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<mods:title id="17F7E77B21B028B685F0A0A593053233">A key to European species of Hyposoter Förster, 1869 (Ichneumonidae: Campopleginae) with descriptions of 18 new species, and notes on all included species</mods:title>
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<mods:namePart id="5EAD41668B63E37999C97DFCBDEA9F35">Galsworthy, Anthony</mods:namePart>
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<mods:namePart id="8154815C77399268E31D83790FBC2C54">Shaw, Mark R.</mods:namePart>
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<subSubSection id="26966522FFBAFFD9FF7E40F3909860F0" box="[151,764,1224,1251]" pageId="36" pageNumber="37" type="nomenclature">
<paragraph id="6E3336A9FFBAFFD9FF7E40F3909860F0" blockId="36.[151,764,1224,1287]" box="[151,764,1224,1251]" pageId="36" pageNumber="37">
<heading id="357B81C5FFBAFFD9FF7E40F3909860F0" bold="true" box="[151,764,1224,1251]" fontSize="11" level="2" pageId="36" pageNumber="37" reason="4">
<emphasis id="5CF8EABBFFBAFFD9FF7E40F3909860F0" bold="true" box="[151,764,1224,1251]" pageId="36" pageNumber="37">
<taxonomicName id="A98C4D2AFFBAFFD9FF7E40F390F260F1" authority="Galsworthy &amp; Shaw" authorityName="Galsworthy &amp; Shaw &amp; Haraldseide" authorityYear="2023" box="[151,662,1224,1251]" class="Insecta" family="Ichneumonidae" genus="Hyposoter" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="castaneus" status="sp. nov.">
<emphasis id="5CF8EABBFFBAFFD9FF7E40F393F560F0" bold="true" box="[151,401,1225,1251]" italics="true" pageId="36" pageNumber="37">Hyposoter castaneus</emphasis>
Galsworthy &amp; Shaw
</taxonomicName>
<taxonomicNameLabel id="47CB57C0FFBAFFD9FD7740F3909860F0" box="[670,764,1225,1251]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="26966522FFBAFFD9FF7E40D693766114" box="[151,274,1260,1287]" pageId="36" pageNumber="37" type="description">
<paragraph id="6E3336A9FFBAFFD9FF7E40D693766114" blockId="36.[151,764,1224,1287]" box="[151,274,1260,1287]" pageId="36" pageNumber="37">
(
<figureCitation id="F6B72A2CFFBAFFD9FF7640D6936D6114" box="[159,265,1260,1287]" captionStart="FIGURES 15" captionStartId="32.[152,270,1370,1394]" captionTargetBox="[151,1436,181,1346]" captionTargetId="figure-22@32.[151,1436,181,1346]" captionTargetPageId="32" captionText="FIGURES 15. 1,2: Hyposoter aglyphus sp. n. Habitus: 1 lateral, 2 dorsal. 35: Hyposoter castaneus sp. n. Habitus: 3 lateral, 4 dorsal; 5 detail of mesopleuron." figureDoi="http://doi.org/10.5281/zenodo.7959260" httpUri="https://zenodo.org/record/7959260/files/figure.png" pageId="36" pageNumber="37">Figs. 35</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="26966522FFBAFFD9FF7E410E963C61CD" pageId="36" pageNumber="37" type="diagnosis">
<paragraph id="6E3336A9FFBAFFD9FF7E410E963C61CD" blockId="36.[151,1437,1332,2042]" pageId="36" pageNumber="37">
<emphasis id="5CF8EABBFFBAFFD9FF7E410E9376615D" bold="true" box="[151,274,1332,1358]" pageId="36" pageNumber="37">Diagnosis.</emphasis>
Most specimens have the metasoma darkish brown, which is unusual in the genus and an immediately obvious distinguishing feature; but the metasoma can be more orange, and the species is therefore included twice in the key. Apart from the key characters, the species is distinguished by an unusually broad clypeus and face, which separate it readily from the otherwise quite similar (in some colour forms)
<emphasis id="5CF8EABBFFBAFFD9FC37419B960D61A9" box="[990,1129,1440,1466]" italics="true" pageId="36" pageNumber="37">
<taxonomicName id="A98C4D2AFFBAFFD9FC37419B960161A9" authorityName="Gravenhorst" authorityYear="1829" box="[990,1125,1440,1466]" class="Insecta" family="Ichneumonidae" genus="Campoplex" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="caedator">H. caedator</taxonomicName>
.
</emphasis>
The sculpture of the mesopleuron, with clear transverse rugae in front of a smooth speculum, is also distinctive.
</paragraph>
</subSubSection>
<subSubSection id="26966522FFBAFFD8FF2E41D29060672A" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37" type="description">
<paragraph id="6E3336A9FFBAFFD9FF2E41D290A962A5" blockId="36.[151,1437,1332,2042]" pageId="36" pageNumber="37">
<emphasis id="5CF8EABBFFBAFFD9FF2E41D2933D6211" bold="true" box="[199,345,1512,1538]" pageId="36" pageNumber="37">Description.</emphasis>
Fore wing
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. Head: face twice as broad as long, rugosely sculptured throughout, without evident punctures; clypeus short, curved at sides, similarly sculptured, with narrow, sharp, unsculptured apex; malar space equal to basal width of mandible; mandible tapering, with ventral flange narrow; genal carina complete, joining oral carina just short of base of mandible; frons and vertex rugosely sculptured, but less strongly than face; temples sharply narrowed behind eyes, very short, similarly sculptured; antenna slender, flagellum with 2425 flagellomeres, first flagellomere 2.5x as long as wide.
</paragraph>
<paragraph id="6E3336A9FFBAFFD9FF2E42FB939B63A1" blockId="36.[151,1437,1332,2042]" pageId="36" pageNumber="37">Mesosoma: epomia present; mesoscutum centrally with close-set deep punctures on a granulate background, lateral and posterior parts becoming strongly rugose; mesopleuron shining, but with faint linear sculpture throughout; groove in front of fovea strongly striate; anterior and ventral areas, and metapleuron, with strong, evenly spaced punctures on a granulate background; propodeum rather short and rounded; area basalis narrow, V-shaped; area superomedia closed, broader than long, 5-sided, with front lateral edges much longer than rear ones; costulae complete; area petiolaris broadly triangular; whole propodeum rugosely sculptured, most strongly on petiolar area; propodeal spiracle slightly oval.</paragraph>
<paragraph id="6E3336A9FFBAFFD9FF2E438690D163E9" blockId="36.[151,1437,1332,2042]" pageId="36" pageNumber="37">
Wings narrow, 1
<emphasis id="5CF8EABBFFBAFFD9FE6A438793D163C5" box="[387,437,1981,2006]" italics="true" pageId="36" pageNumber="37">cu-a</emphasis>
interstitial, areolet almost sessile to short-stalked with vein 2
<emphasis id="5CF8EABBFFBAFFD9FB98438796C063C5" box="[1137,1188,1981,2007]" italics="true" pageId="36" pageNumber="37">rs -m</emphasis>
entering well beyond middle, nervellus gently curved, almost vertical.
</paragraph>
<paragraph id="6E3336A9FFBBFFD8FF2E44AD976A6552" blockId="37.[151,1437,151,825]" pageId="37" pageNumber="38">Metasoma: first tergite with petiole narrowed before postpetiole, and ventral edge sinuate in lateral view; glymma obvious, but shallow; dorsolateral carinae present, but weak; postpetiole triangular, faintly granulate, with symmetrical depressions above and behind spiracles; tergite 2 strongly granulate to two thirds, its posterior part and other tergites only faintly sculptured, and with scattered fine punctures; tergite 6 with posterior border entire; ovipositor slightly longer than apical depth of metasoma, almost straight, with strong subapical notch.</paragraph>
<paragraph id="6E3336A9FFBBFFD8FF2E4571918965C2" blockId="37.[151,1437,151,825]" pageId="37" pageNumber="38">Colour: black; apex of underside of scape, mandibles other than extreme base and teeth (which are brown), tegulae, 2nd metasomal sternite yellow; femora, tibiae and fore and mid tarsi orange; hind tarsus brown, basitarsus pale at base; tergite 1 black; tergite 2 black, with posterior edge sometimes orange, tergite 3 orange to brown with orange wash; tergite 4 orange to pale brown; tergites 5 onwards dark brown.</paragraph>
<paragraph id="6E3336A9FFBBFFD8FF2E45E193B46706" blockId="37.[151,1437,151,825]" pageId="37" pageNumber="38">
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<typeStatus id="B137880BFFBBFFD8FF2E45E1935065E6" box="[199,308,475,501]" pageId="37" pageNumber="38" type="holotype">Holotype</typeStatus>
:
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, [UK]
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,
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, TL818883,
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: heath with birch and pine, 25.7 5.8.83,
<collectorName id="C379537FFFBBFFD8FF0545C5935E660A" box="[236,314,511,537]" pageId="37" pageNumber="38">J.Field</collectorName>
, RMSNH 1986.021 (
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);
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<typeStatus id="B137880BFFBBFFD8FD6845C5909A660A" box="[641,766,511,537]" pageId="37" pageNumber="38" type="paratype">paratypes</typeStatus>
:
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,
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,
<collectorName id="C379537FFFBBFFD8FC1C45C59610660A" box="[1013,1140,510,537]" pageId="37" pageNumber="38">Cambs, TL</collectorName>
650693,
<collectingMethod id="B7CD4EBEFFBBFFD8FB3C45C59702660A" box="[1237,1382,511,537]" pageId="37" pageNumber="38">Malaise trap</collectingMethod>
: carr at reedbed edge, 1624.6.83,
<collectorName id="C379537FFFBBFFD8FE024619905D662E" box="[491,569,547,573]" pageId="37" pageNumber="38">J.Field</collectorName>
, RMSNH 1986.021;
</materialsCitation>
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<specimenCount id="788AFD20FFBBFFD8FCC746199135662E" box="[814,849,547,573]" pageId="37" pageNumber="38" type="female">1♀</specimenCount>
,
<location id="6B536072FFBBFFD8FC8B461991DB662E" LSID="urn:lsid:plazi:treatment:E62587BFFFBAFFD8FF7E40F39060672A:6B536072FFBBFFD8FC8B461991DB662E" box="[866,959,547,573]" country="Germany" county="Matthias Riedel" municipality="Augsburg" name="Catfield" pageId="37" pageNumber="38" stateProvince="Norfolk">Catfield</location>
,
<collectingRegion id="AC48F84BFFBBFFD8FC244619964D662E" box="[973,1065,547,573]" country="United Kingdom" name="Norfolk" pageId="37" pageNumber="38">Norfolk</collectingRegion>
, TG379201,
<collectingMethod id="B7CD4EBEFFBBFFD8FB5646199735662E" box="[1215,1361,547,573]" pageId="37" pageNumber="38">Malaise trap</collectingMethod>
: abandoned wet meadow,
<collectorName id="C379537FFFBBFFD8FE94467D90666672" box="[381,514,583,609]" pageId="37" pageNumber="38">R.T.J.Jarvis</collectorName>
, 27.7.10.8.84 RSMNH 1983.102 (all in
<collectionCode id="089DAE6CFFBBFFD8FC38467D966B6672" box="[977,1039,583,609]" country="United Kingdom" httpUri="http://grbio.org/cool/h8hj-4rxx" name="National Museum of Scotland - Natural Sciences" pageId="37" pageNumber="38" type="Museum">NMS</collectionCode>
);
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<materialsCitation id="DEE43CF4FFBBFFD8FBCF467D91896696" ID-GBIF-Occurrence="4116948327" collectingDate="1959-07-04" collectingDateMax="1969-09-01" collectingDateMin="1959-07-04" collectionCode="NMS, NHMUK, ZSM" collectorName="A. C. Galsworthy" country="Germany" county="Matthias Riedel" determinerName="Riedel" location="Santon Downham" municipality="Augsburg" pageId="37" pageNumber="38" specimenCount="12" specimenCount-female="12" stateProvince="Norfolk" typeStatus="paratype">
<specimenCount id="788AFD20FFBBFFD8FBCF467D962E6672" box="[1062,1098,583,609]" pageId="37" pageNumber="38" type="female">2♀</specimenCount>
,
<location id="6B536072FFBBFFD8FBBE467D96826672" LSID="urn:lsid:plazi:treatment:E62587BFFFBAFFD8FF7E40F39060672A:6B536072FFBBFFD8FBBE467D96826672" box="[1111,1254,583,609]" country="Germany" county="Matthias Riedel" municipality="Augsburg" name="Merton Park" pageId="37" pageNumber="38" stateProvince="Norfolk">Merton Park</location>
,
<collectorName id="C379537FFFBBFFD8FB18467D97196672" box="[1265,1405,583,609]" pageId="37" pageNumber="38">London SW</collectorName>
19, TQ2469,
<collectingMethod id="B7CD4EBEFFBBFFD8FEEA465193EB6696" box="[259,399,619,645]" pageId="37" pageNumber="38">Malaise trap</collectingMethod>
in garden, 1219.vii.14,
<collectorName id="C379537FFFBBFFD8FD43465191056696" box="[682,865,619,645]" pageId="37" pageNumber="38">A.C.Galsworthy</collectorName>
(
<collectionCode id="089DAE6CFFBBFFD8FC99465191B96696" box="[880,989,619,645]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="37" pageNumber="38" type="Museum">NHMUK</collectionCode>
);
</materialsCitation>
<materialsCitation id="DEE43CF4FFBBFFD8FC1C465197F866BA" ID-GBIF-Occurrence="4116948369" collectingDate="1959-07-04" collectingDateMax="1969-09-01" collectingDateMin="1959-07-04" collectionCode="NMS, NHMUK, ZSM" country="Germany" county="Matthias Riedel" determinerName="Riedel" location="Santon Downham" municipality="Augsburg" pageId="37" pageNumber="38" specimenCount="12" specimenCount-female="12" stateProvince="Norfolk" typeStatus="paratype">
<specimenCount id="788AFD20FFBBFFD8FC1C465196736696" box="[1013,1047,619,645]" pageId="37" pageNumber="38" type="female">1♀</specimenCount>
, [
<collectingCountry id="169B7639FFBBFFD8FBC4465196FF6696" box="[1069,1179,619,645]" name="Germany" pageId="37" pageNumber="38">Germany</collectingCountry>
]
<collectingMunicipality id="8E57ACD3FFBBFFD8FB4E4651977C6696" box="[1191,1304,619,645]" pageId="37" pageNumber="38">Augsburg</collectingMunicipality>
,
<date id="1A321069FFBBFFD8FACD465097FC6696" box="[1316,1432,618,645]" pageId="37" pageNumber="38" value="1959-07-04">
<collectingDate id="0A76E981FFBBFFD8FACD465097FC6696" box="[1316,1432,618,645]" pageId="37" pageNumber="38" value="1959-07-04">4.vii.1959</collectingDate>
</date>
,
<taxonomicName id="A98C4D2AFFBBFFD8FF7E46B593FA66BA" baseAuthorityName="Ratzeburg" baseAuthorityYear="1844" box="[151,414,655,681]" class="Insecta" family="Ichneumonidae" genus="Hyposoter" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="carbonarius">
<emphasis id="5CF8EABBFFBBFFD8FF7E46B593FA66BA" box="[151,414,655,681]" italics="true" pageId="37" pageNumber="38">Hyposoter carbonarius</emphasis>
</taxonomicName>
<specimenCount id="788AFD20FFBBFFD8FE4E46B593D866BA" box="[423,444,655,681]" pageId="37" pageNumber="38" type="female"></specimenCount>
det
<collectorName id="C379537FFFBBFFD8FE1846B5902B66BA" box="[497,591,655,681]" pageId="37" pageNumber="38">R.Bauer</collectorName>
, [in Horstmanns hand]
<emphasis id="5CF8EABBFFBBFFD8FC8E46B5919466BA" box="[871,1008,655,681]" italics="true" pageId="37" pageNumber="38">
<taxonomicName id="A98C4D2AFFBBFFD8FC8E46B591B866BB" authorityName="Forster" authorityYear="1869" box="[871,988,655,680]" class="Insecta" family="Ichneumonidae" genus="Hyposoter" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus">Hyposoter</taxonomicName>
?
</emphasis>
<taxonomicName id="A98C4D2AFFBBFFD8FC1946B5975B66BA" authority="Aub. Horstm. 2013" authorityName="Aub. Horstm." authorityYear="2013" box="[1008,1343,655,681]" class="Insecta" family="Ichneumonidae" genus="Hyposoter" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="insulator">
<emphasis id="5CF8EABBFFBBFFD8FC1946B5963D66BA" box="[1008,1113,655,681]" italics="true" pageId="37" pageNumber="38">insulator</emphasis>
Aub. Horstm. 2013
</taxonomicName>
(
<collectionCode id="089DAE6CFFBBFFD8FAB946B597EB66BA" box="[1360,1423,655,681]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="37" pageNumber="38" type="Museum">ZSM</collectionCode>
);
</materialsCitation>
<materialsCitation id="DEE43CF4FFBBFFD8FF7E468997F866DE" ID-GBIF-Occurrence="4116948304" box="[151,1436,690,717]" collectingDate="1959-07-04" collectingDateMax="1969-09-01" collectingDateMin="1959-07-04" collectionCode="NMS, NHMUK, ZSM" country="Germany" county="Matthias Riedel" determinerName="Riedel" location="Santon Downham" municipality="Augsburg" pageId="37" pageNumber="38" specimenCount="12" specimenCount-female="12" stateProvince="Norfolk" typeStatus="paratype">
<specimenCount id="788AFD20FFBBFFD8FF7E468992D866DE" box="[151,188,691,717]" pageId="37" pageNumber="38" type="female">4♀</specimenCount>
, Nürnberg [
<collectingCountry id="169B7639FFBBFFD8FEA8468993D666DE" box="[321,434,691,717]" name="Germany" pageId="37" pageNumber="38">Germany</collectingCountry>
], 7 &amp; 11.7.76, 23.7.88, and 27.5.89, all labelled
<taxonomicName id="A98C4D2AFFBBFFD8FC25468997FC66DE" authority="Brke, R. Bauer (ZSM)" box="[972,1432,691,717]" class="Insecta" family="Ichneumonidae" genus="Hyposoter" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="alienus">
<emphasis id="5CF8EABBFFBBFFD8FC25468996FD66DE" box="[972,1177,691,717]" italics="true" pageId="37" pageNumber="38">Hyposoter alienus</emphasis>
Brke, R.Bauer (ZSM)
</taxonomicName>
;
</materialsCitation>
<materialsCitation id="DEE43CF4FFBBFFD8FF7E46ED93B46706" ID-GBIF-Occurrence="4116948390" collectingDate="1959-07-04" collectingDateMax="1969-09-01" collectingDateMin="1959-07-04" collectionCode="NMS, NHMUK, ZSM" country="Germany" county="Matthias Riedel" determinerName="Riedel" location="Santon Downham" municipality="Augsburg" pageId="37" pageNumber="38" specimenCount="12" specimenCount-female="12" stateProvince="Norfolk" typeStatus="paratype">
<specimenCount id="788AFD20FFBBFFD8FF7E46ED92DE66E2" box="[151,186,727,753]" pageId="37" pageNumber="38" type="female">1♀</specimenCount>
,
<collectorName id="C379537FFFBBFFD8FF2046ED932566E2" box="[201,321,727,753]" pageId="37" pageNumber="38">Messkirch</collectorName>
,
<collectorName id="C379537FFFBBFFD8FEA546ED93F066E2" box="[332,404,727,753]" pageId="37" pageNumber="38">Baden</collectorName>
[
<collectingCountry id="169B7639FFBBFFD8FE4846ED907666E2" box="[417,530,727,753]" name="Germany" pageId="37" pageNumber="38">Germany</collectingCountry>
],
<date id="1A321069FFBBFFD8FDCD46ED90ED66E2" box="[548,649,727,753]" pageId="37" pageNumber="38" value="1969-09-01">
<collectingDate id="0A76E981FFBBFFD8FDCD46ED90ED66E2" box="[548,649,727,753]" pageId="37" pageNumber="38" value="1969-09-01">1.9.1969</collectingDate>
</date>
,
<collectorName id="C379537FFFBBFFD8FD7D46ED90BF66E2" box="[660,731,727,753]" pageId="37" pageNumber="38">I.Wall</collectorName>
,
<emphasis id="5CF8EABBFFBBFFD8FD0E46ED910966E2" box="[743,877,727,753]" italics="true" pageId="37" pageNumber="38">
<taxonomicName id="A98C4D2AFFBBFFD8FD0E46ED913866E3" authorityName="Forster" authorityYear="1869" box="[743,860,727,752]" class="Insecta" family="Ichneumonidae" genus="Hyposoter" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus">Hyposoter</taxonomicName>
?
</emphasis>
<taxonomicName id="A98C4D2AFFBBFFD8FC8746ED961266E2" authority="Aubert" authorityName="Aubert" authorityYear="1965" box="[878,1142,727,753]" class="Insecta" family="Ichneumonidae" genus="Hyposoter" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="meridionellator">
<emphasis id="5CF8EABBFFBBFFD8FC8746ED964666E2" box="[878,1058,727,753]" italics="true" pageId="37" pageNumber="38">meridionellator</emphasis>
Aubert
</taxonomicName>
det.
<determinerName id="9BEFDA6DFFBBFFD8FB4746ED969E66E2" box="[1198,1274,727,753]" pageId="37" pageNumber="38">Riedel</determinerName>
, 2014 (private collection
<collectingCounty id="87524E25FFBBFFD8FEE446C193A06706" box="[269,452,763,789]" pageId="37" pageNumber="38">Matthias Riedel</collectingCounty>
).
</materialsCitation>
</paragraph>
<paragraph id="6E3336A9FFBBFFD8FF2E47259060672A" blockId="37.[151,1437,151,825]" box="[199,516,799,825]" pageId="37" pageNumber="38">Male and biology unknown.</paragraph>
</subSubSection>
</treatment>
</document>