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<document id="2282B837499097D4DE4A331312CFAC8D" ID-DOI="10.5281/zenodo.6391857" ID-ISSN="1942-1354" ID-Zenodo-Dep="6391857" ID-ZooBank="E80F6A2A-2E1A-479F-BBEF-A5E3BB84E7C9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1648503991815" checkinUser="felipe" docAuthor="Muona, Jyrki" docDate="2022" docId="03F5F45FFF9A6305FF1CFD5AFD2D5A2C" docLanguage="en" docName="InsectaMundi.2022.0912.1-11.pdf" docOrigin="Insecta Mundi 2022 (912)" docStyle="DocumentStyle:F9976E9B7F8A094D890004515A7A0592.3:InsectaMundi.2020-.journal_article" docStyleId="F9976E9B7F8A094D890004515A7A0592" docStyleName="InsectaMundi.2020-.journal_article" docStyleVersion="3" docTitle="Thambus woodruffi Muona 2022, new species" docType="treatment" docVersion="3" lastPageNumber="9" masterDocId="FFCC8C27FF9D630FFFDCFFCDFFAD5B51" masterDocTitle="New fossil false click-beetles from the Americas (Coleoptera: Eucnemidae)" masterLastPageNumber="11" masterPageNumber="1" pageNumber="6" updateTime="1699308786968" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
<mods:mods id="7956E482E7474042EF7EB08CB95F7BBB" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C819BB6AB983C9ACF9E948438A5CAD88">
<mods:title id="C4A5AC596E6C5C560B5CA2D1607BD2B4">New fossil false click-beetles from the Americas (Coleoptera: Eucnemidae)</mods:title>
</mods:titleInfo>
<mods:name id="CA9E2B574B4F74CEC81921E44D1463D2" type="personal">
<mods:role id="595183B0FA0050BA47B4013AEC3DBCC9">
<mods:roleTerm id="CFB618250D2897A60937D6CC19B7C104">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9F8E903E299E93AA3111EEDFF2D1B03D">Muona, Jyrki</mods:namePart>
<mods:affiliation id="6F92D79721A85B3AC6F79C752D20F9DF">Finnish Museum of Natural History P. O. Box 17 FIN- 00014 University of Helsinki Helsinki, Finland</mods:affiliation>
</mods:name>
<mods:typeOfResource id="E2FCD543572650D28C186BD9165DD461">text</mods:typeOfResource>
<mods:relatedItem id="82E8AA9D8A0E8876EC42623A25B7FD06" type="host">
<mods:titleInfo id="945A1A23D0B0F6157EDC1A9BE36B4BE8">
<mods:title id="7BD87036D7B2C35286D99D8B0F7EB747">Insecta Mundi</mods:title>
</mods:titleInfo>
<mods:part id="ADEBE8D2AFBDCE50E30546585CD6F86A">
<mods:date id="120216A55F3A1E6DEA4EE5C8CA2043A1">2022</mods:date>
<mods:detail id="2B76411226B20F14C1E797829E386C25" type="pubDate">
<mods:number id="52B907FEE6960C1BD5716CA852035BF5">2022-02-24</mods:number>
</mods:detail>
<mods:detail id="C6EADD8A5D95FE473B7B235E3DE8FEE4" type="volume">
<mods:number id="66570F0306BDF1EBFB14A032B5649F6F">2022</mods:number>
</mods:detail>
<mods:detail id="6AAA27B0370B14C95935C3A557683B24" type="issue">
<mods:number id="9BC07156BA30E39D8F3C834EBE6D839F">912</mods:number>
</mods:detail>
<mods:extent id="0A7A378396A043028DC8E345676F186C" unit="page">
<mods:start id="8AF93C05D1424A8EB9AC89EA9B570551">1</mods:start>
<mods:end id="D4127C20014BBC66E2C32D266261B7D6">11</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="5FF7FF5F45207E960EE005B10D916E09">journal article</mods:classification>
<mods:identifier id="E856EA20B141EEF21521CA14AE5897CD" type="DOI">10.5281/zenodo.6391857</mods:identifier>
<mods:identifier id="9E7C7724A2A3955B70657058E8E1C341" type="ISSN">1942-1354</mods:identifier>
<mods:identifier id="0E29CCD775FE8B9621AB1AAFE9AF7400" type="Zenodo-Dep">6391857</mods:identifier>
<mods:identifier id="F02BDC0C05E0ED1BF592207AEB93B452" type="ZooBank">E80F6A2A-2E1A-479F-BBEF-A5E3BB84E7C9</mods:identifier>
</mods:mods>
<treatment id="03F5F45FFF9A6305FF1CFD5AFD2D5A2C" ID-DOI="http://doi.org/10.5281/zenodo.6391863" ID-Zenodo-Dep="6391863" LSID="urn:lsid:plazi:treatment:03F5F45FFF9A6305FF1CFD5AFD2D5A2C" httpUri="http://treatment.plazi.org/id/03F5F45FFF9A6305FF1CFD5AFD2D5A2C" lastPageId="10" lastPageNumber="9" pageId="7" pageNumber="6">
<subSubSection id="C34616C2FF9A6308FF1CFD5AFD1459E5" box="[192,697,663,692]" pageId="7" pageNumber="6" type="nomenclature">
<paragraph id="8BE34549FF9A6308FF1CFD5AFD1459E5" blockId="7.[192,697,663,725]" box="[192,697,663,692]" pageId="7" pageNumber="6">
<heading id="D0ABF225FF9A6308FF1CFD5AFD1459E5" box="[192,697,663,692]" fontSize="12" level="2" pageId="7" pageNumber="6" reason="2">
<emphasis id="B928995BFF9A6308FF1CFD5AFD1459E5" bold="true" box="[192,697,663,692]" pageId="7" pageNumber="6">
<taxonomicName id="4C5C3ECAFF9A6308FF1CFD5AFDBA59E2" authority="Muona" authorityName="Muona" authorityYear="2022" box="[192,535,663,692]" class="Insecta" family="Eucnemidae" genus="Thambus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="woodruffi" status="sp. nov.">
<emphasis id="B928995BFF9A6308FF1CFD5AFE1959E5" bold="true" box="[192,436,663,692]" italics="true" pageId="7" pageNumber="6">Thambus woodruffi</emphasis>
Muona
</taxonomicName>
,
<taxonomicNameLabel id="A21B2420FF9A6308FDF8FD5AFD1459E5" box="[548,697,663,692]" pageId="7" pageNumber="6" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34616C2FF9A6308FF1CFD77FEEA5984" box="[192,327,698,725]" pageId="7" pageNumber="6" type="description">
<paragraph id="8BE34549FF9A6308FF1CFD77FEEA5984" blockId="7.[192,697,663,725]" box="[192,327,698,725]" pageId="7" pageNumber="6">
<figureCitation id="136759CCFF9A6308FF1CFD77FE855985" box="[192,296,698,724]" captionStart="Figure 6" captionStartId="7.[194,270,1900,1926]" captionTargetBox="[192,1440,1205,1871]" captionTargetId="figure-280@7.[192,1440,1205,1871]" captionTargetPageId="7" captionText="Figure 6. Thambus woodruffi n. sp., holotype, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.6391873" httpUri="https://zenodo.org/record/6391873/files/figure.png" pageId="7" pageNumber="6">Figures 6</figureCitation>
,
<figureCitation id="136759CCFF9A6308FEEFFD76FEEE5984" box="[307,323,699,725]" captionStart="Figure 7" captionStartId="8.[192,268,1140,1166]" captionTargetBox="[192,1440,213,1110]" captionTargetId="figure-336@8.[192,1440,213,1110]" captionTargetPageId="8" captionText="Figure 7. Thambus woodruffi n. sp., holotype, ventral view." figureDoi="http://doi.org/10.5281/zenodo.6391875" httpUri="https://zenodo.org/record/6391875/files/figure.png" pageId="7" pageNumber="6">7</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34616C2FF9A6308FF1CFD29FDD85871" pageId="7" pageNumber="6" type="etymology">
<paragraph id="8BE34549FF9A6308FF1CFD29FDD85871" blockId="7.[192,1440,740,800]" pageId="7" pageNumber="6">
<emphasis id="B928995BFF9A6308FF1CFD29FEE959AF" bold="true" box="[192,324,740,766]" pageId="7" pageNumber="6">Etymology.</emphasis>
Named in honor of the late Robert E. Woodruff, whose many interests included fossil resins and especially
<collectingCountry id="F34B05D9FF9A6308FEEEFCCBFE765871" box="[306,475,774,800]" name="Dominican Republic" pageId="7" pageNumber="6">the Dominican</collectingCountry>
amber fauna.
</paragraph>
</subSubSection>
<subSubSection id="C34616C2FF9A6308FF1CFCE2FB72583A" pageId="7" pageNumber="6" type="materials_examined">
<paragraph id="8BE34549FF9A6308FF1CFCE2FB72583A" blockId="7.[192,1440,815,875]" pageId="7" pageNumber="6">
<emphasis id="B928995BFF9A6308FF1CFCE2FE995818" bold="true" box="[192,308,815,841]" pageId="7" pageNumber="6">
<typeStatus id="54E7FBEBFF9A6308FF1CFCE2FE825818" box="[192,303,815,841]" pageId="7" pageNumber="6" type="holotype">Holotype</typeStatus>
.
</emphasis>
Embedded in flat, rectangular Dominican resin piece with broadly rounder corners,
<quantity id="4CA4E8ACFF9A6308FB3CFCE2FA82581B" box="[1248,1327,815,842]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="7" pageNumber="6" unit="mm" value="17.0">17 mm</quantity>
×
<quantity id="4CA4E8ACFF9A6308FA97FCE2FA31581B" box="[1355,1436,815,842]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="7" pageNumber="6" unit="mm" value="11.0">11 mm</quantity>
, female. The same piece includes an unidentified throscid beetle as well as other insects (
<figureCitation id="136759CCFF9A6308FB53FC9CFB7C583A" box="[1167,1233,849,875]" captionStart="Figure 7" captionStartId="8.[192,268,1140,1166]" captionTargetBox="[192,1440,213,1110]" captionTargetId="figure-336@8.[192,1440,213,1110]" captionTargetPageId="8" captionText="Figure 7. Thambus woodruffi n. sp., holotype, ventral view." figureDoi="http://doi.org/10.5281/zenodo.6391875" httpUri="https://zenodo.org/record/6391875/files/figure.png" pageId="7" pageNumber="6">Fig. 7</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34616C2FF9A6308FF1CFCB7FA9458C4" box="[192,1337,890,917]" pageId="7" pageNumber="6" type="diagnosis">
<paragraph id="8BE34549FF9A6308FF1CFCB7FA9458C4" blockId="7.[192,1337,890,917]" box="[192,1337,890,917]" pageId="7" pageNumber="6">
<emphasis id="B928995BFF9A6308FF1CFCB7FE9158C5" bold="true" box="[192,316,890,916]" pageId="7" pageNumber="6">Diagnosis.</emphasis>
The length/width ratio of all other
<taxonomicName id="4C5C3ECAFF9A6308FD19FCB7FCD258C5" box="[709,895,890,916]" pageId="7" pageNumber="6">
<emphasis id="B928995BFF9A6308FD19FCB7FC8758C5" box="[709,810,890,916]" italics="true" pageId="7" pageNumber="6">Thambus</emphasis>
species
</taxonomicName>
is 2.4–2.8, in this species it is about 2.1.
</paragraph>
</subSubSection>
<subSubSection id="C34616C2FF9A6307FF1CFC69FC735DDF" lastPageId="8" lastPageNumber="7" pageId="7" pageNumber="6" type="description">
<paragraph id="8BE34549FF9A6308FF1CFC69FCF15F14" blockId="7.[192,1441,932,1161]" pageId="7" pageNumber="6">
<emphasis id="B928995BFF9A6308FF1CFC69FEFF58EF" bold="true" box="[192,338,932,958]" pageId="7" pageNumber="6">Description.</emphasis>
<taxonomicName id="4C5C3ECAFF9A6308FE82FC69FE7158EF" authorityName="Leach" authorityYear="1815" box="[350,476,932,958]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="2" phylum="Arthropoda" rank="superFamily" superFamily="Elateroidea">Elateroidea</taxonomicName>
synapomorphy: pro-mesocoxal clicking mechanism present.
<taxonomicName id="4C5C3ECAFF9A6308FB73FC69FA9758EF" authorityName="Eschscholtz" authorityYear="1829" box="[1199,1338,932,958]" class="Insecta" family="Eucnemidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="family">Eucnemidae</taxonomicName>
synapomorphies: pedicel attached subapically to scape, labrum hidden, elytral striae apically grooved with excretory punctures (
<figureCitation id="136759CCFF9A6308FE9CFC2AFE2E5F53" box="[320,387,999,1026]" captionStart="Figure 6" captionStartId="7.[194,270,1900,1926]" captionTargetBox="[192,1440,1205,1871]" captionTargetId="figure-280@7.[192,1440,1205,1871]" captionTargetPageId="7" captionText="Figure 6. Thambus woodruffi n. sp., holotype, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.6391873" httpUri="https://zenodo.org/record/6391873/files/figure.png" pageId="7" pageNumber="6">Fig. 6</figureCitation>
).
<taxonomicName id="4C5C3ECAFF9A6308FE45FC2AFD895F50" authorityName="Eschscholtz" authorityYear="1829" box="[409,548,999,1025]" class="Insecta" family="Eucnemidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="family">Eucnemidae</taxonomicName>
other characters: abdominal ventrites connate (
<figureCitation id="136759CCFF9A6308FB9DFC2AFB285F53" box="[1089,1157,999,1026]" captionStart="Figure 7" captionStartId="8.[192,268,1140,1166]" captionTargetBox="[192,1440,213,1110]" captionTargetId="figure-336@8.[192,1440,213,1110]" captionTargetPageId="8" captionText="Figure 7. Thambus woodruffi n. sp., holotype, ventral view." figureDoi="http://doi.org/10.5281/zenodo.6391875" httpUri="https://zenodo.org/record/6391875/files/figure.png" pageId="7" pageNumber="6">Fig. 7</figureCitation>
).
<taxonomicName id="4C5C3ECAFF9A6308FB47FC2AFA925F50" authorityName="Fleutiaux" authorityYear="1922" box="[1179,1343,999,1025]" class="Insecta" family="Eucnemidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="9" phylum="Arthropoda" rank="subFamily" subFamily="Macraulacinae">Macraulacinae</taxonomicName>
synapomorphy: hypomera with basally open lateral antennal grooves (
<figureCitation id="136759CCFF9A6308FCA3FBC4FC6C5F72" box="[895,961,1033,1059]" captionStart="Figure 7" captionStartId="8.[192,268,1140,1166]" captionTargetBox="[192,1440,213,1110]" captionTargetId="figure-336@8.[192,1440,213,1110]" captionTargetPageId="8" captionText="Figure 7. Thambus woodruffi n. sp., holotype, ventral view." figureDoi="http://doi.org/10.5281/zenodo.6391875" httpUri="https://zenodo.org/record/6391875/files/figure.png" pageId="7" pageNumber="6">Fig. 7</figureCitation>
).
<taxonomicName id="4C5C3ECAFF9A6308FC08FBC4FBCA5F72" authorityName="Fleutiaux" authorityYear="1922" box="[980,1127,1033,1059]" class="Insecta" family="Eucnemidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="9" phylum="Arthropoda" rank="tribe" tribe="Macraulacini">Macraulacini</taxonomicName>
other characters: hypomera without excretory pits, metasternum without tarsal grooves.
</paragraph>
<paragraph id="8BE34549FF9A6307FEDCFB81FCAD5D3A" blockId="7.[192,1441,932,1161]" lastBlockId="8.[191,1442,1238,1678]" lastPageId="8" lastPageNumber="7" pageId="7" pageNumber="6">
Length 3.0 mm. Form very wide, ratio L/W about 2.1 (
<figureCitation id="136759CCFF9A6308FCB5FB81FC015F36" box="[873,940,1100,1127]" captionStart="Figure 6" captionStartId="7.[194,270,1900,1926]" captionTargetBox="[192,1440,1205,1871]" captionTargetId="figure-280@7.[192,1440,1205,1871]" captionTargetPageId="7" captionText="Figure 6. Thambus woodruffi n. sp., holotype, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.6391873" httpUri="https://zenodo.org/record/6391873/files/figure.png" pageId="7" pageNumber="6">Fig. 6</figureCitation>
). Pronotum wider than long and wider than elytra with slightly expanded, acute non-carinate hind angles, evenly and densely punctate (
<figureCitation id="136759CCFF9A6308FB0BFBA3FAB65FD8" box="[1239,1307,1134,1161]" captionStart="Figure 6" captionStartId="7.[194,270,1900,1926]" captionTargetBox="[192,1440,1205,1871]" captionTargetId="figure-280@7.[192,1440,1205,1871]" captionTargetPageId="7" captionText="Figure 6. Thambus woodruffi n. sp., holotype, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.6391873" httpUri="https://zenodo.org/record/6391873/files/figure.png" pageId="7" pageNumber="6">Fig. 6</figureCitation>
). Scutellum transverse, widely triangular, apex cut off, elytra with sharp, complete sutural striae with deep apical excretory groove continuing around to lateralmost striae, densely and finely punctate, other striae faint but present (
<figureCitation id="136759CCFF956307FAAAFB34FF7F5E67" captionStart="Figure 6" captionStartId="7.[194,270,1900,1926]" captionTargetBox="[192,1440,1205,1871]" captionTargetId="figure-280@7.[192,1440,1205,1871]" captionTargetPageId="7" captionText="Figure 6. Thambus woodruffi n. sp., holotype, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.6391873" httpUri="https://zenodo.org/record/6391873/files/figure.png" pageId="8" pageNumber="7">Fig. 6</figureCitation>
). Head rounded, densely punctate, eyes large, flat, frons wide, without carinae, frontoclypeus apically about 1.3 times as wide as between antennal sockets, mandibles short, bifid (
<figureCitation id="136759CCFF956307FC40FAF3FC705E09" box="[924,989,1342,1368]" captionStart="Figure 7" captionStartId="8.[192,268,1140,1166]" captionTargetBox="[192,1440,213,1110]" captionTargetId="figure-336@8.[192,1440,213,1110]" captionTargetPageId="8" captionText="Figure 7. Thambus woodruffi n. sp., holotype, ventral view." figureDoi="http://doi.org/10.5281/zenodo.6391875" httpUri="https://zenodo.org/record/6391875/files/figure.png" pageId="8" pageNumber="7">Fig. 7</figureCitation>
). Antennae fairly short, reaching slightly past the hind angles, scape as long as pedicel plus f1 and f2 combined, f2–f3 feebly serrate, f4–f7 not visible, f9 1.4 times as long as f8, both longer than wide. Hypomeron with sharply defined, basally open lateral antennal grooves, about triangular, sparsely punctate, without excretory pits (
<figureCitation id="136759CCFF956307FC71FA68FC405EEE" box="[941,1005,1445,1471]" captionStart="Figure 7" captionStartId="8.[192,268,1140,1166]" captionTargetBox="[192,1440,213,1110]" captionTargetId="figure-336@8.[192,1440,213,1110]" captionTargetPageId="8" captionText="Figure 7. Thambus woodruffi n. sp., holotype, ventral view." figureDoi="http://doi.org/10.5281/zenodo.6391875" httpUri="https://zenodo.org/record/6391875/files/figure.png" pageId="8" pageNumber="7">Fig. 7</figureCitation>
). Prosternum wide, moderately densely punctate, prosternal peg long, triangular, acute, deeply marginate, metasternum wide and long, densely punctate, without tarsal grooves, elytral epipleura simple. Metacoxal plates triangular, widest close to midline, angle tightly rounded and there less than times as wide as on sides, densely punctate. Legs long and delicate, meso- and metatarsi shorter than tibiae, meso- and metatarsomere 1 slightly shorter than the others combined, fourth apically narrowly bilobed, excavated, claw simple (
<figureCitation id="136759CCFF956307FD4AF99CFD7A5D3A" box="[662,727,1617,1643]" captionStart="Figure 6" captionStartId="7.[194,270,1900,1926]" captionTargetBox="[192,1440,1205,1871]" captionTargetId="figure-280@7.[192,1440,1205,1871]" captionTargetPageId="7" captionText="Figure 6. Thambus woodruffi n. sp., holotype, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.6391873" httpUri="https://zenodo.org/record/6391873/files/figure.png" pageId="8" pageNumber="7">Fig. 6</figureCitation>
,
<figureCitation id="136759CCFF956307FD3FF99CFD5E5D3A" box="[739,755,1617,1643]" captionStart="Figure 7" captionStartId="8.[192,268,1140,1166]" captionTargetBox="[192,1440,213,1110]" captionTargetId="figure-336@8.[192,1440,213,1110]" captionTargetPageId="8" captionText="Figure 7. Thambus woodruffi n. sp., holotype, ventral view." figureDoi="http://doi.org/10.5281/zenodo.6391875" httpUri="https://zenodo.org/record/6391875/files/figure.png" pageId="8" pageNumber="7">7</figureCitation>
).
</paragraph>
<caption id="DF2315C1FF9A6308FF1EF8A1FCEA5CD7" ID-DOI="http://doi.org/10.5281/zenodo.6391873" ID-Zenodo-Dep="6391873" box="[194,839,1900,1927]" httpUri="https://zenodo.org/record/6391873/files/figure.png" pageId="7" pageNumber="6" startId="7.[194,270,1900,1926]" targetBox="[192,1440,1205,1871]" targetPageId="7">
<paragraph id="8BE34549FF9A6308FF1EF8A1FCEA5CD7" blockId="7.[194,839,1900,1927]" box="[194,839,1900,1927]" pageId="7" pageNumber="6">
<emphasis id="B928995BFF9A6308FF1EF8A1FE875CD7" bold="true" box="[194,298,1900,1926]" pageId="7" pageNumber="6">Figure 6.</emphasis>
<taxonomicName id="4C5C3ECAFF9A6308FEF2F8A1FDA95CD7" authority="Muona, 2022" authorityName="Muona" authorityYear="2022" box="[302,516,1900,1926]" class="Insecta" family="Eucnemidae" genus="Thambus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="woodruffi" status="sp. nov.">
<emphasis id="B928995BFF9A6308FEF2F8A1FDA95CD7" box="[302,516,1900,1926]" italics="true" pageId="7" pageNumber="6">Thambus woodruffi</emphasis>
</taxonomicName>
<emphasis id="B928995BFF9A6308FDD6F8A1FDEA5CD7" bold="true" box="[522,583,1900,1926]" pageId="7" pageNumber="6">
<taxonomicNameLabel id="A21B2420FF9A6308FDD6F8A1FDEA5CD7" box="[522,583,1900,1926]" pageId="7" pageNumber="6" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, dorsal view.
</paragraph>
</caption>
<caption id="DF2315C1FF956307FF1CFBB9FCFD5FDF" ID-DOI="http://doi.org/10.5281/zenodo.6391875" ID-Zenodo-Dep="6391875" box="[192,848,1140,1167]" httpUri="https://zenodo.org/record/6391875/files/figure.png" pageId="8" pageNumber="7" startId="8.[192,268,1140,1166]" targetBox="[192,1440,213,1110]" targetPageId="8">
<paragraph id="8BE34549FF956307FF1CFBB9FCFD5FDF" blockId="8.[192,848,1140,1167]" box="[192,848,1140,1167]" pageId="8" pageNumber="7">
<emphasis id="B928995BFF956307FF1CFBB9FE855FDF" bold="true" box="[192,296,1140,1166]" pageId="8" pageNumber="7">Figure 7.</emphasis>
<taxonomicName id="4C5C3ECAFF956307FEF2FBB9FDA95FDF" authority="Muona, 2022" authorityName="Muona" authorityYear="2022" box="[302,516,1140,1166]" class="Insecta" family="Eucnemidae" genus="Thambus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="woodruffi" status="sp. nov.">
<emphasis id="B928995BFF956307FEF2FBB9FDA95FDF" box="[302,516,1140,1166]" italics="true" pageId="8" pageNumber="7">Thambus woodruffi</emphasis>
</taxonomicName>
<emphasis id="B928995BFF956307FDD6FBB9FDEA5FDF" bold="true" box="[522,583,1140,1166]" pageId="8" pageNumber="7">
<taxonomicNameLabel id="A21B2420FF956307FDD6FBB9FDEA5FDF" box="[522,583,1140,1166]" pageId="8" pageNumber="7" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, ventral view.
</paragraph>
</caption>
<paragraph id="8BE34549FF956307FEDCF9BEFC735DDF" blockId="8.[191,1442,1238,1678]" box="[256,990,1651,1678]" pageId="8" pageNumber="7">
Abdominal ventrites connate, densely and finely punctate (
<figureCitation id="136759CCFF956307FC53F9BEFC7C5DDF" box="[911,977,1651,1678]" captionStart="Figure 7" captionStartId="8.[192,268,1140,1166]" captionTargetBox="[192,1440,213,1110]" captionTargetId="figure-336@8.[192,1440,213,1110]" captionTargetPageId="8" captionText="Figure 7. Thambus woodruffi n. sp., holotype, ventral view." figureDoi="http://doi.org/10.5281/zenodo.6391875" httpUri="https://zenodo.org/record/6391875/files/figure.png" pageId="8" pageNumber="7">Fig. 7</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34616C2FF956305FF1CF953FD2D5A2C" lastPageId="10" lastPageNumber="9" pageId="8" pageNumber="7" type="discussion">
<paragraph id="8BE34549FF956307FF1CF953FED05CD6" blockId="8.[192,1442,1694,1927]" pageId="8" pageNumber="7">
<emphasis id="B928995BFF956307FF1CF953FE9D5DE9" bold="true" box="[192,304,1694,1720]" pageId="8" pageNumber="7">Remarks.</emphasis>
<taxonomicName id="4C5C3ECAFF956307FEE4F953FD855DE9" authority="Bonvouloir" authorityName="Bonvouloir" authorityYear="1871" box="[312,552,1694,1720]" class="Insecta" family="Eucnemidae" genus="Thambus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B928995BFF956307FEE4F953FE335DE9" box="[312,414,1694,1720]" italics="true" pageId="8" pageNumber="7">Thambus</emphasis>
Bonvouloir
</taxonomicName>
is a primarily tropical American genus, with additional species known from Europe (1),
<collectingCountry id="F34B05D9FF956307FE95F90DFE235D8B" box="[329,398,1728,1754]" name="China" pageId="8" pageNumber="7">China</collectingCountry>
(1),
<collectingCountry id="F34B05D9FF956307FE1AF90CFDA85D8A" box="[454,517,1729,1755]" name="Japan" pageId="8" pageNumber="7">Japan</collectingCountry>
(1) and North America (1). The species are small, have a basally wide pronotum with sides strongly converging craniad. In addition to their shape and small size,
<taxonomicName id="4C5C3ECAFF956307FB98F92FFB525DAD" box="[1092,1279,1762,1788]" pageId="8" pageNumber="7">
<emphasis id="B928995BFF956307FB98F92FFB075DAD" box="[1092,1194,1762,1788]" italics="true" pageId="8" pageNumber="7">Thambus</emphasis>
species
</taxonomicName>
are characterized by round pit-like areas laterally on the prosternum. This feature varies between species from faint (e.g.
<taxonomicName id="4C5C3ECAFF956307FA55F8CBFE045C13" authority="Bonvouloir" authorityName="Bonvouloir" class="Insecta" family="Eucnemidae" genus="Thambus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="deyrollei">
<emphasis id="B928995BFF956307FA55F8CBFA375C4E" box="[1417,1434,1798,1823]" italics="true" pageId="8" pageNumber="7">T</emphasis>
.
<emphasis id="B928995BFF956307FF1CF8EAFEB35C10" box="[192,286,1831,1857]" italics="true" pageId="8" pageNumber="7">deyrollei</emphasis>
Bonvouloir
</taxonomicName>
) to conspicuous (e.g.
<taxonomicName id="4C5C3ECAFF956307FD7AF8E5FC615C13" authority="Bonvouloir" authorityName="Bonvouloir" box="[678,972,1831,1858]" class="Insecta" family="Eucnemidae" genus="Thambus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="friwaldskyi">
<emphasis id="B928995BFF956307FD7AF8E5FD185C10" box="[678,693,1832,1857]" italics="true" pageId="8" pageNumber="7">T</emphasis>
.
<emphasis id="B928995BFF956307FD18F8EAFC935C10" box="[708,830,1831,1857]" italics="true" pageId="8" pageNumber="7">friwaldskyi</emphasis>
Bonvouloir
</taxonomicName>
). These pits are not visible in the present sample, but the other characters listed place it to
<taxonomicName id="4C5C3ECAFF956307FD27F887FC3D5C35" authorityName="Fleutiaux" authorityYear="1922" box="[763,912,1866,1892]" class="Insecta" family="Eucnemidae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="tribe" tribe="Macraulacini">Macraulacini</taxonomicName>
and within this tribe the correct placement is clearly
<taxonomicName id="4C5C3ECAFF956307FECCF8A1FEDB5CD7" authorityName="Bonvouloir" authorityYear="1871" box="[272,374,1900,1926]" class="Insecta" family="Eucnemidae" genus="Thambus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B928995BFF956307FECCF8A1FEDB5CD7" box="[272,374,1900,1926]" italics="true" pageId="8" pageNumber="7">Thambus</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="DF2315C1FF946306FF1CFCEBFCC15810" ID-DOI="http://doi.org/10.5281/zenodo.6391877" ID-Zenodo-Dep="6391877" box="[192,876,806,833]" httpUri="https://zenodo.org/record/6391877/files/figure.png" pageId="9" pageNumber="8" startId="9.[192,268,806,832]" targetBox="[192,1440,213,777]" targetPageId="9">
<paragraph id="8BE34549FF946306FF1CFCEBFCC15810" blockId="9.[192,876,806,833]" box="[192,876,806,833]" pageId="9" pageNumber="8">
<emphasis id="B928995BFF946306FF1CFCEBFE855811" bold="true" box="[192,296,806,832]" pageId="9" pageNumber="8">Figure 8.</emphasis>
<taxonomicName id="4C5C3ECAFF946306FEF2FCEAFD845810" authority="Muona, 2022" authorityName="Muona" authorityYear="2022" box="[302,553,807,833]" class="Insecta" family="Eucnemidae" genus="Asiocnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="colombicus" status="sp. nov.">
<emphasis id="B928995BFF946306FEF2FCEAFD845810" box="[302,553,807,833]" italics="true" pageId="9" pageNumber="8">Asiocnemis colombicus</emphasis>
</taxonomicName>
<emphasis id="B928995BFF946306FDF3FCEAFDC15810" bold="true" box="[559,620,807,833]" pageId="9" pageNumber="8">
<taxonomicNameLabel id="A21B2420FF946306FDF3FCEAFDC15810" box="[559,620,807,833]" pageId="9" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, dorsal view.
</paragraph>
</caption>
<caption id="DF2315C1FF946306FF1CF964FCF85DB7" ID-DOI="http://doi.org/10.5281/zenodo.6391879" ID-Zenodo-Dep="6391879" httpUri="https://zenodo.org/record/6391879/files/figure.png" pageId="9" pageNumber="8" startId="9.[192,268,1705,1731]" targetBox="[192,1440,885,1676]" targetPageId="9">
<paragraph id="8BE34549FF946306FF1CF964FCF85DB7" blockId="9.[191,1440,1705,1766]" pageId="9" pageNumber="8">
<emphasis id="B928995BFF946306FF1CF964FE8A5D92" bold="true" box="[192,295,1705,1731]" pageId="9" pageNumber="8">Figure 9.</emphasis>
<taxonomicName id="4C5C3ECAFF946306FEF1F967FD8A5D95" authority="Muona, 2022" authorityName="Muona" authorityYear="2022" box="[301,551,1706,1732]" class="Insecta" family="Eucnemidae" genus="Asiocnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="colombicus" status="sp. nov.">
<emphasis id="B928995BFF946306FEF1F967FD8A5D95" box="[301,551,1706,1732]" italics="true" pageId="9" pageNumber="8">Asiocnemis colombicus</emphasis>
</taxonomicName>
<emphasis id="B928995BFF946306FDF0F967FDC55D95" bold="true" box="[556,616,1706,1732]" pageId="9" pageNumber="8">
<taxonomicNameLabel id="A21B2420FF946306FDF0F967FDC55D95" box="[556,616,1706,1732]" pageId="9" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, ventral view of side of fore body showing flagellomeres 2–9 and the basally open, medially sharply defined antennal groove.
</paragraph>
</caption>
<paragraph id="8BE34549FF976305FF1CFF1BFD0B5BA2" blockId="10.[192,678,214,243]" box="[192,678,214,243]" pageId="10" pageNumber="9">
<emphasis id="B928995BFF976305FF1CFF1BFD0B5BA2" bold="true" box="[192,678,214,243]" pageId="10" pageNumber="9">
Family
<taxonomicName id="4C5C3ECAFF976305FEC3FF1BFD0B5BA2" authority="Eschscholtz, 1829" authorityName="Eschscholtz" authorityYear="1829" box="[287,678,214,243]" class="Insecta" family="Eucnemidae" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="family">Eucnemidae Eschscholtz, 1829</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8BE34549FF976305FF1CFED6FD7D5A69" blockId="10.[192,720,283,312]" box="[192,720,283,312]" pageId="10" pageNumber="9">
<emphasis id="B928995BFF976305FF1CFED6FD7D5A69" bold="true" box="[192,720,283,312]" pageId="10" pageNumber="9">
Subfamily
<taxonomicName id="4C5C3ECAFF976305FE95FED6FD7D5A69" authority="Fleutiaux, 1922" authorityName="Fleutiaux" authorityYear="1922" box="[329,720,283,312]" class="Insecta" family="Eucnemidae" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="subFamily" subFamily="Macraulacinae">Macraulacinae Fleutiaux, 1922</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8BE34549FF976305FF1CFEADFD2D5A2C" blockId="10.[192,640,352,381]" box="[192,640,352,381]" pageId="10" pageNumber="9">
<emphasis id="B928995BFF976305FF1CFEADFD2D5A2C" bold="true" box="[192,640,352,381]" pageId="10" pageNumber="9">
Tribe
<taxonomicName id="4C5C3ECAFF976305FED7FEADFD2D5A2C" authority="Fleutiaux, 1922" authorityName="Fleutiaux" authorityYear="1922" box="[267,640,352,381]" class="Insecta" family="Eucnemidae" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="tribe" tribe="Macraulacini">Macraulacini Fleutiaux, 1922</taxonomicName>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>
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