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<document ID-DOI="10.5281/zenodo.163451" ID-GBIF-Dataset="f9566778-1df4-44d6-9a4c-849de39a4b6c" ID-GBIF-Taxon="124955175" ID-Zenodo-Dep="163451" checkinTime="1480975261095" checkinUser="plazi" docAuthor="Vincent J. Kalkman" docDate="2016" docId="03938796FFDDFFE926B08C99FBC0FB87" docLanguage="en" docName="10.5281zenodo.163451_Kalkman.pdf" docOrigin="Odonatologica 45 (3)" docStyle="DocumentStyle{}" docTitle="Celebargiolestes Kennedy 1925" docType="treatment" docVersion="11" lastPageId="7" lastPageNumber="242" masterDocId="FFAAFFEEFFDEFFEE27258934FFCBFF99" masterDocTitle="Revision of the genus Celebargiolestes Kennedy, 1925 (Odonata: Argiolestidae)" masterLastPageNumber="269" masterPageNumber="235" pageId="3" pageNumber="238" updateTime="1643502553511" updateUser="ExternalLinkService">
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<mods:title>Revision of the genus Celebargiolestes Kennedy, 1925 (Odonata: Argiolestidae)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Vincent J. Kalkman</mods:namePart>
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<mods:title>Odonatologica</mods:title>
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<mods:date>2016</mods:date>
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<mods:number>2016-12-01</mods:number>
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<mods:number>45</mods:number>
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<mods:number>3</mods:number>
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<mods:identifier type="DOI">10.5281/zenodo.163451</mods:identifier>
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<treatment ID-GBIF-Taxon="124955175" LSID="urn:lsid:plazi:treatment:03938796FFDDFFE926B08C99FBC0FB87" httpUri="http://treatment.plazi.org/id/03938796FFDDFFE926B08C99FBC0FB87" lastPageId="7" lastPageNumber="242" pageId="3" pageNumber="238">
<subSubSection box="[405,789,1453,1481]" pageId="3" pageNumber="238" type="nomenclature">
<paragraph blockId="3.[159,1036,1453,1549]" box="[405,789,1453,1481]" pageId="3" pageNumber="238">
<heading bold="true" box="[405,789,1453,1481]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="238" reason="0">
<taxonomicName authority="Kennedy, 1925" authorityName="Kennedy" authorityYear="1925" box="[405,789,1453,1481]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="238" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[405,789,1453,1481]" pageId="3" pageNumber="238">
<emphasis bold="true" box="[405,604,1453,1481]" italics="true" pageId="3" pageNumber="238">Celebargiolestes</emphasis>
<bibRefCitation author="Kennedy C. H." box="[611,789,1453,1481]" firstAuthor="Kennedy" journalOrPublisher="Bulletin of the Museum of Comparative Zoology" pageId="3" pageNumber="238" pagination="291 - 311" part="67" refId="ref12786" refString="Kennedy C. H. 1925. New genera of Mega- podagrioninae, with notes on the subfamily. Bulletin of the Museum of Comparative Zoology 67: 291 - 311" title="New genera of Mega- podagrioninae, with notes on the subfamily" type="journal article" year="1925">Kennedy, 1925</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="238" type="reference_group">
<paragraph blockId="3.[159,1036,1453,1549]" pageId="3" pageNumber="238">
<taxonomicName authority="Kennedy, 1925: 295" authorityName="Kennedy" authorityPageNumber="295" authorityYear="1925" box="[159,548,1494,1518]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="238" phylum="Arthropoda" rank="genus">
<emphasis box="[159,322,1494,1518]" italics="true" pageId="3" pageNumber="238">Celebargiolestes</emphasis>
<bibRefCitation author="Kennedy C. H." box="[330,548,1494,1518]" firstAuthor="Kennedy" journalOrPublisher="Bulletin of the Museum of Comparative Zoology" pageId="3" pageNumber="238" pagination="291 - 311" part="67" refId="ref12786" refString="Kennedy C. H. 1925. New genera of Mega- podagrioninae, with notes on the subfamily. Bulletin of the Museum of Comparative Zoology 67: 291 - 311" title="New genera of Mega- podagrioninae, with notes on the subfamily" type="journal article" year="1925">Kennedy, 1925: 295</bibRefCitation>
</taxonomicName>
f. [type species:
<taxonomicName authority="Selys, 1886" authorityName="Selys" authorityYear="1886" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="238" phylum="Arthropoda" rank="species" species="cinctus">
<emphasis box="[725,969,1494,1518]" italics="true" pageId="3" pageNumber="238">Celebargiolestes cinctus</emphasis>
Selys, 1886
</taxonomicName>
; by original designation]
</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="241" pageId="4" pageNumber="239" type="description">
<paragraph blockId="4.[159,1036,254,542]" box="[159,220,254,280]" pageId="4" pageNumber="239">
<heading bold="true" box="[159,220,254,280]" fontSize="11" level="3" pageId="4" pageNumber="239" reason="0">
<emphasis bold="true" box="[159,220,254,280]" pageId="4" pageNumber="239">Male</emphasis>
</heading>
</paragraph>
<paragraph blockId="4.[159,1036,254,542]" pageId="4" pageNumber="239">
Head Labium dull yellow or dull yellow with the anterior half to ⅔ dark brown to black, slightly wider than long, median cleft almost ²/₅ the length of labium and about half as wide as deep. Pattern on front of head varies between species, labrum is either pale yellow-brown or black, front of head largely dull yellow-brown with or without a slight reddish tinge (ochre), dorsum of head mat black, antennae yellow brown getting darker towards apex. Head seen from dorsal elongated (
<figureCitation box="[626,710,515,542]" captionStart="Figures 15 18" captionStartId="14.[159,238,1441,1466]" captionTargetBox="[203,991,248,1401]" captionTargetId="figure@14.[203,991,248,1401]" captionTargetPageId="14" captionText="Figures 15 18. Body of Celebargiolestes males. 15 C. cinctus (immature), Ranotongkor, 02 - viii- 1940, JvT 6476; 16 C. cinctus (old adult), Mapanget, 23 - vi- 1940, JvT 6638; 17 C. cinctus ( red legs form), 25 km NE of Luwuk, 30 - i- 1989, JvT 6670; 18 C. askewi sp. nov., holotype." httpUri="https://zenodo.org/record/192856/files/figure.png" pageId="4" pageNumber="239">Fig. 15</figureCitation>
).
</paragraph>
<paragraph blockId="4.[159,1036,589,1101]" pageId="4" pageNumber="239">
Thorax Shape of prothorax and pronotum shown in
<figureCitation box="[827,928,589,616]" captionStart="Figures 1 6" captionStartId="10.[159,238,1412,1437]" captionTargetBox="[173,1024,597,1348]" captionTargetId="figure@10.[159,1036,578,1371]" captionTargetPageId="10" captionText="Figures 1 6. Details of Celebargiolestes cinctus. 1 Prothorax and pronotum, male; 2 Venation, male, Ranotongkor, 02 - viii- 1940, JvT 6476; 3 Pterostigma of fore and hind wing, male, Mapanget, 23 - vi- 1940, JvT 6638; 4 Pterostigma of fore and hind wing, Mapanget, 23 - vi- 1940, female, JvT 6639; 5 Body, female, Ton- dano, 03 - ix- 1940, JvT 6494; 6 Ovipositor, female, Tondano, 03 - ix- 1940, JvT 6494." httpUri="https://zenodo.org/record/192854/files/figure.png" pageId="4" pageNumber="239">Figure 1</figureCitation>
. Posteri- or lobe broad with rounded margin, seen from lateral gently sloping and with the outer border steeply raised. Sides of prothorax dull yellow-brown; dorso-median part of pronotum black, the black becomes more extensive with age and can cover most of the pronotum. Metepimeron largely yellowbrown, mesepimeron and dorsum of thorax largely black. Mesepimeron has a pale line running diagonally from the coxa of the second leg to near the hind margin of the mesepimeron, in older specimens the mesepimeron can be largely black making the pale line invisible. The front of the synthorax has in the outer anterior corners an incomplete ante-humeral stripe which at most reaches slightly halfway the dorsum of the thorax. The size of this stripe varies between species but also with age and can be largely or completely absent in fully mature specimens. Old specimens of at least some species develop greyish blue pruinosity on thorax.
</paragraph>
<paragraph blockId="4.[159,1036,1149,1288]" pageId="4" pageNumber="239">
Legs Coxae, trochanters and femora dull yellow to ochre, colour of tibiae and tarsi yellow-brown to brown. Spines on tibiae and femora long. Femora with two broad but poorly defined black rings (absent in
<taxonomicName authorityName="Selys" authorityYear="1886" box="[870,983,1225,1251]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="239" phylum="Arthropoda" rank="species" species="cinctus">
<emphasis box="[870,983,1225,1251]" italics="true" pageId="4" pageNumber="239">C. cinctus</emphasis>
</taxonomicName>
red legs), knees black, spines orange-brown.
</paragraph>
<paragraph blockId="4.[159,1036,1336,1549]" lastBlockId="5.[159,1036,253,466]" lastPageId="5" lastPageNumber="240" pageId="4" pageNumber="239">
Wings Venation dark brown to black (
<figureCitation box="[645,716,1336,1362]" captionStart="Figures 1 6" captionStartId="10.[159,238,1412,1437]" captionTargetBox="[173,1024,597,1348]" captionTargetId="figure@10.[159,1036,578,1371]" captionTargetPageId="10" captionText="Figures 1 6. Details of Celebargiolestes cinctus. 1 Prothorax and pronotum, male; 2 Venation, male, Ranotongkor, 02 - viii- 1940, JvT 6476; 3 Pterostigma of fore and hind wing, male, Mapanget, 23 - vi- 1940, JvT 6638; 4 Pterostigma of fore and hind wing, Mapanget, 23 - vi- 1940, female, JvT 6639; 5 Body, female, Ton- dano, 03 - ix- 1940, JvT 6494; 6 Ovipositor, female, Tondano, 03 - ix- 1940, JvT 6494." httpUri="https://zenodo.org/record/192854/files/figure.png" pageId="4" pageNumber="239">Fig. 2</figureCitation>
). Wings hyaline in young- er specimens but a smoky outer border develops with age while some old specimens have wings with a strong brown tinge throughout. Petiolation long. Fw and Hw with 2 Ax; Fw with 29 to 40 Px and Hw with 25 to 36 Px. R4 originating basal to subnodus and closer to subnodus than to arculus. Discoidal cell in Fw and Hw with costal side about two and a half times as long as distal side, sharpest angle about 40°. Ac clearly closer to Ax1 then to Ax2. Brace vein absent. Pterostigma dark brown and shaped as in Figures 3, 4. Mostly up to three rows of cells between anal vein and border of wing in Fw and up to four in Hw. Fields between Ir2 and R3, between R3 and IR3, and between IR3 and R4 at distal end containing 3 or more rows of cells.
</paragraph>
<paragraph blockId="5.[159,1036,515,1250]" pageId="5" pageNumber="240">
Abdomen Pattern on abdomen as in
<figureCitation box="[606,772,515,542]" captionStart-0="Figures 15 18" captionStart-1="Figures 19 22" captionStartId-0="14.[159,238,1441,1466]" captionStartId-1="18.[159,238,1470,1495]" captionTargetBox-0="[203,991,248,1401]" captionTargetBox-1="[212,983,244,1431]" captionTargetId-0="figure@14.[203,991,248,1401]" captionTargetId-1="figure@18.[212,983,244,1431]" captionTargetPageId-0="14" captionTargetPageId-1="18" captionText-0="Figures 15 18. Body of Celebargiolestes males. 15 C. cinctus (immature), Ranotongkor, 02 - viii- 1940, JvT 6476; 16 C. cinctus (old adult), Mapanget, 23 - vi- 1940, JvT 6638; 17 C. cinctus ( red legs form), 25 km NE of Luwuk, 30 - i- 1989, JvT 6670; 18 C. askewi sp. nov., holotype." captionText-1="Figures 19 22. Body of Celebargiolestes males. 19 C. toli sp. nov., holotype; 20 C. toli sp. nov., Peleng Island, 05 - viii- 1941, JvT 6453; 21 C. orri sp. nov., holotype; 22 C. orri sp. nov., 10 km WNW Palopo, 29 - x- 1993, JvT 11910." httpUri-0="https://zenodo.org/record/192856/files/figure.png" httpUri-1="https://zenodo.org/record/192857/files/figure.png" pageId="5" pageNumber="240">Figures 1522</figureCitation>
; dark pattern elements are dark brown to black, pale pattern elements are yellow-brown. S1 yellowbrown with posterior half to ⅔ of dorsum brown-black, S2 dorsally dark brown, laterally yellow-brown, S36 with dorsum brown with a poorly defined pale anterior ring, S710 dark brown to black, S79 with an indication of a pale ring. S10 strongly depressed in the middle, hindborder of S10 without modifications. Upper appendages black, general shape as shown in
<figureCitation box="[159,272,776,802]" captionStart="Figures 34 39" captionStartId="27.[159,238,1413,1438]" captionTargetBox="[156,1038,480,1372]" captionTargetId="figure@27.[156,1038,480,1372]" captionTargetPageId="27" captionText="Figures 34 39. Appendages of Celebargiolestes males. 34 C. cinctus, Ranotongkor, 02 - viii- 1940, JvT 6476, dorsal view; 35 C. cinctus, Ranotongkor, 02 - viii- 1940, JvT 6476, lateral view; 36 Ranotongkor, 02 - viii- 1940, JvT 6476, ventral view; 37 C. askewi sp. nov., holotype, dorsal view; 38 C. askewi sp. nov., holotype, lateral view; 39 C. askewi sp. nov., holotype, ventral view." httpUri="https://zenodo.org/record/192860/files/figure.png" pageId="5" pageNumber="240">Figure 34</figureCitation>
. Upper appendages with two poorly developed flanges just before the apex, the upper carries a short row of denticles; the lower flange runs towards the base of the appendages ending at one third of lengths of the appendages. Inferior appendages with rounded and swollen base, abruptly constricted into a slender and medium long to long apex which is slightly directed upwards reaching to ¼ or just surpassing half of the upper appendages. Genital ligula as in
<figureCitation box="[466,637,1000,1026]" captionStart-0="Figures 23 27" captionStart-1="Figures 28 33" captionStartId-0="25.[159,237,1412,1437]" captionStartId-1="26.[159,237,1353,1378]" captionTargetBox-0="[208,986,249,1370]" captionTargetBox-1="[156,1036,247,1313]" captionTargetId-0="figure@25.[208,986,249,1370]" captionTargetId-1="figure@26.[156,1036,247,1313]" captionTargetPageId-0="25" captionTargetPageId-1="26" captionText-0="Figures 23 27. Genital ligula of Celebargiolestes males. 23 C. cinctus, Dumoga Bone NP, 04 - vi- 1985, JvT 6486, lateral view; 24 C. cinctus, Dumoga Bone NP, 04 - vi- 1985, JvT 6486, ventral view; 25 C. cinctus red leg, Banggai Peninsula, 08 - x- 1989, latero-ventral view; 26 C. askewi sp. nov., Polewali, 23 - x- 1940, JvT 6636, lateral view; 27 C. askewi sp. nov., Polewali, 23 - x- 1940, JvT 6636, ventral view." captionText-1="Figures 28 33. Genital ligula of Celebargiolestes males. 28 C. toli sp. nov., Gunung Watuwila, 02 04 - xi- 1989, JvT 6636, lateral view, lateral lobe deformed and bent inwards; 29 C. toli sp. nov., Gunung Watuwila, 02 04 - xi- 1989, JvT 6636, ventral view; 30 C. toli sp. nov., Banggai Archipelago, Peleng Island, 05 - viii- 1941, JvT 6501, lateral view; 31 C. toli sp. nov., Banggai Archipelago, Peleng Island, 05 - viii- 1941, JvT 6501, ventral view; 32 C. orri sp. nov., Lore Lindu, 17 - xii- 1985, JvT 6444, lateral view; 33 C. orri sp. nov., Lore Lindu, 17 - xii- 1985, JvT 6444, ventral view." httpUri-0="https://zenodo.org/record/192858/files/figure.png" httpUri-1="https://zenodo.org/record/192859/files/figure.png" pageId="5" pageNumber="240">Figures 2333</figureCitation>
, setae on the shaft of the genital ligula lacking, ligula head with internal flange, apical lobe either with a deep narrow cleft or with a shallow, wide cleft, lateral lobes large and broadly rounded and largely covering the opening of head of the ligula, the lateral lobes become easily deformed. The median part of the head of the ligula is, in longitudinal direction, only lightly sclerotized, this is well visible on
<figureCitation box="[159,272,1224,1250]" captionStart="Figures 28 33" captionStartId="26.[159,237,1353,1378]" captionTargetBox="[156,1036,247,1313]" captionTargetId="figure@26.[156,1036,247,1313]" captionTargetPageId="26" captionText="Figures 28 33. Genital ligula of Celebargiolestes males. 28 C. toli sp. nov., Gunung Watuwila, 02 04 - xi- 1989, JvT 6636, lateral view, lateral lobe deformed and bent inwards; 29 C. toli sp. nov., Gunung Watuwila, 02 04 - xi- 1989, JvT 6636, ventral view; 30 C. toli sp. nov., Banggai Archipelago, Peleng Island, 05 - viii- 1941, JvT 6501, lateral view; 31 C. toli sp. nov., Banggai Archipelago, Peleng Island, 05 - viii- 1941, JvT 6501, ventral view; 32 C. orri sp. nov., Lore Lindu, 17 - xii- 1985, JvT 6444, lateral view; 33 C. orri sp. nov., Lore Lindu, 17 - xii- 1985, JvT 6444, ventral view." httpUri="https://zenodo.org/record/192859/files/figure.png" pageId="5" pageNumber="240">Figure 28</figureCitation>
.
</paragraph>
<paragraph blockId="5.[159,1036,1299,1400]" box="[159,247,1299,1325]" pageId="5" pageNumber="240">
<heading bold="true" box="[159,247,1299,1325]" fontSize="11" level="3" pageId="5" pageNumber="240" reason="0">
<emphasis bold="true" box="[159,247,1299,1325]" pageId="5" pageNumber="240">Female</emphasis>
</heading>
</paragraph>
<paragraph blockId="5.[159,1036,1299,1400]" pageId="5" pageNumber="240">Pattern and colours of females are largely identical to those of males. The main differences from males are given below.</paragraph>
<paragraph blockId="5.[159,1036,1448,1549]" pageId="5" pageNumber="240">Wings Pt becomes cream white-yellow often with the outer border mottled greyish-black. Fw and Hw with 2 Ax; Fw with 25 sclerotized 33 Px and Hw with 23 sclerotized 31 Px.</paragraph>
<paragraph blockId="6.[159,1036,253,429]" pageId="6" pageNumber="241">
Abdomen Short and stocky, subequal or shorter than Hw (
<figureCitation box="[856,924,253,280]" captionStart="Figures 1 6" captionStartId="10.[159,238,1412,1437]" captionTargetBox="[173,1024,597,1348]" captionTargetId="figure@10.[159,1036,578,1371]" captionTargetPageId="10" captionText="Figures 1 6. Details of Celebargiolestes cinctus. 1 Prothorax and pronotum, male; 2 Venation, male, Ranotongkor, 02 - viii- 1940, JvT 6476; 3 Pterostigma of fore and hind wing, male, Mapanget, 23 - vi- 1940, JvT 6638; 4 Pterostigma of fore and hind wing, Mapanget, 23 - vi- 1940, female, JvT 6639; 5 Body, female, Ton- dano, 03 - ix- 1940, JvT 6494; 6 Ovipositor, female, Tondano, 03 - ix- 1940, JvT 6494." httpUri="https://zenodo.org/record/192854/files/figure.png" pageId="6" pageNumber="241">Fig. 5</figureCitation>
). S37 all short, S78 with broad intersegmental annulae, S8 shorter than S9. Ovipositor large, its tip surpassing that of the cerci (
<figureCitation box="[659,727,328,355]" captionStart="Figures 1 6" captionStartId="10.[159,238,1412,1437]" captionTargetBox="[173,1024,597,1348]" captionTargetId="figure@10.[159,1036,578,1371]" captionTargetPageId="10" captionText="Figures 1 6. Details of Celebargiolestes cinctus. 1 Prothorax and pronotum, male; 2 Venation, male, Ranotongkor, 02 - viii- 1940, JvT 6476; 3 Pterostigma of fore and hind wing, male, Mapanget, 23 - vi- 1940, JvT 6638; 4 Pterostigma of fore and hind wing, Mapanget, 23 - vi- 1940, female, JvT 6639; 5 Body, female, Ton- dano, 03 - ix- 1940, JvT 6494; 6 Ovipositor, female, Tondano, 03 - ix- 1940, JvT 6494." httpUri="https://zenodo.org/record/192854/files/figure.png" pageId="6" pageNumber="241">Fig. 6</figureCitation>
). Cerci pointed, ovipositor with a dense row (30 or more) of small spines. Cerci brown, base of valvifer yellow becoming brown to black towards the tip.
</paragraph>
<paragraph blockId="6.[159,1036,477,914]" box="[159,433,477,503]" pageId="6" pageNumber="241">
<heading bold="true" box="[159,433,477,503]" fontSize="11" level="3" pageId="6" pageNumber="241" reason="0">
<emphasis bold="true" box="[159,433,477,503]" pageId="6" pageNumber="241">Age-related variability</emphasis>
</heading>
</paragraph>
<paragraph blockId="6.[159,1036,477,914]" pageId="6" pageNumber="241">
In addition to the regional variability within species there is also age-related variability in both male and female. This is illustrated in
<figureCitation box="[811,946,552,579]" captionStart="Figures 7 14" captionStartId="13.[159,238,1384,1409]" captionTargetBox="[196,998,249,1341]" captionTargetId="figure@13.[196,998,249,1342]" captionTargetPageId="13" captionText="Figures 7 14. Head of Celebargiolestes males. 7 C. cinctus (immature), Ranotongkor, 02 - viii- 1940, JvT 6476; 8 C. cinctus (old adult), Mapanget, 23 - vi- 1940, JvT 6638; 9 C. cinctus ( red legs form), 25 km NE of Luwuk, 30 - i- 1989, JvT 6670; 10 C. askewi sp. nov., holotype; 11 C. toli sp. nov., holotype; 12 C. toli sp. nov., Peleng Island, 05 - viii- 1941, JvT 6453; 13 C. orri sp. nov., holotype; 14 C. orri sp. nov., 10 km WNW Palopo, 29 - x- 1993, JvT 11910." httpUri="https://zenodo.org/record/192855/files/figure.png" pageId="6" pageNumber="241">Figures 7, 8</figureCitation>
,
<figureCitation captionStart="Figures 15 18" captionStartId="14.[159,238,1441,1466]" captionTargetBox="[203,991,248,1401]" captionTargetId="figure@14.[203,991,248,1401]" captionTargetPageId="14" captionText="Figures 15 18. Body of Celebargiolestes males. 15 C. cinctus (immature), Ranotongkor, 02 - viii- 1940, JvT 6476; 16 C. cinctus (old adult), Mapanget, 23 - vi- 1940, JvT 6638; 17 C. cinctus ( red legs form), 25 km NE of Luwuk, 30 - i- 1989, JvT 6670; 18 C. askewi sp. nov., holotype." httpUri="https://zenodo.org/record/192856/files/figure.png" pageId="6" pageNumber="241">15 and 16</figureCitation>
. In older specimens the pale parts of prothorax, synthorax and abdomen become darker and largely obscured, looking nearly completely and uniformly black in some old specimens. Some old specimens develop a greyish blue pruinosity on thorax and, in males, the base of the upper appendages. The wings are hyaline in younger specimens but a smoky outer border of the wing develops with age, whereas some old specimens have wings with a strong brown tinge throughout. In males the Pt becomes darker with age but in females the Pt becomes cream white-yellow often with the outer bor- der mottled greyish-black (
<figureCitation box="[475,544,888,914]" captionStart="Figures 1 6" captionStartId="10.[159,238,1412,1437]" captionTargetBox="[173,1024,597,1348]" captionTargetId="figure@10.[159,1036,578,1371]" captionTargetPageId="10" captionText="Figures 1 6. Details of Celebargiolestes cinctus. 1 Prothorax and pronotum, male; 2 Venation, male, Ranotongkor, 02 - viii- 1940, JvT 6476; 3 Pterostigma of fore and hind wing, male, Mapanget, 23 - vi- 1940, JvT 6638; 4 Pterostigma of fore and hind wing, Mapanget, 23 - vi- 1940, female, JvT 6639; 5 Body, female, Ton- dano, 03 - ix- 1940, JvT 6494; 6 Ovipositor, female, Tondano, 03 - ix- 1940, JvT 6494." httpUri="https://zenodo.org/record/192854/files/figure.png" pageId="6" pageNumber="241">Fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="242" pageId="6" pageNumber="241" type="diagnosis">
<paragraph blockId="6.[159,1036,963,1550]" box="[159,281,963,989]" pageId="6" pageNumber="241">
<heading bold="true" box="[159,281,963,989]" fontSize="11" level="3" pageId="6" pageNumber="241" reason="0">
<emphasis bold="true" box="[159,281,963,989]" pageId="6" pageNumber="241">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph blockId="6.[159,1036,963,1550]" pageId="6" pageNumber="241">
<taxonomicName box="[159,313,1000,1026]" class="Insecta" family="Argiolestidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="241" phylum="Arthropoda" rank="family">Argiolestidae</taxonomicName>
occurring from the Philippines and Sulawesi eastwards to Australia and New Caledonia are characterised by the males having a lower and upper flange near the apex of the upper appendage of which the upper flange always has a row of small and black denticles. In
<taxonomicName authorityName="Kennedy" authorityYear="1925" box="[718,895,1112,1138]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="241" phylum="Arthropoda" rank="genus">
<emphasis box="[718,895,1112,1138]" italics="true" pageId="6" pageNumber="241">Celebargiolestes</emphasis>
</taxonomicName>
these flanges and the denticles are relatively poorly developed but are still always visible. Within this group,
<taxonomicName authorityName="Kennedy" authorityYear="1925" box="[380,557,1187,1213]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="241" phylum="Arthropoda" rank="genus">
<emphasis box="[380,557,1187,1213]" italics="true" pageId="6" pageNumber="241">Celebargiolestes</emphasis>
</taxonomicName>
is characterised by the lower appendages being broad at the base and abruptly constricted forming a slender finger like apex, running to about a third to half the length of the upper appendages. In addition to this the genital ligula has a distinct shape, the lateral lobes are directed in such a way that they cover the opening of the genital ligula. Females characters separating
<taxonomicName authorityName="Kennedy" authorityYear="1925" box="[509,686,1373,1399]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="241" phylum="Arthropoda" rank="genus">
<emphasis box="[509,686,1373,1399]" italics="true" pageId="6" pageNumber="241">Celebargiolestes</emphasis>
</taxonomicName>
from related genera are (i) abdomen subequal to the wings (abdomen clearly longer than wings in most other genera) and (ii) Pt white on both sides in mature individuals (shared with
<taxonomicName authorityName="Tillyard" authorityYear="1908" box="[217,495,1485,1511]" class="Insecta" family="Megapodagrionidae" genus="Miniargiolestes" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="241" phylum="Arthropoda" rank="species" species="minimus">
<emphasis box="[217,495,1485,1511]" italics="true" pageId="6" pageNumber="241">Miniargiolestes minimus</emphasis>
</taxonomicName>
, some species of
<taxonomicName authorityName="Theischinger" authorityYear="1998" box="[694,869,1485,1511]" class="Insecta" family="Megapodagrionidae" genus="Griseargiolestes" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="241" phylum="Arthropoda" rank="genus">
<emphasis box="[694,869,1485,1511]" italics="true" pageId="6" pageNumber="241">Griseargiolestes</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Kennedy" authorityYear="1925" class="Insecta" family="Megapodagrionidae" genus="Austroargiolestes" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="241" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="6" pageNumber="241">Austroargiolestes</emphasis>
</taxonomicName>
(all from Australia) and
<taxonomicName authorityName="Montrousier" authorityYear="1864" box="[534,771,1523,1549]" class="Insecta" family="Megapodagrionidae" genus="Argiolestes" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="241" phylum="Arthropoda" rank="species" species="ochraceus">
<emphasis box="[534,771,1523,1549]" italics="true" pageId="6" pageNumber="241">Argiolestes ochraceus</emphasis>
</taxonomicName>
from New Caledonia).
</paragraph>
<paragraph blockId="7.[159,1036,253,391]" pageId="7" pageNumber="242">
The only other species of
<taxonomicName box="[456,614,253,279]" class="Insecta" family="Argiolestidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="242" phylum="Arthropoda" rank="family">Argiolestidae</taxonomicName>
occurring on Sulawesi is
<taxonomicName authorityName="Kalkman" authorityYear="2007" class="Insecta" family="Megapodagrionidae" genus="Argiolestes" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="242" phylum="Arthropoda" rank="species" species="celebensis">
<emphasis italics="true" pageId="7" pageNumber="242">Argiolestes celebensis</emphasis>
</taxonomicName>
. This species is however easily distinguished from
<taxonomicName authorityName="Kennedy" authorityYear="1925" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="242" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="7" pageNumber="242">Celebargiolestes</emphasis>
</taxonomicName>
by having less than 24 Px and very long and slender apices of the ligula.
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="242" type="key">
<paragraph blockId="7.[401,793,466,494]" box="[401,793,466,494]" pageId="7" pageNumber="242">
<heading bold="true" box="[401,793,466,494]" centered="true" fontSize="11" level="2" pageId="7" pageNumber="242" reason="0">
<emphasis bold="true" box="[401,793,466,494]" pageId="7" pageNumber="242">
Key to males of
<materialsCitation ID-GBIF-Occurrence="1413786765" box="[594,793,466,494]" pageId="7" pageNumber="242">
<taxonomicName authorityName="Kennedy" authorityYear="1925" box="[594,793,466,494]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="242" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[594,793,466,494]" italics="true" pageId="7" pageNumber="242">Celebargiolestes</emphasis>
</taxonomicName>
</materialsCitation>
</emphasis>
</heading>
</paragraph>
<paragraph blockId="7.[159,1036,511,1054]" box="[159,1035,511,538]" pageId="7" pageNumber="242">
1a Labrum yellow to yellow-ochre (
<figureCitation box="[599,728,512,538]" captionStart="Figures 7 14" captionStartId="13.[159,238,1384,1409]" captionTargetBox="[196,998,249,1341]" captionTargetId="figure@13.[196,998,249,1342]" captionTargetPageId="13" captionText="Figures 7 14. Head of Celebargiolestes males. 7 C. cinctus (immature), Ranotongkor, 02 - viii- 1940, JvT 6476; 8 C. cinctus (old adult), Mapanget, 23 - vi- 1940, JvT 6638; 9 C. cinctus ( red legs form), 25 km NE of Luwuk, 30 - i- 1989, JvT 6670; 10 C. askewi sp. nov., holotype; 11 C. toli sp. nov., holotype; 12 C. toli sp. nov., Peleng Island, 05 - viii- 1941, JvT 6453; 13 C. orri sp. nov., holotype; 14 C. orri sp. nov., 10 km WNW Palopo, 29 - x- 1993, JvT 11910." httpUri="https://zenodo.org/record/192855/files/figure.png" pageId="7" pageNumber="242">Figs 1112</figureCitation>
) ................
<emphasis bold="true" box="[865,1035,511,538]" pageId="7" pageNumber="242">
<taxonomicName authority="Vincent J. Kalkman, 2016" authorityName="Vincent J. Kalkman" authorityYear="2016" box="[865,938,511,538]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="242" phylum="Arthropoda" rank="species" species="toli" status="sp. nov.">
<emphasis bold="true" box="[865,938,511,538]" italics="true" pageId="7" pageNumber="242">C. toli</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[945,1035,512,538]" pageId="7" pageNumber="242" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph blockId="7.[159,1036,511,1054]" box="[159,1036,556,583]" pageId="7" pageNumber="242">
1b Labrum black or brownish black (
<figureCitation box="[614,815,556,583]" captionStart="Figures 7 14" captionStartId="13.[159,238,1384,1409]" captionTargetBox="[196,998,249,1341]" captionTargetId="figure@13.[196,998,249,1342]" captionTargetPageId="13" captionText="Figures 7 14. Head of Celebargiolestes males. 7 C. cinctus (immature), Ranotongkor, 02 - viii- 1940, JvT 6476; 8 C. cinctus (old adult), Mapanget, 23 - vi- 1940, JvT 6638; 9 C. cinctus ( red legs form), 25 km NE of Luwuk, 30 - i- 1989, JvT 6670; 10 C. askewi sp. nov., holotype; 11 C. toli sp. nov., holotype; 12 C. toli sp. nov., Peleng Island, 05 - viii- 1941, JvT 6453; 13 C. orri sp. nov., holotype; 14 C. orri sp. nov., 10 km WNW Palopo, 29 - x- 1993, JvT 11910." httpUri="https://zenodo.org/record/192855/files/figure.png" pageId="7" pageNumber="242">Figs 710, 1314</figureCitation>
) ..........................
<emphasis bold="true" box="[1021,1036,557,583]" pageId="7" pageNumber="242">2</emphasis>
</paragraph>
<paragraph blockId="7.[159,1036,511,1054]" pageId="7" pageNumber="242">
2a Postclypeus and frons yellow-brown (
<figureCitation box="[657,762,617,643]" captionStart="Figures 7 14" captionStartId="13.[159,238,1384,1409]" captionTargetBox="[196,998,249,1341]" captionTargetId="figure@13.[196,998,249,1342]" captionTargetPageId="13" captionText="Figures 7 14. Head of Celebargiolestes males. 7 C. cinctus (immature), Ranotongkor, 02 - viii- 1940, JvT 6476; 8 C. cinctus (old adult), Mapanget, 23 - vi- 1940, JvT 6638; 9 C. cinctus ( red legs form), 25 km NE of Luwuk, 30 - i- 1989, JvT 6670; 10 C. askewi sp. nov., holotype; 11 C. toli sp. nov., holotype; 12 C. toli sp. nov., Peleng Island, 05 - viii- 1941, JvT 6453; 13 C. orri sp. nov., holotype; 14 C. orri sp. nov., 10 km WNW Palopo, 29 - x- 1993, JvT 11910." httpUri="https://zenodo.org/record/192855/files/figure.png" pageId="7" pageNumber="242">Figs 79</figureCitation>
). Head of genital ligula with a deep, narrow cleft (
<figureCitation box="[593,724,654,681]" captionStart="Figures 23 27" captionStartId="25.[159,237,1412,1437]" captionTargetBox="[208,986,249,1370]" captionTargetId="figure@25.[208,986,249,1370]" captionTargetPageId="25" captionText="Figures 23 27. Genital ligula of Celebargiolestes males. 23 C. cinctus, Dumoga Bone NP, 04 - vi- 1985, JvT 6486, lateral view; 24 C. cinctus, Dumoga Bone NP, 04 - vi- 1985, JvT 6486, ventral view; 25 C. cinctus red leg, Banggai Peninsula, 08 - x- 1989, latero-ventral view; 26 C. askewi sp. nov., Polewali, 23 - x- 1940, JvT 6636, lateral view; 27 C. askewi sp. nov., Polewali, 23 - x- 1940, JvT 6636, ventral view." httpUri="https://zenodo.org/record/192858/files/figure.png" pageId="7" pageNumber="242">Figs 2325</figureCitation>
) .........................
<taxonomicName authorityName="Selys" authorityYear="1886" box="[918,1035,654,680]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="242" phylum="Arthropoda" rank="species" species="cinctus">
<emphasis bold="true" box="[918,1035,654,680]" italics="true" pageId="7" pageNumber="242">C. cinctus</emphasis>
</taxonomicName>
</paragraph>
<paragraph blockId="7.[159,1036,511,1054]" pageId="7" pageNumber="242">
2b Postclypeus and frons largely black (
<figureCitation box="[659,841,699,725]" captionStart="Figures 7 14" captionStartId="13.[159,238,1384,1409]" captionTargetBox="[196,998,249,1341]" captionTargetId="figure@13.[196,998,249,1342]" captionTargetPageId="13" captionText="Figures 7 14. Head of Celebargiolestes males. 7 C. cinctus (immature), Ranotongkor, 02 - viii- 1940, JvT 6476; 8 C. cinctus (old adult), Mapanget, 23 - vi- 1940, JvT 6638; 9 C. cinctus ( red legs form), 25 km NE of Luwuk, 30 - i- 1989, JvT 6670; 10 C. askewi sp. nov., holotype; 11 C. toli sp. nov., holotype; 12 C. toli sp. nov., Peleng Island, 05 - viii- 1941, JvT 6453; 13 C. orri sp. nov., holotype; 14 C. orri sp. nov., 10 km WNW Palopo, 29 - x- 1993, JvT 11910." httpUri="https://zenodo.org/record/192855/files/figure.png" pageId="7" pageNumber="242">Figs 10, 13, 14</figureCitation>
). Head of genital ligula with a shallow, wide cleft (
<figureCitation box="[682,810,736,762]" captionStart="Figures 23 27" captionStartId="25.[159,237,1412,1437]" captionTargetBox="[208,986,249,1370]" captionTargetId="figure@25.[208,986,249,1370]" captionTargetPageId="25" captionText="Figures 23 27. Genital ligula of Celebargiolestes males. 23 C. cinctus, Dumoga Bone NP, 04 - vi- 1985, JvT 6486, lateral view; 24 C. cinctus, Dumoga Bone NP, 04 - vi- 1985, JvT 6486, ventral view; 25 C. cinctus red leg, Banggai Peninsula, 08 - x- 1989, latero-ventral view; 26 C. askewi sp. nov., Polewali, 23 - x- 1940, JvT 6636, lateral view; 27 C. askewi sp. nov., Polewali, 23 - x- 1940, JvT 6636, ventral view." httpUri="https://zenodo.org/record/192858/files/figure.png" pageId="7" pageNumber="242">Figs 26, 27</figureCitation>
,
<figureCitation box="[822,893,736,762]" captionStart="Figures 28 33" captionStartId="26.[159,237,1353,1378]" captionTargetBox="[156,1036,247,1313]" captionTargetId="figure@26.[156,1036,247,1313]" captionTargetPageId="26" captionText="Figures 28 33. Genital ligula of Celebargiolestes males. 28 C. toli sp. nov., Gunung Watuwila, 02 04 - xi- 1989, JvT 6636, lateral view, lateral lobe deformed and bent inwards; 29 C. toli sp. nov., Gunung Watuwila, 02 04 - xi- 1989, JvT 6636, ventral view; 30 C. toli sp. nov., Banggai Archipelago, Peleng Island, 05 - viii- 1941, JvT 6501, lateral view; 31 C. toli sp. nov., Banggai Archipelago, Peleng Island, 05 - viii- 1941, JvT 6501, ventral view; 32 C. orri sp. nov., Lore Lindu, 17 - xii- 1985, JvT 6444, lateral view; 33 C. orri sp. nov., Lore Lindu, 17 - xii- 1985, JvT 6444, ventral view." httpUri="https://zenodo.org/record/192859/files/figure.png" pageId="7" pageNumber="242">32, 33</figureCitation>
) ..............
<emphasis bold="true" box="[1021,1036,736,762]" pageId="7" pageNumber="242">3</emphasis>
</paragraph>
<paragraph blockId="7.[159,1036,511,1054]" pageId="7" pageNumber="242">
3a Lower appendages short, about ¼ the length of the upper appendages, slightly down-curved and, when seen from lateral, with the tip not reaching the level of the upper appendages (
<figureCitation box="[228,310,908,934]" captionStart="Figures 34 39" captionStartId="27.[159,238,1413,1438]" captionTargetBox="[156,1038,480,1372]" captionTargetId="figure@27.[156,1038,480,1372]" captionTargetPageId="27" captionText="Figures 34 39. Appendages of Celebargiolestes males. 34 C. cinctus, Ranotongkor, 02 - viii- 1940, JvT 6476, dorsal view; 35 C. cinctus, Ranotongkor, 02 - viii- 1940, JvT 6476, lateral view; 36 Ranotongkor, 02 - viii- 1940, JvT 6476, ventral view; 37 C. askewi sp. nov., holotype, dorsal view; 38 C. askewi sp. nov., holotype, lateral view; 39 C. askewi sp. nov., holotype, ventral view." httpUri="https://zenodo.org/record/192860/files/figure.png" pageId="7" pageNumber="242">Fig. 38</figureCitation>
) .........................................................................
<emphasis bold="true" box="[823,1035,908,935]" pageId="7" pageNumber="242">
<taxonomicName authority="Vincent J. Kalkman, 2016" authorityName="Vincent J. Kalkman" authorityYear="2016" box="[823,938,908,935]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="242" phylum="Arthropoda" rank="species" species="askewi" status="sp. nov.">
<emphasis bold="true" box="[823,938,908,935]" italics="true" pageId="7" pageNumber="242">C. askewi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[945,1035,908,934]" pageId="7" pageNumber="242" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph blockId="7.[159,1036,511,1054]" pageId="7" pageNumber="242">
3b Lower appendages long, about ½ the length of the upper appendages and, when seen from lateral, with the tip reaching the level of the upper appendages (
<figureCitation box="[624,706,1028,1054]" captionStart="Figures 40 45" captionStartId="28.[159,238,1442,1467]" captionTargetBox="[156,1039,522,1404]" captionTargetId="figure@28.[156,1039,522,1404]" captionTargetPageId="28" captionText="Figures 40 45. Appendages of Celebargiolestes males. 40 C. toli sp. nov., holotype, dorsal view; 41 C. toli sp. nov., holotype, lateral view; 42 C. toli sp. nov., holotype, ventral view; 43 C. orri sp. nov., holotype, dorsal view; 44 C. orri sp. nov., holotype, lateral view; 45 C. orri sp. nov., JvT 6646, ventral view." httpUri="https://zenodo.org/record/192861/files/figure.png" pageId="7" pageNumber="242">Fig. 44</figureCitation>
) ..................
<emphasis bold="true" box="[858,1035,1028,1054]" pageId="7" pageNumber="242">
<taxonomicName authority="Vincent J. Kalkman, 2016" authorityName="Vincent J. Kalkman" authorityYear="2016" box="[858,939,1028,1054]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="242" phylum="Arthropoda" rank="species" species="orri" status="sp. nov.">
<emphasis bold="true" box="[858,939,1028,1054]" italics="true" pageId="7" pageNumber="242">C. orri</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[945,1035,1028,1054]" pageId="7" pageNumber="242" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>