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<mods:title id="A51C222222C5CF0AA4733257CCB976E8">Nomenclatural notes on the Mediterranean firs (Abies, Pinaceae)</mods:title>
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<mods:namePart id="93677165A7E592FE01F1802E29ED016C">Ferrer-Gallego, P. Pablo</mods:namePart>
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<mods:namePart id="D7A1EFBC8554C3E6BDAC11CEAB357FDE">Laguna, Emilio</mods:namePart>
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<mods:affiliation id="9703A7E1F265F2C84EECB1F86C78F4F5">Bodega Ferrer-Gallego, 46311 Jaraguas, Valencia, Spain &amp; raul. ferrer @ vitec. wine; https: // orcid. org / 0000 - 0002 - 0301 - 5334</mods:affiliation>
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<treatment id="03FF903F4B19FF9214F9D43FFBF1FDCE" ID-DOI="http://doi.org/10.5281/zenodo.6608721" ID-GBIF-Taxon="195728681" ID-Zenodo-Dep="6608721" LSID="urn:lsid:plazi:treatment:03FF903F4B19FF9214F9D43FFBF1FDCE" httpUri="http://treatment.plazi.org/id/03FF903F4B19FF9214F9D43FFBF1FDCE" lastPageId="7" lastPageNumber="38" pageId="2" pageNumber="33">
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<paragraph id="8BE921294B19FF9714F9D43FFE1AFA6E" blockId="2.[136,506,1382,1409]" box="[136,506,1382,1409]" pageId="2" pageNumber="33">
<heading id="D0A196454B19FF9714F9D43FFE1AFA6E" box="[136,506,1382,1409]" fontSize="11" level="2" pageId="2" pageNumber="33" reason="3">
<taxonomicName id="4C565AAA4B19FF9714F9D43FFE1AFA6E" ID-CoL="8KFM" authority="Boissier (1838: 8)" authorityName="Boissier, G" authorityPageNumber="8" authorityYear="1838" box="[136,506,1382,1409]" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="2" pageNumber="33" phylum="Tracheophyta" rank="species" species="pinsapo">
<emphasis id="B922FD3B4B19FF9714F9D43FFEC9FA6E" bold="true" box="[136,297,1382,1409]" italics="true" pageId="2" pageNumber="33">Abies pinsapo</emphasis>
Boissier (1838: 8)
</taxonomicName>
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<subSubSection id="C34C72A24B19FF9214F9D4F7FBF1FDCE" lastPageId="7" lastPageNumber="38" pageId="2" pageNumber="33" type="discussion">
<paragraph id="8BE921294B19FF9714F9D4F7FAC5F9DA" blockId="2.[136,1452,1454,1985]" pageId="2" pageNumber="33">
The
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of
<taxonomicName id="4C565AAA4B19FF97153BD4F7FE09FA26" authorityName="Boissier, G" authorityYear="1838" box="[330,489,1454,1480]" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="2" pageNumber="33" phylum="Tracheophyta" rank="species" species="pinsapo">
<emphasis id="B922FD3B4B19FF97153BD4F7FE09FA26" box="[330,489,1454,1480]" italics="true" pageId="2" pageNumber="33">Abies pinsapo</emphasis>
</taxonomicName>
was designated by
<bibRefCitation id="EFC75CD84B19FF9716B9D4F7FC02FA27" author="Burdet, H. M. &amp; Charpin, A. &amp; Jacquemoud, F." box="[712,994,1454,1481]" pageId="2" pageNumber="33" pagination="543 - 584" refId="ref7771" refString="Burdet, H. M., Charpin, A. &amp; Jacquemoud, F. (1981) Types nomenclaturaux des taxa iberiques decrits par Boissier ou Reuter. 1. Gymnospermes a Graminees. Candollea 36: 543 - 584." type="journal article" year="1981">
Burdet
<emphasis id="B922FD3B4B19FF97176CD4F6FCB6FA26" box="[797,854,1454,1480]" italics="true" pageId="2" pageNumber="33">et al.</emphasis>
(1981: 546)
</bibRefCitation>
from a specimen preserved at G, as: “le
<typeStatus id="54ED9F8B4B19FF9714F9D48BFF0FFA02" box="[136,239,1490,1516]" pageId="2" pageNumber="33" type="lectotype">lectotype</typeStatus>
est le rameau de droite sur la feuille munie de létiquette a” [the
<typeStatus id="54ED9F8B4B19FF9717AAD48BFBA2FA02" box="[987,1090,1490,1516]" pageId="2" pageNumber="33" type="lectotype">lectotype</typeStatus>
is the right branch on the sheet with the label a]. This specimen is now barcoded as G00047732.
<bibRefCitation id="EFC75CD84B19FF97172DD4AFFBCDF9FE" author="Farjon, A." box="[860,1069,1526,1552]" pageId="2" pageNumber="33" refId="ref8781" refString="Farjon, A. (2010) A handbook of the world's conifers, vol. 1. Brill Academic Publishers, Leiden &amp; Boston, 1111 pp. https: // doi. org / 10.1163 / 9789047430629" type="book" year="2010">Farjon (2010: 112)</bibRefCitation>
mentioned “Type:
<collectingCountry id="F34161B94B19FF971173D4AFFAA6F9FE" box="[1282,1350,1526,1552]" name="Spain" pageId="2" pageNumber="33">Spain</collectingCountry>
: Malaga, Sierra Bermeja, N. of Estepona [“Sierra de la Nieve”], E. Boissier s.n. (
<typeStatus id="54ED9F8B4B19FF9717D6D743FBEBF9DA" box="[935,1035,1562,1588]" pageId="2" pageNumber="33" type="holotype">holotype</typeStatus>
not located, isotype K)”.
</paragraph>
<paragraph id="8BE921294B19FF9714CCD766FD76F92A" blockId="2.[136,1452,1454,1985]" pageId="2" pageNumber="33">
In the Boissier herbarium at G (G-BOIS) there are two relevant specimens. The specimen with barcode G00047732 bears material collected in 1837 and is mounted on six herbarium sheets (
<figureCitation id="136D3DAC4B19FF9717BED73BFBF5F992" box="[975,1045,1634,1660]" captionStart="FIGURE 2" captionStartId="4.[136,229,1982,2004]" captionTargetBox="[392,1195,497,1958]" captionTargetId="figure-174@4.[392,1195,497,1958]" captionTargetPageId="4" captionText="FIGURE 2. Lectotype of Abies pinsapo Boissier, G (barcode G00047732 “le lectotype est le rameau de droite sur la feuille munie de létiquette a” [the lectotype is the right branch on the sheet with the label a]). Photography courtesy of Herbarium G; reproduced with permission." figureDoi="http://doi.org/10.5281/zenodo.6605308" httpUri="https://zenodo.org/record/6605308/files/figure.png" pageId="2" pageNumber="33">Fig. 2</figureCitation>
), with a label handwritten by Pablo Prolongo García (
<figureCitation id="136D3DAC4B19FF971524D7DFFE7CF94E" box="[341,412,1670,1696]" captionStart="FIGURE 3" captionStartId="5.[136,229,1685,1707]" captionTargetBox="[164,1423,190,1660]" captionTargetId="figure-22@5.[164,1423,190,1660]" captionTargetPageId="5" captionText="FIGURE 3. Handwritten label by Pablo Prolongo García in the sheet that bears the lectotype of Abies pinsapo Boissier. Photography courtesy of Herbarium G; reproduced with permission." figureDoi="http://doi.org/10.5281/zenodo.6605312" httpUri="https://zenodo.org/record/6605312/files/figure.png" pageId="2" pageNumber="33">Fig. 3</figureCitation>
). The label contains a complete diagnosis of this plant. The herbarium sheets also contain several original labels handwritten by Boissier.
</paragraph>
<paragraph id="8BE921294B19FF9714CCD796FE0CF8DF" blockId="2.[136,1452,1454,1985]" pageId="2" pageNumber="33">
However,
<bibRefCitation id="EFC75CD84B19FF971541D797FDA0F907" author="Burdet, H. M. &amp; Charpin, A. &amp; Jacquemoud, F." box="[304,576,1742,1769]" pageId="2" pageNumber="33" pagination="543 - 584" refId="ref7771" refString="Burdet, H. M., Charpin, A. &amp; Jacquemoud, F. (1981) Types nomenclaturaux des taxa iberiques decrits par Boissier ou Reuter. 1. Gymnospermes a Graminees. Candollea 36: 543 - 584." type="journal article" year="1981">
Burdet
<emphasis id="B922FD3B4B19FF9715F3D796FE59F906" box="[386,441,1742,1768]" italics="true" pageId="2" pageNumber="33">et al.</emphasis>
(1981: 546)
</bibRefCitation>
mentioned that the specimen G00047732 is mounted on five herbarium sheets: “G, ex herbier Boissier (5 feuilles)”, and the
<typeStatus id="54ED9F8B4B19FF9716F7D7ABFD18F8E2" box="[646,760,1778,1804]" pageId="2" pageNumber="33" type="lectotype">lectotypes</typeStatus>
is only a stem preserved on the sheet with the label “a” (i.e., the first sheet described above).
</paragraph>
<paragraph id="8BE921294B19FF9714CCD663FA7FF82F" blockId="2.[136,1452,1454,1985]" pageId="2" pageNumber="33">
On the other hand, there is another specimen at G, with barcode G00349514, which bears an original specimen of
<taxonomicName id="4C565AAA4B19FF9714F9D606FEE0F896" authorityName="Boissier, G" authorityYear="1838" box="[136,256,1887,1913]" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="2" pageNumber="33" phylum="Tracheophyta" rank="species" species="pinsapo">
<emphasis id="B922FD3B4B19FF9714F9D606FEE0F896" box="[136,256,1887,1913]" italics="true" pageId="2" pageNumber="33">A. pinsapo</emphasis>
</taxonomicName>
and a label handwritten by Reuter: “Herb. Boissier /
<taxonomicName id="4C565AAA4B19FF971723D607FBA0F896" authority="Boiss." authorityName="Boissier, G" authorityYear="1838" box="[850,1088,1886,1912]" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="2" pageNumber="33" phylum="Tracheophyta" rank="species" species="pinsapo">Abies Pinsapo Boiss.</taxonomicName>
El / Sierra dEstepona et Sa de / la Nieve alt 35006000”. In addition, there are two herbarium sheets at K and P that bears original material collected in
<quantity id="4CAE8CCC4B19FF9714D4D6FFFF1AF82E" box="[165,250,1958,1984]" metricMagnitude="1" metricUnit="m" metricValue="4.66598" pageId="2" pageNumber="33" unit="in" value="1837.0">1837 in</quantity>
“In Sierra dEstepona et / Sª de la Nieve” (K barcoded K000288176 (
<figureCitation id="136D3DAC4B19FF971078D6FFFBADF82E" box="[1033,1101,1958,1984]" captionStart="FIGURE 4" captionStartId="6.[136,229,1517,1539]" captionTargetBox="[340,1247,190,1493]" captionTargetId="figure-20@6.[340,1247,190,1493]" captionTargetPageId="6" captionText="FIGURE 4. Original specimen of A. pinsapo Boissier, K (barcode K000288176). Photography courtesy of Herbarium K; reproduced with permission." figureDoi="http://doi.org/10.5281/zenodo.6605316" httpUri="https://zenodo.org/record/6605316/files/figure.png" pageId="2" pageNumber="33">Fig. 4</figureCitation>
) and P barcoded P01635323).
</paragraph>
<caption id="DF2971A14B18FF9614F9D762FDDBF99B" ID-DOI="http://doi.org/10.5281/zenodo.6605306" ID-Zenodo-Dep="6605306" httpUri="https://zenodo.org/record/6605306/files/figure.png" pageId="3" pageNumber="34" startId="3.[136,229,1595,1617]" targetBox="[340,1247,187,1568]" targetPageId="3">
<paragraph id="8BE921294B18FF9614F9D762FDDBF99B" blockId="3.[136,1451,1595,1653]" pageId="3" pageNumber="34">
<emphasis id="B922FD3B4B18FF9614F9D762FF1BF9BE" bold="true" box="[136,251,1595,1617]" pageId="3" pageNumber="34">FIGURE 1</emphasis>
. Lectotype of
<taxonomicName id="4C565AAA4B18FF9615E5D762FD04F9BF" authority="Miller, Herb. Clifford" authorityName="Miller, Herb. Clifford" authorityYear="1768" box="[404,740,1595,1617]" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="34" phylum="Tracheophyta" rank="species" species="alba">
<emphasis id="B922FD3B4B18FF9615E5D762FE1EF9BE" box="[404,510,1595,1616]" italics="true" pageId="3" pageNumber="34">Abies alba</emphasis>
Miller, Herb. Clifford
</taxonomicName>
: 449,
<taxonomicName id="4C565AAA4B18FF961754D762FCBAF9BE" box="[805,858,1595,1616]" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="34" phylum="Tracheophyta" rank="genus">
<emphasis id="B922FD3B4B18FF961754D762FCBAF9BE" box="[805,858,1595,1616]" italics="true" pageId="3" pageNumber="34">Abies</emphasis>
</taxonomicName>
2, BM (barcode BM000647435). Photography courtesy of Herbarium BM; reproduced with permission.
</paragraph>
</caption>
<paragraph id="8BE921294B18FF9614CCD7FDFE8CF878" blockId="3.[136,1452,1700,2086]" pageId="3" pageNumber="34">
According to the protologue, Boissier visited two populations in 1837. On the 15
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of May, they visited Sierra Bermeja (Estepona) (see
<bibRefCitation id="EFC75CD84B18FF9615D0D791FD63F90C" author="Boissier, P. E." box="[417,643,1736,1762]" pageId="3" pageNumber="34" pagination="401 - 410" refId="ref7637" refString="Boissier, P. E. (1838 a) Notice sur l' Abies pinsapo. Biblioth. Universelle Geneve ser. 2, 13: 401 - 410." type="journal article" year="1838">Boissier 1838a: 34</bibRefCitation>
, Boissier 18391845(vol. 1): 41, 51); at the end of September, Yunquera (Sierra de las
<collectingCountry id="F34161B94B18FF961553D7B5FE99F8E9" box="[290,377,1772,1799]" name="Saint Kitts and Nevis" pageId="3" pageNumber="34">Nieves)</collectingCountry>
together with David Ravey, Pablo Prolongo, and Félix Haenseler (
<bibRefCitation id="EFC75CD84B18FF961010D7B5FAA4F8E8" author="Boissier, P. E." box="[1121,1348,1772,1798]" pageId="3" pageNumber="34" pagination="401 - 410" refId="ref7637" refString="Boissier, P. E. (1838 a) Notice sur l' Abies pinsapo. Biblioth. Universelle Geneve ser. 2, 13: 401 - 410." type="journal article" year="1838">Boissier 1838a: 34</bibRefCitation>
, Boissier 18391845(vol. 1): 155,
<bibRefCitation id="EFC75CD84B18FF9615D1D649FD58F8C4" author="Gonzalez Bueno, A." box="[416,696,1808,1834]" pageId="3" pageNumber="34" pagination="5 - 21" refId="ref9515" refString="Gonzalez Bueno, A. (2010) El viaje botanico de Pierre Edmond Boissier por el sur de Espana (1837). Acta Botanica Malacitana 35: 5 - 21. https: // doi. org / 10.24310 / abm. v 35 i 0.2872" type="journal article" year="2010">González Bueno 2010: 9</bibRefCitation>
). Therefore, according to the protologue and the annotations on the labels of the sheets at G, K, and P, there is a mixture in the provenance of the original material. Consequently, it is not posible to know the concrete provenance of the fragment of the specimen that was designated as the
<typeStatus id="54ED9F8B4B18FF961150D601FA68F89C" box="[1313,1416,1880,1906]" pageId="3" pageNumber="34" type="lectotype">lectotype</typeStatus>
by
<bibRefCitation id="EFC75CD84B18FF9614F9D625FE88F878" author="Burdet, H. M. &amp; Charpin, A. &amp; Jacquemoud, F." box="[136,360,1916,1942]" pageId="3" pageNumber="34" pagination="543 - 584" refId="ref7771" refString="Burdet, H. M., Charpin, A. &amp; Jacquemoud, F. (1981) Types nomenclaturaux des taxa iberiques decrits par Boissier ou Reuter. 1. Gymnospermes a Graminees. Candollea 36: 543 - 584." type="journal article" year="1981">
Burdet
<emphasis id="B922FD3B4B18FF9614AAD624FEF4F878" box="[219,276,1916,1942]" italics="true" pageId="3" pageNumber="34">et al.</emphasis>
(1981)
</bibRefCitation>
.
</paragraph>
<paragraph id="8BE921294B18FF9114CCD6F9FCBEFE37" blockId="3.[136,1452,1700,2086]" lastBlockId="4.[136,1452,158,473]" lastPageId="4" lastPageNumber="35" pageId="3" pageNumber="34">
However, according to Boissier (18391845(vol. 2): 586) “Grâce à lamitié de MM. Haenseler et Prolongo, qui ont bien voulu gravir, au printemps, la Sierra de la Nieve, et qui mont envoyé de bons échantillons et des notes prècieuses sur les parties florales de cet arbre, je puis donner ici sa description complète et le figurer dans tous ses états […]”. Thus, Haenseler and Prolongo collected material after Boissiers visit in 1837 (maybe in the spring of 1838) and sent it to Boissier. Boissier used this material to complete the description of the plant and make the drawings of the tables that accompany the new description of the plant (Boissier 18391845(vol. 2): 584586, Tab. CLXVII, CLXVIII and CLXIX). In addition, the material sent by Felix Haenseler and Pablo Prolongo bears some handwritten notes [“et des notes prècieuses sur les parties florales de cet arbre” (Boissier (18391845(vol. 2): 586)]. In the herbarium sheet barcoded G00047732 there are some handwritten labels. A label bears a complete and extense note handwritten by Prolongo (
<figureCitation id="136D3DAC4B1FFF91148FD00BFEA2FE82" box="[254,322,338,364]" captionStart="FIGURE 3" captionStartId="5.[136,229,1685,1707]" captionTargetBox="[164,1423,190,1660]" captionTargetId="figure-22@5.[164,1423,190,1660]" captionTargetPageId="5" captionText="FIGURE 3. Handwritten label by Pablo Prolongo García in the sheet that bears the lectotype of Abies pinsapo Boissier. Photography courtesy of Herbarium G; reproduced with permission." figureDoi="http://doi.org/10.5281/zenodo.6605312" httpUri="https://zenodo.org/record/6605312/files/figure.png" pageId="4" pageNumber="35">Fig. 3</figureCitation>
). In contrast, the material barcoded G00349514, K000288176 (
<figureCitation id="136D3DAC4B1FFF911076D00BFBABFE82" box="[1031,1099,338,364]" captionStart="FIGURE 4" captionStartId="6.[136,229,1517,1539]" captionTargetBox="[340,1247,190,1493]" captionTargetId="figure-20@6.[340,1247,190,1493]" captionTargetPageId="6" captionText="FIGURE 4. Original specimen of A. pinsapo Boissier, K (barcode K000288176). Photography courtesy of Herbarium K; reproduced with permission." figureDoi="http://doi.org/10.5281/zenodo.6605316" httpUri="https://zenodo.org/record/6605316/files/figure.png" pageId="4" pageNumber="35">Fig. 4</figureCitation>
), and P01635323 only contains labels that attributes the material to the first two collections, in which Boissier himself participated (i.e.,
<date id="FFE807E94B1FFF911167D02FFA4BFE7E" box="[1302,1451,374,401]" pageId="4" pageNumber="35" value="1837-05-15">
15 May
<quantity id="4CAE8CCC4B1FFF911102D02FFA4BFE7E" box="[1395,1451,374,400]" metricMagnitude="1" metricUnit="m" metricValue="4.66598" pageId="4" pageNumber="35" unit="in" value="1837.0">1837</quantity>
</date>
in Sierra Bermeja [“Sierra dEstepona” handwritten by Boissier on the labels], and
<date id="FFE807E94B1FFF91104ED0C3FB17FE5A" box="[1087,1271,410,436]" pageId="4" pageNumber="35" value="1837-09">
September
<quantity id="4CAE8CCC4B1FFF9110CED0C3FB17FE5A" box="[1215,1271,410,436]" metricMagnitude="1" metricUnit="m" metricValue="4.66598" pageId="4" pageNumber="35" unit="in" value="1837.0">1837</quantity>
</date>
in Sierra de las
<collectingCountry id="F34161B94B1FFF9114F9D0E7FF36FE36" box="[136,214,446,472]" name="Saint Kitts and Nevis" pageId="4" pageNumber="35">Nieves</collectingCountry>
[“Sª de la Nieve” handwritten by Boissier on the labels]).
</paragraph>
<caption id="DF2971A14B1FFF9114F9D6E7FF19F7F2" ID-DOI="http://doi.org/10.5281/zenodo.6605308" ID-Zenodo-Dep="6605308" httpUri="https://zenodo.org/record/6605308/files/figure.png" pageId="4" pageNumber="35" startId="4.[136,229,1982,2004]" targetBox="[392,1195,497,1958]" targetPageId="4">
<paragraph id="8BE921294B1FFF9114F9D6E7FF19F7F2" blockId="4.[136,1452,1982,2076]" pageId="4" pageNumber="35">
<emphasis id="B922FD3B4B1FFF9114F9D6E7FF1FF83D" bold="true" box="[136,255,1982,2004]" pageId="4" pageNumber="35">FIGURE 2.</emphasis>
Lectotype of
<taxonomicName id="4C565AAA4B1FFF9115FBD6E7FD67F83D" authority="Boissier, G" authorityName="Boissier, G" authorityYear="1838" box="[394,647,1982,2004]" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="35" phylum="Tracheophyta" rank="species" species="pinsapo">
<emphasis id="B922FD3B4B1FFF9115FBD6E7FDF3F83A" box="[394,531,1982,2004]" italics="true" pageId="4" pageNumber="35">Abies pinsapo</emphasis>
Boissier, G
</taxonomicName>
(barcode G00047732 “le lectotype est le rameau de droite sur la feuille munie de létiquette a” [the lectotype is the right branch on the sheet with the label a]). Photography courtesy of Herbarium G; reproduced with permission.
</paragraph>
</caption>
<caption id="DF2971A14B1EFF9014F9D7CCFD78F921" ID-DOI="http://doi.org/10.5281/zenodo.6605312" ID-Zenodo-Dep="6605312" httpUri="https://zenodo.org/record/6605312/files/figure.png" pageId="5" pageNumber="36" startId="5.[136,229,1685,1707]" targetBox="[164,1423,190,1660]" targetPageId="5">
<paragraph id="8BE921294B1EFF9014F9D7CCFD78F921" blockId="5.[136,1452,1685,1743]" pageId="5" pageNumber="36">
<emphasis id="B922FD3B4B1EFF9014F9D7CCFF1FF944" bold="true" box="[136,255,1685,1707]" pageId="5" pageNumber="36">FIGURE 3.</emphasis>
Handwritten label by Pablo Prolongo García in the sheet that bears the lectotype of
<taxonomicName id="4C565AAA4B1EFF901032D7CCFA4CF945" authority="Boissier. Photography" authorityName="Boissier. Photography" box="[1091,1452,1685,1707]" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="5" pageNumber="36" phylum="Tracheophyta" rank="species" species="pinsapo">
<emphasis id="B922FD3B4B1EFF901032D7CCFB2CF945" box="[1091,1228,1685,1707]" italics="true" pageId="5" pageNumber="36">Abies pinsapo</emphasis>
Boissier. Photography
</taxonomicName>
courtesy of Herbarium G; reproduced with permission.
</paragraph>
</caption>
<paragraph id="8BE921294B1EFF9014CCD7A7FF46F86D" blockId="5.[136,1452,1789,2068]" pageId="5" pageNumber="36">
In conclusion, it seems that the material barcoded G00047732 could be considered a mixed gathering. However,
<bibRefCitation id="EFC75CD84B1EFF9014F9D678FE86F8D5" author="Burdet, H. M. &amp; Charpin, A. &amp; Jacquemoud, F." box="[136,358,1825,1851]" pageId="5" pageNumber="36" pagination="543 - 584" refId="ref7771" refString="Burdet, H. M., Charpin, A. &amp; Jacquemoud, F. (1981) Types nomenclaturaux des taxa iberiques decrits par Boissier ou Reuter. 1. Gymnospermes a Graminees. Candollea 36: 543 - 584." type="journal article" year="1981">
Burdet
<emphasis id="B922FD3B4B1EFF9014ADD67BFEF4F8D5" box="[220,276,1825,1851]" italics="true" pageId="5" pageNumber="36">et al.</emphasis>
(1981)
</bibRefCitation>
selected only a concrete fragment as the
<typeStatus id="54ED9F8B4B1EFF901747D678FC7DF8D5" box="[822,925,1825,1851]" pageId="5" pageNumber="36" type="lectotype">lectotype</typeStatus>
(“the
<typeStatus id="54ED9F8B4B1EFF901795D678FBABF8D5" box="[996,1099,1825,1851]" pageId="5" pageNumber="36" type="lectotype">lectotype</typeStatus>
is the right branch on the sheet with the label a”), and this fragment (very poorly preserved) is labeled as “Sª de la Nieve / 1837” by Boissier (
<figureCitation id="136D3DAC4B1EFF90110FD61CFF79F86D" captionStart="FIGURE 2" captionStartId="4.[136,229,1982,2004]" captionTargetBox="[392,1195,497,1958]" captionTargetId="figure-174@4.[392,1195,497,1958]" captionTargetPageId="4" captionText="FIGURE 2. Lectotype of Abies pinsapo Boissier, G (barcode G00047732 “le lectotype est le rameau de droite sur la feuille munie de létiquette a” [the lectotype is the right branch on the sheet with the label a]). Photography courtesy of Herbarium G; reproduced with permission." figureDoi="http://doi.org/10.5281/zenodo.6605308" httpUri="https://zenodo.org/record/6605308/files/figure.png" pageId="5" pageNumber="36">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="8BE921294B1EFF9014CCD6D4FA7EF7FD" blockId="5.[136,1452,1789,2068]" pageId="5" pageNumber="36">
Unfortunately, the rest of the material on the sheets that contains the specimen (or gatherings) barcoded G00047732 cannot be identified as Boissiers gatherings collected in 1837 (see
<emphasis id="B922FD3B4B1EFF90171DD6EBFC7CF825" box="[876,924,1970,1995]" italics="true" pageId="5" pageNumber="36">note</emphasis>
below), or a Haensler and Prolongos gathering sent to Boissier post-protologue. Therefore, it cannot be considered original material. On the other hand, the specimens G00349514, K000288176 (
<figureCitation id="136D3DAC4B1EFF9015CDD6A0FDE0F7FD" box="[444,512,2041,2067]" captionStart="FIGURE 4" captionStartId="6.[136,229,1517,1539]" captionTargetBox="[340,1247,190,1493]" captionTargetId="figure-20@6.[340,1247,190,1493]" captionTargetPageId="6" captionText="FIGURE 4. Original specimen of A. pinsapo Boissier, K (barcode K000288176). Photography courtesy of Herbarium K; reproduced with permission." figureDoi="http://doi.org/10.5281/zenodo.6605316" httpUri="https://zenodo.org/record/6605316/files/figure.png" pageId="5" pageNumber="36">Fig. 4</figureCitation>
), P01635323 certainly belong to the same gathering collected by Boissier in 1837.
</paragraph>
<caption id="DF2971A14B1DFF9314F9D4B4FF19F9C9" ID-DOI="http://doi.org/10.5281/zenodo.6605316" ID-Zenodo-Dep="6605316" httpUri="https://zenodo.org/record/6605316/files/figure.png" pageId="6" pageNumber="37" startId="6.[136,229,1517,1539]" targetBox="[340,1247,190,1493]" targetPageId="6">
<paragraph id="8BE921294B1DFF9314F9D4B4FF19F9C9" blockId="6.[136,1452,1517,1575]" pageId="6" pageNumber="37">
<emphasis id="B922FD3B4B1DFF9314F9D4B4FF1DF9EC" bold="true" box="[136,253,1517,1539]" pageId="6" pageNumber="37">FIGURE 4.</emphasis>
Original specimen of
<taxonomicName id="4C565AAA4B1DFF9315A1D4B7FD46F9ED" authority="Boissier, K" authorityName="Boissier, K" box="[464,678,1517,1539]" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="37" phylum="Tracheophyta" rank="species" species="pinsapo">
<emphasis id="B922FD3B4B1DFF9315A1D4B7FDD6F9ED" box="[464,566,1518,1539]" italics="true" pageId="6" pageNumber="37">A. pinsapo</emphasis>
Boissier, K
</taxonomicName>
(barcode K000288176). Photography courtesy of Herbarium K; reproduced with permission.
</paragraph>
</caption>
<paragraph id="8BE921294B1DFF9214CCD70FFC44FF32" blockId="6.[136,1452,1622,2080]" lastBlockId="7.[136,1452,158,545]" lastPageId="7" lastPageNumber="38" pageId="6" pageNumber="37">
On the other hand, as for the authors of the name,
<taxonomicName id="4C565AAA4B1DFF931775D70FFC45F99E" box="[772,933,1622,1648]" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="37" phylum="Tracheophyta" rank="species" species="pinsapo">
<emphasis id="B922FD3B4B1DFF931775D70FFC45F99E" box="[772,933,1622,1648]" italics="true" pageId="6" pageNumber="37">Abies pinsapo</emphasis>
</taxonomicName>
was described for the first time by Clemente in the first decade of the 19th century (
<bibRefCitation id="EFC75CD84B1DFF931635D723FCFDF97A" author="Arista, M. &amp; Herrera, J. &amp; Talavera, S." box="[580,797,1658,1684]" pageId="6" pageNumber="37" pagination="427 - 436" refId="ref7101" refString="Arista, M., Herrera, J. &amp; Talavera, S. (1997 b) Abies pinsapo Boiss.: a protected species in a protected area. Bocconea 7: 427 - 436." type="journal article" year="1997">
Arista
<emphasis id="B922FD3B4B1DFF9316E2D722FD2CF97A" box="[659,716,1658,1684]" italics="true" pageId="6" pageNumber="37">et al.</emphasis>
1997b
</bibRefCitation>
,
<bibRefCitation id="EFC75CD84B1DFF931758D723FC5AF97A" author="Vargas, M. A." box="[809,954,1658,1684]" pageId="6" pageNumber="37" pagination="95 - 120" refId="ref12202" refString="Vargas, M. A. (2020) Los bosques de la Serrania de Ronda: La mirada ilustrada de un botanico decimononico, Simon de Rojas Clemente Rubio. In: Gomez Zotano, J. &amp; Olmedo Cobo, J. A. (Eds.) Los bosques de la Serrania de Ronda. Una perspectiva espacio-temporal. Alcala del Valle, Editorial La Serrania, pp. 95 - 120." type="book chapter" year="2020">Vargas 2020</bibRefCitation>
). However, this description was lost. In this sense,
<bibRefCitation id="EFC75CD84B1DFF9314A2D7C7FE37F956" author="Cabezudo, B. &amp; Perez Latorre, A. V. &amp; Casimiro-Soriguer Solanas, F. &amp; Gavia Romero, O. &amp; Garcia Rojas, J. A. &amp; Pavon, M." box="[211,471,1694,1720]" pageId="6" pageNumber="37" pagination="195 - 204" refId="ref7814" refString="Cabezudo, B., Perez Latorre, A. V., Casimiro-Soriguer Solanas, F., Gavia Romero, O., Garcia Rojas, J. A. &amp; Pavon, M. (2010) Nota sobre Abies pinsapo Clemente ex Boiss. y otras plantas andaluzas. Acta Botanica Malacitana 35: 195 - 204. https: // doi. org / 10.24310 / abm. v 35 i 0.2867" type="journal article" year="2010">
Cabezudo
<emphasis id="B922FD3B4B1DFF93153AD7C6FE64F956" box="[331,388,1694,1720]" italics="true" pageId="6" pageNumber="37">et al.</emphasis>
(2010)
</bibRefCitation>
suggest that this authorship of the Spanish botanist Simón de Rojas Clemente y Rubio should be recognized in the species name, as “
<taxonomicName id="4C565AAA4B1DFF9316EED79BFBC5F932" authority="Clemente ex Boiss." authorityName="Boiss." box="[671,1061,1730,1756]" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="37" phylum="Tracheophyta" rank="species" species="pinsapo">
<emphasis id="B922FD3B4B1DFF9316EED79BFCDEF932" box="[671,830,1730,1756]" italics="true" pageId="6" pageNumber="37">Abies pinsapo</emphasis>
Clemente ex Boiss.
</taxonomicName>
”, since there is a Clementes work (see Clemente 1818, 2002) in which the epithet “
<taxonomicName id="4C565AAA4B1DFF9316C1D7BEFCE9F8EE" box="[688,777,1767,1792]" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="37" phylum="Tracheophyta" rank="species" species="pinsapo">
<emphasis id="B922FD3B4B1DFF9316C1D7BEFCE9F8EE" box="[688,777,1767,1792]" italics="true" pageId="6" pageNumber="37">pinsapo</emphasis>
</taxonomicName>
” was first mentioned.
<bibRefCitation id="EFC75CD84B1DFF931079D7BFFAE9F8EE" author="Cabezudo, B. &amp; Perez Latorre, A. V. &amp; Casimiro-Soriguer Solanas, F. &amp; Gavia Romero, O. &amp; Garcia Rojas, J. A. &amp; Pavon, M." box="[1032,1289,1766,1792]" pageId="6" pageNumber="37" pagination="195 - 204" refId="ref7814" refString="Cabezudo, B., Perez Latorre, A. V., Casimiro-Soriguer Solanas, F., Gavia Romero, O., Garcia Rojas, J. A. &amp; Pavon, M. (2010) Nota sobre Abies pinsapo Clemente ex Boiss. y otras plantas andaluzas. Acta Botanica Malacitana 35: 195 - 204. https: // doi. org / 10.24310 / abm. v 35 i 0.2867" type="journal article" year="2010">
Cabezudo
<emphasis id="B922FD3B4B1DFF93100ED7BEFB58F8EE" box="[1151,1208,1766,1792]" italics="true" pageId="6" pageNumber="37">et al.</emphasis>
(2010)
</bibRefCitation>
also indicated “Con toda seguridad Clemente comunicó estos datos a sus discípulos malagueños Haenseler y Prolongo, que a su vez lo comunicaron a Boissier, seguramente olvidándose de la paternidad de Clemente sobre este asunto” [Clemente certainly communicated these data to his Malaga disciples Haenseler and Prolongo, who in turn communicated it to Boissier, surely forgetting about Clementes paternity on this matter]. Unfortunately, Clementes work was not cited in the protologue, and nor does it seem that there is an ascription of the name
<taxonomicName id="4C565AAA4B1DFF93179CD6C3FB85F85A" box="[1005,1125,1946,1972]" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="37" phylum="Tracheophyta" rank="species" species="pinsapo">
<emphasis id="B922FD3B4B1DFF93179CD6C3FB85F85A" box="[1005,1125,1946,1972]" italics="true" pageId="6" pageNumber="37">A. pinsapo</emphasis>
</taxonomicName>
to Clemente, therefore is not possible to demonstrate this affirmation. According to
<bibRefCitation id="EFC75CD84B1DFF931680D6E7FBCAF836" author="Gonzalez Bueno, A." box="[753,1066,1982,2008]" pageId="6" pageNumber="37" pagination="5 - 21" refId="ref9515" refString="Gonzalez Bueno, A. (2010) El viaje botanico de Pierre Edmond Boissier por el sur de Espana (1837). Acta Botanica Malacitana 35: 5 - 21. https: // doi. org / 10.24310 / abm. v 35 i 0.2872" type="journal article" year="2010">González Bueno (2010: 19)</bibRefCitation>
“No parece que Edmond Boissier tuviera acceso a los manuscritos de Simón de Rojas Clemente que compusieron su Historia Natural del Reino de
<collectingCountry id="F34161B94B1DFF9314F9D95FFF0AF7CE" box="[136,234,2054,2080]" name="Grenada" pageId="6" pageNumber="37">Granada</collectingCountry>
” [It does not seem that Edmond Boissier had access to the manuscripts of Simón de Rojas Clemente that composed his Natural History of the Kingdom of
<collectingCountry id="F34161B94B1CFF9216C2D1C7FCF2FF56" box="[691,786,158,184]" name="Grenada" pageId="7" pageNumber="38">Granada</collectingCountry>
(i.e., Clemente 2002)]. In conclusion, the species was cited without ascription to Clemente (
<bibRefCitation id="EFC75CD84B1CFF921580D19BFC77FF32" author="Galan de Mera, A. &amp; Perez Latorre, A. V." box="[497,919,194,221]" pageId="7" pageNumber="38" pagination="127 - 129" refId="ref9348" refString="Galan de Mera, A. &amp; Perez Latorre, A. V. (2021) Sobre la autoria de Abies pinsapo Boiss. Acta Botanica Malacitana 46: 127 - 129. https: // doi. org / 10.24310 / abm. v 46 i. 12827" type="journal article" year="2021">Galán de Mera &amp; Pérez Latorre 2021</bibRefCitation>
).
</paragraph>
<paragraph id="8BE921294B1CFF9214CCD1BFFBF1FDCE" blockId="7.[136,1452,158,545]" pageId="7" pageNumber="38">
Finally, a relevant comment was included in the protologue referring to the original material used by Boissier to describe his
<taxonomicName id="4C565AAA4B1CFF92154AD053FE3CFECA" box="[315,476,266,292]" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="7" pageNumber="38" phylum="Tracheophyta" rank="species" species="pinsapo">
<emphasis id="B922FD3B4B1CFF92154AD053FE3CFECA" box="[315,476,266,292]" italics="true" pageId="7" pageNumber="38">Abies pinsapo</emphasis>
</taxonomicName>
: “Examinant, en
<date id="FFE807E94B1CFF9216D8D053FCC2FECA" box="[681,802,266,292]" pageId="7" pageNumber="38" value="1837-04">abril 1837</date>
, quelques plantes recueillies sur la Sierra Bermeja, près dEstepona, par M. Haenseler, qui cultive avec succès la botanique à Malaga, je trouvai, parmi elles, une branche de conifère dont les feuilles, extrêmement courtes, épaisses, presque sédiformes, me frappèrent. Il ny avait pas moyen, à cause de labsence du fruit, de déterminer le genre de cette plante, qui avait cependant le port dun abies.” (
<bibRefCitation id="EFC75CD84B1CFF92113FD02FFF0AFE5A" author="Boissier, P. E." pageId="7" pageNumber="38" pagination="401 - 410" refId="ref7637" refString="Boissier, P. E. (1838 a) Notice sur l' Abies pinsapo. Biblioth. Universelle Geneve ser. 2, 13: 401 - 410." type="journal article" year="1838">Boissier 1838a: 3</bibRefCitation>
, see also
<bibRefCitation id="EFC75CD84B1CFF921524D0C3FDCAFE5A" author="Boissier, P. E." box="[341,554,410,436]" pageId="7" pageNumber="38" refId="ref7667" refString="Boissier, P. E. (1838 b) Elenchus plantarum novarum minusque cognitarum quas in itinere Hispanico. Typographia Lador et Ramboz, Genevae, 94 pp. https: // doi. org / 10.5962 / bhl. title. 52301" type="book" year="1838">Boissier 1838b: 84</bibRefCitation>
, Boissier 18391845(vol. 1): 49, (vol. 2): 585). In this sentence, Boissier clearly mentioned a
<typeStatus id="54ED9F8B4B1CFF921568D0E6FE93FE37" box="[281,371,447,473]" pageId="7" pageNumber="38" type="syntype">syntype</typeStatus>
, collected by Haenseler in “Sierra Bermeja, près dEstepona [without date]”. Unfortunately, this material is lost (
<bibRefCitation id="EFC75CD84B1CFF921535D0BBFDA7FE12" author="Gonzalez Bueno, A." box="[324,583,482,509]" pageId="7" pageNumber="38" pagination="5 - 21" refId="ref9515" refString="Gonzalez Bueno, A. (2010) El viaje botanico de Pierre Edmond Boissier por el sur de Espana (1837). Acta Botanica Malacitana 35: 5 - 21. https: // doi. org / 10.24310 / abm. v 35 i 0.2872" type="journal article" year="2010">González Bueno 2010</bibRefCitation>
,
<bibRefCitation id="EFC75CD84B1CFF921624D0BBFCC7FE12" author="Perez-Rubin, J." box="[597,807,482,508]" pageId="7" pageNumber="38" pagination="141 - 162" refId="ref10690" refString="Perez-Rubin, J. (2012) El naturalista y farmaceutico germane-espanol Felix Hanseler Jeger (1780 - 1841) en la Malaga de su epoca. Acta Botanica Malacitana 37: 141 - 162. https: // doi. org / 10.24310 / abm. v 37 i 0.2685" type="journal article" year="2012">Pérez-Rubín 2012</bibRefCitation>
), but if it were located, it would have preference in the
<typeStatus id="54ED9F8B4B1CFF9214F9D35FFF0FFDCE" box="[136,239,518,544]" pageId="7" pageNumber="38" type="lectotype">lectotype</typeStatus>
designation according to Art. 9.12 of the
<emphasis id="B922FD3B4B1CFF9216B0D35EFD10FDCF" box="[705,752,519,545]" italics="true" pageId="7" pageNumber="38">ICN</emphasis>
(see
<bibRefCitation id="EFC75CD84B1CFF92175AD35FFBE5FDCE" author="Turland, N. J. &amp; Wiersema, J. H. &amp; Barrie, F. R. &amp; Greuter, W. &amp; Hawksworth, D. L. &amp; Herendeen, P. S. &amp; Knapp, S. &amp; Kusber, W. - H. &amp; Li, D. - Z. &amp; Marhold, K. &amp; May, T. W. &amp; McNeill, J. &amp; Monro, A. M. &amp; Prado, J. &amp; Price, M. J. &amp; Smith, G. F." box="[811,1029,518,544]" pageId="7" pageNumber="38" refId="ref11897" refString="Turland, N. J., Wiersema, J. H., Barrie, F. R., Greuter, W., Hawksworth, D. L., Herendeen, P. S., Knapp, S., Kusber, W. - H., Li, D. - Z., Marhold, K., May, T. W., McNeill, J., Monro, A. M., Prado, J., Price, M. J. &amp; Smith, G. F. (eds.) (2018) International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017. Regnum Vegetabile 159. Koeltz Botanical Books, Glashutten. https: // doi. org / 10.12705 / Code. 2018" type="book" year="2018">
Turland
<emphasis id="B922FD3B4B1CFF9217F8D35EFC21FDCE" box="[905,961,518,544]" italics="true" pageId="7" pageNumber="38">et al.</emphasis>
2018
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>