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<document id="AC9AF8A990BA55AD848296D8687B570F" ID-CLB-Dataset="308564" ID-DOI="10.3897/mycokeys.115.145774" ID-GBIF-Dataset="e32a8f6b-dbe4-4c70-ad1e-37e2dd49da60" ID-publisher-id="145774" URI-arpha="84E228B6-3D1B-55CC-9A93-8533E8894BFA" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" article-type="research-article" checkinTime="1741792473941" checkinUser="pensoft" docAuthor="Yan, Jun, Xiong, Li, Yang, Li-Xun, He, Zheng-Mi, Zhang, Ping &amp; Liao, Ke" docDate="2025" docId="03C72319818355D98A847886304EEAF3" docLanguage="en" docName="MycoKeys 115: 137-153" docOrigin="MycoKeys 115" docSource="https://mycokeys.pensoft.net/article/145774/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.mycokeys" docStyleName="PensoftTaxPub.0000.journal_article.mycokeys" docTitle="Clavaria orientalis P. Zhang &amp; Ju. Yan 2025, sp. nov." docType="treatment" docVersion="5" dtd-version="3.0" lastPageNumber="153" masterDocId="84E228B63D1B55CC9A938533E8894BFA" masterDocTitle="Morphological and multi-locus phylogenetic analyses reveal three new branched species of Clavaria (Clavariaceae, Agaricales) from China" masterLastPageNumber="153" masterPageNumber="137" pageNumber="137" updateTime="1741798939028" updateUser="admin">
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<mods:title id="2749A9386C7AFB9DD2F10CF75E552F31">Morphological and multi-locus phylogenetic analyses reveal three new branched species of Clavaria (Clavariaceae, Agaricales) from China</mods:title>
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<mods:namePart id="8A517081CCB2CA78C15380B0E83E8784">Yan, Jun</mods:namePart>
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<mods:namePart id="1ECFA9A7DBF10E86D77713A5A17841A7">Xiong, Li</mods:namePart>
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<mods:affiliation id="B8658E94FE40531360E800D65C6FF08E">Hunan Provincial Demonstration Center of Forestry Seeding Breeding, Changsha 410329, China</mods:affiliation>
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<mods:affiliation id="12CB4D7A5B4E72D61230362AE40C1D39">Bureau of Forestry, Tongdao Dong Autonomous County, Huaihua 418500, China</mods:affiliation>
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<mods:namePart id="61F0E41D2145441A60C2E4BA9FE86DA1">Zhang, Ping</mods:namePart>
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<mods:namePart id="5E1C376D38F7D965161FA1E6F7CF4C16">Liao, Ke</mods:namePart>
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<mods:date id="590C690765E3E8A4F9CAD82D0B6E4E6E">2025</mods:date>
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<mods:identifier id="9B9DA3121F59F28FD2644A78C0D7EC4C" type="DOI">10.3897/mycokeys.115.145774</mods:identifier>
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<treatment id="03C72319818355D98A847886304EEAF3" ID-DOI="http://doi.org/10.5281/zenodo.15013465" ID-GBIF-Taxon="255440943" ID-Zenodo-Dep="15013465" ID-arpha="03C72319-8183-55D9-8A84-7886304EEAF3" ID-mycobank="857601" LSID="urn:lsid:plazi:treatment:03C72319818355D98A847886304EEAF3" httpUri="http://treatment.plazi.org/id/03C72319818355D98A847886304EEAF3" scope_family="Clavariaceae" scope_order="Agaricales">
<subSubSection id="A19D1DA2D9196158ED1E81062BE6A7F2" type="nomenclature">
<paragraph id="785B03C04FEFDB774138CF9933126BC2">
<taxonomicName id="6BE4F339E585A9BA11F55EDB84B4F657" authority="P. Zhang &amp; Ju. Yan" authorityName="P. Zhang &amp; Ju. Yan" authorityYear="2025" class="Agaricomycetes" family="Clavariaceae" genus="Clavaria" kingdom="Fungi" order="Agaricales" phylum="Basidiomycota" rank="species" species="orientalis" status="sp. nov.">
<emphasis id="02395B4F9DEAFCFE48AC72A347290B41" italics="true">Clavaria orientalis</emphasis>
P. Zhang &amp; Ju. Yan
</taxonomicName>
<taxonomicNameLabel id="EF61D4F0E979DA4EE61F27C104C8A7DF" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="BA7DA93CAFC5761936FAF2C7D16A8304" type="description">
<paragraph id="F513EF5C7F8CF5CF1E478EB7EBF408B6">
<figureCitation id="E0B189CC3FE462D4BAA53EF61F47BC1C" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Basidiomata of Clavaria orientalis a MHHNU 7767 b MHHNU 32116. Scale bars: 2 cm." figureDoi="10.3897/mycokeys.115.145774.figure4" httpUri="https://binary.pensoft.net/fig/1278626">Figs 4</figureCitation>
,
<figureCitation id="FF3A96401958FB6285213BC85B43AAD6" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Microscopic features of Clavaria orientalis (MHHNU 7767) a basidiospores b basidia." figureDoi="10.3897/mycokeys.115.145774.figure5" httpUri="https://binary.pensoft.net/fig/1278627">5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EWJCI" type="etymology">
<paragraph id="8121D73C40D392FB175C49E3139C083E">
<heading id="B0306864162A1FB9709FBEB318DE3306" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="9763787629381B412141D70AE0C78FE4">
<taxonomicName id="A6143239A77DCA865C2C9DBCBE9F544D" authorityName="P. Zhang &amp; Ju. Yan" authorityYear="2025" class="Agaricomycetes" family="Clavariaceae" genus="Clavaria" kingdom="Fungi" order="Agaricales" phylum="Basidiomycota" rank="species" species="orientalis">
<emphasis id="87E0F69993F004CA82EA0C13FDB78244" italics="true">orientalis</emphasis>
</taxonomicName>
(Latin), meaning eastern, refers to the occurrence of the species in
<collectingCountry id="08C312389DE1F697D57A52EA78D0EDED" name="China">East Asia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EGKCI" type="Holotype">
<paragraph id="55AA488BCB164DE669B4530A00310345">
<heading id="E8C8B9339FFA2B8D543EBA5629D6A116" reason="title">
<typeStatus id="6DE32AFAC01C438A5A6DA5C6DAB04AFB" type="holotype">Holotype</typeStatus>
.
</heading>
</paragraph>
<paragraph id="1271D00E9CB198CDFF482AADED5FD872">
<materialsCitation id="79E29487E6C78E7262E0510E0630AAD8" ID-GBIF-Occurrence="5082333310" collectingDate="2012-09-11" collectionCode="MHHNU" collectorName="P. Zhang" country="China" county="Shimen County" elevation="1828" latitude="30.049583" location="Hupingshan Nature Reserve" longLatPrecision="1" longitude="110.52359" specimenCode="MHHNU 7767" specimenCount="1" stateProvince="Hunan Province" typeStatus="holotype">
<collectingCountry id="FD755C5D4AE6ED62AC2C0F63D002AA81" name="China">China</collectingCountry>
<collectingRegion id="AFD6A473AF4434E3E0FDA6C3B7C9D3A7" country="China" name="Hunan">Hunan Province</collectingRegion>
:
<collectingCounty id="1B1936CED307379D5240269926B63739">Shimen County</collectingCounty>
,
<location id="3C3257C7E5CC77E0CC14D3F86BCC03A0" LSID="urn:lsid:plazi:treatment:03C72319818355D98A847886304EEAF3:3C3257C7E5CC77E0CC14D3F86BCC03A0" country="China" county="Shimen County" latitude="30.049583" longLatPrecision="1" longitude="110.52359" name="Hupingshan Nature Reserve" stateProvince="Hunan Province">Hupingshan Nature Reserve</location>
, alt.
<quantity id="9F4314661BB60782D1903AAD2B7183CC" metricMagnitude="3" metricUnit="m" metricValue="1.828" unit="m" value="1828.0">
<elevation id="EF345EB4C6195C54F463B24A78CF5C8B" metricMagnitude="3" metricUnit="m" metricValue="1.828" unit="m" value="1828.0">1828 m</elevation>
</quantity>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EPKCI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[110.523583,30.049583]}">
<geoCoordinate id="564F4C4D4D724328AD0BB17A1EBD1417" degrees="30" direction="north" minutes="02" orientation="latitude" precision="1" seconds="58.50" value="30.049583">30°02'58.50&quot;N</geoCoordinate>
,
<geoCoordinate id="0ED22F51ED9AA5557B060C6C1B3EAD98" degrees="110" direction="east" minutes="31" orientation="longitude" precision="1" seconds="24.90" value="110.52359">110°31'24.90&quot;E</geoCoordinate>
</named-content>
,
<date id="4D0780EBB23EB34E25BEFD65280E73CE" value="2012-09-11">
<collectingDate id="1D33E9BC5A4040CE7CD92FFE4E64D2EB" value="2012-09-11">11 September 2012</collectingDate>
</date>
,
<collectorName id="AF83D4E169F3B761581F7D0E8B122A41">P. Zhang</collectorName>
(
<specimenCode id="B98380C20C696C64A803B88403A60A6B" collectionCode="MHHNU">MHHNU 7767</specimenCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EUKCI" type="diagnosis">
<paragraph id="01E54BDF734FA9193DEC62D12B594340">
<heading id="393EE58822478D73591868EE92D46037" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="18332F10095D26A2799BB9F28A970029">
Differs from
<taxonomicName id="92CFF71C1CA9CE379F28FB2C0B5182E7" class="Agaricomycetes" family="Clavariaceae" genus="Clavaria" kingdom="Fungi" order="Agaricales" phylum="Basidiomycota" rank="species" species="zollingeri">
<emphasis id="DB2E2812EE7B3F097598A4AD2FC0FAAC" italics="true">Clavaria zollingeri</emphasis>
</taxonomicName>
in its stout branches, lesser degree of branching, and shorter basidia.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EFLCI" type="description">
<paragraph id="B6869B7AFD9E5E242ED1F236FB85F918">
<heading id="55875F1D286BEE4FB1CE30F923C6735F" reason="title">Description.</heading>
</paragraph>
<paragraph id="72E56757BD1CF4B4F23577337B0FA004">
Basidiomata (Fig.
<figureCitation id="6AF4B69AB97C56B81EF6B01BBB985DB6" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Basidiomata of Clavaria orientalis a MHHNU 7767 b MHHNU 32116. Scale bars: 2 cm." figureDoi="10.3897/mycokeys.115.145774.figure4" httpUri="https://binary.pensoft.net/fig/1278626">4 a, b</figureCitation>
) branched, brittle, gregarious to caespitose clusters; clusters
<quantity id="56D081211C75F39C08FB1D67D2B69E3C" metricMagnitude="-2" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" unit="mm" value="65.0" valueMax="80.0" valueMin="50.0">5080 mm</quantity>
high,
<quantity id="0F7E7B1A6C56FEBD7D9EA069D88EA407" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" unit="mm" value="20.0" valueMax="30.0" valueMin="10.0">1030 mm</quantity>
broad; branches terete,
<quantity id="2B0CC1773FBC35E8CD8A7CB97BAA7A88" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" unit="mm" value="2.0" valueMax="3.0" valueMin="1.0">13 mm</quantity>
wide, 14 times, dichotomous; branch tips obtuse, broadly rounded, or narrowly rounded. Fertile part coralloid, smooth, obviously curved or slightly twisted, deep amethyst [15 A 4-7, 15 B 4-6, 16 A 4-6; Amparo Purple, Lobelia Violet, Vinaceous Purple] to lilac [14 A 2-3, 15 A 2-3; Pale Vinaceous Purple, Pale Lobelia Violet], and changing to pale greyish purple [16 A 2, 17 A 2; Lavender Gray, Pale Paynes Gray, Pale Verbena Violet] with age. Apex concolorous with lower part, becoming yellowish or tawny with age. Stipe distinct, sterile, smooth, often terete, semi-translucent, hygrophanous, and darker than the fertile part, sometimes flattened and pallid. Flesh concolorous or slightly paler than surface of basidiomata.
</paragraph>
<caption id="70DF8A40E25CA99B0947A6EC6732E147" ID-DOI="10.3897/mycokeys.115.145774.figure4" ID-arpha="171B2F58-D36F-573D-BFE1-B7E43A64D1F0" httpUri="https://binary.pensoft.net/fig/1278626" startId="F4">
<paragraph id="53ECD997C3426D9FE8B5D24AA2AD01F8">
<label id="C0DF911386E4FD0EF38E73E3479262AF">Figure 4.</label>
</paragraph>
<paragraph id="104799059116CA6C9A99723293EC2E9F">
Basidiomata of
<taxonomicName id="8165AD2C03A7386D213D890DBD103D3A" authorityName="P. Zhang &amp; Ju. Yan" authorityYear="2025" class="Agaricomycetes" family="Clavariaceae" genus="Clavaria" kingdom="Fungi" order="Agaricales" phylum="Basidiomycota" rank="species" species="orientalis">
<emphasis id="6CB02E52AA12A346A3E655D90D5BDF5D" italics="true">Clavaria orientalis</emphasis>
</taxonomicName>
<emphasis id="62A158BE7851A7B14E440F5E8BA26E3D" bold="true">a</emphasis>
MHHNU 7767
<emphasis id="2141309F282682B7A7C08D478D9B48E8" bold="true">b</emphasis>
MHHNU 32116. Scale bars: 2 cm.
</paragraph>
</caption>
<paragraph id="6779B1241E850560FA38A190E78F0C9E">
Basidiospores (Fig.
<figureCitation id="C907C813536A668AE375D8C5AA917CFB" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Microscopic features of Clavaria orientalis (MHHNU 7767) a basidiospores b basidia." figureDoi="10.3897/mycokeys.115.145774.figure5" httpUri="https://binary.pensoft.net/fig/1278627">5 a</figureCitation>
) [100 / 5 / 5] (4.8) 5.06.0 × 4.05.0 (5.5) μm [Q = (1.09) 1.131.38 (1.43), Q = 1.21 ± 0.11], mostly broadly ellipsoid, sometimes ellipsoid or subglobose, smooth, hyaline, nonamyloid, thin-walled; hilar appendage present (&lt;2.0 μm in length). Basidia (Fig.
<figureCitation id="389117AFAE750C2CBF5E95CF31BA4E92" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Microscopic features of Clavaria orientalis (MHHNU 7767) a basidiospores b basidia." figureDoi="10.3897/mycokeys.115.145774.figure5" httpUri="https://binary.pensoft.net/fig/1278627">5 b</figureCitation>
) 3448 × 5.08.0 μm, clavate, 4 - spored, hyaline, thin-walled; sterigmata below 5.0 μm long. Incrustations or crystals absent. Hyphae of the context parallel, thin-walled, hyaline, cylindrical to inflated, secondarily septated. Clamp connections absent in all parts of basidiomata.
</paragraph>
<caption id="F3D1470E7C92B3863594181C4BE25179" ID-DOI="10.3897/mycokeys.115.145774.figure5" ID-arpha="2EF94FAB-711F-5ADF-887D-8C9A1E229B4C" httpUri="https://binary.pensoft.net/fig/1278627" startId="F5">
<paragraph id="A402842CA0C44FC4DCFE2D57716ECD67">
<label id="757E01692816837E2260F67485BEAE2E">Figure 5.</label>
</paragraph>
<paragraph id="1AC11BF191F74F852723BF97857A23C8">
Microscopic features of
<taxonomicName id="90394AFA95C66911BBE7F9A032C9E24D" authorityName="P. Zhang &amp; Ju. Yan" authorityYear="2025" class="Agaricomycetes" family="Clavariaceae" genus="Clavaria" kingdom="Fungi" order="Agaricales" phylum="Basidiomycota" rank="species" species="orientalis">
<emphasis id="BD478D219379B52CD7EA06EC9DE01F21" italics="true">Clavaria orientalis</emphasis>
</taxonomicName>
(MHHNU 7767)
<emphasis id="B5D500BE08F87FC5EF3793101C83092D" bold="true">a</emphasis>
basidiospores
<emphasis id="A484BE569626D22F5BE6AB0B811536A2" bold="true">b</emphasis>
basidia.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0ERNCI" type="habitat">
<paragraph id="1773C9599B13678057B7F9A354BB8BBE">
<heading id="42CCA4CBD440871818E68176D581B1B0" reason="title">Habitat, ecology, and distribution.</heading>
</paragraph>
<paragraph id="278D0C3FEB537B0296671B7D2DED2E56">
Gregarious to caespitose in humus layers of soil under broadleaved forests, coniferous forests, or mixed coniferous broadleaved forests. Basidiomata produced in summer or autumn, usually throughout the months of July to September; known from subtropical zones of southern
<collectingCountry id="C31DED3CEE7F3587B0113D73DE966414" name="China">China</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EWNCI" type="material">
<paragraph id="5E46240287670FBD0405FC5A87EEB9DF">
<heading id="D76C7A24BA805120C20C362A08EB16BA" reason="title">Additional specimens examined.</heading>
</paragraph>
<paragraph id="E06CB2FE50AF356EE75A1593BBFBA7CF">
<materialsCitation id="AED60E83BE106BED3A4F3CB3FA368415" ID-GBIF-Occurrence="5082333302" collectingDate="2003-07-22" collectionCode="MHHNU" collectorName="P. Zhang" country="China" county="Sangzhi County" elevation="1500" latitude="29.782825" location="Badagongshan National Nature Reserve" longLatPrecision="1" longitude="110.081024" specimenCode="MHHNU 6801" specimenCount="1" stateProvince="Hunan Province">
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:
<collectingCounty id="2F0013DA242BE2A5E55DCC4F07530475">Sangzhi County</collectingCounty>
,
<location id="0968844231A2DCD7089DB35C77B19860" LSID="urn:lsid:plazi:treatment:03C72319818355D98A847886304EEAF3:0968844231A2DCD7089DB35C77B19860" country="China" county="Sangzhi County" latitude="29.782825" longLatPrecision="1" longitude="110.081024" name="Badagongshan National Nature Reserve" stateProvince="Hunan Province">Badagongshan National Nature Reserve</location>
, alt.
<quantity id="C9CBA1A20FEE1F6333A36079447BE8E3" metricMagnitude="3" metricUnit="m" metricValue="1.5" unit="m" value="1500.0">
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<geoCoordinate id="6A3E1839FFDEC83882C20DCE69722545" degrees="29" direction="north" minutes="46" orientation="latitude" precision="1" seconds="58.17" value="29.782825">29°46'58.17&quot;N</geoCoordinate>
,
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,
<date id="22B284E98E15AE68CA2A41CACDA64FBD" value="2003-07-22">
<collectingDate id="61C03C3D5D656A1E75AA79DFD1D134F6" value="2003-07-22">22 July 2003</collectingDate>
</date>
,
<collectorName id="3C0F26B6FC348419FEB8F67E38EBE360">P. Zhang</collectorName>
(
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</materialsCitation>
; •
<materialsCitation id="1D80612F0D86E01615786133D6A65796" ID-GBIF-Occurrence="5082333308" collectingDate="2010-08-31" collectionCode="MHHNU" collectorName="P. Zhang" country="China" county="Shimen County" elevation="1828" latitude="30.049583" location="Hupingshan Nature Reserve" longLatPrecision="1" longitude="110.52359" specimenCode="MHHNU 7352" specimenCount="1" stateProvince="Hunan">
<collectingCounty id="178BA5FDD5844D152D0D0B4AE990D172">Shimen County</collectingCounty>
,
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, alt.
<quantity id="01EB3AD5ECCB633C4A8117BDFA95B1D4" metricMagnitude="3" metricUnit="m" metricValue="1.828" unit="m" value="1828.0">
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<date id="6D679FA865E5682341854D39E3D32FDE" value="2010-08-31">
<collectingDate id="275A0FA4C50BCFC9B68BAE097819059D" value="2010-08-31">31 August 2010</collectingDate>
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,
<collectorName id="6EC042F63EE86B8B2DBFBF02BB7DB7BA">P. Zhang</collectorName>
(
<specimenCode id="40C24F04069870F6283BD8E293940C1C" collectionCode="MHHNU">MHHNU 7352</specimenCode>
)
</materialsCitation>
; •
<materialsCitation id="EAA4BC0CC3E117FB4681FE6F7631360C" ID-GBIF-Occurrence="5082333307" collectingDate="2011-09-26" collectionCode="MHHNU" collectorName="P. Zhang" country="China" county="Shimen County" elevation="1828" latitude="30.049583" location="Hupingshan Nature Reserve" longLatPrecision="1" longitude="110.52359" specimenCode="MHHNU 7586" specimenCount="1" stateProvince="Hunan">
<date id="5B5AF59BAA955ED3DCF5659329D0F30A" value="2011-09-26">
<collectingDate id="066C009B2B925DAFAF4472A13B37C3CA" value="2011-09-26">26 September 2011</collectingDate>
</date>
,
<collectorName id="C59E05F83F09499691101173EE94994B">P. Zhang</collectorName>
(
<specimenCode id="767515C9C16A35E09288634BDBB4D535" collectionCode="MHHNU">MHHNU 7586</specimenCode>
)
</materialsCitation>
. •
<materialsCitation id="B9A48D1F6B406FF9A227C86588987A98" ID-GBIF-Occurrence="5082333309" collectingDate="2020-08-01" collectionCode="MHHNU" collectorName="Z. H. Chen" country="China" county="Hefeng County" elevation="1413" latitude="30.058935" location="Mulinzi National Nature Reserve" longLatPrecision="1" longitude="110.20954" specimenCode="MHHNU 32116" specimenCount="1" stateProvince="Hubei Province">
<collectingRegion id="6C2A2EF31DEF01611067B700DD0CAA68" country="China" name="Hubei">Hubei Province</collectingRegion>
:
<collectingCounty id="DAD41754E0F99F7825D2B6EC346213B1">Hefeng County</collectingCounty>
,
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, alt.
<quantity id="68E69ECE4A1DDCF612D59336BBB7B6E1" metricMagnitude="3" metricUnit="m" metricValue="1.413" unit="m" value="1413.0">
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,
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,
<geoCoordinate id="B3382DD6F9115F16A35ED640C2AF59D7" degrees="110" direction="east" minutes="12" orientation="longitude" precision="1" seconds="34.35" value="110.20954">110°12'34.35&quot;E</geoCoordinate>
</named-content>
,
<date id="F3E857079CAEE03581AF3522057FE824" value="2020-08-01">
<collectingDate id="A4C54C57671C257BD4AD25D81265372A" value="2020-08-01">1 August 2020</collectingDate>
</date>
,
<collectorName id="1A56486D4A59D8109C14C3A77BB4E49D">Z. H. Chen</collectorName>
(
<specimenCode id="F7DA67555137E9BF95ACD37EBE2FEEB6" collectionCode="MHHNU">MHHNU 32116</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>