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<document id="7A774343448733F7A2043BC5CC903A11" ID-CLB-Dataset="8143" ID-DOI="10.2478/aemnp-2019-0020" ID-GBIF-Dataset="a03ecce2-b526-4f94-aa65-620016956099" ID-ISSN="1804-6487" ID-Zenodo-Dep="4488918" ID-ZooBank="9F309FCC-A2ED-47B9-BC37-D0C4A3B482E5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612219281920" checkinUser="felipe" docAuthor="Fikáček, Martin &amp; Liu, Hsing-Che" docDate="2019" docId="03F387C57A21FFB1FF5C079737B0B3DC" docLanguage="en" docName="ActaEntMusNatPra.59.1.229-252.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 59 (1)" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Thysanarthria hongsonensis Hebauer 2001" docType="treatment" docVersion="9" lastPageNumber="244" masterDocId="FFCAFFBD7A2EFFBEFF8901283431B709" masterDocTitle="A review of Thysanarthria with description of seven new species and comments on its relationship to Chaetarthria (Hydrophilidae: Chaetarthriini)" masterLastPageNumber="252" masterPageNumber="229" pageNumber="244" updateTime="1698926077238" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="CA4ACCBEEE795DEACA3A65ED412B6F74">A review of Thysanarthria with description of seven new species and comments on its relationship to Chaetarthria (Hydrophilidae: Chaetarthriini)</mods:title>
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<mods:namePart id="83E80C08AF6D6574F708A61546692852">Fikáček, Martin</mods:namePart>
<mods:affiliation id="B8DA6BF7970CAA0D95E123D520991409">Department of Entomology, National Museum, Cirkusová 1740, CZ- 19100 Praha 9 Horní Počernice, Czech Republic &amp; Department of Zoology, Faculty of Science, Charles University, Viničná 7, CZ- 12844 Praha 2, Czech Republic</mods:affiliation>
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<mods:namePart id="E434B206DE45E014721C1811A7A7C599">Liu, Hsing-Che</mods:namePart>
<mods:affiliation id="1DF31C891F7F7FC4B9A85E9C06BB6D7C">Department of Environmental Engineering and Management, Chaoyang University of Technology, No. 168, Jifeng E. Rd., Taichung City 413, Taiwan</mods:affiliation>
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<mods:title id="7567D4F024E32BE68AFD156F46EFD677">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<treatment id="03F387C57A21FFB1FF5C079737B0B3DC" ID-DOI="http://doi.org/10.5281/zenodo.4548837" ID-GBIF-Taxon="177423425" ID-Zenodo-Dep="4548837" LSID="urn:lsid:plazi:treatment:03F387C57A21FFB1FF5C079737B0B3DC" httpUri="http://treatment.plazi.org/id/03F387C57A21FFB1FF5C079737B0B3DC" lastPageNumber="244" pageId="15" pageNumber="244">
<subSubSection id="C34065587A21FFB1FF5C0797368BB1DF" box="[213,698,1727,1750]" pageId="15" pageNumber="244" type="nomenclature">
<paragraph id="8BE536D37A21FFB1FF5C0797368BB1DF" blockId="15.[213,698,1727,1750]" box="[213,698,1727,1750]" pageId="15" pageNumber="244">
<heading id="D0AD81BF7A21FFB1FF5C0797368BB1DF" box="[213,698,1727,1750]" centered="true" fontSize="10" level="2" pageId="15" pageNumber="244" reason="3">
<taxonomicName id="4C5A4D507A21FFB1FF5C0797368BB1DF" ID-CoL="56TH5" authority="Hebauer, 2001" authorityName="Hebauer" authorityYear="2001" box="[213,698,1727,1750]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="244" phylum="Arthropoda" rank="species" species="hongsonensis">
<emphasis id="B92EEAC17A21FFB1FF5C0797368BB1DF" bold="true" box="[213,698,1727,1750]" pageId="15" pageNumber="244">
<emphasis id="B92EEAC17A21FFB1FF5C07973621B1DF" bold="true" box="[213,528,1727,1750]" italics="true" pageId="15" pageNumber="244">Thysanarthria hongsonensis</emphasis>
<bibRefCitation id="EFCB4B227A21FFB1FD9F0797368BB1DF" author="Hebauer" box="[534,698,1727,1750]" firstAuthor="Hebauer" pageId="15" pageNumber="244" pagination="393 - 400" refId="ref18120" refString="HEBAUER F. 2001: The species of the genus Thysanarthria d'Orchymont, 1926 (Coleoptera: Hydrophilidae). Beitrage zur Entomologie 51: 393 - 400." type="journal article" year="2001">Hebauer, 2001</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34065587A21FFB1FEFD07F5362BB1F9" box="[372,538,1757,1776]" pageId="15" pageNumber="244" type="description">
<paragraph id="8BE536D37A21FFB1FEFD07F5362BB1F9" blockId="15.[151,761,1757,1985]" box="[372,538,1757,1776]" pageId="15" pageNumber="244">
(
<figureCitation id="13612A567A21FFB1FEF307F535F0B1F9" box="[378,449,1757,1776]" captionStart="Fig" captionStartId="2.[151,182,2004,2023]" captionTargetBox="[165,1409,612,1979]" captionTargetId="figure-13@2.[151,1437,604,1996]" captionTargetPageId="2" captionText="Fig. 1. Habitus of the Thysanarthria species and related Old Word Chaetarthriini.A Thysanarthria bengalensis Hebauer, 2001, holotype; B T. persica sp. nov., paratype; C T. siamensis Hebauer, 2001, holotype; D T. hongsonensis Hebauer, 2001, holotype; E European Chaetarthria: C. seminulum (Herbst, 1797); F large Old World Chaetarthria: C. nigerrima (Blackburn, 1891)." figureDoi="http://doi.org/10.5281/zenodo.4488920" httpUri="https://zenodo.org/record/4488920/files/figure.png" pageId="15" pageNumber="244">Figs 1D</figureCitation>
,
<figureCitation id="13612A567A21FFB1FE4307F535E2B1F9" box="[458,467,1757,1776]" captionStart="Fig" captionStartId="16.[151,182,1976,1995]" captionTargetBox="[204,1420,202,1944]" captionTargetId="figure-108@16.[162,1439,189,1956]" captionTargetPageId="16" captionText="Fig. 8. Male genitalia of Thysanarthria species, holotypes.AE T. chui sp. nov.(AC whole aedeagus in dorsal, lateral and ventral view; DE detail of median lobe and parameres in dorsal and ventral view). FJ T. hongsonensis Hebauer, 2001 (FH whole aedeagus in dorsal, lateral and ventral view; IJ details of parameres and median lobe in dorsal and ventral view). KO T. madurensis Hebauer, 2001 (KM whole aedeagus in dorsal, lateral and ventral view; NO details of median lobe and parameres in dorsal and ventral view)." figureDoi="http://doi.org/10.5281/zenodo.4488940" httpUri="https://zenodo.org/record/4488940/files/figure.png" pageId="15" pageNumber="244">8</figureCitation>
FJ,
<figureCitation id="13612A567A21FFB1FE7607F53625B1F9" box="[511,532,1757,1776]" captionStart="Fig" captionStartId="20.[151,182,1949,1968]" captionTargetBox="[151,1436,401,1931]" captionTargetId="figure-13@20.[151,1437,401,1931]" captionTargetPageId="20" captionText="Fig. 11. Distribution of Thysanarthria and American Chaetarthria in Asia (AB) and Near East (C), and examples of habitats of Thysanarthria (DF). Localities: D India, Madhya Pradesh, Bhadhua Chora stream ca. 10 km E of Matkuli Jhir, habitat of T. ceylonensis Hebauer, 2001; E Republic of South Africa, Western Cape, 8 km NEE of Stanford, habitat of T. atriceps (Régimber, 1903), collecting spot marked by an arrow; FG Taiwan, Wufeng distr., 4.3 km SEE of Chaoyang University of Technology, habitat of T. chui sp. nov. (G detail of sandy shore with one alive beetle). Photo D by M. Jäch, photo E by M. Fikáček, FG by H.-C. Liu." figureDoi="http://doi.org/10.5281/zenodo.4488946" httpUri="https://zenodo.org/record/4488946/files/figure.png" pageId="15" pageNumber="244">11</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34065587A21FFB1FF1E07D7360CB01B" box="[151,573,1791,1810]" pageId="15" pageNumber="244" type="reference_group">
<paragraph id="8BE536D37A21FFB1FF1E07D7360CB01B" blockId="15.[151,761,1757,1985]" box="[151,573,1791,1810]" pageId="15" pageNumber="244">
<treatmentCitationGroup id="AB4A11FD7A21FFB1FF1E07D7360CB01B" box="[151,573,1791,1810]" pageId="15" pageNumber="244">
<treatmentCitation id="0AFB10C27A21FFB1FF1E07D7360BB01B" author="HEBAUER F." box="[151,570,1791,1810]" page="398" pageId="15" pageNumber="244" year="2001">
<taxonomicName id="4C5A4D507A21FFB1FF1E07D7360BB01B" ID-CoL="56TH5" authority="Hebauer, 2001: 398" authorityName="Hebauer" authorityPageNumber="398" authorityYear="2001" box="[151,570,1791,1810]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="244" phylum="Arthropoda" rank="species" species="hongsonensis">
<emphasis id="B92EEAC17A21FFB1FF1E07D735BBB01B" box="[151,394,1791,1810]" italics="true" pageId="15" pageNumber="244">Thysanarthria hongsonensis</emphasis>
<bibRefCitation id="EFCB4B227A21FFB1FE0607D7360BB01B" author="Hebauer" box="[399,570,1791,1810]" firstAuthor="Hebauer" page="398" pageId="15" pageNumber="244" pagination="393 - 400" refId="ref18120" refString="HEBAUER F. 2001: The species of the genus Thysanarthria d'Orchymont, 1926 (Coleoptera: Hydrophilidae). Beitrage zur Entomologie 51: 393 - 400." type="journal article" year="2001">Hebauer, 2001: 398</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C34065587A21FFB1FF1E060136ECB0C8" pageId="15" pageNumber="244" type="materials_examined">
<paragraph id="8BE536D37A21FFB1FF1E06013668B078" blockId="15.[151,761,1757,1985]" pageId="15" pageNumber="244">
<emphasis id="B92EEAC17A21FFB1FF1E06013544B035" bold="true" box="[151,373,1833,1852]" pageId="15" pageNumber="244">Type material exmined.</emphasis>
<materialsCitation id="3B323C8E7A21FFB1FEF506013668B078" collectingDate="1999-05-28" collectingDateMax="1999-06-02" collectingDateMin="1999-05-28" collectionCode="SMNS" collectorName="D. Hauck" country="Thailand" elevation="600" latitude="19.450000" longitude="98.333333" pageId="15" pageNumber="244" specimenCount="1" specimenCount-male="1" stateProvince="MAE HONG SON" typeStatus="holotype">
<typeStatus id="54E188717A21FFB1FEF5060135E1B033" box="[380,464,1833,1852]" pageId="15" pageNumber="244" type="holotype">HOLOTYPE</typeStatus>
:
<specimenCount id="9D5CFD5A7A21FFB1FE57060A35DFB032" box="[478,494,1826,1851]" pageId="15" pageNumber="244" type="male"></specimenCount>
(
<collectionCode id="ED4BAE167A21FFB1FE750601360BB035" box="[508,570,1833,1852]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="15" pageNumber="244">SMNS</collectionCode>
),
<emphasis id="B92EEAC17A21FFB1FDC5060134CDB05E" bold="true" pageId="15" pageNumber="244">
<collectingCountry id="F34D76437A21FFB1FDC5060136F2B035" box="[588,707,1833,1852]" name="Thailand" pageId="15" pageNumber="244">THAILAND</collectingCountry>
:
<collectingRegion id="499EF8317A21FFB1FD46060134C7B05C" pageId="15" pageNumber="244">MAE HONG SON</collectingRegion>
:
</emphasis>
ʻThai, N. Mae Hong Son / prov. Soppong env.,
<quantity id="4CA29B367A21FFB1FD31066C36D8B05E" box="[696,745,1860,1879]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="15" pageNumber="244" unit="m" value="600.0">
<elevation id="0077D1E07A21FFB1FD31066C36D8B05E" box="[696,745,1860,1879]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="15" pageNumber="244" unit="m" value="600.0">600m</elevation>
</quantity>
/ 19´27´´N 98´20´´E, 28.5.- /
<date id="FFE410137A21FFB1FE0F067635E2B078" box="[390,467,1886,1905]" pageId="15" pageNumber="244" value="1999-06-02">
<collectingDate id="EFA0E9FB7A21FFB1FE0F067635E2B078" box="[390,467,1886,1905]" pageId="15" pageNumber="244" value="1999-06-02">2.6.1999</collectingDate>
</date>
,
<collectorName id="26AF53057A21FFB1FE520676361CB078" box="[475,557,1886,1905]" pageId="15" pageNumber="244">D. Hauck</collectorName>
leg.
</materialsCitation>
</paragraph>
<paragraph id="8BE536D37A21FFB1FF1E065136ECB0C8" blockId="15.[151,761,1757,1985]" pageId="15" pageNumber="244">
<emphasis id="B92EEAC17A21FFB1FF1E06513581B085" bold="true" box="[151,432,1913,1932]" pageId="15" pageNumber="244">Additional material examined.</emphasis>
<materialsCitation id="3B323C8E7A21FFB1FE3E065136C8B0AE" ID-GBIF-Occurrence="3756360351" pageId="15" pageNumber="244">
<emphasis id="B92EEAC17A21FFB1FE3E065136FBB085" bold="true" box="[439,714,1913,1932]" pageId="15" pageNumber="244">
<collectingCountry id="F34D76437A21FFB1FE3E0651361CB085" box="[439,557,1913,1932]" name="Thailand" pageId="15" pageNumber="244">THAILAND</collectingCountry>
:
<collectingRegion id="499EF8317A21FFB1FDB106513650B083" box="[568,609,1913,1932]" country="Thailand" name="Mae Hong Son" pageId="15" pageNumber="244">MAE</collectingRegion>
HONG SON:
</emphasis>
1 ♂,
<specimenCount id="9D5CFD5A7A21FFB1FF1E06BC34E4B0AE" box="[151,213,1940,1959]" pageId="15" pageNumber="244" type="generic">2 spec.</specimenCount>
(
<collectionCode id="ED4BAE167A21FFB1FF6906BC3519B0AE" box="[224,296,1940,1959]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="15" pageNumber="244" type="Museum">NHMW</collectionCode>
,
<collectionCode id="ED4BAE167A21FFB1FEBB06BC3542B0AE" box="[306,371,1940,1959]" country="Czech Republic" name="National Museum Prague" pageId="15" pageNumber="244">NMPC</collectionCode>
): Mae Ping, at light,
<date id="FFE410137A21FFB1FDA206BC364CB0AE" box="[555,637,1940,1959]" pageId="15" pageNumber="244" value="1991-09-06">6.ix.1991</date>
, lgt. Malicky;
</materialsCitation>
<materialsCitation id="3B323C8E7A21FFB1FF1E068636ECB0C8" ID-GBIF-Occurrence="3756360339" box="[151,733,1960,1985]" pageId="15" pageNumber="244">
1 ♂,
<specimenCount id="9D5CFD5A7A21FFB1FF48068634CFB0C8" box="[193,254,1966,1985]" pageId="15" pageNumber="244" type="generic">1 spec.</specimenCount>
(
<collectionCode id="ED4BAE167A21FFB1FE8106863565B0C8" box="[264,340,1966,1985]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="15" pageNumber="244" type="Museum">NHMW</collectionCode>
): same locality and collector,
<date id="FFE410137A21FFB1FDD9068636E8B0C8" box="[592,729,1966,1985]" pageId="15" pageNumber="244" value="1991-06-24" valueMax="1991-06-25" valueMin="1991-06-24">24.25.vi. 1991</date>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34065587A21FFB1FF1E06F231ACB41C" pageId="15" pageNumber="244" type="description">
<paragraph id="8BE536D37A21FFB1FF1E06F231ACB51C" blockId="15.[151,760,2010,2065]" lastBlockId="15.[828,1437,190,1237]" pageId="15" pageNumber="244">
<emphasis id="B92EEAC17A21FFB1FF1E06F23542B0F8" bold="true" box="[151,371,2010,2033]" pageId="15" pageNumber="244">
Redescription.
<emphasis id="B92EEAC17A21FFB1FEB506F23542B0F8" bold="true" box="[316,371,2010,2033]" italics="true" pageId="15" pageNumber="244">Body</emphasis>
</emphasis>
length
<quantity id="4CA29B367A21FFB1FE3406F2363FB0F8" box="[445,526,2010,2033]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="15" pageNumber="244" unit="mm" value="1.5">1.5 mm</quantity>
, maximum body width
<quantity id="4CA29B367A21FFB1FF1E06D234DABF18" box="[151,235,2042,2065]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="15" pageNumber="244" unit="mm" value="0.9">0.9 mm</quantity>
. Head and labrum black; pronotum dark brown in the middle, becoming weakly paler towards margins; elytra uniformly yellowish; legs yellowish.
<emphasis id="B92EEAC17A21FFB1FB3001F630C2B7FC" bold="true" box="[1209,1267,222,245]" italics="true" pageId="15" pageNumber="244">Head</emphasis>
without microsculpture on interstices; punctation sparse, each puncture bearing pointed seta. Eyes separated by 3.3× the width of one eye in dorsal view.
<emphasis id="B92EEAC17A21FFB1FBB50016309AB65C" bold="true" box="[1084,1195,318,341]" italics="true" pageId="15" pageNumber="244">Pronotum</emphasis>
with sparse setiferous punctation similar to that on head; interstices without microsculpture.
<emphasis id="B92EEAC17A21FFB1FC620056301EB69C" bold="true" box="[1003,1071,382,405]" italics="true" pageId="15" pageNumber="244">Elytra</emphasis>
with 10 striae sharply impressed except anteromedially (near scutellar shield) where neither striae nor serial punctures are visible; intervals weakly convex at midlength and near apex; interval punctation sparse, setiferous; interstices without microsculpture.
</paragraph>
<paragraph id="8BE536D37A21FFB1FCB5033631ACB41C" blockId="15.[828,1437,190,1237]" pageId="15" pageNumber="244">
<emphasis id="B92EEAC17A21FFB1FCB503363796B53C" bold="true" box="[828,935,542,565]" italics="true" pageId="15" pageNumber="244">Aedeagus</emphasis>
(
<figureCitation id="13612A567A21FFB1FC31033637CDB53C" box="[952,1020,542,565]" captionStart="Fig" captionStartId="16.[151,182,1976,1995]" captionTargetBox="[204,1420,202,1944]" captionTargetId="figure-108@16.[162,1439,189,1956]" captionTargetPageId="16" captionText="Fig. 8. Male genitalia of Thysanarthria species, holotypes.AE T. chui sp. nov.(AC whole aedeagus in dorsal, lateral and ventral view; DE detail of median lobe and parameres in dorsal and ventral view). FJ T. hongsonensis Hebauer, 2001 (FH whole aedeagus in dorsal, lateral and ventral view; IJ details of parameres and median lobe in dorsal and ventral view). KO T. madurensis Hebauer, 2001 (KM whole aedeagus in dorsal, lateral and ventral view; NO details of median lobe and parameres in dorsal and ventral view)." figureDoi="http://doi.org/10.5281/zenodo.4488940" httpUri="https://zenodo.org/record/4488940/files/figure.png" pageId="15" pageNumber="244">Figs 8</figureCitation>
FH)
<quantity id="4CA29B367A21FFB1FBC8033630A6B53C" box="[1089,1175,542,565]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="15" pageNumber="244" unit="mm" value="0.5">0.5 mm</quantity>
long. Phallobase at the base of parameres slightly widened, but not wider than parameres combined; very slightly constricted more basally, strongly arcuate in lateral view. Parameres narrow and elongate, narrow basally, gradually narrowing into a rather acute apex, outer face of parameres sinuate. Median lobe nearly reaching apex of parameres, narrow, membranous and rounded apically, without paired subapical projections.
</paragraph>
</subSubSection>
<subSubSection id="C34065587A21FFB1FCB5023630F5B33C" pageId="15" pageNumber="244" type="diagnosis">
<paragraph id="8BE536D37A21FFB1FCB5023630F5B33C" blockId="15.[828,1437,190,1237]" pageId="15" pageNumber="244">
<emphasis id="B92EEAC17A21FFB1FCB502363007B43C" bold="true" box="[828,1078,798,821]" pageId="15" pageNumber="244">Differential diagnosis.</emphasis>
The species is characterized by a tiny body size, absence of dorsal microsculpture and very characteristic aedeagus with narrow elongate parameres. In narrow parameres it may resemble
<taxonomicName id="4C5A4D507A21FFB1FB2702563170B49C" authorityName="Hebauer" authorityYear="2001" box="[1198,1345,894,917]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="244" phylum="Arthropoda" rank="species" species="madurensis">
<emphasis id="B92EEAC17A21FFB1FB2702563170B49C" box="[1198,1345,894,917]" italics="true" pageId="15" pageNumber="244">T. madurensis</emphasis>
</taxonomicName>
but may be distinguished from it by sinuate lateral face of parameres (arcuate in
<taxonomicName id="4C5A4D507A21FFB1FC3202963063B4DC" authorityName="Hebauer" authorityYear="2001" box="[955,1106,958,981]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="244" phylum="Arthropoda" rank="species" species="madurensis">
<emphasis id="B92EEAC17A21FFB1FC3202963063B4DC" box="[955,1106,958,981]" italics="true" pageId="15" pageNumber="244">T. madurensis</emphasis>
</taxonomicName>
), median lobe without paired subapical projections (with the projections in
<taxonomicName id="4C5A4D507A21FFB1FA9502F63769B31C" authorityName="Hebauer" authorityYear="2001" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="244" phylum="Arthropoda" rank="species" species="madurensis">
<emphasis id="B92EEAC17A21FFB1FA9502F63769B31C" italics="true" pageId="15" pageNumber="244">T. madurensis</emphasis>
</taxonomicName>
) and only indistinctly widened base of the phallobase (abruptly widened in
<taxonomicName id="4C5A4D507A21FFB1FBA905363084B33C" authorityName="Hebauer" authorityYear="2001" box="[1056,1205,1054,1077]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="244" phylum="Arthropoda" rank="species" species="madurensis">
<emphasis id="B92EEAC17A21FFB1FBA905363084B33C" box="[1056,1205,1054,1077]" italics="true" pageId="15" pageNumber="244">T. madurensis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C34065587A21FFB1FCB5051637E5B37C" pageId="15" pageNumber="244" type="biology_ecology">
<paragraph id="8BE536D37A21FFB1FCB5051637E5B37C" blockId="15.[828,1437,190,1237]" pageId="15" pageNumber="244">
<emphasis id="B92EEAC17A21FFB1FCB5051637A6B35C" bold="true" box="[828,919,1086,1109]" pageId="15" pageNumber="244">Biology.</emphasis>
Unknown, most examined specimens were collected at light.
</paragraph>
</subSubSection>
<subSubSection id="C34065587A21FFB1FCB5055637B0B3DC" pageId="15" pageNumber="244" type="distribution">
<paragraph id="8BE536D37A21FFB1FCB5055637B0B3DC" blockId="15.[828,1437,190,1237]" pageId="15" pageNumber="244">
<emphasis id="B92EEAC17A21FFB1FCB5055637FBB39C" bold="true" box="[828,970,1150,1173]" pageId="15" pageNumber="244">Distribution.</emphasis>
So far only known from two nearby localities in northern
<collectingCountry id="F34D76437A21FFB1FC3F05B63024B3BC" box="[950,1045,1182,1205]" name="Thailand" pageId="15" pageNumber="244">Thailand</collectingCountry>
(
<collectingRegion id="499EF8317A21FFB1FBA805B63118B3BC" box="[1057,1321,1182,1205]" country="Thailand" name="Mae Hong Son" pageId="15" pageNumber="244">Mae Hong Son Province</collectingRegion>
) (
<bibRefCitation id="EFCB4B227A21FFB1FAB405B63747B3DC" author="HEBAUER" firstAuthor="HEBAUER" pageId="15" pageNumber="244" pagination="393 - 400" refId="ref18120" refString="HEBAUER F. 2001: The species of the genus Thysanarthria d'Orchymont, 1926 (Coleoptera: Hydrophilidae). Beitrage zur Entomologie 51: 393 - 400." type="journal article" year="2001">HEBAUER 2001</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>