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<document id="E08F41974C5FFCFE6AC1507E08ECB750" ID-DOI="10.26107/RBZ-2019-0003" ID-ISSN="2345-7600" ID-Zenodo-Dep="5343178" ID-ZooBank="026EF5FA-B910-41A8-9D86-3086D24319E9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630195483579" checkinUser="diego" docAuthor="Tan, Ji, Chan, Zi Yang, Ong, Ching Ang &amp; Yong, Hoi Sen" docDate="2019" docId="9A6DF50C104A35248A67E7C9226CF988" docLanguage="en" docName="RBZ-2019-0003.pdf" docOrigin="Raffles Bulletin of Zoology 67" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Actinacantha globulata" docType="treatment" docVersion="3" lastPageNumber="43" masterDocId="66548D741040352F8912E27D2164FFED" masterDocTitle="Phylogenetic relationships of Actinacantha Simon, Gasteracantha Sundevall, Macracantha Hasselt and Thelacantha Simon spiny orbweavers (Araneae: Araneidae) in Peninsular Malaysia" masterLastPageNumber="55" masterPageNumber="32" pageNumber="42" updateTime="1699182970771" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="0949672686886731C2A4EFC1B4EEC02C">Phylogenetic relationships of Actinacantha Simon, Gasteracantha Sundevall, Macracantha Hasselt and Thelacantha Simon spiny orbweavers (Araneae: Araneidae) in Peninsular Malaysia</mods:title>
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<subSubSection id="5ADE1791104A35258A67E7C92407FA20" box="[885,1379,1460,1485]" pageId="10" pageNumber="42" type="nomenclature">
<paragraph id="127B441A104A35258A67E7C92407FA20" blockId="10.[885,1379,1460,1517]" box="[885,1379,1460,1485]" pageId="10" pageNumber="42">
<heading id="4933F376104A35258A67E7C92407FA20" bold="true" box="[885,1379,1460,1485]" centered="true" fontSize="10" level="2" pageId="10" pageNumber="42" reason="2">
<taxonomicName id="D5C43F99104A35258A67E7C92407FA20" ID-CoL="9SVQ" authority="(Walckenaer, 1841)" baseAuthorityName="Walckenaer" baseAuthorityYear="1841" box="[885,1379,1460,1485]" class="Arachnida" family="Araneidae" genus="Actinacantha" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="42" phylum="Arthropoda" rank="species" species="globulata">
<emphasis id="20B09808104A35258A67E7C92407FA20" bold="true" box="[885,1379,1460,1485]" pageId="10" pageNumber="42">
<emphasis id="20B09808104A35258A67E7C9251EFA21" bold="true" box="[885,1146,1460,1484]" italics="true" pageId="10" pageNumber="42">Actinacantha globulata</emphasis>
(
<bibRefCitation id="765539EB104A35258D98E7C8243FFA20" author="Walckenaer CA" box="[1162,1371,1461,1485]" pageId="10" pageNumber="42" refId="ref13006" refString="Walckenaer CA (1841) Histoire naturelle des Insects. Apteres. Paris, 2 (for 1837), 549 pp." type="book" year="1841">Walckenaer, 1841</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="5ADE1791104A35258D1EE7A825A8FA00" box="[1036,1228,1493,1517]" pageId="10" pageNumber="42" type="description">
<paragraph id="127B441A104A35258D1EE7A825A8FA00" blockId="10.[885,1379,1460,1517]" box="[1036,1228,1493,1517]" pageId="10" pageNumber="42">
(
<figureCitation id="8AFF589F104A35258D06E7A82510FA00" box="[1044,1140,1493,1517]" captionStart="Fig" captionStartId="6.[113,148,1294,1316]" captionTargetBox="[114,1451,190,1277]" captionTargetId="figure-420@6.[113,1451,189,1278]" captionTargetPageId="6" captionText="Fig. 1. General habitus. A, Actinacantha globulata (Walckenaer, 1841), sub-female; B, Gasteracantha diardi (Lucas, 1835), female; C, G. diardi yellow morph, female; D, G. diardi white morph, sub-female; E, G. kuhli, female; F, Gasteracantha hasselti C. L. Koch, 1837, female; G, Macracantha arcuata (Fabricius, 1793), female; H, Thelacantha brevispina (Doleschall, 1857), female; I, G. mengei Keyserling, 1864, female. Photos by Tan Ji." figureDoi="http://doi.org/10.5281/zenodo.5343182" httpUri="https://zenodo.org/record/5343182/files/figure.png" pageId="10" pageNumber="42">Figs. 1A</figureCitation>
,
<figureCitation id="8AFF589F104A35258D90E7A825EAFA00" box="[1154,1166,1493,1517]" captionStart="Fig" captionStartId="9.[136,171,650,672]" captionTargetBox="[138,774,192,634]" captionTargetId="figure-8@9.[120,786,174,644]" captionTargetPageId="9" captionText="Fig. 5. Sub-female Actinacantha globulata (Walckenaer, 1841). A, Dorsal habitus; B, Ventral habitus; C, Lateral habitus. Scale = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5343194" httpUri="https://zenodo.org/record/5343194/files/figure.png" pageId="10" pageNumber="42">5</figureCitation>
AC)
</paragraph>
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<subSubSection id="5ADE1791104A35258A3FE4692598F98F" pageId="10" pageNumber="42" type="reference_group">
<paragraph id="127B441A104A35258A3FE46925B4F9C4" blockId="10.[813,1276,1555,1635]" box="[813,1232,1555,1578]" pageId="10" pageNumber="42">
<treatmentCitationGroup id="32D46334104A35258A3FE46925B4F9C4" box="[813,1232,1555,1578]" pageId="10" pageNumber="42">
<treatmentCitation id="9365620B104A35258A3FE46925B4F9C4" author="Walckenaer CA" box="[813,1232,1555,1578]" page="151" pageId="10" pageNumber="42" year="1841">
<taxonomicName id="D5C43F99104A35258A3FE46925B4F9C4" authority="Walckenaer, 1841: 151" authorityName="Walckenaer" authorityPageNumber="151" authorityYear="1841" box="[813,1232,1555,1578]" class="Arachnida" family="Araneidae" genus="Plectana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="42" phylum="Arthropoda" rank="species" species="globulata">
<emphasis id="20B09808104A35258A3FE4692282F9C4" box="[813,998,1556,1577]" italics="true" pageId="10" pageNumber="42">Plectana globulata</emphasis>
<bibRefCitation id="765539EB104A35258AFCE46925B4F9C4" author="Walckenaer CA" box="[1006,1232,1555,1578]" pageId="10" pageNumber="42" refId="ref13006" refString="Walckenaer CA (1841) Histoire naturelle des Insects. Apteres. Paris, 2 (for 1837), 549 pp." type="book" year="1841">Walckenaer, 1841: 151</bibRefCitation>
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<paragraph id="127B441A104A35258A3FE44D25B4F9AB" blockId="10.[813,1276,1555,1635]" box="[813,1232,1584,1606]" pageId="10" pageNumber="42">
<treatmentCitationGroup id="32D46334104A35258A3FE44D25B4F9AB" box="[813,1232,1584,1606]" pageId="10" pageNumber="42">
<treatmentCitation id="9365620B104A35258A3FE44D25A8F9AB" author="Simon E" box="[813,1228,1584,1606]" page="286" pageId="10" pageNumber="42" year="1864">
<taxonomicName id="D5C43F99104A35258A3FE44D25A8F9AB" ID-CoL="9SVQ" authority="Simon, 1864: 286" authorityName="Simon" authorityPageNumber="286" authorityYear="1864" box="[813,1228,1584,1606]" class="Arachnida" family="Araneidae" genus="Actinacantha" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="42" phylum="Arthropoda" rank="species" species="globulata">
<emphasis id="20B09808104A35258A3FE44D2574F9A8" box="[813,1040,1584,1605]" italics="true" pageId="10" pageNumber="42">Actinacantha globulata</emphasis>
<bibRefCitation id="765539EB104A35258D08E44D25A8F9AB" author="Simon E" box="[1050,1228,1584,1606]" pageId="10" pageNumber="42" refId="ref12544" refString="Simon E (1864) Histoire naturelle des araignees (araneides). Paris, 540 pp." type="book" year="1864">Simon, 1864: 286</bibRefCitation>
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.
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</paragraph>
<paragraph id="127B441A104A35258A3FE4302598F98F" blockId="10.[813,1276,1555,1635]" box="[813,1276,1613,1635]" pageId="10" pageNumber="42">
<treatmentCitationGroup id="32D46334104A35258A3FE4302598F98F" box="[813,1276,1613,1635]" pageId="10" pageNumber="42">
<treatmentCitation id="9365620B104A35258A3FE4302598F98F" author="Hasselt AWM van" box="[813,1276,1613,1635]" page="12" pageId="10" pageNumber="42" year="1882">
<taxonomicName id="D5C43F99104A35258A3FE4302598F98F" authority="van Hasselt, 1882: 12" authorityName="van Hasselt" authorityPageNumber="12" authorityYear="1882" box="[813,1276,1613,1635]" class="Arachnida" family="Araneidae" genus="Gasteracantha" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="42" phylum="Arthropoda" rank="species" species="globulata">
<emphasis id="20B09808104A35258A3FE430257BF98F" box="[813,1055,1613,1634]" italics="true" pageId="10" pageNumber="42">Gasteracantha globulata</emphasis>
van
<bibRefCitation id="765539EB104A35258D43E4302598F98F" author="Hasselt AWM van" box="[1105,1276,1613,1635]" pageId="10" pageNumber="42" pagination="1 - 56" refId="ref11461" refString="Hasselt AWM van (1882) Araneae. In: Veth PJ (ed.) Midden Sumatra IV. 3 de Aflev. Naturlijke Historie. Leiden, 11 A. Pp. 1 - 56." type="journal article" year="1882">Hasselt, 1882: 12</bibRefCitation>
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</paragraph>
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<subSubSection id="5ADE1791104A35258A3FE4F622D3F90E" pageId="10" pageNumber="42" type="materials_examined">
<paragraph id="127B441A104A35258A3FE4F622D3F90E" blockId="10.[813,1452,1675,1763]" pageId="10" pageNumber="42">
<emphasis id="20B09808104A35258A3FE4F6257BF94E" bold="true" box="[813,1055,1675,1699]" pageId="10" pageNumber="42">Material examined.</emphasis>
<materialsCitation id="A2AC4E47104A35258D38E4F622D6F90E" collectingDate="2016-01-15" collectionCode="AGLO" collectorName="Semenyih. &amp; Chan ZY" country="Malaysia" location="Selangor" pageId="10" pageNumber="42" specimenCode="AGLO1" specimenCount="1" stateProvince="Selangor">
<collectingCountry id="6AD3048A104A35258D38E4F625A6F94E" box="[1066,1218,1675,1699]" name="Malaysia" pageId="10" pageNumber="42">MALAYSIA</collectingCountry>
. 1 sub-female (reg. UIR150116-
<specimenCode id="4262EC61104A35258AABE4D62574F92E" box="[953,1040,1707,1731]" collectionCode="AGLO" pageId="10" pageNumber="42">AGLO1</specimenCode>
),
<collectorName id="BF3121CC104A35258D30E4D625F3F92E" box="[1058,1175,1707,1731]" pageId="10" pageNumber="42">Semenyih.</collectorName>
,
<collectingRegion id="D0008AF8104A35258DB3E4D62467F92E" box="[1185,1283,1707,1731]" country="Malaysia" name="Selangor" pageId="10" pageNumber="42">Selangor</collectingRegion>
, coll.
<collectorName id="BF3121CC104A35258C51E4D624CCF92E" box="[1347,1448,1707,1731]" pageId="10" pageNumber="42">Chan ZY</collectorName>
,
<date id="667A62DA104A35258A3FE4B622D6F90E" box="[813,946,1739,1763]" pageId="10" pageNumber="42" value="2016-01-15">
<collectingDate id="763E9B32104A35258A3FE4B622D6F90E" box="[813,946,1739,1763]" pageId="10" pageNumber="42" value="2016-01-15">15 Jan.2016</collectingDate>
</date>
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.
</paragraph>
</subSubSection>
<subSubSection id="5ADE1791104A35258A3FE57622F8F7CE" pageId="10" pageNumber="42" type="diagnosis">
<paragraph id="127B441A104A35258A3FE57622BAF86E" blockId="10.[813,1452,1803,1923]" pageId="10" pageNumber="42">
<emphasis id="20B09808104A35258A3FE57622C5F8CE" bold="true" box="[813,929,1803,1827]" pageId="10" pageNumber="42">Diagnosis.</emphasis>
Adult female
<taxonomicName id="D5C43F99104A35258D2FE57125A5F8CE" authorityName="van Hasselt" authorityYear="1882" box="[1085,1217,1803,1827]" class="Arachnida" family="Araneidae" genus="Gasteracantha" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="42" phylum="Arthropoda" rank="species" species="globulata">
<emphasis id="20B09808104A35258D2FE57125A5F8CE" box="[1085,1217,1803,1827]" italics="true" pageId="10" pageNumber="42">A. globulata</emphasis>
</taxonomicName>
is differentiated from those of
<taxonomicName id="D5C43F99104A35258A9FE55625E2F8AE" authorityName="C. L. Koch" authorityYear="1837" box="[909,1158,1835,1859]" class="Arachnida" family="Araneidae" genus="Gasteracantha" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="42" phylum="Arthropoda" rank="species" species="hasselti">
<emphasis id="20B09808104A35258A9FE55625E2F8AE" box="[909,1158,1835,1859]" italics="true" pageId="10" pageNumber="42">Gasteracantha hasselti</emphasis>
</taxonomicName>
and
<taxonomicName id="D5C43F99104A35258DD2E55624CFF8AE" baseAuthorityName="Fabricius" baseAuthorityYear="1793" box="[1216,1451,1835,1859]" class="Arachnida" family="Araneidae" genus="Macracantha" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="42" phylum="Arthropoda" rank="species" species="arcuata">
<emphasis id="20B09808104A35258DD2E55624CFF8AE" box="[1216,1451,1835,1859]" italics="true" pageId="10" pageNumber="42">Macracantha arcuata</emphasis>
</taxonomicName>
based on the unique bulbous bases from which the middle spines originate.
</paragraph>
<paragraph id="127B441A104A35258A3FE5D622F8F7CE" blockId="10.[813,1453,1963,2083]" pageId="10" pageNumber="42">
<emphasis id="20B09808104A35258A3FE5D62282F82E" bold="true" box="[813,998,1963,1987]" pageId="10" pageNumber="42">Natural history.</emphasis>
A sub-female was collected from a web in the understorey of a primary tropical forest. The species appears to be uncommon and very localised in the Malay Peninsula.
</paragraph>
</subSubSection>
<caption id="46BB1492104B3524899AE10225F4FC5F" ID-DOI="http://doi.org/10.5281/zenodo.5343202" ID-Zenodo-Dep="5343202" httpUri="https://zenodo.org/record/5343202/files/figure.png" pageId="11" pageNumber="43" startId="11.[136,171,895,917]" targetBox="[140,1476,189,879]" targetPageId="11">
<paragraph id="127B441A104B3524899AE10225F4FC5F" blockId="11.[136,1474,895,946]" pageId="11" pageNumber="43">
Fig. 8. Epigynal structure of
<taxonomicName id="D5C43F99104B352488B3E102227EFC79" authority="C. L. Koch, 1837" authorityName="C. L. Koch" authorityYear="1837" box="[417,794,895,917]" class="Arachnida" family="Araneidae" genus="Gasteracantha" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="43" phylum="Arthropoda" rank="species" species="kuhli">
<emphasis id="20B09808104B352488B3E1022303FC79" box="[417,615,895,916]" italics="true" pageId="11" pageNumber="43">Gasteracantha kuhli</emphasis>
C. L.
<bibRefCitation id="765539EB104B35248BB4E102227EFC79" author="Koch CL" box="[678,794,895,917]" pageId="11" pageNumber="43" pagination="1 - 108" refId="ref11717" refString="Koch CL (1837) Die Arachniden. Dritter Band [6]. C. H. Zeh'sche Buchhandlung, Nurnberg, pp. 105 - 119, pls. CVI-CVIII; Vierter Band, pp. 1 - 108." type="book chapter" year="1837">Koch, 1837</bibRefCitation>
</taxonomicName>
. A, Ventral view; B, Posterior view; C, Lateral view; D, Dorsal view; E, Anterior view. Scale = 100 µm. CO = Copulatory opening; FD = Fertilisation duct; S = Spermatheca.
</paragraph>
</caption>
<subSubSection id="5ADE1791104B3524899AE1912005FBC8" pageId="11" pageNumber="43" type="distribution">
<paragraph id="127B441A104B3524899AE1912005FBC8" blockId="11.[136,775,1004,1061]" pageId="11" pageNumber="43">
<emphasis id="20B09808104B3524899AE191207EFBE9" bold="true" box="[136,282,1004,1028]" pageId="11" pageNumber="43">Distribution.</emphasis>
Peninsular
<collectingCountry id="6AD3048A104B3524888CE1902366FBE8" box="[414,514,1005,1029]" name="Malaysia" pageId="11" pageNumber="43">Malaysia</collectingCountry>
, Borneo, and
<collectingCountry id="6AD3048A104B35248B8DE1902263FBE8" box="[671,775,1005,1029]" name="Indonesia" pageId="11" pageNumber="43">Indonesia</collectingCountry>
(
<collectingRegion id="D0008AF8104B3524899DE670218EFBC8" box="[143,234,1037,1061]" country="Indonesia" name="Sumatera Utara" pageId="11" pageNumber="43">Sumatra</collectingRegion>
and
<collectingRegion id="D0008AF8104B35248830E6702032FBC8" box="[290,342,1037,1061]" country="Indonesia" name="Jawa Timur" pageId="11" pageNumber="43">Java</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="5ADE1791104B3524899AE630226CF988" pageId="11" pageNumber="43" type="discussion">
<paragraph id="127B441A104B3524899AE630226CF988" blockId="11.[135,776,1101,1637]" pageId="11" pageNumber="43">
<emphasis id="20B09808104B3524899AE6302193FB88" bold="true" box="[136,247,1101,1125]" pageId="11" pageNumber="43">Remarks.</emphasis>
The similarities in the morphological descriptions and geographical distribution of
<taxonomicName id="D5C43F99104B35248B20E61023ABFB68" authorityName="van Hasselt" authorityYear="1882" box="[562,719,1133,1157]" class="Arachnida" family="Araneidae" genus="Gasteracantha" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="43" phylum="Arthropoda" rank="species" species="globulata">
<emphasis id="20B09808104B35248B20E61023ABFB68" box="[562,719,1133,1157]" italics="true" pageId="11" pageNumber="43">A. globulata</emphasis>
</taxonomicName>
and
<taxonomicName id="D5C43F99104B3524899AE6F020F8FB49" authorityName="Bradley" authorityYear="1877" box="[136,412,1165,1189]" class="Arachnida" family="Araneidae" genus="Gasteracantha" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="43" phylum="Arthropoda" rank="species" species="crucigera">
<emphasis id="20B09808104B3524899AE6F020F8FB49" box="[136,412,1165,1189]" italics="true" pageId="11" pageNumber="43">Gasteracantha crucigera</emphasis>
</taxonomicName>
suggest that they may likely be synonymous (
<bibRefCitation id="765539EB104B35248837E6D02093FB28" author="Walckenaer CA" box="[293,503,1197,1221]" pageId="11" pageNumber="43" refId="ref13006" refString="Walckenaer CA (1841) Histoire naturelle des Insects. Apteres. Paris, 2 (for 1837), 549 pp." type="book" year="1841">Walckenaer, 1841</bibRefCitation>
;
<bibRefCitation id="765539EB104B35248B14E6D023FDFB28" author="Simon E" box="[518,665,1197,1221]" pageId="11" pageNumber="43" refId="ref12544" refString="Simon E (1864) Histoire naturelle des araignees (araneides). Paris, 540 pp." type="book" year="1864">Simon, 1864</bibRefCitation>
;
<bibRefCitation id="765539EB104B35248BBBE6D021A5FB08" author="Bradley HB" pageId="11" pageNumber="43" pagination="115 - 120" refId="ref10513" refString="Bradley HB (1877) Araneides of the Chevert Expedition. Part II. Proceedings of the Linnean Society of New South Wales, 2: 115 - 120." type="journal article" year="1877">Bradley, 1877</bibRefCitation>
;
<bibRefCitation id="765539EB104B352489DEE6B020F3FB08" author="Chrysanthus P" box="[204,407,1229,1253]" pageId="11" pageNumber="43" pagination="1 - 52" refId="ref10731" refString="Chrysanthus P (1971) Further notes on the spiders of New Guinea I (Argyopidae). Zoologische Verhandelingen, 113: 1 - 52." type="journal article" year="1971">Chrysanthus, 1971</bibRefCitation>
;
<bibRefCitation id="765539EB104B352488B1E6B0237AFB08" author="Dahl F" box="[419,542,1229,1253]" pageId="11" pageNumber="43" pagination="235 - 301" refId="ref10831" refString="Dahl F (1914) Die Gasteracanthen des Berliner Zoologischen Museums und deren geographische Verbreitung. Mitteilungen aus dem Zoologischen Museum in Berlin, 7: 235 - 301." type="journal article" year="1914">Dahl, 1914</bibRefCitation>
;
<bibRefCitation id="765539EB104B35248B3BE6B02398FB08" author="Yong HS &amp; Ono H" box="[553,764,1229,1253]" pageId="11" pageNumber="43" pagination="101 - 103" refId="ref13350" refString="Yong HS &amp; Ono H (2009) Gasteracantha crucigera (Araneae: Araneidae) from Maliau Basin, Sabah, Malaysia: an overlooked spiny-backed orb-weaver spider from Peninsular Malaysia. Journal of Science and Technology in the Tropics 5: 101 - 103." type="journal article" year="2009">Yong &amp; Ono, 2009</bibRefCitation>
). However, a solid conclusion cannot be made based on the morphology and DNA sequence of only one
<taxonomicName id="D5C43F99104B35248B46E77023B0FAC8" authorityName="van Hasselt" authorityYear="1882" box="[596,724,1293,1317]" class="Arachnida" family="Araneidae" genus="Gasteracantha" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="43" phylum="Arthropoda" rank="species" species="globulata">
<emphasis id="20B09808104B35248B46E77023B0FAC8" box="[596,724,1293,1317]" italics="true" pageId="11" pageNumber="43">A. globulata</emphasis>
</taxonomicName>
subadult female in its penultimate molt. This limited information may overlook cryptic or sibling species existing between Peninsular
<collectingCountry id="6AD3048A104B35248813E7102001FA68" box="[257,357,1389,1413]" name="Malaysia" pageId="11" pageNumber="43">Malaysia</collectingCountry>
, Borneo, and
<collectingCountry id="6AD3048A104B352488ECE71023B6FA68" box="[510,722,1389,1413]" name="Papua New Guinea" pageId="11" pageNumber="43">Papua New Guinea</collectingCountry>
. The recent employment of molecular data has already revealed the existence of species complexes in spider and insect taxa which may well be possible among geographically isolated
<taxonomicName id="D5C43F99104B35248BFDE7B0218AF9E8" authorityName="van Hasselt" authorityYear="1882" class="Arachnida" family="Araneidae" genus="Gasteracantha" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="43" phylum="Arthropoda" rank="species" species="globulata">
<emphasis id="20B09808104B35248BFDE7B0218AF9E8" italics="true" pageId="11" pageNumber="43">A. globulata</emphasis>
</taxonomicName>
and
<taxonomicName id="D5C43F99104B3524880FE79020C7F9E9" authorityName="Bradley" authorityYear="1877" box="[285,419,1517,1540]" class="Arachnida" family="Araneidae" genus="Gasteracantha" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="43" phylum="Arthropoda" rank="species" species="crucigera">
<emphasis id="20B09808104B3524880FE79020C7F9E9" box="[285,419,1517,1540]" italics="true" pageId="11" pageNumber="43">G. crucigera</emphasis>
</taxonomicName>
(
<bibRefCitation id="765539EB104B352488A3E79023E2F9E8" author="Franzini PZN &amp; Dippenaar-Schoeman AS &amp; Yessoufou K &amp; Van der Bank FH" box="[433,646,1517,1541]" pageId="11" pageNumber="43" pagination="21 - 34" refId="ref11126" refString="Franzini PZN, Dippenaar-Schoeman AS, Yessoufou K &amp; Van der Bank FH (2013) Combined analyses of genetic and morphological data indicate more than one species of Cyrtophora (Araneae: Araneidae) in South Africa. International Journal of Modern Biological Research, 1: 21 - 34." type="journal article" year="2013">Franzini et al., 2013</bibRefCitation>
;
<bibRefCitation id="765539EB104B35248B82E7902181F9C8" author="Gregoric M &amp; Blackledge TA &amp; Agnarsson I &amp; Kuntner M" pageId="11" pageNumber="43" pagination="293 - 312" refId="ref11381" refString="Gregoric M, Blackledge TA, Agnarsson I &amp; Kuntner M (2015) A molecular phylogeny of bark spiders reveals new species from Africa and Madagascar (Araneae: Araneidae: Caerostris). Journal of Arachnology, 43: 293 - 312." type="journal article" year="2015">Gregorič et al., 2015</bibRefCitation>
;
<bibRefCitation id="765539EB104B352489FCE47020C0F9C8" author="Yong HS &amp; Lim PE &amp; Tan J &amp; Song SL &amp; Suana IW &amp; Eamsobhana P" box="[238,420,1549,1573]" pageId="11" pageNumber="43" pagination="0134581" refId="ref13396" refString="Yong HS, Lim PE, Tan J, Song SL, Suana IW, Eamsobhana P (2015) Multigene phylogeography of Bactrocera caudata (Insecta: Tephritidae): Distinct genetic lineages in Northern and Southern hemispheres. PLoS ONE 31, 10 (7): e 0134581." type="journal article" year="2015">Yong et al., 2015</bibRefCitation>
). Nevertheless, the morphological and molecular characterisation of more specimens of both species are expected to resolve this taxonomic uncertainty.
</paragraph>
</subSubSection>
</treatment>
</document>