331 lines
42 KiB
XML
331 lines
42 KiB
XML
<document id="D0102AF31A06873C6F9560BE5777E01D" ID-CLB-Dataset="8143" ID-DOI="10.2478/aemnp-2019-0020" ID-GBIF-Dataset="a03ecce2-b526-4f94-aa65-620016956099" ID-ISSN="1804-6487" ID-Zenodo-Dep="4488918" ID-ZooBank="9F309FCC-A2ED-47B9-BC37-D0C4A3B482E5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612219281920" checkinUser="felipe" docAuthor="Fikáček, Martin & Liu, Hsing-Che" docDate="2019" docId="03F387C57A29FFB7FC0A01963680B69C" docLanguage="en" docName="ActaEntMusNatPra.59.1.229-252.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 59 (1)" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Thysanarthria atriceps" docType="treatment" docVersion="9" lastPageNumber="238" masterDocId="FFCAFFBD7A2EFFBEFF8901283431B709" masterDocTitle="A review of Thysanarthria with description of seven new species and comments on its relationship to Chaetarthria (Hydrophilidae: Chaetarthriini)" masterLastPageNumber="252" masterPageNumber="229" pageNumber="236" updateTime="1698926077238" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="061694D6CB0CE6DFD64EC940D0A7921D">A review of Thysanarthria with description of seven new species and comments on its relationship to Chaetarthria (Hydrophilidae: Chaetarthriini)</mods:title>
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<mods:namePart id="C1C08CAADF40B91FCE06B7F8982D3E81">Fikáček, Martin</mods:namePart>
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<mods:affiliation id="C095EB0274CE19DF89978CE80ACF228B">Department of Entomology, National Museum, Cirkusová 1740, CZ- 19100 Praha 9 – Horní Počernice, Czech Republic & Department of Zoology, Faculty of Science, Charles University, Viničná 7, CZ- 12844 Praha 2, Czech Republic</mods:affiliation>
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<mods:namePart id="B04BD8E36E52FD02529F7AF63B615F37">Liu, Hsing-Che</mods:namePart>
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<mods:affiliation id="CB38681C0F634AA8E7246D23D3158FB1">Department of Environmental Engineering and Management, Chaoyang University of Technology, No. 168, Jifeng E. Rd., Taichung City 413, Taiwan</mods:affiliation>
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<mods:nameIdentifier id="2A1CA535D41AD8817D1FEE1CB05792FD" type="email">td965771@gmail.com</mods:nameIdentifier>
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<mods:title id="A692DA12C0CA3879F7D131A35E776F29">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="046433D4439238865FD5FD221BAEB605">2019</mods:date>
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<treatment id="03F387C57A29FFB7FC0A01963680B69C" ID-DOI="http://doi.org/10.5281/zenodo.4548821" ID-GBIF-Taxon="177423417" ID-Zenodo-Dep="4548821" LSID="urn:lsid:plazi:treatment:03F387C57A29FFB7FC0A01963680B69C" httpUri="http://treatment.plazi.org/id/03F387C57A29FFB7FC0A01963680B69C" lastPageId="9" lastPageNumber="238" pageId="7" pageNumber="236">
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<subSubSection id="C34065587A29FFB9FC0A01963164B7DC" box="[899,1365,190,213]" pageId="7" pageNumber="236" type="nomenclature">
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<paragraph id="8BE536D37A29FFB9FC0A01963164B7DC" blockId="7.[899,1365,190,213]" box="[899,1365,190,213]" pageId="7" pageNumber="236">
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<heading id="D0AD81BF7A29FFB9FC0A01963164B7DC" box="[899,1365,190,213]" centered="true" fontSize="10" level="2" pageId="7" pageNumber="236" reason="3">
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<taxonomicName id="4C5A4D507A29FFB9FC0A01963164B7DC" ID-CoL="56TGX" authority="(Regimbart, 1903)" baseAuthorityName="Regimbart" baseAuthorityYear="1903" box="[899,1365,190,213]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="236" phylum="Arthropoda" rank="species" species="atriceps">
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<emphasis id="B92EEAC17A29FFB9FC0A01963164B7DC" bold="true" box="[899,1365,190,213]" pageId="7" pageNumber="236">
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<emphasis id="B92EEAC17A29FFB9FC0A0196304EB7DC" bold="true" box="[899,1151,190,213]" italics="true" pageId="7" pageNumber="236">Thysanarthria atriceps</emphasis>
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(
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<bibRefCitation id="EFCB4B227A29FFB9FB050196317CB7DC" author="Regimbart" box="[1164,1357,190,213]" firstAuthor="Regimbart" pageId="7" pageNumber="236" pagination="1 - 51" refId="ref18592" refString="REGIMBART M. 1903: Coleopteres aquatiques (Haliplidae, Dytiscidae, Gyrinidae et Hydrophilidae) recueillis dans le Sud de Madagascar par M. Ch. Alluaud (Juillet 1900 - mai 1901). Annales de la Societe Entomologique de France 72: 1 - 51." type="journal article" year="1903">Régimbart, 1903</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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||
</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34065587A29FFB9FB8A01F430E4B7E6" box="[1027,1237,220,239]" pageId="7" pageNumber="236" type="description">
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<paragraph id="8BE536D37A29FFB9FB8A01F430E4B7E6" blockId="7.[828,1437,220,513]" box="[1027,1237,220,239]" pageId="7" pageNumber="236">
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(
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<figureCitation id="13612A567A29FFB9FB8001F4307BB7E6" box="[1033,1098,220,239]" captionStart="Fig" captionStartId="3.[151,182,1959,1978]" captionTargetBox="[157,1426,396,1946]" captionTargetId="figure-14@3.[151,1436,395,1949]" captionTargetPageId="3" captionText="Fig. 2. Morphological differences among Thysanarthria and principal groups of Chaetarthria. A–Z – male genitalia and surrounding sclerites; a–e – details of elytral pubescence. A – Chaetarthria similis Wollaston, 1864; B–G, c – Chaetarthria seminulum (Herbst, 1797); H–M, b – Thysanarthria championi (Knisch, 1924); N–S, d – Chaetarthria panda Orchymont, 1939; T–Z, e – Chaetarthria malickyi Hebauer, 1995; a – Thysanarthria atriceps (Régimbart, 1903). A–D, K–M, Q–S, X–Z – aedeagus in dorsal, lateral and ventral view; E, J, P, V – male sternite 9; F, H, N, T – male sternite 8; G, I, O, U – male tergite 8. Figures a–e are all to the same scale." figureDoi="http://doi.org/10.5281/zenodo.4488924" httpUri="https://zenodo.org/record/4488924/files/figure.png" pageId="7" pageNumber="236">Figs 2a</figureCitation>
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,
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<figureCitation id="13612A567A29FFB9FBDA01F430A9B7E6" box="[1107,1176,220,239]" captionStart="Fig" captionStartId="6.[151,182,1970,1989]" captionTargetBox="[178,1407,394,1955]" captionTargetId="figure-13@6.[176,1412,393,1956]" captionTargetPageId="6" captionText="Fig. 3. Morphology of Thysanarthria. A – mentum and maxillary palps; B – meso- and metaventrite and meso- and metafemora; C – head and prothorax in ventral view; D – antenna; E – ultimate maxillary palpomere with basal peg-like setae; F – metatarsus; G – ventral view with gelatinous substrance on the base of abdomen; H – detail of abdomen with gelatinous substance in the cavity on ventrites 1–2. A, C–F – Thysanarthria championi (Knisch, 1924); B, G–H – T. atriceps (Régimbart, 1903)." figureDoi="http://doi.org/10.5281/zenodo.4488928" httpUri="https://zenodo.org/record/4488928/files/figure.png" pageId="7" pageNumber="236">3B,G,H</figureCitation>
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,
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<figureCitation id="13612A567A29FFB9FB2901F4309BB7E6" box="[1184,1194,220,239]" captionStart="Fig" captionStartId="8.[151,182,1941,1960]" captionTargetBox="[165,1398,197,1882]" captionTargetId="figure-13@8.[157,1431,189,1911]" captionTargetPageId="8" captionText="Fig. 4. Male genitalia of Thysanarthria species. A–E – T. atriceps (Régimbart, 1903) from Zambia: 20 km W of Gwanda, coll. NMPC (A–C – whole aedeagus in dorsal, lateral and ventral view; D–E – detail of median lobe and parameres in dorsal and ventral view); F–J – T. atriceps from Republic of South Africa: 8 km NEE of Stanford, coll. NMPC (F–H – whole aedeagus in dorsal, lateral and ventral view; I–J – detail of median lobe and parameres in dorsal and ventral view); K–O – T. brittoni Balfour-Browne, 1951, holotype (K–M – whole aedeagus in dorsal, lateral and ventral view; N–O – detail of median lobe and parameres in dorsal and ventral view)." figureDoi="http://doi.org/10.5281/zenodo.4488930" httpUri="https://zenodo.org/record/4488930/files/figure.png" pageId="7" pageNumber="236">4</figureCitation>
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A–J)
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</paragraph>
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<subSubSection id="C34065587A29FFB9FCC801D63081B508" pageId="7" pageNumber="236" type="reference_group">
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<paragraph id="8BE536D37A29FFB9FCC801D63091B618" blockId="7.[828,1437,220,513]" box="[833,1184,254,273]" pageId="7" pageNumber="236">
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<treatmentCitationGroup id="AB4A11FD7A29FFB9FCC801D63091B618" box="[833,1184,254,273]" pageId="7" pageNumber="236">
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<treatmentCitation id="0AFB10C27A29FFB9FCC801D630ADB618" author="REGIMBART M." box="[833,1180,254,273]" page="33" pageId="7" pageNumber="236" year="1903">
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<taxonomicName id="4C5A4D507A29FFB9FCC801D630ADB618" ID-CoL="6MJVW" authority="Regimbart, 1903: 33" authorityName="Regimbart" authorityPageNumber="33" authorityYear="1903" box="[833,1180,254,273]" class="Insecta" family="Hydrophilidae" genus="Hydrobius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="236" phylum="Arthropoda" rank="species" species="atriceps">
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<emphasis id="B92EEAC17A29FFB9FCC801D637D4B618" box="[833,997,254,273]" italics="true" pageId="7" pageNumber="236">Hydrobius atriceps</emphasis>
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<bibRefCitation id="EFCB4B227A29FFB9FC6301D630ADB618" author="Regimbart" box="[1002,1180,254,273]" firstAuthor="Regimbart" page="33" pageId="7" pageNumber="236" pagination="1 - 51" refId="ref18592" refString="REGIMBART M. 1903: Coleopteres aquatiques (Haliplidae, Dytiscidae, Gyrinidae et Hydrophilidae) recueillis dans le Sud de Madagascar par M. Ch. Alluaud (Juillet 1900 - mai 1901). Annales de la Societe Entomologique de France 72: 1 - 51." type="journal article" year="1903">Régimbart, 1903: 33</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BE536D37A29FFB9FCB5003137D5B64F" blockId="7.[828,1437,220,513]" pageId="7" pageNumber="236">
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<treatmentCitationGroup id="AB4A11FD7A29FFB9FCB5003137D5B64F" pageId="7" pageNumber="236">
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<treatmentCitation id="0AFB10C27A29FFB9FCB500313095B625" author="KNISCH A." box="[828,1188,281,300]" page="169" pageId="7" pageNumber="236" year="1924">
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<taxonomicName id="4C5A4D507A29FFB9FCB500313095B625" ID-CoL="6MJVW" authority=": KNISCH (1924 a: 169" authorityName="KNISCH" authorityYear="1924" box="[828,1188,281,300]" class="Insecta" family="Hydrophilidae" genus="Hydrobius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="236" phylum="Arthropoda" rank="species" species="atriceps">
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<emphasis id="B92EEAC17A29FFB9FCB5003137D5B625" box="[828,996,281,300]" italics="true" pageId="7" pageNumber="236">Hydrobius atriceps</emphasis>
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:
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<bibRefCitation id="EFCB4B227A29FFB9FC7B00313095B625" author="KNISCH" box="[1010,1188,281,300]" firstAuthor="KNISCH" page="169" pageId="7" pageNumber="236" refId="ref18391" refString="KNISCH A. 1924 a: Hydrophilidae. In: JUNK W. & SCHENKLING S. (eds): Coleopterorum Catalogus. Vol. 14, part 79. W. Junk, Berlin, 306 pp." type="book" year="1924" yearSuffix="a">KNISCH (1924a: 169</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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, catalogue);
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<treatmentCitation id="0AFB10C27A29FFB9FA910031374DB64F" author="ZAITZEV F. A." page="373" pageId="7" pageNumber="236" year="1908">
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<bibRefCitation id="EFCB4B227A29FFB9FA910031374DB64F" author="ZAITZEV" firstAuthor="ZAITZEV" page="373" pageId="7" pageNumber="236" pagination="283 - 420" refId="ref18868" refString="ZAITZEV F. A. 1908: Catalogue des Coleopteres aquatiques des familles Dryopidae, Georyssidae, Cyathoceridae, Heteroceridae et Hydrophilidae. Horae Societatis Entomologicae Rossicae 38: 283 - 420." type="journal article" year="1908">ZAITZEV (1908: 373</bibRefCitation>
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</treatmentCitation>
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, catalogue).
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BE536D37A29FFB9FCB500663081B508" blockId="7.[828,1437,220,513]" pageId="7" pageNumber="236">
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<treatmentCitationGroup id="AB4A11FD7A29FFB9FCB500663081B508" pageId="7" pageNumber="236">
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<treatmentCitation id="0AFB10C27A29FFB9FCB5006630C0B668" author="ORCHYMONT A. d'" box="[828,1265,334,353]" page="195" pageId="7" pageNumber="236" year="1926">
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<taxonomicName id="4C5A4D507A29FFB9FCB5006630C0B668" ID-CoL="56TGX" authority=": ORCHYMONT (1926 a: 195" authorityName=": ORCHYMONT" authorityYear="1926" box="[828,1265,334,353]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="236" phylum="Arthropoda" rank="species" species="atriceps">
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<emphasis id="B92EEAC17A29FFB9FCB500663037B668" box="[828,1030,334,353]" italics="true" pageId="7" pageNumber="236">Thysanarthria atriceps</emphasis>
|
||
:
|
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<bibRefCitation id="EFCB4B227A29FFB9FB9C006630C0B668" author="ORCHYMONT" box="[1045,1265,334,353]" firstAuthor="ORCHYMONT" page="195" pageId="7" pageNumber="236" pagination="195 - 196" refId="ref18492" refString="ORCHYMONT A. d' 1926 a: Note sur Thysanarthria n. g. Bulletin et Annales de la Societe Entomologique de Belgique 66: 195 - 196." type="journal article" year="1926" yearSuffix="a">ORCHYMONT (1926a: 195</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
|
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, transfer to
|
||
<taxonomicName id="4C5A4D507A29FFB9FAE800663797B675" ID-CoL="8X7X2" authorityName="Orchymnont" authorityYear="1926" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="236" phylum="Arthropoda" rank="genus">
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<emphasis id="B92EEAC17A29FFB9FAE800663797B675" italics="true" pageId="7" pageNumber="236">Thysanarthria</emphasis>
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</taxonomicName>
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);
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<treatmentCitation id="0AFB10C27A29FFB9FC3400413095B675" author="ORCHYMONT A. d'" box="[957,1188,361,380]" page="242" pageId="7" pageNumber="236" year="1926">
|
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<bibRefCitation id="EFCB4B227A29FFB9FC3400413095B675" author="ORCHYMONT" box="[957,1188,361,380]" firstAuthor="ORCHYMONT" page="242" pageId="7" pageNumber="236" pagination="201 - 248" refId="ref18521" refString="ORCHYMONT A. d' 1926 b: Contribution a l'etude des Hydrophilides VI. Bulletin et Annales de la Societe Entomologique de Belgique 66: 201 - 248." type="journal article" year="1926" yearSuffix="b">ORCHYMONT (1926b: 242</bibRefCitation>
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</treatmentCitation>
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||
, transfer to
|
||
<taxonomicName id="4C5A4D507A29FFB9FA93004131AAB675" ID-CoL="8X7X2" authorityName="Orchymnont" authorityYear="1926" box="[1306,1435,361,380]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="236" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B92EEAC17A29FFB9FA93004131AAB675" box="[1306,1435,361,380]" italics="true" pageId="7" pageNumber="236">Thysanarthria</emphasis>
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||
</taxonomicName>
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||
explained in more detail, comparison with
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||
<taxonomicName id="4C5A4D507A29FFB9FB4000AB3109B69F" ID-CoL="56TH4" baseAuthorityName="Knisch" baseAuthorityYear="1924" box="[1225,1336,387,406]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="236" phylum="Arthropoda" rank="species" species="championi">
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||
<emphasis id="B92EEAC17A29FFB9FB4000AB3109B69F" box="[1225,1336,387,406]" italics="true" pageId="7" pageNumber="236">T. championi</emphasis>
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||
</taxonomicName>
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||
); BALFOUR--BROWNE (1952: 134, distribution); BALFOUR- BROWNE (1957: 21, distribution);
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||
<treatmentCitation id="0AFB10C27A29FFB9FC5900913049B6C5" author="HANSEN M." box="[976,1144,441,460]" page="105" pageId="7" pageNumber="236" year="1999">
|
||
<bibRefCitation id="EFCB4B227A29FFB9FC5900913049B6C5" author="HANSEN" box="[976,1144,441,460]" firstAuthor="HANSEN" page="105" pageId="7" pageNumber="236" pagination="1 - 416" refId="ref18053" refString="HANSEN M. 1999: Hydrophiloidea (s. str.) (Coleoptera). World Catalogue of Insects 2: 1 - 416." type="journal article" year="1999">HANSEN (1999: 105</bibRefCitation>
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||
</treatmentCitation>
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||
, catalogue);
|
||
<treatmentCitation id="0AFB10C27A29FFB9FB6E009131A8B6C5" author="HEBAUER F." box="[1255,1433,441,460]" page="394" pageId="7" pageNumber="236" year="2001">
|
||
<bibRefCitation id="EFCB4B227A29FFB9FB6E009131A8B6C5" author="HEBAUER" box="[1255,1433,441,460]" firstAuthor="HEBAUER" page="394" pageId="7" pageNumber="236" pagination="393 - 400" refId="ref18120" refString="HEBAUER F. 2001: The species of the genus Thysanarthria d'Orchymont, 1926 (Coleoptera: Hydrophilidae). Beitrage zur Entomologie 51: 393 - 400." type="journal article" year="2001">HEBAUER (2001: 394</bibRefCitation>
|
||
</treatmentCitation>
|
||
, redescription and update of distribution);
|
||
<treatmentCitation id="0AFB10C27A29FFB9FB3500FB3151B6EF" author="HEBAUER F." box="[1212,1376,467,486]" page="39" pageId="7" pageNumber="236" year="2005">
|
||
<bibRefCitation id="EFCB4B227A29FFB9FB3500FB3151B6EF" author="HEBAUER" box="[1212,1376,467,486]" firstAuthor="HEBAUER" page="39" pageId="7" pageNumber="236" pagination="37 - 40" refId="ref18149" refString="HEBAUER F. 2005: Contribution to the knowledge of the Hydrophilidae of Malawi (Coleoptera: Hydrophilidae). Acta Coleopterologica 21 (1): 37 - 40." type="journal article" year="2005">HEBAUER (2005: 39</bibRefCitation>
|
||
</treatmentCitation>
|
||
, distribution);
|
||
<treatmentCitation id="0AFB10C27A29FFB9FC2A00C63076B508" author="HEBAUER F." box="[931,1095,494,513]" page="24" pageId="7" pageNumber="236" year="2006">
|
||
<bibRefCitation id="EFCB4B227A29FFB9FC2A00C63076B508" author="HEBAUER" box="[931,1095,494,513]" firstAuthor="HEBAUER" page="24" pageId="7" pageNumber="236" pagination="19 - 36" refId="ref18180" refString="HEBAUER F. 2006: Checklist of the Hydrophiloidea of Africa and adjacent archipelagos (Coleoptera: Epimetopidae, Georissidae, Helophoridae, Hydrochidae, Hydrophilidae, Spercheidae). Entomologische Zeitschrift 116: 19 - 36." type="journal article" year="2006">HEBAUER (2006: 24</bibRefCitation>
|
||
</treatmentCitation>
|
||
, catalogue).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34065587A29FFB9FCB503303101B409" pageId="7" pageNumber="236" type="materials_examined">
|
||
<paragraph id="8BE536D37A29FFB9FCB503303075B522" blockId="7.[828,1437,536,768]" box="[828,1092,536,555]" pageId="7" pageNumber="236">
|
||
<emphasis id="B92EEAC17A29FFB9FCB5033037F1B522" bold="true" box="[828,960,536,555]" pageId="7" pageNumber="236">
|
||
<typeStatus id="54E188717A29FFB9FCB503303758B522" box="[828,873,536,555]" pageId="7" pageNumber="236">Type</typeStatus>
|
||
material.
|
||
</emphasis>
|
||
Not examined.
|
||
</paragraph>
|
||
<paragraph id="8BE536D37A29FFB9FCB5031A3101B409" blockId="7.[828,1437,536,768]" pageId="7" pageNumber="236">
|
||
<emphasis id="B92EEAC17A29FFB9FCB5031A307AB54C" bold="true" box="[828,1099,562,581]" pageId="7" pageNumber="236">Additional material examined.</emphasis>
|
||
<materialsCitation id="3B323C8E7A29FFB9FBC6031A3116B569" ID-GBIF-Occurrence="3756360340" latitude="-12.92716" longitude="34.2831" pageId="7" pageNumber="236">
|
||
<emphasis id="B92EEAC17A29FFB9FBC6031A3083B54C" bold="true" box="[1103,1202,562,581]" pageId="7" pageNumber="236">
|
||
<collectingCountry id="F34D76437A29FFB9FBC6031A309FB54C" box="[1103,1198,562,581]" name="Malawi" pageId="7" pageNumber="236">MALAWI</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="9D5CFD5A7A29FFB9FB3C031A30D3B54F" box="[1205,1250,559,582]" pageId="7" pageNumber="236" type="female">2 ♀♀</specimenCount>
|
||
(
|
||
<collectionCode id="ED4BAE167A29FFB9FB65031A311CB54C" box="[1260,1325,562,581]" country="Czech Republic" name="National Museum Prague" pageId="7" pageNumber="236">NMPC</collectionCode>
|
||
):
|
||
<collectingRegion id="499EF8317A29FFB9FAB3031A31ADB54C" box="[1338,1436,562,581]" country="Malawi" name="Nkhotakota" pageId="7" pageNumber="236">Nkhotakota</collectingRegion>
|
||
env.,
|
||
<geoCoordinate id="EE6E50147A29FFB9FCE5036537E0B569" box="[876,977,589,608]" degrees="12.92716" direction="south" orientation="latitude" pageId="7" pageNumber="236" precision="1" value="-12.92716">12.92716°S</geoCoordinate>
|
||
<geoCoordinate id="EE6E50147A29FFB9FC5003653006B569" box="[985,1079,589,608]" degrees="34.2831" direction="east" orientation="longitude" pageId="7" pageNumber="236" precision="5" value="34.2831">34.2831°E</geoCoordinate>
|
||
,
|
||
<date id="FFE410137A29FFB9FBCB03653091B569" box="[1090,1184,589,608]" pageId="7" pageNumber="236" value="2002-01-02" valueMax="2002-01-03" valueMin="2002-01-02">2–3.i.2002</date>
|
||
, J. Bezděk lgt.
|
||
</materialsCitation>
|
||
<materialsCitation id="3B323C8E7A29FFB9FAA7036537BFB59C" ID-GBIF-Occurrence="3756360313" pageId="7" pageNumber="236">
|
||
<emphasis id="B92EEAC17A29FFB9FAA7036530AEB572" bold="true" pageId="7" pageNumber="236">
|
||
<collectingCountry id="F34D76437A29FFB9FAA703653036B572" name="South Africa" pageId="7" pageNumber="236">REPUBLIC OF SOUTH AFRICA</collectingCountry>
|
||
:
|
||
<collectingRegion id="499EF8317A29FFB9FB9B03403052B570" box="[1042,1123,616,635]" country="South Africa" name="Western Cape" pageId="7" pageNumber="236">WESTERN</collectingRegion>
|
||
CAPE:
|
||
</emphasis>
|
||
3 ♂♂, 2 unsexed specimens (
|
||
<collectionCode id="ED4BAE167A29FFB9FCCB03AA37B5B59C" box="[834,900,642,661]" country="Czech Republic" name="National Museum Prague" pageId="7" pageNumber="236">NMPC</collectionCode>
|
||
):
|
||
</materialsCitation>
|
||
<materialsCitation id="3B323C8E7A29FFB9FC1803AA37BEB5EC" ID-GBIF-Occurrence="3756360321" latitude="-34.416668" longitude="19.54" pageId="7" pageNumber="236">
|
||
<quantity id="4CA29B367A29FFB9FC1803AA378AB59C" box="[913,955,642,661]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="7" pageNumber="236" unit="km" value="8.0">8 km</quantity>
|
||
NEE of Stanford, in gravel/sand and small isolated pools at the sandy stream and on/in sandy banks along the stream,
|
||
<geoCoordinate id="EE6E50147A29FFB9FAC103B531ADB5B9" box="[1352,1436,669,688]" degrees="34" direction="south" minutes="25.0" orientation="latitude" pageId="7" pageNumber="236" precision="92" value="-34.416668">34°25.0′S</geoCoordinate>
|
||
<geoCoordinate id="EE6E50147A29FFB9FCB5039037A4B5C2" box="[828,917,696,715]" degrees="19" direction="east" minutes="32.4" orientation="longitude" pageId="7" pageNumber="236" precision="92" value="19.54">19°32.4′E</geoCoordinate>
|
||
,
|
||
<date id="FFE410137A29FFB9FC2803903023B5C2" box="[929,1042,696,715]" pageId="7" pageNumber="236" value="2015-12-04" valueMax="2015-12-05" valueMin="2015-12-04">4–5.xii.2015</date>
|
||
, Arriaga, Fikáček, Seidel & Vondráček lgt. (
|
||
<collectingCountry id="F34D76437A29FFB9FCCB03FA375EB5EC" box="[834,879,722,741]" name="South Africa" pageId="7" pageNumber="236">RSA</collectingCountry>
|
||
49).
|
||
</materialsCitation>
|
||
<materialsCitation id="3B323C8E7A29FFB9FC1A03FA3101B409" ID-GBIF-Occurrence="3756360337" pageId="7" pageNumber="236">
|
||
<emphasis id="B92EEAC17A29FFB9FC1A03FA3025B5EC" bold="true" box="[915,1044,722,741]" pageId="7" pageNumber="236">
|
||
<collectingCountry id="F34D76437A29FFB9FC1A03FA3021B5EC" box="[915,1040,722,741]" name="Zimbabwe" pageId="7" pageNumber="236">ZIMBABWE</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
1 ♂, 7 unsexed specimens (
|
||
<collectionCode id="ED4BAE167A29FFB9FB7603FA3170B5EC" box="[1279,1345,722,741]" country="Czech Republic" name="National Museum Prague" pageId="7" pageNumber="236">NMPC</collectionCode>
|
||
):
|
||
<quantity id="4CA29B367A29FFB9FAC603FA31B5B5EC" box="[1359,1412,722,741]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="236" unit="km" value="20.0">20 km</quantity>
|
||
W Gwanda,
|
||
<quantity id="4CA29B367A29FFB9FC0403C537FFB409" box="[909,974,749,768]" metricMagnitude="5" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="236" unit="km" value="120.0">120 km</quantity>
|
||
SE
|
||
<collectingRegion id="499EF8317A29FFB9FC7B03C5307AB409" box="[1010,1099,749,768]" country="Zimbabwe" name="Bulawayo" pageId="7" pageNumber="236">Bulawayo</collectingRegion>
|
||
,
|
||
<date id="FFE410137A29FFB9FBDA03C5309DB409" box="[1107,1196,749,768]" pageId="7" pageNumber="236" value="1999-12-06">6.xii.1999</date>
|
||
, F. Kantner lgt.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34065587A29FFB9FCB5023130E7B1D9" pageId="7" pageNumber="236" type="description">
|
||
<paragraph id="8BE536D37A29FFB9FCB50231301AB299" blockId="7.[828,1437,793,2064]" pageId="7" pageNumber="236">
|
||
<emphasis id="B92EEAC17A29FFB9FCB50231302FB439" bold="true" box="[828,1054,793,816]" pageId="7" pageNumber="236">
|
||
Redescription.
|
||
<emphasis id="B92EEAC17A29FFB9FC6F0231302FB439" bold="true" box="[998,1054,793,816]" italics="true" pageId="7" pageNumber="236">Body</emphasis>
|
||
</emphasis>
|
||
length 1.4–2.0 mm, maximum body width
|
||
<quantity id="4CA29B367A29FFB9FCF702113031B459" box="[894,1024,825,848]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" metricValueMax="1.3" metricValueMin="1.1" pageId="7" pageNumber="236" unit="mm" value="1.2" valueMax="1.3" valueMin="1.1">1.1–1.3 mm</quantity>
|
||
. Head and labrum black, pronotum and elytra uniformly yellowish; legs reddish to yellowish.
|
||
<emphasis id="B92EEAC17A29FFB9FAEB027131ADB479" bold="true" box="[1378,1436,857,880]" italics="true" pageId="7" pageNumber="236">Head</emphasis>
|
||
with strongly granulate microsculpture on interstices; punctation sparse. Eyes separated by 2.7× the width of one eye in dorsal view.
|
||
<emphasis id="B92EEAC17A29FFB9FBB50291309AB4D9" bold="true" box="[1084,1195,953,976]" italics="true" pageId="7" pageNumber="236">Pronotum</emphasis>
|
||
with sparse setiferous punctation similar to that on head; interstices with strongly granulate microsculpture.
|
||
<emphasis id="B92EEAC17A29FFB9FBD202D13091B319" bold="true" box="[1115,1184,1017,1040]" italics="true" pageId="7" pageNumber="236">Elytra</emphasis>
|
||
with 10 striae sharply impressed except anteromedially (near scutellar shield) where neither striae nor serial punctures are visible; interval punctation sparse, setiferous; interstices without distinct microsculpture.
|
||
<emphasis id="B92EEAC17A29FFB9FBB705513098B399" bold="true" box="[1086,1193,1145,1168]" italics="true" pageId="7" pageNumber="236">Aedeagus</emphasis>
|
||
(
|
||
<figureCitation id="13612A567A29FFB9FB3F055130C8B399" box="[1206,1273,1145,1168]" captionStart="Fig" captionStartId="8.[151,182,1941,1960]" captionTargetBox="[165,1398,197,1882]" captionTargetId="figure-13@8.[157,1431,189,1911]" captionTargetPageId="8" captionText="Fig. 4. Male genitalia of Thysanarthria species. A–E – T. atriceps (Régimbart, 1903) from Zambia: 20 km W of Gwanda, coll. NMPC (A–C – whole aedeagus in dorsal, lateral and ventral view; D–E – detail of median lobe and parameres in dorsal and ventral view); F–J – T. atriceps from Republic of South Africa: 8 km NEE of Stanford, coll. NMPC (F–H – whole aedeagus in dorsal, lateral and ventral view; I–J – detail of median lobe and parameres in dorsal and ventral view); K–O – T. brittoni Balfour-Browne, 1951, holotype (K–M – whole aedeagus in dorsal, lateral and ventral view; N–O – detail of median lobe and parameres in dorsal and ventral view)." figureDoi="http://doi.org/10.5281/zenodo.4488930" httpUri="https://zenodo.org/record/4488930/files/figure.png" pageId="7" pageNumber="236">Figs 4</figureCitation>
|
||
A–J) c.
|
||
<quantity id="4CA29B367A29FFB9FAC505513754B3B9" metricMagnitude="-4" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="7" pageNumber="236" unit="mm" value="0.55" valueMax="0.6" valueMin="0.5">0.5–0.6 mm</quantity>
|
||
long. Phallobase slightly widened at base of parameres, slightly narrower than bases of parameres combined, weakly constricted at c. midlength, slightly bent in lateral view. Paremere widely rounded basally, narrowing in apical third, apex rounded. Median lobe narrow, membranous apically, without subapical projections; apex reaching c. level of apex of parameres; gonopore transversely oval, situated in distal third.
|
||
</paragraph>
|
||
<paragraph id="8BE536D37A29FFB9FCB504B130E7B1D9" blockId="7.[828,1437,793,2064]" pageId="7" pageNumber="236">
|
||
<emphasis id="B92EEAC17A29FFB9FCB504B1378BB2B9" bold="true" box="[828,954,1433,1456]" pageId="7" pageNumber="236">Variability.</emphasis>
|
||
The examined specimens from the
|
||
<collectingCountry id="F34D76437A29FFB9FAB504B137EFB2D9" name="South Africa" pageId="7" pageNumber="236">Republic of South Africa</collectingCountry>
|
||
and
|
||
<collectingCountry id="F34D76437A29FFB9FB99049130B3B2D9" box="[1040,1154,1465,1488]" name="Zimbabwe" pageId="7" pageNumber="236">Zimbabwe</collectingCountry>
|
||
differ slightly in the shape of the basal part of the parameres (compare
|
||
<figureCitation id="13612A567A29FFB9FA8904F13171B2F9" box="[1280,1344,1497,1520]" captionStart="Fig" captionStartId="8.[151,182,1941,1960]" captionTargetBox="[165,1398,197,1882]" captionTargetId="figure-13@8.[157,1431,189,1911]" captionTargetPageId="8" captionText="Fig. 4. Male genitalia of Thysanarthria species. A–E – T. atriceps (Régimbart, 1903) from Zambia: 20 km W of Gwanda, coll. NMPC (A–C – whole aedeagus in dorsal, lateral and ventral view; D–E – detail of median lobe and parameres in dorsal and ventral view); F–J – T. atriceps from Republic of South Africa: 8 km NEE of Stanford, coll. NMPC (F–H – whole aedeagus in dorsal, lateral and ventral view; I–J – detail of median lobe and parameres in dorsal and ventral view); K–O – T. brittoni Balfour-Browne, 1951, holotype (K–M – whole aedeagus in dorsal, lateral and ventral view; N–O – detail of median lobe and parameres in dorsal and ventral view)." figureDoi="http://doi.org/10.5281/zenodo.4488930" httpUri="https://zenodo.org/record/4488930/files/figure.png" pageId="7" pageNumber="236">Figs 4</figureCitation>
|
||
A–E and 4F
|
||
<emphasis id="B92EEAC17A29FFB9FCD104D13754B119" bold="true" box="[856,869,1529,1552]" pageId="7" pageNumber="236">–</emphasis>
|
||
J) but seem to be identical in all other aspects including the morphology of the median lobe. Examination of much larger material from Africa covering the known distribution would be needed to reveal whether these differences may be constant and correlated with geography; without such a study we consider the observed differences to be intraspecific variation for the moment.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34065587A29FFB9FCB507F13000B099" pageId="7" pageNumber="236" type="diagnosis">
|
||
<paragraph id="8BE536D37A29FFB9FCB507F13000B099" blockId="7.[828,1437,793,2064]" pageId="7" pageNumber="236">
|
||
<emphasis id="B92EEAC17A29FFB9FCB507F13006B1F9" bold="true" box="[828,1079,1753,1776]" pageId="7" pageNumber="236">Differential diagnosis.</emphasis>
|
||
<taxonomicName id="4C5A4D507A29FFB9FBB607F13107B1F9" authorityName=": ORCHYMONT" authorityYear="1926" box="[1087,1334,1753,1776]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="236" phylum="Arthropoda" rank="species" species="atriceps">
|
||
<emphasis id="B92EEAC17A29FFB9FBB607F13107B1F9" box="[1087,1334,1753,1776]" italics="true" pageId="7" pageNumber="236">Thysanarthria atriceps</emphasis>
|
||
</taxonomicName>
|
||
seems to be the only species occurring in Africa and is hence easy to identify. In form of the median lobe and parameres it resembles only the Arabian
|
||
<taxonomicName id="4C5A4D507A29FFB9FBE0061130E5B059" authorityName="Balfour-Browne" authorityYear="1951" box="[1129,1236,1849,1872]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="236" phylum="Arthropoda" rank="species" species="brittoni">
|
||
<emphasis id="B92EEAC17A29FFB9FBE0061130E5B059" box="[1129,1236,1849,1872]" italics="true" pageId="7" pageNumber="236">T. brittoni</emphasis>
|
||
</taxonomicName>
|
||
from which it differs in relatively longer and narrower parameres and less constricted phallobase.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34065587A29FFB7FCB506B13629B7FC" lastPageId="9" lastPageNumber="238" pageId="7" pageNumber="236" type="biology_ecology">
|
||
<paragraph id="8BE536D37A29FFB7FCB506B13629B7FC" blockId="7.[828,1437,793,2064]" lastBlockId="9.[151,760,190,405]" lastPageId="9" lastPageNumber="238" pageId="7" pageNumber="236">
|
||
<emphasis id="B92EEAC17A29FFB9FCB506B137A6B0B9" bold="true" box="[828,919,1945,1968]" pageId="7" pageNumber="236">Biology.</emphasis>
|
||
The species seems to be usually collected at light. Examined South African specimens were collected from wet sandy banks of a small lowland stream (
|
||
<figureCitation id="13612A567A29FFB9FAA706F131BEB0F9" box="[1326,1423,2009,2032]" captionStart="Fig" captionStartId="20.[151,182,1949,1968]" captionTargetBox="[151,1436,401,1931]" captionTargetId="figure-13@20.[151,1437,401,1931]" captionTargetPageId="20" captionText="Fig. 11. Distribution of Thysanarthria and American Chaetarthria in Asia (A–B) and Near East (C), and examples of habitats of Thysanarthria (D–F). Localities: D – India, Madhya Pradesh, Bhadhua Chora stream ca. 10 km E of Matkuli Jhir, habitat of T. ceylonensis Hebauer, 2001; E – Republic of South Africa, Western Cape, 8 km NEE of Stanford, habitat of T. atriceps (Régimber, 1903), collecting spot marked by an arrow; F–G – Taiwan, Wufeng distr., 4.3 km SEE of Chaoyang University of Technology, habitat of T. chui sp. nov. (G – detail of sandy shore with one alive beetle). Photo D by M. Jäch, photo E by M. Fikáček, F–G by H.-C. Liu." figureDoi="http://doi.org/10.5281/zenodo.4488946" httpUri="https://zenodo.org/record/4488946/files/figure.png" pageId="7" pageNumber="236">Fig. 11E</figureCitation>
|
||
), the beetles were found when the sandy parts were flooded with water or pressed to get submerged, which caused the beetles to float on the water surface.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF25665B7A26FFB6FF1E06BD36B8BF1A" ID-DOI="http://doi.org/10.5281/zenodo.4488930" ID-Zenodo-Dep="4488930" httpUri="https://zenodo.org/record/4488930/files/figure.png" pageId="8" pageNumber="237" startId="8.[151,182,1941,1960]" targetBox="[165,1398,197,1882]" targetPageId="8">
|
||
<paragraph id="8BE536D37A26FFB6FF1E06BD36B8BF1A" blockId="8.[151,1437,1941,2067]" pageId="8" pageNumber="237">
|
||
Fig. 4. Male genitalia of
|
||
<taxonomicName id="4C5A4D507A26FFB6FEE706BD35D9B0A1" authorityName="Orchymnont" authorityYear="1926" box="[366,488,1941,1960]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="237" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92EEAC17A26FFB6FEE706BD35D9B0A1" box="[366,488,1941,1960]" italics="true" pageId="8" pageNumber="237">Thysanarthria</emphasis>
|
||
</taxonomicName>
|
||
species. A–E –
|
||
<taxonomicName id="4C5A4D507A26FFB6FDFA06BD3743B0A1" authority="(Regimbart, 1903)" baseAuthorityName="Regimbart" baseAuthorityYear="1903" box="[627,882,1941,1960]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="237" phylum="Arthropoda" rank="species" species="atriceps">
|
||
<emphasis id="B92EEAC17A26FFB6FDFA06BD36FFB0A1" box="[627,718,1941,1960]" italics="true" pageId="8" pageNumber="237">T. atriceps</emphasis>
|
||
(
|
||
<bibRefCitation id="EFCB4B227A26FFB6FD5006BD375DB0A1" author="Regimbart" box="[729,876,1941,1960]" firstAuthor="Regimbart" pageId="8" pageNumber="237" pagination="1 - 51" refId="ref18592" refString="REGIMBART M. 1903: Coleopteres aquatiques (Haliplidae, Dytiscidae, Gyrinidae et Hydrophilidae) recueillis dans le Sud de Madagascar par M. Ch. Alluaud (Juillet 1900 - mai 1901). Annales de la Societe Entomologique de France 72: 1 - 51." type="journal article" year="1903">Régimbart, 1903</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
from Zambia: 20 km W of Gwanda, coll. NMPC (A–C – whole aedeagus in dorsal, lateral and ventral view; D–E – detail of median lobe and parameres in dorsal and ventral view); F–J –
|
||
<taxonomicName id="4C5A4D507A26FFB6FB2206983134B0CA" baseAuthorityName="Regimbart" baseAuthorityYear="1903" box="[1195,1285,1968,1987]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="237" phylum="Arthropoda" rank="species" species="atriceps">
|
||
<emphasis id="B92EEAC17A26FFB6FB2206983134B0CA" box="[1195,1285,1968,1987]" italics="true" pageId="8" pageNumber="237">T. atriceps</emphasis>
|
||
</taxonomicName>
|
||
from Republic of South Africa: 8 km NEE of Stanford, coll. NMPC (F–H – whole aedeagus in dorsal, lateral and ventral view; I–J – detail of median lobe and parameres in dorsal and ventral view); K–O –
|
||
<taxonomicName id="4C5A4D507A26FFB6FE4B06CD36EFB0F1" authority="Balfour-Browne, 1951" authorityName="Balfour-Browne" authorityYear="1951" box="[450,734,2021,2040]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="237" phylum="Arthropoda" rank="species" species="brittoni">
|
||
<emphasis id="B92EEAC17A26FFB6FE4B06CD3626B0F1" box="[450,535,2021,2040]" italics="true" pageId="8" pageNumber="237">T. brittoni</emphasis>
|
||
<bibRefCitation id="EFCB4B227A26FFB6FD9206CD36EFB0F1" author="Balfour-Browne" box="[539,734,2021,2040]" firstAuthor="Balfour-Browne" pageId="8" pageNumber="237" pagination="1937 - 1938" refId="ref17638" refString="BALFOUR-BROWNE J. 1951: Coleoptera: Haliplidae, Dytiscidae, Gyrinidae, Hydraenidae, Hydrophilidae. Pp 179 - 220 + pls 10 - 11. In: Expedition to South-west Arabia 1937 - 1938. British Museum (Natural History), London, xiv + 504 pp." type="book chapter" year="1951">Balfour-Browne, 1951</bibRefCitation>
|
||
</taxonomicName>
|
||
, holotype (K–M – whole aedeagus in dorsal, lateral and ventral view; N–O – detail of median lobe and parameres in dorsal and ventral view).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C34065587A27FFB7FF1E01D63680B69C" pageId="9" pageNumber="238" type="distribution">
|
||
<paragraph id="8BE536D37A27FFB7FF1E01D63680B69C" blockId="9.[151,760,190,405]" pageId="9" pageNumber="238">
|
||
<emphasis id="B92EEAC17A27FFB7FF1E01D6351BB61C" bold="true" box="[151,298,254,277]" pageId="9" pageNumber="238">Distribution.</emphasis>
|
||
Central and southern part of Africa and
|
||
<collectingCountry id="F34D76437A27FFB7FD4201D634C5B63C" name="Madagascar" pageId="9" pageNumber="238">Madagascar</collectingCountry>
|
||
(where the
|
||
<typeStatus id="54E188717A27FFB7FEFC00363592B63C" box="[373,419,286,309]" pageId="9" pageNumber="238">type</typeStatus>
|
||
locality is situated); on African continent so far recorded from
|
||
<collectingCountry id="F34D76437A27FFB7FE5400163627B65C" box="[477,534,318,341]" name="Togo" pageId="9" pageNumber="238">Togo</collectingCountry>
|
||
, the
|
||
<collectingCountry id="F34D76437A27FFB7FDCE001636F4B65C" box="[583,709,318,341]" name="Ivory Coast" pageId="9" pageNumber="238">Ivory Coast</collectingCountry>
|
||
,
|
||
<collectingCountry id="F34D76437A27FFB7FD47001634F7B67C" name="Angola" pageId="9" pageNumber="238">Angola</collectingCountry>
|
||
,
|
||
<collectingCountry id="F34D76437A27FFB7FF4600763519B67C" box="[207,296,350,373]" name="Rwanda" pageId="9" pageNumber="238">Rwanda</collectingCountry>
|
||
,
|
||
<collectingCountry id="F34D76437A27FFB7FEB9007635B9B67C" box="[304,392,350,373]" name="Burundi" pageId="9" pageNumber="238">Burundi</collectingCountry>
|
||
,
|
||
<collectingCountry id="F34D76437A27FFB7FE18007635D5B67C" box="[401,484,350,373]" name="Malawi" pageId="9" pageNumber="238">Malawi</collectingCountry>
|
||
,
|
||
<collectingCountry id="F34D76437A27FFB7FE650076366EB67C" box="[492,607,350,373]" name="Zimbabwe" pageId="9" pageNumber="238">Zimbabwe</collectingCountry>
|
||
,
|
||
<collectingCountry id="F34D76437A27FFB7FDEE007636C5B67C" box="[615,756,350,373]" name="Mozambique" pageId="9" pageNumber="238">Mozambique</collectingCountry>
|
||
, and the
|
||
<collectingCountry id="F34D76437A27FFB7FF65005635C6B69C" box="[236,503,382,405]" name="South Africa" pageId="9" pageNumber="238">Republic of South Africa</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFCB4B227A27FFB7FD8C00563694B69C" author="HEBAUER" box="[517,677,382,405]" firstAuthor="HEBAUER" pageId="9" pageNumber="238" pagination="19 - 36" refId="ref18180" refString="HEBAUER F. 2006: Checklist of the Hydrophiloidea of Africa and adjacent archipelagos (Coleoptera: Epimetopidae, Georissidae, Helophoridae, Hydrochidae, Hydrophilidae, Spercheidae). Entomologische Zeitschrift 116: 19 - 36." type="journal article" year="2006">HEBAUER 2006</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |