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<document id="1A7CB0A658D714607FA46940C40EE90A" ID-DOI="10.11646/zootaxa.3698.1.1" ID-GBIF-Dataset="9d9508f0-a878-4283-86fd-1a7d9da9f3a4" ID-ISSN="1175-5326" ID-Zenodo-Dep="284413" ID-ZooBank="AD070B0D-967E-4AE5-8260-EC30432D67BC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460661232869" checkinUser="plazi" docAuthor="Gerstmeier, Roland &amp; Seitner, Martin" docDate="2013" docId="51116F777852D64FFF5DFDE3FE8A589E" docLanguage="en" docName="zt03698p077.pdf" docOrigin="Zootaxa 3698 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Eunatalis Schenkling 1909" docType="treatment" docVersion="6" lastPageNumber="6" masterDocId="AD28170F7857D64AFFCAFFB8FFE85F6A" masterDocTitle="Revision of the checkered beetle genus Eunatalis Schenkling, 1909 (Coleoptera: Cleridae: Clerinae)" masterLastPageNumber="77" masterPageNumber="1" pageNumber="6" updateTime="1698347555672" updateUser="plazi">
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<mods:title id="9DCE9DD9246D347BAF9FC9FE65D9CA20">Revision of the checkered beetle genus Eunatalis Schenkling, 1909 (Coleoptera: Cleridae: Clerinae)</mods:title>
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<mods:namePart id="2BB2565F64CF6CAF172523AAA8D75CBB">Gerstmeier, Roland</mods:namePart>
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<mods:namePart id="20ED1EFE3B4454E3A3BFFCA853F94242">Seitner, Martin</mods:namePart>
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<mods:date id="2D9691448E0D4E0E41E9059E1DD41BD8">2013</mods:date>
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<treatment id="51116F777852D64FFF5DFDE3FE8A589E" ID-DOI="http://doi.org/10.5281/zenodo.6164244" ID-GBIF-Taxon="127665251" ID-Zenodo-Dep="6164244" LSID="urn:lsid:plazi:treatment:51116F777852D64FFF5DFDE3FE8A589E" httpUri="http://treatment.plazi.org/id/51116F777852D64FFF5DFDE3FE8A589E" lastPageNumber="6" pageId="5" pageNumber="6">
<subSubSection id="91A28DEA7852D64FFF5DFDE3FAFD5C82" pageId="5" pageNumber="6" type="nomenclature">
<paragraph id="D907DE617852D64FFF5DFDE3FE0E5D1C" blockId="5.[151,486,603,630]" box="[151,486,603,630]" pageId="5" pageNumber="6">
<heading id="824F690D7852D64FFF5DFDE3FE0E5D1C" bold="true" box="[151,486,603,630]" fontSize="11" level="1" pageId="5" pageNumber="6" reason="1">
<taxonomicName id="1EB8A5E27852D64FFF5DFDE3FE0E5D1C" authority="Schenkling, 1909" authorityName="Schenkling" authorityYear="1909" box="[151,486,603,630]" class="Insecta" family="Cleridae" genus="Eunatalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="EBCC02737852D64FFF5DFDE3FE0E5D1C" bold="true" box="[151,486,603,630]" pageId="5" pageNumber="6">
<emphasis id="EBCC02737852D64FFF5DFDE3FEE45D1F" bold="true" box="[151,268,603,629]" italics="true" pageId="5" pageNumber="6">Eunatalis</emphasis>
Schenkling, 1909
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="D907DE617852D64FFF5DFD26FD5D5DD9" blockId="5.[151,995,669,721]" box="[151,693,669,691]" pageId="5" pageNumber="6">
<typeStatus id="060360C37852D64FFF5DFD26FF215DD9" box="[151,201,670,691]" pageId="5" pageNumber="6">Type</typeStatus>
species:
<taxonomicName id="1EB8A5E27852D64FFEECFD26FD595DD9" authority="Fabricius, 1787: 127" authorityName="Fabricius" authorityPageNumber="127" authorityYear="1787" box="[294,689,669,691]" class="Insecta" family="Anthicidae" genus="Notoxus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="porcatus">
<emphasis id="EBCC02737852D64FFEECFD26FE3F5DD9" box="[294,471,670,691]" italics="true" pageId="5" pageNumber="6">Notoxus porcatus</emphasis>
Fabricius, 1787: 127
</taxonomicName>
.
</paragraph>
<paragraph id="D907DE617852D64FFF5DFD03FC0B5DBB" blockId="5.[151,995,669,721]" box="[151,995,699,721]" pageId="5" pageNumber="6">Schenkling 1909: 163; Elston 1921: 148; Kolibáč 1998: 191; Solervicens 2007: 57.</paragraph>
<paragraph id="D907DE617852D64FFF5DFD41FAFD5C82" blockId="5.[151,1437,761,2036]" pageId="5" pageNumber="6">
<emphasis id="EBCC02737852D64FFF5DFD41FF085C78" bold="true" box="[151,224,761,786]" pageId="5" pageNumber="6">Head:</emphasis>
Dark brown to black; vested with whitish to grey setae; labrum broadly emarginate, V-shaped, light brown to dark brown; terminal maxillary palpomeres with sides parallel to very weakly divergent and apex obliquely truncate, terminal labial palpomeres securiform (
<figureCitation id="4183C2E47852D64FFD06FCF8FCFA5C33" box="[716,786,832,857]" captionStart="FIGURES 1 10" captionStartId="6.[151,264,1860,1883]" captionTargetBox="[262,1336,198,1830]" captionTargetId="figure@6.[248,1336,179,1830]" captionTargetPageId="6" captionText="FIGURES 1 10. Morphological characters of the genus Eunatalis. (1) Head, lower surface (Scale bar 1 mm), (2) antennae (Scale bar 1 mm), (3) Pro- and mesothorax, lower surface (Scale bar 1 mm), (4) Metendosternite (Scale bar 1 mm), (5) Nodules of elytral punctation, arrow indicates the basal seta (Scale bar 20 µm), (6) Protarsus, with spine, spur and pulvilli (Scale bar 1 mm), (7) Wing venation (Scale bar 5 mm), (8) Longitudinal groove between eyes of E. ninae n. sp. (Scale bar 1 mm), (9) Elytral punctation with two lateral nodules, E. foveata (Scale bar 1 mm), (10) Elytral punctation with two lateral nodules and one basal nodule, E. pernodulosa n. sp. (Scale bar 1 mm)." httpUri="https://zenodo.org/record/284414/files/figure.png" pageId="5" pageNumber="6">Fig. 1</figureCitation>
); eyes coarsely facetted, protruding laterally, margined; head including eyes as broad as or broader than anterior width of pronotum; interocular space from one to double eye width (as viewed from above); gular sutures converging, gular process conspicuous, with two papilles. Antennae with 11 antennomeres, filiform, light brown to black; antennomeres with dense, fine punctation, terminal three antennomeres densely tomentose, forming a loose club, A
<quantity id="1E4073847852D64FFCABFC68FC7F5C82" box="[865,919,975,1000]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" pageId="5" pageNumber="6" unit="in" value="11.0">11 in</quantity>
<taxonomicName id="1EB8A5E27852D64FFC54FC68FBD25C8D" box="[926,1082,976,999]" class="Insecta" family="Cleridae" genus="Eunatalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="spinicornis">
<emphasis id="EBCC02737852D64FFC54FC68FBD25C8D" box="[926,1082,976,999]" italics="true" pageId="5" pageNumber="6">E. spinicornis</emphasis>
</taxonomicName>
acuminate (
<figureCitation id="4183C2E47852D64FFB09FC77FAE05C82" box="[1219,1288,975,1000]" captionStart="FIGURES 1 10" captionStartId="6.[151,264,1860,1883]" captionTargetBox="[262,1336,198,1830]" captionTargetId="figure@6.[248,1336,179,1830]" captionTargetPageId="6" captionText="FIGURES 1 10. Morphological characters of the genus Eunatalis. (1) Head, lower surface (Scale bar 1 mm), (2) antennae (Scale bar 1 mm), (3) Pro- and mesothorax, lower surface (Scale bar 1 mm), (4) Metendosternite (Scale bar 1 mm), (5) Nodules of elytral punctation, arrow indicates the basal seta (Scale bar 20 µm), (6) Protarsus, with spine, spur and pulvilli (Scale bar 1 mm), (7) Wing venation (Scale bar 5 mm), (8) Longitudinal groove between eyes of E. ninae n. sp. (Scale bar 1 mm), (9) Elytral punctation with two lateral nodules, E. foveata (Scale bar 1 mm), (10) Elytral punctation with two lateral nodules and one basal nodule, E. pernodulosa n. sp. (Scale bar 1 mm)." httpUri="https://zenodo.org/record/284414/files/figure.png" pageId="5" pageNumber="6">Fig. 2</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="91A28DEA7852D64FFF0DFC4BFE8A589E" pageId="5" pageNumber="6" type="description">
<paragraph id="D907DE617852D64FFF0DFC4BFD465A27" blockId="5.[151,1437,761,2036]" pageId="5" pageNumber="6">
<emphasis id="EBCC02737852D64FFF0DFC4BFEC15B66" bold="true" box="[199,297,1011,1036]" pageId="5" pageNumber="6">Thorax:</emphasis>
Light brown to black; punctation and pilosity variable, base and apex with fringe of setae, pilosity mostly grey to whitish; pronotum longer than wide, sides more or less sinuate, with wrinkles, middle always with an elongated groove (sometimes missing or weak); grooves of sub-basal collar on each side of pronotum fused or separated by a weak or distinct bar. Pro-intercoxal process narrow, broadly dilated distally; procoxal cavities opened to almost closed posteriorly (sometimes variable within a species); hypomeron tapering apically towards intercoxal process; mesoventrite with variable sculpture and punctation, anterior mesoventral process distinct, sinuate, pit-shaped, slightly opened apically; posterior part of mesepisternon mostly smooth or slightly punctate (
<figureCitation id="4183C2E47852D64FFF55FB55FF0B5A6C" box="[159,227,1261,1286]" captionStart="FIGURES 1 10" captionStartId="6.[151,264,1860,1883]" captionTargetBox="[262,1336,198,1830]" captionTargetId="figure@6.[248,1336,179,1830]" captionTargetPageId="6" captionText="FIGURES 1 10. Morphological characters of the genus Eunatalis. (1) Head, lower surface (Scale bar 1 mm), (2) antennae (Scale bar 1 mm), (3) Pro- and mesothorax, lower surface (Scale bar 1 mm), (4) Metendosternite (Scale bar 1 mm), (5) Nodules of elytral punctation, arrow indicates the basal seta (Scale bar 20 µm), (6) Protarsus, with spine, spur and pulvilli (Scale bar 1 mm), (7) Wing venation (Scale bar 5 mm), (8) Longitudinal groove between eyes of E. ninae n. sp. (Scale bar 1 mm), (9) Elytral punctation with two lateral nodules, E. foveata (Scale bar 1 mm), (10) Elytral punctation with two lateral nodules and one basal nodule, E. pernodulosa n. sp. (Scale bar 1 mm)." httpUri="https://zenodo.org/record/284414/files/figure.png" pageId="5" pageNumber="6">Fig. 3</figureCitation>
); metaventrite with a conspicuous discriminal line, more or less punctate and wrinkled. Scutellum round to oblong-ovate, light brown to black, with isolated lateral setae, slightly punctate. Metendosternite with conspicuous furcal arms, laminae broadly triangular (
<figureCitation id="4183C2E47852D64FFD96FA8CFD495A27" box="[604,673,1332,1357]" captionStart="FIGURES 1 10" captionStartId="6.[151,264,1860,1883]" captionTargetBox="[262,1336,198,1830]" captionTargetId="figure@6.[248,1336,179,1830]" captionTargetPageId="6" captionText="FIGURES 1 10. Morphological characters of the genus Eunatalis. (1) Head, lower surface (Scale bar 1 mm), (2) antennae (Scale bar 1 mm), (3) Pro- and mesothorax, lower surface (Scale bar 1 mm), (4) Metendosternite (Scale bar 1 mm), (5) Nodules of elytral punctation, arrow indicates the basal seta (Scale bar 20 µm), (6) Protarsus, with spine, spur and pulvilli (Scale bar 1 mm), (7) Wing venation (Scale bar 5 mm), (8) Longitudinal groove between eyes of E. ninae n. sp. (Scale bar 1 mm), (9) Elytral punctation with two lateral nodules, E. foveata (Scale bar 1 mm), (10) Elytral punctation with two lateral nodules and one basal nodule, E. pernodulosa n. sp. (Scale bar 1 mm)." httpUri="https://zenodo.org/record/284414/files/figure.png" pageId="5" pageNumber="6">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="D907DE617852D64FFF0DFAE0FAE359E5" blockId="5.[151,1437,761,2036]" pageId="5" pageNumber="6">
<emphasis id="EBCC02737852D64FFF0DFAE0FEF45A1B" bold="true" box="[199,284,1368,1393]" pageId="5" pageNumber="6">Elytra:</emphasis>
Light brown to black; pilosity variable, mostly densely vested with anteriorly directed setae at the lower margin of elytrae, whitish to grey; elytral base not margined, lateral margins parallel or broadest behind middle, apices broadly rounded or with a conspicuous tooth, elytral punctation arranged into ten striae (
<taxonomicName id="1EB8A5E27852D64FFAF0FA19FA745AD2" box="[1338,1436,1439,1464]" class="Insecta" family="Cleridae" genus="Eunatalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="laevis">
<emphasis id="EBCC02737852D64FFAF0FA19FA745AD2" box="[1338,1436,1439,1464]" italics="true" pageId="5" pageNumber="6">E. laevis</emphasis>
</taxonomicName>
with hidden punctures), the ninth stria is partly or completely vestigial, interstices with a fine microsculpture (wrinkled to punctate); diameter of punctations from base to apex similar, or diameter of punctations behind the middle decreasing or obsolete; punctation round or oblong, with two or three nodules (a pair of lateral nodules, and one posterior nodule), nodules more or less sinuate, with a single seta beneath the basal nodule (
<figureCitation id="4183C2E47852D64FFB05F996FAFC592D" box="[1231,1300,1582,1607]" captionStart="FIGURES 1 10" captionStartId="6.[151,264,1860,1883]" captionTargetBox="[262,1336,198,1830]" captionTargetId="figure@6.[248,1336,179,1830]" captionTargetPageId="6" captionText="FIGURES 1 10. Morphological characters of the genus Eunatalis. (1) Head, lower surface (Scale bar 1 mm), (2) antennae (Scale bar 1 mm), (3) Pro- and mesothorax, lower surface (Scale bar 1 mm), (4) Metendosternite (Scale bar 1 mm), (5) Nodules of elytral punctation, arrow indicates the basal seta (Scale bar 20 µm), (6) Protarsus, with spine, spur and pulvilli (Scale bar 1 mm), (7) Wing venation (Scale bar 5 mm), (8) Longitudinal groove between eyes of E. ninae n. sp. (Scale bar 1 mm), (9) Elytral punctation with two lateral nodules, E. foveata (Scale bar 1 mm), (10) Elytral punctation with two lateral nodules and one basal nodule, E. pernodulosa n. sp. (Scale bar 1 mm)." httpUri="https://zenodo.org/record/284414/files/figure.png" pageId="5" pageNumber="6">Fig. 5</figureCitation>
); interstices in basal half flattened, posterior of middle sometimes more or less ridged. Hindwing with closed wedge cell, margin of radial cell strongly sclerotized, MP3, MP4, AA3+4 and CuA+AA reaching margin (
<figureCitation id="4183C2E47852D64FFB72F9CEFB1459E5" box="[1208,1276,1654,1679]" captionStart="FIGURES 1 10" captionStartId="6.[151,264,1860,1883]" captionTargetBox="[262,1336,198,1830]" captionTargetId="figure@6.[248,1336,179,1830]" captionTargetPageId="6" captionText="FIGURES 1 10. Morphological characters of the genus Eunatalis. (1) Head, lower surface (Scale bar 1 mm), (2) antennae (Scale bar 1 mm), (3) Pro- and mesothorax, lower surface (Scale bar 1 mm), (4) Metendosternite (Scale bar 1 mm), (5) Nodules of elytral punctation, arrow indicates the basal seta (Scale bar 20 µm), (6) Protarsus, with spine, spur and pulvilli (Scale bar 1 mm), (7) Wing venation (Scale bar 5 mm), (8) Longitudinal groove between eyes of E. ninae n. sp. (Scale bar 1 mm), (9) Elytral punctation with two lateral nodules, E. foveata (Scale bar 1 mm), (10) Elytral punctation with two lateral nodules and one basal nodule, E. pernodulosa n. sp. (Scale bar 1 mm)." httpUri="https://zenodo.org/record/284414/files/figure.png" pageId="5" pageNumber="6">Fig. 7</figureCitation>
).
</paragraph>
<paragraph id="D907DE617852D64FFF0DF922FDFD580F" blockId="5.[151,1437,761,2036]" pageId="5" pageNumber="6">
<emphasis id="EBCC02737852D64FFF0DF922FEE059D9" bold="true" box="[199,264,1690,1715]" pageId="5" pageNumber="6">Legs:</emphasis>
Light brown to black; more or less pilose, inner edge of protibiae and sometimes mesotibiae with a row of short, thick bristles, with more or less conspicuous punctation; profemora conspicuously swollen, protibiae slightly bent, mesotibiae more or less bent (except
<taxonomicName id="1EB8A5E27852D64FFD3AF95AFC7F5993" box="[752,919,1762,1785]" class="Insecta" family="Cleridae" genus="Eunatalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="semicostata">
<emphasis id="EBCC02737852D64FFD3AF95AFC7F5993" box="[752,919,1762,1785]" italics="true" pageId="5" pageNumber="6">E. semicostata</emphasis>
</taxonomicName>
with almost straight mesotibiae), metatibiae straight to very slightly bent, all tibiae without longitudinal carinae, protibiae terminating in a short spine and a single spur, tibial spur formula 1-2-2; tarsal pulvillar formula 3-3-3, tarsal pulvilli broadly lobed, but not emarginate; claws simple (
<figureCitation id="4183C2E47852D64FFE08F8F4FDEE580F" box="[450,518,1868,1893]" captionStart="FIGURES 1 10" captionStartId="6.[151,264,1860,1883]" captionTargetBox="[262,1336,198,1830]" captionTargetId="figure@6.[248,1336,179,1830]" captionTargetPageId="6" captionText="FIGURES 1 10. Morphological characters of the genus Eunatalis. (1) Head, lower surface (Scale bar 1 mm), (2) antennae (Scale bar 1 mm), (3) Pro- and mesothorax, lower surface (Scale bar 1 mm), (4) Metendosternite (Scale bar 1 mm), (5) Nodules of elytral punctation, arrow indicates the basal seta (Scale bar 20 µm), (6) Protarsus, with spine, spur and pulvilli (Scale bar 1 mm), (7) Wing venation (Scale bar 5 mm), (8) Longitudinal groove between eyes of E. ninae n. sp. (Scale bar 1 mm), (9) Elytral punctation with two lateral nodules, E. foveata (Scale bar 1 mm), (10) Elytral punctation with two lateral nodules and one basal nodule, E. pernodulosa n. sp. (Scale bar 1 mm)." httpUri="https://zenodo.org/record/284414/files/figure.png" pageId="5" pageNumber="6">Fig. 6</figureCitation>
).
</paragraph>
<paragraph id="D907DE617852D64FFF0DF8C8FE8A589E" blockId="5.[151,1437,761,2036]" pageId="5" pageNumber="6">
<emphasis id="EBCC02737852D64FFF0DF8C8FEAD58E3" bold="true" box="[199,325,1904,1929]" pageId="5" pageNumber="6">Abdomen:</emphasis>
Six visible abdominal ventrites, with a more or less dense pilosity and punctation. Ventrite VI in males straight or emarginate, in females rounded. Tegmen ventrally closed, phallobasic apodeme and struts distally more or less spoon-like, branches of spicular fork distally strongly broadened. Female copulatory organs have not been investigated.
</paragraph>
</subSubSection>
</treatment>
</document>