221 lines
26 KiB
XML
221 lines
26 KiB
XML
<document id="8FE44EFA29C562C3291F0C6A610B4492" ID-CLB-Dataset="8143" ID-DOI="10.2478/aemnp-2019-0020" ID-GBIF-Dataset="a03ecce2-b526-4f94-aa65-620016956099" ID-ISSN="1804-6487" ID-Zenodo-Dep="4488918" ID-ZooBank="9F309FCC-A2ED-47B9-BC37-D0C4A3B482E5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612219281920" checkinUser="felipe" docAuthor="Fikáček, Martin & Liu, Hsing-Che" docDate="2019" docId="03F387C57A27FFB7FF4200C334EFB0F4" docLanguage="en" docName="ActaEntMusNatPra.59.1.229-252.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 59 (1)" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Thysanarthria brittoni Balfour-Browne 1951" docType="treatment" docVersion="9" lastPageNumber="238" masterDocId="FFCAFFBD7A2EFFBEFF8901283431B709" masterDocTitle="A review of Thysanarthria with description of seven new species and comments on its relationship to Chaetarthria (Hydrophilidae: Chaetarthriini)" masterLastPageNumber="252" masterPageNumber="229" pageNumber="238" updateTime="1698926077238" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="594C8A5D54A56709B92EFEEFA5AD0881">A review of Thysanarthria with description of seven new species and comments on its relationship to Chaetarthria (Hydrophilidae: Chaetarthriini)</mods:title>
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<mods:name id="3281F3ADB614AD790674F36A82CAF002" type="personal">
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<mods:namePart id="A32770ED4A6AD3A9142E352BF90558FE">Fikáček, Martin</mods:namePart>
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<mods:affiliation id="6CB50F5FB58429F156A08F27E62C31A0">Department of Entomology, National Museum, Cirkusová 1740, CZ- 19100 Praha 9 – Horní Počernice, Czech Republic & Department of Zoology, Faculty of Science, Charles University, Viničná 7, CZ- 12844 Praha 2, Czech Republic</mods:affiliation>
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<mods:nameIdentifier id="749C50628DF3C55A842ACF4FCDD28A72" type="email">mfikacek@gmail.com</mods:nameIdentifier>
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<mods:namePart id="4B6D94949E9AA0B218988F3812A36ED1">Liu, Hsing-Che</mods:namePart>
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<mods:affiliation id="809214B5630326D34ADEBF746D58E3A4">Department of Environmental Engineering and Management, Chaoyang University of Technology, No. 168, Jifeng E. Rd., Taichung City 413, Taiwan</mods:affiliation>
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<mods:nameIdentifier id="6905030A2D972EF8140C150CF05F9FED" type="email">td965771@gmail.com</mods:nameIdentifier>
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<mods:title id="5F5550B303A8A5CAAC5C40D06D337745">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="DE31982BA8638B0EA44103644B33D375">2019</mods:date>
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<mods:number id="7D99C1B18FD841865B2C36909909E5DE">2019-06-20</mods:number>
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<mods:number id="CF07DF236C3D62985F83107D120CDB41">59</mods:number>
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<mods:identifier id="F85408BE935360590300D476D1F80631" type="DOI">10.2478/aemnp-2019-0020</mods:identifier>
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<treatment id="03F387C57A27FFB7FF4200C334EFB0F4" ID-DOI="http://doi.org/10.5281/zenodo.4548823" ID-GBIF-Taxon="177423418" ID-Zenodo-Dep="4548823" LSID="urn:lsid:plazi:treatment:03F387C57A27FFB7FF4200C334EFB0F4" httpUri="http://treatment.plazi.org/id/03F387C57A27FFB7FF4200C334EFB0F4" lastPageNumber="238" pageId="9" pageNumber="238">
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<subSubSection id="C34065587A27FFB7FF4200C336F2B50B" box="[203,707,491,514]" pageId="9" pageNumber="238" type="nomenclature">
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<paragraph id="8BE536D37A27FFB7FF4200C336F2B50B" blockId="9.[203,707,491,514]" box="[203,707,491,514]" pageId="9" pageNumber="238">
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<heading id="D0AD81BF7A27FFB7FF4200C336F2B50B" box="[203,707,491,514]" centered="true" fontSize="10" level="2" pageId="9" pageNumber="238" reason="3">
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<taxonomicName id="4C5A4D507A27FFB7FF4200C336F2B50B" ID-CoL="56TH2" authority="Balfour-Browne, 1951" authorityName="Balfour-Browne" authorityYear="1951" box="[203,707,491,514]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="238" phylum="Arthropoda" rank="species" species="brittoni">
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<emphasis id="B92EEAC17A27FFB7FF4200C336F2B50B" bold="true" box="[203,707,491,514]" pageId="9" pageNumber="238">
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<emphasis id="B92EEAC17A27FFB7FF4200C335F2B50B" bold="true" box="[203,451,491,514]" italics="true" pageId="9" pageNumber="238">Thysanarthria brittoni</emphasis>
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<bibRefCitation id="EFCB4B227A27FFB7FE4000C336F2B50B" author="Balfour-Browne" box="[457,707,491,514]" firstAuthor="Balfour-Browne" pageId="9" pageNumber="238" pagination="1937 - 1938" refId="ref17638" refString="BALFOUR-BROWNE J. 1951: Coleoptera: Haliplidae, Dytiscidae, Gyrinidae, Hydraenidae, Hydrophilidae. Pp 179 - 220 + pls 10 - 11. In: Expedition to South-west Arabia 1937 - 1938. British Museum (Natural History), London, xiv + 504 pp." type="book chapter" year="1951">Balfour-Browne, 1951</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34065587A27FFB7FE0B0322363CB514" box="[386,525,522,541]" pageId="9" pageNumber="238" type="description">
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<paragraph id="8BE536D37A27FFB7FE0B0322363CB514" blockId="9.[151,761,522,750]" box="[386,525,522,541]" pageId="9" pageNumber="238">
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(
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<figureCitation id="13612A567A27FFB7FE0E0322358FB514" box="[391,446,522,541]" captionStart="Fig" captionStartId="8.[151,182,1941,1960]" captionTargetBox="[165,1398,197,1882]" captionTargetId="figure-13@8.[157,1431,189,1911]" captionTargetPageId="8" captionText="Fig. 4. Male genitalia of Thysanarthria species. A–E – T. atriceps (Régimbart, 1903) from Zambia: 20 km W of Gwanda, coll. NMPC (A–C – whole aedeagus in dorsal, lateral and ventral view; D–E – detail of median lobe and parameres in dorsal and ventral view); F–J – T. atriceps from Republic of South Africa: 8 km NEE of Stanford, coll. NMPC (F–H – whole aedeagus in dorsal, lateral and ventral view; I–J – detail of median lobe and parameres in dorsal and ventral view); K–O – T. brittoni Balfour-Browne, 1951, holotype (K–M – whole aedeagus in dorsal, lateral and ventral view; N–O – detail of median lobe and parameres in dorsal and ventral view)." figureDoi="http://doi.org/10.5281/zenodo.4488930" httpUri="https://zenodo.org/record/4488930/files/figure.png" pageId="9" pageNumber="238">Figs 4</figureCitation>
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K–O,
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<figureCitation id="13612A567A27FFB7FE7B03223636B514" box="[498,519,522,541]" captionStart="Fig" captionStartId="20.[151,182,1949,1968]" captionTargetBox="[151,1436,401,1931]" captionTargetId="figure-13@20.[151,1437,401,1931]" captionTargetPageId="20" captionText="Fig. 11. Distribution of Thysanarthria and American Chaetarthria in Asia (A–B) and Near East (C), and examples of habitats of Thysanarthria (D–F). Localities: D – India, Madhya Pradesh, Bhadhua Chora stream ca. 10 km E of Matkuli Jhir, habitat of T. ceylonensis Hebauer, 2001; E – Republic of South Africa, Western Cape, 8 km NEE of Stanford, habitat of T. atriceps (Régimber, 1903), collecting spot marked by an arrow; F–G – Taiwan, Wufeng distr., 4.3 km SEE of Chaoyang University of Technology, habitat of T. chui sp. nov. (G – detail of sandy shore with one alive beetle). Photo D by M. Jäch, photo E by M. Fikáček, F–G by H.-C. Liu." figureDoi="http://doi.org/10.5281/zenodo.4488946" httpUri="https://zenodo.org/record/4488946/files/figure.png" pageId="9" pageNumber="238">11</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C34065587A27FFB7FF1E03043571B57D" pageId="9" pageNumber="238" type="reference_group">
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<paragraph id="8BE536D37A27FFB7FF1E0304367FB536" blockId="9.[151,761,522,750]" box="[151,590,556,575]" pageId="9" pageNumber="238">
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<treatmentCitationGroup id="AB4A11FD7A27FFB7FF1E0304367FB536" box="[151,590,556,575]" pageId="9" pageNumber="238">
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<treatmentCitation id="0AFB10C27A27FFB7FF1E0304367AB536" author="BALFOUR-BROWNE J." box="[151,587,556,575]" page="215" pageId="9" pageNumber="238" year="1951">
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<taxonomicName id="4C5A4D507A27FFB7FF1E0304367AB536" ID-CoL="56TH2" authority="Balfour-Browne, 1951: 215" authorityName="Balfour-Browne" authorityPageNumber="215" authorityYear="1951" box="[151,587,556,575]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="238" phylum="Arthropoda" rank="species" species="brittoni">
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<emphasis id="B92EEAC17A27FFB7FF1E03043566B536" box="[151,343,556,575]" italics="true" pageId="9" pageNumber="238">Thysanarthria brittoni</emphasis>
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||
<bibRefCitation id="EFCB4B227A27FFB7FED50304367AB536" author="Balfour-Browne" box="[348,587,556,575]" firstAuthor="Balfour-Browne" page="215" pageId="9" pageNumber="238" pagination="1937 - 1938" refId="ref17638" refString="BALFOUR-BROWNE J. 1951: Coleoptera: Haliplidae, Dytiscidae, Gyrinidae, Hydraenidae, Hydrophilidae. Pp 179 - 220 + pls 10 - 11. In: Expedition to South-west Arabia 1937 - 1938. British Museum (Natural History), London, xiv + 504 pp." type="book chapter" year="1951">Balfour-Browne, 1951: 215</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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.
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||
</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BE536D37A27FFB7FF1E036E3571B57D" blockId="9.[151,761,522,750]" pageId="9" pageNumber="238">
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<treatmentCitationGroup id="AB4A11FD7A27FFB7FF1E036E3571B57D" pageId="9" pageNumber="238">
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<treatmentCitation id="0AFB10C27A27FFB7FF1E036E363EB550" author="HEBAUER F." box="[151,527,582,601]" page="267" pageId="9" pageNumber="238" year="1997">
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<taxonomicName id="4C5A4D507A27FFB7FF1E036E363EB550" ID-CoL="56TH2" authority=": HEBAUER (1997: 267" authorityName=": HEBAUER" authorityYear="1997" box="[151,527,582,601]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="238" phylum="Arthropoda" rank="species" species="brittoni">
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<emphasis id="B92EEAC17A27FFB7FF1E036E3567B550" box="[151,342,582,601]" italics="true" pageId="9" pageNumber="238">Thysanarthria brittoni</emphasis>
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:
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<bibRefCitation id="EFCB4B227A27FFB7FEE9036E363EB550" author="HEBAUER" box="[352,527,582,601]" firstAuthor="HEBAUER" page="267" pageId="9" pageNumber="238" pagination="255 - 276" refId="ref18079" refString="HEBAUER F. 1997: Annotated checklist of the Hydrophilidae and Helophoridae (Insecta: Coleoptera) of the Arabian Peninsula with a description of a new genus and species. Fauna of Saudi Arabia 16: 255 - 276." type="journal article" year="1997">HEBAUER (1997: 267</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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||
, catalogue);
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||
<treatmentCitation id="0AFB10C27A27FFB7FDF3036E34E6B57D" author="HANSEN M." page="105" pageId="9" pageNumber="238" year="1999">
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<bibRefCitation id="EFCB4B227A27FFB7FDF3036E34E6B57D" author="HANSEN" firstAuthor="HANSEN" page="105" pageId="9" pageNumber="238" pagination="1 - 416" refId="ref18053" refString="HANSEN M. 1999: Hydrophiloidea (s. str.) (Coleoptera). World Catalogue of Insects 2: 1 - 416." type="journal article" year="1999">HANSEN (1999: 105</bibRefCitation>
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</treatmentCitation>
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, catalogue).
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</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C34065587A27FFB7FF1E03A33651B5E7" pageId="9" pageNumber="238" type="materials_examined">
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<paragraph id="8BE536D37A27FFB7FF1E03A33651B5E7" blockId="9.[151,761,522,750]" pageId="9" pageNumber="238">
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<emphasis id="B92EEAC17A27FFB7FF1E03A3354CB597" bold="true" box="[151,381,651,670]" pageId="9" pageNumber="238">Type material examined.</emphasis>
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<materialsCitation id="3B323C8E7A27FFB7FE0D03A33651B5E7" ID-GBIF-Occurrence="3756360345" collectingDate="1937-11-01" collectingDateMax="1937-11-02" collectingDateMin="1937-11-01" collectionCode="BMNH" collectorName="H. Scott & E. B. Britton" determinerName="J. Balfour-Browne" elevation="1524" location="Wadi at foot of Jebel Harir" pageId="9" pageNumber="238" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<typeStatus id="54E188717A27FFB7FE0D03A335E6B595" box="[388,471,651,670]" pageId="9" pageNumber="238" type="holotype">HOLOTYPE</typeStatus>
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:
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<specimenCount id="9D5CFD5A7A27FFB7FE6C03AC35C4B594" box="[485,501,644,669]" pageId="9" pageNumber="238" type="male">♂</specimenCount>
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||
(
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<collectionCode id="ED4BAE167A27FFB7FD8B03A33677B597" box="[514,582,651,670]" pageId="9" pageNumber="238">BMNH</collectionCode>
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||
), ʻType // W ADEN PROT /
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<location id="8E8560087A27FFB7FF57038E35FCB5B0" LSID="urn:lsid:plazi:treatment:03F387C57A27FFB7FF4200C334EFB0F4:8E8560087A27FFB7FF57038E35FCB5B0" box="[222,461,678,697]" name="Wadi at foot of / Jebel Harir" pageId="9" pageNumber="238">Wadi at foot of / Jebel Harir</location>
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/ ca.
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||
<quantity id="4CA29B367A27FFB7FE75038E360FB5B0" box="[508,574,678,697]" metricMagnitude="3" metricUnit="m" metricValue="1.524" pageId="9" pageNumber="238" unit="ft" value="5000.0">
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||
<elevation id="0077D1E07A27FFB7FE75038E360FB5B0" box="[508,574,678,697]" metricMagnitude="3" metricUnit="m" metricValue="1.524" pageId="9" pageNumber="238" unit="ft" value="5000.0">5,000 ft</elevation>
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</quantity>
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||
/
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||
<collectingDate id="EFA0E9FB7A27FFB7FDC6038E3681B5B0" box="[591,688,678,697]" pageId="9" pageNumber="238">
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||
1,2.
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||
<date id="FFE410137A27FFB7FDE4038E3681B5B0" box="[621,688,678,697]" pageId="9" pageNumber="238" value="1937-11">xi.1937</date>
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</collectingDate>
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||
//
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||
<collectorName id="26AF53057A27FFB7FD4F038E364FB5DA" pageId="9" pageNumber="238">B. M. Exp. to / S. W. Arabia / H. Scott & / E. B. Britton / B. M.</collectorName>
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||
1938-246 //
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<determinerName id="7E39DA177A27FFB7FD6303E83513B5E7" pageId="9" pageNumber="238">J. Balfour-Browne</determinerName>
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||
det. /
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||
<originalDetermination id="A8448E2F7A27FFB7FEDE03F3362FB5E7" box="[343,542,731,750]" pageId="9" pageNumber="238">
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||
<taxonomicName id="4C5A4D507A27FFB7FEDE03F335FCB5E7" authority="Fikáček & Liu, 2019" authorityName="Fikáček & Liu" authorityYear="2019" box="[343,461,731,750]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="252" phylum="Arthropoda" rank="genus" status="gen. nov.">Thysanarthria</taxonomicName>
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/ brittoni
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</originalDetermination>
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Type!ʼ.
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</materialsCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C34065587A27FFB7FF1E022E35D5B134" pageId="9" pageNumber="238" type="description">
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<paragraph id="8BE536D37A27FFB7FF1E022E35BCB294" blockId="9.[151,760,774,2045]" pageId="9" pageNumber="238">
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<emphasis id="B92EEAC17A27FFB7FF1E022E35B4B414" bold="true" box="[151,389,774,797]" pageId="9" pageNumber="238">
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||
Redescription.
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||
<emphasis id="B92EEAC17A27FFB7FEC5022E35B4B414" bold="true" box="[332,389,774,797]" italics="true" pageId="9" pageNumber="238">Body</emphasis>
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||
</emphasis>
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||
length
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||
<quantity id="4CA29B367A27FFB7FE56022E3608B414" box="[479,569,774,797]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="9" pageNumber="238" unit="mm" value="1.6">1.6 mm</quantity>
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, maximum body width 1.0 mm. Head and labrum black, pronotum and elytra uniformly yellowish; legs reddish to yellowish.
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||
<emphasis id="B92EEAC17A27FFB7FF1E024E34E3B474" bold="true" box="[151,210,870,893]" italics="true" pageId="9" pageNumber="238">Head</emphasis>
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||
with weak microsculpture on interstices; punctation sparse, each puncture bearing pointed seta. Eyes separated by 3.1× the width of one eye in dorsal view.
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<emphasis id="B92EEAC17A27FFB7FD0E028E36C9B4B4" bold="true" box="[647,760,934,957]" italics="true" pageId="9" pageNumber="238">Pronotum</emphasis>
|
||
with sparse setiferous punctation similar to that on head; interstices with weak microsculpture.
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||
<emphasis id="B92EEAC17A27FFB7FDC702CE36A4B4F4" bold="true" box="[590,661,998,1021]" italics="true" pageId="9" pageNumber="238">Elytra</emphasis>
|
||
with 10 striae sharply impressed except anteromedially (near scutellar shield) where neither striae nor serial punctures are visible; interval punctation sparse, setiferous; interstices without distinct microsculpture.
|
||
<emphasis id="B92EEAC17A27FFB7FE73054E3656B374" bold="true" box="[506,615,1126,1149]" italics="true" pageId="9" pageNumber="238">Aedeagus</emphasis>
|
||
(
|
||
<figureCitation id="13612A567A27FFB7FDFC054E3688B374" box="[629,697,1126,1149]" captionStart="Fig" captionStartId="8.[151,182,1941,1960]" captionTargetBox="[165,1398,197,1882]" captionTargetId="figure-13@8.[157,1431,189,1911]" captionTargetPageId="8" captionText="Fig. 4. Male genitalia of Thysanarthria species. A–E – T. atriceps (Régimbart, 1903) from Zambia: 20 km W of Gwanda, coll. NMPC (A–C – whole aedeagus in dorsal, lateral and ventral view; D–E – detail of median lobe and parameres in dorsal and ventral view); F–J – T. atriceps from Republic of South Africa: 8 km NEE of Stanford, coll. NMPC (F–H – whole aedeagus in dorsal, lateral and ventral view; I–J – detail of median lobe and parameres in dorsal and ventral view); K–O – T. brittoni Balfour-Browne, 1951, holotype (K–M – whole aedeagus in dorsal, lateral and ventral view; N–O – detail of median lobe and parameres in dorsal and ventral view)." figureDoi="http://doi.org/10.5281/zenodo.4488930" httpUri="https://zenodo.org/record/4488930/files/figure.png" pageId="9" pageNumber="238">Figs 4</figureCitation>
|
||
K–O) c.
|
||
<quantity id="4CA29B367A27FFB7FF3B05AE3534B394" box="[178,261,1158,1181]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="238" unit="mm" value="0.5">0.5 mm</quantity>
|
||
long. Phallobase strongly widened at base of parameres, c. as wide as bases of parameres combined, strongly constricted at c. midlength, slightly bent in lateral view. Paremere widely rounded basally, slightly narrowing in apical third, apex rounded, apices divergent from each other. Median lobe narrow, membranous apically, without subapical projections; apex reaching c. level of apex of parameres; gonopore transversely oval, situated in distal third.
|
||
</paragraph>
|
||
<paragraph id="8BE536D37A27FFB7FF1E048E35D5B134" blockId="9.[151,760,774,2045]" pageId="9" pageNumber="238">
|
||
<emphasis id="B92EEAC17A27FFB7FF1E048E352BB2B4" bold="true" box="[151,282,1446,1469]" pageId="9" pageNumber="238">Variability.</emphasis>
|
||
BALFOUR- BROWNE (1951) mentions that the dorsal microsculpture of the head and pronotum, which is very weakly developed in the
|
||
<typeStatus id="54E188717A27FFB7FE7204CE366CB2F4" box="[507,605,1510,1533]" pageId="9" pageNumber="238" type="holotype">holotype</typeStatus>
|
||
, is stronger in some of the
|
||
<typeStatus id="54E188717A27FFB7FE96072E35B6B114" box="[287,391,1542,1565]" pageId="9" pageNumber="238" type="paratype">paratypes</typeStatus>
|
||
which are hence externally undistinguishable from
|
||
<taxonomicName id="4C5A4D507A27FFB7FEE3070E35ECB134" baseAuthorityName="Regimbart" baseAuthorityYear="1903" box="[362,477,1574,1597]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="238" phylum="Arthropoda" rank="species" species="atriceps">
|
||
<emphasis id="B92EEAC17A27FFB7FEE3070E35ECB134" box="[362,477,1574,1597]" italics="true" pageId="9" pageNumber="238">T. atriceps</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34065587A27FFB7FF1E076E3649B074" pageId="9" pageNumber="238" type="diagnosis">
|
||
<paragraph id="8BE536D37A27FFB7FF1E076E3649B074" blockId="9.[151,760,774,2045]" pageId="9" pageNumber="238">
|
||
<emphasis id="B92EEAC17A27FFB7FF1E076E3660B154" bold="true" box="[151,593,1606,1629]" pageId="9" pageNumber="238">Differential diagnosis and discussion.</emphasis>
|
||
<taxonomicName id="4C5A4D507A27FFB7FDD5076E34D8B174" authorityName=": HEBAUER" authorityYear="1997" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="238" phylum="Arthropoda" rank="species" species="brittoni">
|
||
<emphasis id="B92EEAC17A27FFB7FDD5076E34D8B174" italics="true" pageId="9" pageNumber="238">Thysanarthria brittoni</emphasis>
|
||
</taxonomicName>
|
||
is very similar to
|
||
<taxonomicName id="4C5A4D507A27FFB7FE4B074E3609B174" baseAuthorityName="Regimbart" baseAuthorityYear="1903" box="[450,568,1638,1661]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="238" phylum="Arthropoda" rank="species" species="atriceps">
|
||
<emphasis id="B92EEAC17A27FFB7FE4B074E3609B174" box="[450,568,1638,1661]" italics="true" pageId="9" pageNumber="238">T. atriceps</emphasis>
|
||
</taxonomicName>
|
||
in all characters including male genitalia, which only differ in the proportions of the parameres including their slightly diverging apices, and by the more strongly constricted phallobase (see under
|
||
<taxonomicName id="4C5A4D507A27FFB7FE9907CE35B5B1F4" baseAuthorityName="Regimbart" baseAuthorityYear="1903" box="[272,388,1766,1789]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="238" phylum="Arthropoda" rank="species" species="atriceps">
|
||
<emphasis id="B92EEAC17A27FFB7FE9907CE35B5B1F4" box="[272,388,1766,1789]" italics="true" pageId="9" pageNumber="238">T. atriceps</emphasis>
|
||
</taxonomicName>
|
||
for details). The difference of the genitalia of
|
||
<taxonomicName id="4C5A4D507A27FFB7FE92062E35B8B014" authorityName=": HEBAUER" authorityYear="1997" box="[283,393,1798,1821]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="238" phylum="Arthropoda" rank="species" species="brittoni">
|
||
<emphasis id="B92EEAC17A27FFB7FE92062E35B8B014" box="[283,393,1798,1821]" italics="true" pageId="9" pageNumber="238">T. brittoni</emphasis>
|
||
</taxonomicName>
|
||
from the examined specimens of
|
||
<taxonomicName id="4C5A4D507A27FFB7FF1E060E3536B034" baseAuthorityName="Regimbart" baseAuthorityYear="1903" box="[151,263,1830,1853]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="238" phylum="Arthropoda" rank="species" species="atriceps">
|
||
<emphasis id="B92EEAC17A27FFB7FF1E060E3536B034" box="[151,263,1830,1853]" italics="true" pageId="9" pageNumber="238">T. atriceps</emphasis>
|
||
</taxonomicName>
|
||
is bigger than the observed intraspecific variability of
|
||
<taxonomicName id="4C5A4D507A27FFB7FF7B066E3555B054" baseAuthorityName="Regimbart" baseAuthorityYear="1903" box="[242,356,1862,1885]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="238" phylum="Arthropoda" rank="species" species="atriceps">
|
||
<emphasis id="B92EEAC17A27FFB7FF7B066E3555B054" box="[242,356,1862,1885]" italics="true" pageId="9" pageNumber="238">T. atriceps</emphasis>
|
||
</taxonomicName>
|
||
, which is the reason why we consider
|
||
<taxonomicName id="4C5A4D507A27FFB7FF1E064E3535B074" authorityName=": HEBAUER" authorityYear="1997" box="[151,260,1894,1917]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="238" phylum="Arthropoda" rank="species" species="brittoni">
|
||
<emphasis id="B92EEAC17A27FFB7FF1E064E3535B074" box="[151,260,1894,1917]" italics="true" pageId="9" pageNumber="238">T. brittoni</emphasis>
|
||
</taxonomicName>
|
||
a separate species at the moment.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34065587A27FFB7FF1E06AE355FB094" box="[151,366,1926,1949]" pageId="9" pageNumber="238" type="biology_ecology">
|
||
<paragraph id="8BE536D37A27FFB7FF1E06AE355FB094" blockId="9.[151,760,774,2045]" box="[151,366,1926,1949]" pageId="9" pageNumber="238">
|
||
<emphasis id="B92EEAC17A27FFB7FF1E06AE34C5B094" bold="true" box="[151,244,1926,1949]" pageId="9" pageNumber="238">Biology.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34065587A27FFB7FF1E068E34EFB0F4" pageId="9" pageNumber="238" type="distribution">
|
||
<paragraph id="8BE536D37A27FFB7FF1E068E34EFB0F4" blockId="9.[151,760,774,2045]" pageId="9" pageNumber="238">
|
||
<emphasis id="B92EEAC17A27FFB7FF1E068E3503B0B4" bold="true" box="[151,306,1958,1981]" pageId="9" pageNumber="238">Distribution.</emphasis>
|
||
Only known from the
|
||
<typeStatus id="54E188717A27FFB7FDCF068E3647B0B4" box="[582,630,1958,1981]" pageId="9" pageNumber="238">type</typeStatus>
|
||
locality in western
|
||
<collectingCountry id="F34D76437A27FFB7FF7E06EE3577B0D4" box="[247,326,1990,2013]" name="Yemen" pageId="9" pageNumber="238">Yemen</collectingCountry>
|
||
, Arabian Peninsula (BALFOUR- BROWNE 1951).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |