." type="journal article" year="2001">George, 2001</bibRefCitation>
: 1840;
<bibRefCitation id="EFAB4B7FFFD3FFC0FF43F8B6766F4F4A" author="Brenke" box="[151,555,1800,1826]" pageId="1" pageNumber="492" refString="Brenke, N., Brix, S. &amp; Knuschke, T. (2005) A new deep-sea isopod species from the Angola Basin: Prochelator angolensis sp. nov. (Asellota: Desmosomatidae). Organisms Diversity &amp; Evolution, 5, 179 - 188." type="journal article" year="2005">Brenke, Brix and Knuschke, 2005</bibRefCitation>
: 180.
</paragraph>
</subSubSection>
<subSubSection id="C3206505FFD3FFC0FF12F88E74B64F82" pageId="1" pageNumber="483" type="materials_examined">
<paragraph id="8B85368EFFD3FFC0FF12F88E70344F22" blockId="1.[151,1438,1760,2026]" box="[198,1136,1840,1866]" pageId="1" pageNumber="483">
<emphasis id="B94EEA9CFFD3FFC0FF12F88E75274F21" box="[198,355,1840,1865]" italics="true" pageId="1" pageNumber="483">
<typeStatus id="5481882CFFD3FFC0FF12F88E74B94F21" box="[198,253,1840,1865]" pageId="1" pageNumber="483">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="4C3A4D0DFFD3FFC0FEBEF88E76FA4F22" authority="Sars, 1899" authorityName="Sars" authorityYear="1899" box="[362,702,1840,1866]" class="Malacostraca" family="Desmosomatidae" genus="Eugerda" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="483" phylum="Arthropoda" rank="species" species="lateralis">
<emphasis id="B94EEA9CFFD3FFC0FEBEF88E76734F21" box="[362,567,1840,1865]" italics="true" pageId="1" pageNumber="483">Eugerda lateralis</emphasis>
<bibRefCitation id="EFAB4B7FFFD3FFC0FDEAF88E76FA4F22" author="Sars" box="[574,702,1840,1866]" pageId="1" pageNumber="492" refString="Sars, G. O. (1899) An account of the Crustacea of Norway. Volume 2, parts 13 - 14. Isopoda. Bergen 270 pp." type="book chapter" year="1899">Sars, 1899</bibRefCitation>
</taxonomicName>
; original designation
<bibRefCitation id="EFAB4B7FFFD3FFC0FC6FF88E70284F22" author="Hessler" box="[955,1132,1840,1866]" pageId="1" pageNumber="492" refString="Hessler, R. R. (1970) The Desmosomatidae (Isopoda, Asellota) of the Gay Head-Bermuda Transect. Bulletin of the Scripps Institution of Oceanography, 15, 1 - 185." type="journal article" year="1970">Hessler (1970)</bibRefCitation>
.
</paragraph>
<paragraph id="8B85368EFFD3FFC0FF12F8E674B64F82" blockId="1.[151,1438,1760,2026]" pageId="1" pageNumber="483">
<emphasis id="B94EEA9CFFD3FFC0FF12F8E6764B4F19" box="[198,527,1880,1905]" italics="true" pageId="1" pageNumber="483">
Composition:
<taxonomicName id="4C3A4D0DFFD3FFC0FEA5F8E6764B4F19" authority="Brenke, Brix &amp; Knuschke, 2005" authorityName="Brenke, Brix &amp; Knuschke" authorityYear="2005" box="[369,527,1880,1905]" class="Malacostraca" family="Desmosomatidae" genus="Prochelator" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="483" phylum="Arthropoda" rank="species" species="angolensis">P. angolensis</taxonomicName>
</emphasis>
<bibRefCitation id="EFAB4B7FFFD3FFC0FDCFF8E677EF4F1A" author="Brenke" box="[539,939,1880,1906]" pageId="1" pageNumber="492" refString="Brenke, N., Brix, S. &amp; Knuschke, T. (2005) A new deep-sea isopod species from the Angola Basin: Prochelator angolensis sp. nov. (Asellota: Desmosomatidae). Organisms Diversity &amp; Evolution, 5, 179 - 188." type="journal article" year="2005">Brenke, Brix &amp; Knuschke, 2005</bibRefCitation>
;
<taxonomicName id="4C3A4D0DFFD3FFC0FC6EF8E670BD4F1A" authority="Hessler, 1970" authorityName="Hessler" authorityYear="1970" box="[954,1273,1880,1906]" class="Malacostraca" family="Desmosomatidae" genus="Prochelator" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="483" phylum="Arthropoda" rank="species" species="abyssalis">
<emphasis id="B94EEA9CFFD3FFC0FC6EF8E670034F19" box="[954,1095,1880,1905]" italics="true" pageId="1" pageNumber="483">P. abyssalis</emphasis>
<bibRefCitation id="EFAB4B7FFFD3FFC0FB86F8E670BD4F1A" author="Hessler" box="[1106,1273,1880,1906]" pageId="1" pageNumber="492" refString="Hessler, R. R. (1970) The Desmosomatidae (Isopoda, Asellota) of the Gay Head-Bermuda Transect. Bulletin of the Scripps Institution of Oceanography, 15, 1 - 185." type="journal article" year="1970">Hessler, 1970</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C3A4D0DFFD3FFC0FADDF8E6757F4FF2" authority="Hessler, 1970" authorityName="Hessler" authorityYear="1970" class="Malacostraca" family="Desmosomatidae" genus="Prochelator" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="483" phylum="Arthropoda" rank="species" species="hampsoni">
<emphasis id="B94EEA9CFFD3FFC0FADDF8E671D84F19" box="[1289,1436,1880,1905]" italics="true" pageId="1" pageNumber="483">P. hampsoni</emphasis>
<bibRefCitation id="EFAB4B7FFFD3FFC0FF43F83E757F4FF2" author="Hessler" box="[151,315,1920,1946]" pageId="1" pageNumber="492" refString="Hessler, R. R. (1970) The Desmosomatidae (Isopoda, Asellota) of the Gay Head-Bermuda Transect. Bulletin of the Scripps Institution of Oceanography, 15, 1 - 185." type="journal article" year="1970">Hessler, 1970</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C3A4D0DFFD3FFC0FE9DF83E76D94FF2" authority="Hessler, 1970" authorityName="Hessler" authorityYear="1970" box="[329,669,1920,1946]" class="Malacostraca" family="Desmosomatidae" genus="Prochelator" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="483" phylum="Arthropoda" rank="species" species="incomitatus">
<emphasis id="B94EEA9CFFD3FFC0FE9DF83E75AB4FF1" box="[329,495,1920,1945]" italics="true" pageId="1" pageNumber="483">P. incomitatus</emphasis>
<bibRefCitation id="EFAB4B7FFFD3FFC0FE2CF83E76D94FF2" author="Hessler" box="[504,669,1920,1946]" pageId="1" pageNumber="492" refString="Hessler, R. R. (1970) The Desmosomatidae (Isopoda, Asellota) of the Gay Head-Bermuda Transect. Bulletin of the Scripps Institution of Oceanography, 15, 1 - 185." type="journal article" year="1970">Hessler, 1970</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C3A4D0DFFD3FFC0FD7EF83E77834FF2" authority="Sars, 1899" authorityName="Sars" authorityYear="1899" box="[682,967,1920,1946]" class="Malacostraca" family="Desmosomatidae" genus="Eugerda" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="483" phylum="Arthropoda" rank="species" species="lateralis">
<emphasis id="B94EEA9CFFD3FFC0FD7EF83E776E4FF1" box="[682,810,1920,1945]" italics="true" pageId="1" pageNumber="483">P. lateralis</emphasis>
(
<bibRefCitation id="EFAB4B7FFFD3FFC0FCEFF83E77FA4FF2" author="Sars" box="[827,958,1920,1946]" pageId="1" pageNumber="492" refString="Sars, G. O. (1899) An account of the Crustacea of Norway. Volume 2, parts 13 - 14. Isopoda. Bergen 270 pp." type="book chapter" year="1899">Sars, 1899</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="4C3A4D0DFFD3FFC0FC00F83E70964FF2" authority="Hessler, 1970" authorityName="Hessler" authorityYear="1970" box="[980,1234,1920,1946]" class="Malacostraca" family="Desmosomatidae" genus="Prochelator" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="483" phylum="Arthropoda" rank="species" species="litus">
<emphasis id="B94EEA9CFFD3FFC0FC00F83E70614FF1" box="[980,1061,1920,1945]" italics="true" pageId="1" pageNumber="483">P. litus</emphasis>
<bibRefCitation id="EFAB4B7FFFD3FFC0FBF9F83E70964FF2" author="Hessler" box="[1069,1234,1920,1946]" pageId="1" pageNumber="492" refString="Hessler, R. R. (1970) The Desmosomatidae (Isopoda, Asellota) of the Gay Head-Bermuda Transect. Bulletin of the Scripps Institution of Oceanography, 15, 1 - 185." type="journal article" year="1970">Hessler, 1970</bibRefCitation>
</taxonomicName>
;
<emphasis id="B94EEA9CFFD3FFC0FB0BF83E71774FF1" box="[1247,1331,1920,1945]" italics="true" pageId="1" pageNumber="483">P. s a r s i</emphasis>
<bibRefCitation id="EFAB4B7FFFD3FFC0FAEBF83E74914FAA" author="George" pageId="1" pageNumber="492" refString="George, R. Y. (2001) Desmosomatidae and Nannoniscidae (Crustacea, Isopoda, Asellota) from bathyal and abyssal depth off north Carolina and their evolution. Journal of Natural History, 35 No. 12, 1831 - 1859." type="journal article" year="2001">George, 2001</bibRefCitation>
;
<taxonomicName id="4C3A4D0DFFD3FFC0FF36F816764E4FAA" authority="Hessler, 1970" authorityName="Hessler" authorityYear="1970" box="[226,522,1960,1986]" class="Malacostraca" family="Desmosomatidae" genus="Prochelator" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="483" phylum="Arthropoda" rank="species" species="uncatus">
<emphasis id="B94EEA9CFFD3FFC0FF36F816751F4FA9" box="[226,347,1960,1985]" italics="true" pageId="1" pageNumber="483">P. uncatus</emphasis>
<bibRefCitation id="EFAB4B7FFFD3FFC0FEB1F816764E4FAA" author="Hessler" box="[357,522,1960,1986]" pageId="1" pageNumber="492" refString="Hessler, R. R. (1970) The Desmosomatidae (Isopoda, Asellota) of the Gay Head-Bermuda Transect. Bulletin of the Scripps Institution of Oceanography, 15, 1 - 185." type="journal article" year="1970">Hessler, 1970</bibRefCitation>
</taxonomicName>
;
<emphasis id="B94EEA9CFFD3FFC0FDC3F81676DB4FA9" box="[535,671,1960,1985]" italics="true" pageId="1" pageNumber="483">P. k u s s a k i n i</emphasis>
<bibRefCitation id="EFAB4B7FFFD3FFC0FD78F816771C4FAA" author="Mezhov" box="[684,856,1960,1986]" pageId="1" pageNumber="492" refString="Mezhov, B. V. (1986) Bathyal and abyssal Nannoniscidae and Desmosomatidae (Isopoda, Asellota) from Alaska Bay. Archives of Zoological Museum Moscow State University, 24, 126 - 167." type="journal article" year="1986">Mezhov, 1986</bibRefCitation>
;
<taxonomicName id="4C3A4D0DFFD3FFC0FCB1F81671614FAA" authority="Fresi &amp; Schiecke, 1969" authorityName="Fresi &amp; Schiecke" authorityYear="1969" box="[869,1317,1960,1986]" class="Malacostraca" family="Desmosomatidae" genus="Prochelator" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="483" phylum="Arthropoda" rank="species" species="serratum">
<emphasis id="B94EEA9CFFD3FFC0FCB1F81677A94FA9" box="[869,1005,1960,1985]" italics="true" pageId="1" pageNumber="483">P. serratum</emphasis>
(
<bibRefCitation id="EFAB4B7FFFD3FFC0FC2AF81671584FAA" author="Fresi" box="[1022,1308,1960,1986]" pageId="1" pageNumber="492" refString="Fresi, E. &amp; Schiecke, U. (1969) Two new desmosomatids from the Gulf of Naples: Desmosoma serratum n. sp. and Desmosoma thoracicum n. sp. (Isopoda, Parasellidae). Crustaceana, 17 Part 2, 159 - 170." type="journal article" year="1969">Fresi &amp; Schiecke, 1969</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="4C3A4D0DFFD3FFC0FAE7F81671D84FA9" box="[1331,1436,1960,1985]" class="Malacostraca" family="Desmosomatidae" genus="Prochelator" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="483" phylum="Arthropoda" rank="species" species="tupuhi" status="sp. nov.">
<emphasis id="B94EEA9CFFD3FFC0FAE7F81671D84FA9" box="[1331,1436,1960,1985]" italics="true" pageId="1" pageNumber="483">P. tupuhi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27D57E7FFD3FFC0FF43F86E74B64F82" box="[151,242,2000,2026]" pageId="1" pageNumber="483" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C3206505FFD0FFC2FF12FF26756F4A72" lastPageId="3" lastPageNumber="485" pageId="2" pageNumber="484" type="diagnosis">
<paragraph id="8B85368EFFD0FFC3FF12FF2670C9496A" blockId="2.[151,1438,152,2018]" pageId="2" pageNumber="484">
<emphasis id="B94EEA9CFFD0FFC3FF12FF26750348D9" box="[198,327,152,177]" italics="true" pageId="2" pageNumber="484">Diagnosis:</emphasis>
Pereopod 1 large, carpo­chelate, dactylus and propodus forming movable counterpart to large spine­like claw­seta on distal end of carpus. Inferior margin of carpus of pereopod 1 with one midventral seta and a slender seta distally proximal to claw­seta. Pleotelson with posterolateral spines.
</paragraph>
<paragraph id="8B85368EFFD0FFC3FF12FEAE76654B32" blockId="2.[151,1438,152,2018]" pageId="2" pageNumber="484">
<emphasis id="B94EEA9CFFD0FFC3FF12FEAE75C84941" box="[198,396,272,297]" italics="true" pageId="2" pageNumber="484">Generic remarks</emphasis>
: The genus
<taxonomicName id="4C3A4D0DFFD0FFC3FDC9FEAE76EE4941" box="[541,682,272,297]" class="Malacostraca" family="Desmosomatidae" genus="Prochelator" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="484" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA9CFFD0FFC3FDC9FEAE76EE4941" box="[541,682,272,297]" italics="true" pageId="2" pageNumber="484">Prochelator</emphasis>
</taxonomicName>
was erected by
<bibRefCitation id="EFAB4B7FFFD0FFC3FCBEFEAE70534942" author="Hessler" box="[874,1047,272,298]" pageId="2" pageNumber="492" refString="Hessler, R. R. (1970) The Desmosomatidae (Isopoda, Asellota) of the Gay Head-Bermuda Transect. Bulletin of the Scripps Institution of Oceanography, 15, 1 - 185." type="journal article" year="1970">Hessler (1970)</bibRefCitation>
who transferred the
<typeStatus id="5481882CFFD0FFC3FAD9FEAE71044942" box="[1293,1344,272,298]" pageId="2" pageNumber="484">type</typeStatus>
species from
<taxonomicName id="4C3A4D0DFFD0FFC3FF0EFE86751F4939" box="[218,347,312,337]" class="Malacostraca" family="Desmosomatidae" genus="Eugerdella" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="484" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA9CFFD0FFC3FF0EFE86751F4939" box="[218,347,312,337]" italics="true" pageId="2" pageNumber="484">Eugerdella</emphasis>
</taxonomicName>
to
<taxonomicName id="4C3A4D0DFFD0FFC3FE53FE8676574939" box="[391,531,312,337]" class="Malacostraca" family="Desmosomatidae" genus="Prochelator" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="484" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA9CFFD0FFC3FE53FE8676574939" box="[391,531,312,337]" italics="true" pageId="2" pageNumber="484">Prochelator</emphasis>
</taxonomicName>
and described five new species of
<taxonomicName id="4C3A4D0DFFD0FFC3FC10FE86700B4939" box="[964,1103,312,337]" class="Malacostraca" family="Desmosomatidae" genus="Prochelator" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="484" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA9CFFD0FFC3FC10FE86700B4939" box="[964,1103,312,337]" italics="true" pageId="2" pageNumber="484">Prochelator</emphasis>
</taxonomicName>
.
<taxonomicName id="4C3A4D0DFFD0FFC3FBB4FE8671204939" box="[1120,1380,312,337]" class="Malacostraca" family="Desmosomatidae" genus="Desmosoma" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="484" phylum="Arthropoda" rank="species" species="serratum">
<emphasis id="B94EEA9CFFD0FFC3FBB4FE8671204939" box="[1120,1380,312,337]" italics="true" pageId="2" pageNumber="484">Desmosoma serratum</emphasis>
</taxonomicName>
was recently transferred to
<taxonomicName id="4C3A4D0DFFD0FFC3FE71FEDE76764911" box="[421,562,352,377]" class="Malacostraca" family="Desmosomatidae" genus="Prochelator" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="484" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA9CFFD0FFC3FE71FEDE76764911" box="[421,562,352,377]" italics="true" pageId="2" pageNumber="484">Prochelator</emphasis>
</taxonomicName>
by Brenke
<emphasis id="B94EEA9CFFD0FFC3FD6AFEDE76B74911" box="[702,755,352,377]" italics="true" pageId="2" pageNumber="484">et al</emphasis>
. (2005). Five publications have subsequently defined the genus (
<bibRefCitation id="EFAB4B7FFFD0FFC3FF3FFE3675CE49CA" author="Hessler" box="[235,394,392,418]" pageId="2" pageNumber="492" refString="Hessler, R. R. (1970) The Desmosomatidae (Isopoda, Asellota) of the Gay Head-Bermuda Transect. Bulletin of the Scripps Institution of Oceanography, 15, 1 - 185." type="journal article" year="1970">Hessler 1970</bibRefCitation>
,
<bibRefCitation id="EFAB4B7FFFD0FFC3FE43FE36767849CA" author="Mezhov" box="[407,572,392,418]" pageId="2" pageNumber="492" refString="Mezhov, B. V. (1986) Bathyal and abyssal Nannoniscidae and Desmosomatidae (Isopoda, Asellota) from Alaska Bay. Archives of Zoological Museum Moscow State University, 24, 126 - 167." type="journal article" year="1986">Mezhov 1986</bibRefCitation>
,
<bibRefCitation id="EFAB4B7FFFD0FFC3FD9DFE3676BF49CA" author="Kussakin" box="[585,763,392,418]" pageId="2" pageNumber="492" refString="Kussakin, O. G. (1999) Marine and salt-water Asellota (Isopoda) of the cold and temperate Waters of the northern hemisphere (in Russian). Vol. III. Suborder Asellota. Part 2. Nauka, Leningrad - AH SSSR, 383 pp." type="book" year="1999">Kussakin 1999</bibRefCitation>
,
<bibRefCitation id="EFAB4B7FFFD0FFC3FCDCFE3677E549CA" author="George" box="[776,929,392,418]" pageId="2" pageNumber="492" refString="George, R. Y. (2001) Desmosomatidae and Nannoniscidae (Crustacea, Isopoda, Asellota) from bathyal and abyssal depth off north Carolina and their evolution. Journal of Natural History, 35 No. 12, 1831 - 1859." type="journal article" year="2001">George 2001</bibRefCitation>
and Brenke
<emphasis id="B94EEA9CFFD0FFC3FBE3FE36703649C9" box="[1079,1138,392,417]" italics="true" pageId="2" pageNumber="484">et al.</emphasis>
2005). The present diagnosis differs from the diagnosis presented by
<bibRefCitation id="EFAB4B7FFFD0FFC3FD7BFE0E771E49A2" author="George" box="[687,858,432,458]" pageId="2" pageNumber="492" refString="George, R. Y. (2001) Desmosomatidae and Nannoniscidae (Crustacea, Isopoda, Asellota) from bathyal and abyssal depth off north Carolina and their evolution. Journal of Natural History, 35 No. 12, 1831 - 1859." type="journal article" year="2001">George (2001)</bibRefCitation>
in including species with uniramous uropods. In
<taxonomicName id="4C3A4D0DFFD0FFC3FF43FE667520499A" box="[151,356,472,498]" class="Malacostraca" family="Desmosomatidae" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="484" phylum="Arthropoda" rank="family">Desmosomatidae</taxonomicName>
, the uropods are biramous in the plesiomorphic condition but the exopod is reduced and is completely absent in approximately half of the described species. Thinking about
<bibRefCitation id="EFAB4B7FFFD0FFC3FB8AFDBE71654A72" box="[1118,1313,512,538]" pageId="2" pageNumber="492" refString="Hessler, R. R. (1970) The Desmosomatidae (Isopoda, Asellota) of the Gay Head-Bermuda Transect. Bulletin of the Scripps Institution of Oceanography, 15, 1 - 185." type="journal article">Hessler´s (1970)</bibRefCitation>
statement about reduction of the exopod, the loss of the exopod might be homoplasious within the family. Although the phylogenetic significance of these reductions is questioned due to the presence or absence in species of the same genus, the reduction may serve well as species­level diagnostic character, but not as a generic character. Precise definition of the apomorphies is important, as the genera
<taxonomicName id="4C3A4D0DFFD0FFC3FC4AFD1E706E4AD1" box="[926,1066,672,697]" class="Malacostraca" family="Desmosomatidae" genus="Prochelator" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="484" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA9CFFD0FFC3FC4AFD1E706E4AD1" box="[926,1066,672,697]" italics="true" pageId="2" pageNumber="484">Prochelator</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3A4D0DFFD0FFC3FBB2FD1E708A4AD1" box="[1126,1230,672,697]" class="Malacostraca" family="Desmosomatidae" genus="Chelator" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="484" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA9CFFD0FFC3FBB2FD1E708A4AD1" box="[1126,1230,672,697]" italics="true" pageId="2" pageNumber="484">Chelator</emphasis>
</taxonomicName>
closely resemble each other. According to
<bibRefCitation id="EFAB4B7FFFD0FFC3FE14FD7676294A8A" author="Hessler" box="[448,621,712,738]" pageId="2" pageNumber="492" refString="Hessler, R. R. (1970) The Desmosomatidae (Isopoda, Asellota) of the Gay Head-Bermuda Transect. Bulletin of the Scripps Institution of Oceanography, 15, 1 - 185." type="journal article" year="1970">Hessler (1970)</bibRefCitation>
the prefix “Pro­” refers to the fact, that
<taxonomicName id="4C3A4D0DFFD0FFC3FB9CFD7670F44A89" box="[1096,1200,712,737]" class="Malacostraca" family="Desmosomatidae" genus="Chelator" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="484" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA9CFFD0FFC3FB9CFD7670F44A89" box="[1096,1200,712,737]" italics="true" pageId="2" pageNumber="484">Chelator</emphasis>
</taxonomicName>
is probably derived from a species of
<taxonomicName id="4C3A4D0DFFD0FFC3FEAAFD4E764D4B61" box="[382,521,752,777]" class="Malacostraca" family="Desmosomatidae" genus="Prochelator" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="484" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA9CFFD0FFC3FEAAFD4E764D4B61" box="[382,521,752,777]" italics="true" pageId="2" pageNumber="484">Prochelator</emphasis>
</taxonomicName>
. But accepting phylogenetic sytematics and the concept of monophyly, a present­day taxon cannot be assumed to be derived from another present­day taxon, but rather both are derived from a common ancestor.
</paragraph>
<paragraph id="8B85368EFFD0FFC3FF12FCD677F34C4A" blockId="2.[151,1438,152,2018]" pageId="2" pageNumber="484">
To assess the taxonomic position of
<taxonomicName id="4C3A4D0DFFD0FFC3FDB9FCD676BE4BE9" box="[621,762,872,897]" class="Malacostraca" family="Desmosomatidae" genus="Prochelator" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="484" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA9CFFD0FFC3FDB9FCD676BE4BE9" box="[621,762,872,897]" italics="true" pageId="2" pageNumber="484">Prochelator</emphasis>
</taxonomicName>
and
<emphasis id="B94EEA9CFFD0FFC3FCE6FCD677DA4BE9" box="[818,926,872,897]" italics="true" pageId="2" pageNumber="484">
<taxonomicName id="4C3A4D0DFFD0FFC3FCE6FCD677DE4BE9" box="[818,922,872,897]" class="Malacostraca" family="Desmosomatidae" genus="Chelator" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="484" phylum="Arthropoda" rank="genus">Chelator</taxonomicName>
,
</emphasis>
a detailed discussion of all characters (possible apomorphies) and a phylogenetic analysis would be necessary. The principal character shared by
<taxonomicName id="4C3A4D0DFFD0FFC3FA98FC2E74994BB9" class="Malacostraca" family="Desmosomatidae" genus="Prochelator" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="484" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA9CFFD0FFC3FA98FC2E74994BB9" italics="true" pageId="2" pageNumber="484">Prochelator</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3A4D0DFFD0FFC3FEC1FC0675394BB9" box="[277,381,952,977]" class="Malacostraca" family="Desmosomatidae" genus="Chelator" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="484" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA9CFFD0FFC3FEC1FC0675394BB9" box="[277,381,952,977]" italics="true" pageId="2" pageNumber="484">Chelator</emphasis>
</taxonomicName>
is the chela of pereopod 1.
<taxonomicName id="4C3A4D0DFFD0FFC3FD15FC06776D4BB9" box="[705,809,952,977]" class="Malacostraca" family="Desmosomatidae" genus="Chelator" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="484" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA9CFFD0FFC3FD15FC06776D4BB9" box="[705,809,952,977]" italics="true" pageId="2" pageNumber="484">Chelator</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="4C3A4D0DFFD0FFC3FBB3FC0670B04BB9" box="[1127,1268,952,977]" class="Malacostraca" family="Desmosomatidae" genus="Prochelator" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="484" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA9CFFD0FFC3FBB3FC0670B04BB9" box="[1127,1268,952,977]" italics="true" pageId="2" pageNumber="484">Prochelator</emphasis>
</taxonomicName>
by the ventral projection at the base of the claw­seta of the former. This projection is the only distinguishing character that can be identified when comparing generic diagnoses of both genera.
</paragraph>
<paragraph id="8B85368EFFD0FFC3FF12FB8E77754DE2" blockId="2.[151,1438,152,2018]" pageId="2" pageNumber="484">
The ventral setae on the carpus of pereopod 1 were so uniform between the species that
<bibRefCitation id="EFAB4B7FFFD0FFC3FB39FB8E71D84C22" author="Hessler" box="[1261,1436,1072,1098]" pageId="2" pageNumber="492" refString="Hessler, R. R. (1970) The Desmosomatidae (Isopoda, Asellota) of the Gay Head-Bermuda Transect. Bulletin of the Scripps Institution of Oceanography, 15, 1 - 185." type="journal article" year="1970">Hessler (1970)</bibRefCitation>
assigned to
<emphasis id="B94EEA9CFFD0FFC3FEF0FBE675F14C19" box="[292,437,1112,1137]" italics="true" pageId="2" pageNumber="484">
<taxonomicName id="4C3A4D0DFFD0FFC3FEF0FBE675F44C19" box="[292,432,1112,1137]" class="Malacostraca" family="Desmosomatidae" genus="Prochelator" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="484" phylum="Arthropoda" rank="genus">Prochelator</taxonomicName>
,
</emphasis>
that he gave them the formal designation of “accessory setae”.
<bibRefCitation id="EFAB4B7FFFD0FFC3FB7AFBE671194C1A" author="Hessler" box="[1198,1373,1112,1138]" pageId="2" pageNumber="492" refString="Hessler, R. R. (1970) The Desmosomatidae (Isopoda, Asellota) of the Gay Head-Bermuda Transect. Bulletin of the Scripps Institution of Oceanography, 15, 1 - 185." type="journal article" year="1970">Hessler (1970)</bibRefCitation>
used this term for
<taxonomicName id="4C3A4D0DFFD0FFC3FEF9FB3E75FE4CF1" box="[301,442,1152,1177]" class="Malacostraca" family="Desmosomatidae" genus="Prochelator" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="484" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA9CFFD0FFC3FEF9FB3E75FE4CF1" box="[301,442,1152,1177]" italics="true" pageId="2" pageNumber="484">Prochelator</emphasis>
</taxonomicName>
but not for
<taxonomicName id="4C3A4D0DFFD0FFC3FD96FB3E76ED4CF1" box="[578,681,1152,1177]" class="Malacostraca" family="Desmosomatidae" genus="Chelator" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="484" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA9CFFD0FFC3FD96FB3E76ED4CF1" box="[578,681,1152,1177]" italics="true" pageId="2" pageNumber="484">Chelator</emphasis>
</taxonomicName>
. He describes the ventral row of small setae on the lower margin of the carpus in species of
<taxonomicName id="4C3A4D0DFFD0FFC3FE06FB16767E4CA9" box="[466,570,1192,1217]" class="Malacostraca" family="Desmosomatidae" genus="Chelator" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="484" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA9CFFD0FFC3FE06FB16767E4CA9" box="[466,570,1192,1217]" italics="true" pageId="2" pageNumber="484">Chelator</emphasis>
</taxonomicName>
as a row of setae, of which none can be labelled as “accessory seta”. The term “accessory seta” is confusing and may be synonymised with the term “major seta” (
<bibRefCitation id="EFAB4B7FFFD0FFC3FB7AFB6E710B4C82" author="Hessler" box="[1198,1359,1232,1258]" pageId="2" pageNumber="492" refString="Hessler, R. R. (1970) The Desmosomatidae (Isopoda, Asellota) of the Gay Head-Bermuda Transect. Bulletin of the Scripps Institution of Oceanography, 15, 1 - 185." type="journal article" year="1970">Hessler 1970</bibRefCitation>
). Both terms are not used in the present paper; instead the setal
<typeStatus id="5481882CFFD0FFC3FCF2FB4677204D7A" box="[806,868,1272,1298]" pageId="2" pageNumber="484">types</typeStatus>
on the articles of the pereopods are described. A single midventral seta on the lower margin of the carpus is unique to
<taxonomicName id="4C3A4D0DFFD0FFC3FC14FA9E700F4D51" box="[960,1099,1312,1337]" class="Malacostraca" family="Desmosomatidae" genus="Prochelator" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="484" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA9CFFD0FFC3FC14FA9E700F4D51" box="[960,1099,1312,1337]" italics="true" pageId="2" pageNumber="484">Prochelator</emphasis>
</taxonomicName>
. This midventral seta can be used to distinguish
<taxonomicName id="4C3A4D0DFFD0FFC3FEADFAF676424D09" box="[377,518,1352,1377]" class="Malacostraca" family="Desmosomatidae" genus="Prochelator" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="484" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA9CFFD0FFC3FEADFAF676424D09" box="[377,518,1352,1377]" italics="true" pageId="2" pageNumber="484">Prochelator</emphasis>
</taxonomicName>
and
<emphasis id="B94EEA9CFFD0FFC3FDE9FAF676EE4D09" box="[573,682,1352,1377]" italics="true" pageId="2" pageNumber="484">
<taxonomicName id="4C3A4D0DFFD0FFC3FDE9FAF676E14D09" box="[573,677,1352,1377]" class="Malacostraca" family="Desmosomatidae" genus="Chelator" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="484" phylum="Arthropoda" rank="genus">Chelator</taxonomicName>
,
</emphasis>
in addition to the projection at the base of the claw­seta listed as the only distinguishing character in the paragraph above.
</paragraph>
<paragraph id="8B85368EFFD0FFC3FF12FA2676D84F2A" blockId="2.[151,1438,152,2018]" pageId="2" pageNumber="484">
Other desmosomatid species with a chelate pereopod 1 can be distinguished from
<taxonomicName id="4C3A4D0DFFD0FFC3FB53FA2671504DD9" box="[1159,1300,1432,1457]" class="Malacostraca" family="Desmosomatidae" genus="Prochelator" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="484" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA9CFFD0FFC3FB53FA2671504DD9" box="[1159,1300,1432,1457]" italics="true" pageId="2" pageNumber="484">Prochelator</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3A4D0DFFD0FFC3FA98FA2674FD4DB1" class="Malacostraca" family="Desmosomatidae" genus="Chelator" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="484" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA9CFFD0FFC3FA98FA2674FD4DB1" italics="true" pageId="2" pageNumber="484">Chelator</emphasis>
</taxonomicName>
by apomorphies. Species included in
<taxonomicName id="4C3A4D0DFFD0FFC3FDA9FA7E77034DB1" box="[637,839,1472,1497]" class="Malacostraca" family="Desmosomatidae" genus="Oecidiobranchus" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="484" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA9CFFD0FFC3FDA9FA7E77034DB1" box="[637,839,1472,1497]" italics="true" pageId="2" pageNumber="484">Oecidiobranchus</emphasis>
</taxonomicName>
possess a propodus that is more enlarged than the carpus with the dactylus folding against the ventral margin of the propodus. Unique to
<taxonomicName id="4C3A4D0DFFD0FFC3FB58FA5671124E69" box="[1164,1366,1512,1537]" class="Malacostraca" family="Desmosomatidae" genus="Oecidiobranchus" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="484" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA9CFFD0FFC3FB58FA5671124E69" box="[1164,1366,1512,1537]" italics="true" pageId="2" pageNumber="484">Oecidiobranchus</emphasis>
</taxonomicName>
is the small branchial chamber (
<bibRefCitation id="EFAB4B7FFFD0FFC3FE1CF9AE762D4E42" author="Hessler" box="[456,617,1552,1578]" pageId="2" pageNumber="492" refString="Hessler, R. R. (1970) The Desmosomatidae (Isopoda, Asellota) of the Gay Head-Bermuda Transect. Bulletin of the Scripps Institution of Oceanography, 15, 1 - 185." type="journal article" year="1970">Hessler 1970</bibRefCitation>
).
<taxonomicName id="4C3A4D0DFFD0FFC3FDA9F9AE76BE4E41" box="[637,762,1552,1577]" class="Malacostraca" family="Desmosomatidae" genus="Disparella" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="484" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA9CFFD0FFC3FDA9F9AE76BE4E41" box="[637,762,1552,1577]" italics="true" pageId="2" pageNumber="484">Disparella</emphasis>
</taxonomicName>
possess a large cephalic spine at the point of insertion of the antennula. Such a cephalic spine is also found e.g. in
<taxonomicName id="4C3A4D0DFFD0FFC3FCE9F98670704E39" box="[829,1076,1592,1617]" class="Malacostraca" family="Desmosomatidae" genus="Prochelator" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="484" phylum="Arthropoda" rank="species" species="lateralis">
<emphasis id="B94EEA9CFFD0FFC3FCE9F98670704E39" box="[829,1076,1592,1617]" italics="true" pageId="2" pageNumber="484">Prochelator lateralis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3A4D0DFFD0FFC3FBBAF98670A34E39" box="[1134,1255,1592,1617]" class="Malacostraca" family="Desmosomatidae" genus="Prochelator" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="484" phylum="Arthropoda" rank="species" species="uncatus">
<emphasis id="B94EEA9CFFD0FFC3FBBAF98670A34E39" box="[1134,1255,1592,1617]" italics="true" pageId="2" pageNumber="484">P. uncatus</emphasis>
</taxonomicName>
, but not in any species of
<taxonomicName id="4C3A4D0DFFD0FFC3FEC0F9DE753F4E11" box="[276,379,1632,1657]" class="Malacostraca" family="Desmosomatidae" genus="Chelator" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="484" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA9CFFD0FFC3FEC0F9DE753F4E11" box="[276,379,1632,1657]" italics="true" pageId="2" pageNumber="484">Chelator</emphasis>
</taxonomicName>
. The distinguishing apomorphy of
<taxonomicName id="4C3A4D0DFFD0FFC3FCCAF9DE77DE4E11" box="[798,922,1632,1657]" class="Malacostraca" family="Desmosomatidae" genus="Disparella" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="484" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA9CFFD0FFC3FCCAF9DE77DE4E11" box="[798,922,1632,1657]" italics="true" pageId="2" pageNumber="484">Disparella</emphasis>
</taxonomicName>
, which separates the genus from
<taxonomicName id="4C3A4D0DFFD0FFC3FAFCF9DE74FC4EC9" class="Malacostraca" family="Desmosomatidae" genus="Prochelator" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="484" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA9CFFD0FFC3FAFCF9DE74FC4EC9" italics="true" pageId="2" pageNumber="484">Prochelator</emphasis>
</taxonomicName>
, is the elongate slender propodus (more than 3.5 times long as wide) and the setation on the ventral margin of the carpus of pereopod 1.
<taxonomicName id="4C3A4D0DFFD0FFC3FE3EF90E76234EA1" box="[490,615,1712,1737]" class="Malacostraca" family="Desmosomatidae" genus="Disparella" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="484" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA9CFFD0FFC3FE3EF90E76234EA1" box="[490,615,1712,1737]" italics="true" pageId="2" pageNumber="484">Disparella</emphasis>
</taxonomicName>
possess a well defined row of setae behind the claw­seta. Such a row is not found in
<taxonomicName id="4C3A4D0DFFD0FFC3FE99F966759E4E99" box="[333,474,1752,1777]" class="Malacostraca" family="Desmosomatidae" genus="Prochelator" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="484" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA9CFFD0FFC3FE99F966759E4E99" box="[333,474,1752,1777]" italics="true" pageId="2" pageNumber="484">Prochelator</emphasis>
</taxonomicName>
or
<taxonomicName id="4C3A4D0DFFD0FFC3FDD7F966762E4E99" box="[515,618,1752,1777]" class="Malacostraca" family="Desmosomatidae" genus="Chelator" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="484" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA9CFFD0FFC3FDD7F966762E4E99" box="[515,618,1752,1777]" italics="true" pageId="2" pageNumber="484">Chelator</emphasis>
</taxonomicName>
. Pereopods 24 of
<taxonomicName id="4C3A4D0DFFD0FFC3FC86F966778A4E99" box="[850,974,1752,1777]" class="Malacostraca" family="Desmosomatidae" genus="Disparella" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="484" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA9CFFD0FFC3FC86F966778A4E99" box="[850,974,1752,1777]" italics="true" pageId="2" pageNumber="484">Disparella</emphasis>
</taxonomicName>
species are obviously setose (carpus&gt; 20 setae, propodus&gt; 8 setae in adult) while in
<taxonomicName id="4C3A4D0DFFD0FFC3FD6FF8BE770C4F71" box="[699,840,1792,1817]" class="Malacostraca" family="Desmosomatidae" genus="Prochelator" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="484" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA9CFFD0FFC3FD6FF8BE770C4F71" box="[699,840,1792,1817]" italics="true" pageId="2" pageNumber="484">Prochelator</emphasis>
</taxonomicName>
the carpus of the anterior pereopods have no more than 10 setae in a row (except
<taxonomicName id="4C3A4D0DFFD0FFC3FE28F89676CF4F29" box="[508,651,1832,1857]" class="Malacostraca" family="Desmosomatidae" genus="Prochelator" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="484" phylum="Arthropoda" rank="species" species="hampsoni">
<emphasis id="B94EEA9CFFD0FFC3FE28F89676CF4F29" box="[508,651,1832,1857]" italics="true" pageId="2" pageNumber="484">P. hampsoni</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8B85368EFFD0FFC3FF12F8EE71D94F8A" blockId="2.[151,1438,152,2018]" pageId="2" pageNumber="484">
Although the comparison of characters shows that
<taxonomicName id="4C3A4D0DFFD0FFC3FCC3F8EE77E04F01" box="[791,932,1872,1897]" class="Malacostraca" family="Desmosomatidae" genus="Prochelator" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="484" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA9CFFD0FFC3FCC3F8EE77E04F01" box="[791,932,1872,1897]" italics="true" pageId="2" pageNumber="484">Prochelator</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3A4D0DFFD0FFC3FC08F8EE70004F01" box="[988,1092,1872,1897]" class="Malacostraca" family="Desmosomatidae" genus="Chelator" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="484" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA9CFFD0FFC3FC08F8EE70004F01" box="[988,1092,1872,1897]" italics="true" pageId="2" pageNumber="484">Chelator</emphasis>
</taxonomicName>
are very similar,
<taxonomicName id="4C3A4D0DFFD0FFC3FADBF8EE71D84F01" box="[1295,1436,1872,1897]" class="Malacostraca" family="Desmosomatidae" genus="Prochelator" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="484" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA9CFFD0FFC3FADBF8EE71D84F01" box="[1295,1436,1872,1897]" italics="true" pageId="2" pageNumber="484">Prochelator</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3A4D0DFFD0FFC3FF18F8C675704FF9" box="[204,308,1912,1937]" class="Malacostraca" family="Desmosomatidae" genus="Chelator" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="484" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA9CFFD0FFC3FF18F8C675704FF9" box="[204,308,1912,1937]" italics="true" pageId="2" pageNumber="484">Chelator</emphasis>
</taxonomicName>
should be treated as separate genera based on the following reasoning: it is not possible to homologize the setae on the ventral margin of the carpus of pereopod 1, and no transformation series can be postulated explaining how the midventral setae (
<taxonomicName id="4C3A4D0DFFD0FFC3FD0EF87677224F89" box="[730,870,1992,2017]" class="Malacostraca" family="Desmosomatidae" genus="Prochelator" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="484" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA9CFFD0FFC3FD0EF87677224F89" box="[730,870,1992,2017]" italics="true" pageId="2" pageNumber="484">Prochelator</emphasis>
</taxonomicName>
) and the few small setae (
<taxonomicName id="4C3A4D0DFFD0FFC3FB77F876714F4F89" box="[1187,1291,1992,2017]" class="Malacostraca" family="Desmosomatidae" genus="Chelator" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="484" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA9CFFD0FFC3FB77F876714F4F89" box="[1187,1291,1992,2017]" italics="true" pageId="2" pageNumber="484">Chelator</emphasis>
</taxonomicName>
) originated.
</paragraph>
<paragraph id="8B85368EFFD1FFC2FF43FF26756F4A72" blockId="3.[151,1437,152,538]" pageId="3" pageNumber="485">
The uniramous uropod in
<taxonomicName id="4C3A4D0DFFD1FFC2FE12FF26762848D9" box="[454,620,152,177]" class="Malacostraca" family="Desmosomatidae" genus="Prochelator" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="485" phylum="Arthropoda" rank="species" species="incomitatus">
<emphasis id="B94EEA9CFFD1FFC2FE12FF26762848D9" box="[454,620,152,177]" italics="true" pageId="3" pageNumber="485">P. incomitatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3A4D0DFFD1FFC2FD70FF26777948D9" box="[676,829,152,177]" class="Malacostraca" family="Desmosomatidae" genus="Prochelator" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="485" phylum="Arthropoda" rank="species" species="angolensis">
<emphasis id="B94EEA9CFFD1FFC2FD70FF26777948D9" box="[676,829,152,177]" italics="true" pageId="3" pageNumber="485">P. angolensis</emphasis>
</taxonomicName>
may have evolved convergently because the loss of the exopod is an apparently homoplasious character state as it occurs in genera that are otherwise widely disparate; the body form of females differs clearly between the two genera; posterolateral spines on the pleotelson are always present in females of
<taxonomicName id="4C3A4D0DFFD1FFC2FD9FFEAE769C4941" box="[587,728,272,297]" class="Malacostraca" family="Desmosomatidae" genus="Prochelator" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="485" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA9CFFD1FFC2FD9FFEAE769C4941" box="[587,728,272,297]" italics="true" pageId="3" pageNumber="485">Prochelator</emphasis>
</taxonomicName>
while in females of
<taxonomicName id="4C3A4D0DFFD1FFC2FC1FFEAE70774941" box="[971,1075,272,297]" class="Malacostraca" family="Desmosomatidae" genus="Chelator" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="485" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA9CFFD1FFC2FC1FFEAE70774941" box="[971,1075,272,297]" italics="true" pageId="3" pageNumber="485">Chelator</emphasis>
</taxonomicName>
these posterolateral spines are absent. Furthermore, females of
<taxonomicName id="4C3A4D0DFFD1FFC2FDC3FE86763B4939" box="[535,639,312,337]" class="Malacostraca" family="Desmosomatidae" genus="Chelator" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="485" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA9CFFD1FFC2FDC3FE86763B4939" box="[535,639,312,337]" italics="true" pageId="3" pageNumber="485">Chelator</emphasis>
</taxonomicName>
are more compact and the pereonites are rounded, while species of
<taxonomicName id="4C3A4D0DFFD1FFC2FF43FEDE75604911" box="[151,292,352,377]" class="Malacostraca" family="Desmosomatidae" genus="Prochelator" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="485" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA9CFFD1FFC2FF43FEDE75604911" box="[151,292,352,377]" italics="true" pageId="3" pageNumber="485">Prochelator</emphasis>
</taxonomicName>
have a different profile from lateral view. While in some species of
<taxonomicName id="4C3A4D0DFFD1FFC2FBBAFEDE70BF4911" box="[1134,1275,352,377]" class="Malacostraca" family="Desmosomatidae" genus="Prochelator" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="485" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA9CFFD1FFC2FBBAFEDE70BF4911" box="[1134,1275,352,377]" italics="true" pageId="3" pageNumber="485">Prochelator</emphasis>
</taxonomicName>
the posterior pereonites and the pleotelson are flattened, in species of
<taxonomicName id="4C3A4D0DFFD1FFC2FCE6FE3677DE49C9" box="[818,922,392,417]" class="Malacostraca" family="Desmosomatidae" genus="Chelator" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="485" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA9CFFD1FFC2FCE6FE3677DE49C9" box="[818,922,392,417]" italics="true" pageId="3" pageNumber="485">Chelator</emphasis>
</taxonomicName>
the pleotelson is as high as pereonite 5, the rounded pleotelson form giving the body outline a more compact look than in
<taxonomicName id="4C3A4D0DFFD1FFC2FB9DFE0E709049A1" box="[1097,1236,432,457]" class="Malacostraca" family="Desmosomatidae" genus="Prochelator" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="485" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA9CFFD1FFC2FB9DFE0E709049A1" box="[1097,1236,432,457]" italics="true" pageId="3" pageNumber="485">Prochelator</emphasis>
</taxonomicName>
. In
<taxonomicName id="4C3A4D0DFFD1FFC2FAD2FE0E712A49A1" box="[1286,1390,432,457]" class="Malacostraca" family="Desmosomatidae" genus="Chelator" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="485" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA9CFFD1FFC2FAD2FE0E712A49A1" box="[1286,1390,432,457]" italics="true" pageId="3" pageNumber="485">Chelator</emphasis>
</taxonomicName>
the first four body segments are more compact than in
<taxonomicName id="4C3A4D0DFFD1FFC2FD32FE6677374999" box="[742,883,472,497]" class="Malacostraca" family="Desmosomatidae" genus="Prochelator" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="485" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA9CFFD1FFC2FD32FE6677374999" box="[742,883,472,497]" italics="true" pageId="3" pageNumber="485">Prochelator</emphasis>
</taxonomicName>
(the first pereonite in
<taxonomicName id="4C3A4D0DFFD1FFC2FBA1FE6670994999" box="[1141,1245,472,497]" class="Malacostraca" family="Desmosomatidae" genus="Chelator" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="485" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA9CFFD1FFC2FBA1FE6670994999" box="[1141,1245,472,497]" italics="true" pageId="3" pageNumber="485">Chelator</emphasis>
</taxonomicName>
twice as high as pereonite 5).
</paragraph>
</subSubSection>
</treatment>
</document>